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Barkfly - Psocoptera
Barkfly - Psocoptera
Booklice
(Psocoptera)


The booklice (Psocoptera), also called barklice or barkflies, are an order in the class insects (Insecta), the subclass winged insects (Pterygota) and the infraclass Neoptera. Psocoptera comprise approximately 2, 700 species, and are found throughout the world. In Central Europe around 100 booklice species occur.
Booklice reach body lengths of 0. 5 - 10 mm. Most species have brightly coloured bodies. Their antennae are long and thin. Their mouthparts consist of an intermediate form of tools to chew and suck. Some species also have spinning glands. The males’ forewings at their rear edge are reversed like a hook and thus hand along the thicker front edge of the rear wing, while the females have stunted forewings or no forewings at all.
Based on the shape of their segments (antenna limbs) and foot joints, booklice are divided into 3 suborders, namely Trogiomorpha, Troctomorpha and Psocomorpha. Most species found in Germany belong to the Psocomorpha.
The booklice live on different plants, tree trunks, under tree bark, in dead wood, bird nests and even in buildings. They eat fungi, spores, algae and lichens. Occasionally they also eat dead insects. Booklice need a medium to high humidity and temperatures over 15 degrees Celsius and are capable of absorbing water vapour directly from the air. New buildings and buildings with mould infestations are a favourite habitat of booklice. Here the booklice increase the spread of the mould. They can also cause allergies, although this is mainly due to their excretory products.
The pseudoscorpion Chelifer cancroides, which also resides in human environments is a natural enemy of the booklouse. Psocoptera have a life expectancy of up to 12 weeks. A female booklouse is capable of laying up to 120 eggs. Thus, in favourable conditions (in buildings) booklice can produce 8 new generations in a lifetime.
Taxonomy

ClassInsecta
Linnaeus, 1758
Insects, True insects
SubclassPterygota
Lang, 1888
Winged insects
InfraclassNeoptera
van der Wulp, 1890
Modern Wing-folding insects
SuperorderParaneoptera
Martynov, 1923
OrderPsocodea
Yoshizawa & Johnson, 2006
Bark lice, Barklice, Book lice, Booklice, Lice, True lice
Subnodes
SuborderParapsocida Martynov, 1926
SuborderPermopsocida Rasnitsyn & Quicke, 2002
SuborderPsocomorpha Badonnel, 1951
SuborderTroctomorpha Roesler, 1940
SuborderTrogiomorpha Roesler, 1940
FamilyAncistronidae
FamilyAtropidae
FamilyCormopsocidae
FamilyDolabellapsocidae
FamilyPermopsocidae Tillyard, 1926
FamilyPsocidiidae Tillyard, 1926
FamilySurijokopsocidae Becker-Migdisova, 1961
FamilyWestphalopsocidae Azar & al., 2013
FamilyZygopsocidae Tillyard, 1935
GenusActenotarsus Enderlein, 1907
GenusAnisopsocus Ribaga, 1910
GenusAntipsocus Roesler, 1940
GenusBarnola Navas, 1909
GenusBebiosis Enderlein, 1911
GenusBeipiaopsocus Hong, 1992
GenusBurmesopsocus Yoshizawa, 2021
GenusColpostigma Enderlein, 1925
GenusCoryphocopis Enderlein, 1926
GenusDeipnopsocus Enderlein, 1903
GenusEmbidotroctes Enderlein, 1905
GenusEpikodamaius Kuwayama, 1961
GenusEugereon Dohrn, 1866
GenusHormocoria Enderlein, 1926
GenusIdelopsocus Zalessky, 1929
GenusLapithes Bertkau, 1883
GenusLenkoella Machado-Allison & Papavero, 1962
GenusLeptella Reuter, 1893
GenusMaheella Enderlein, 1931
GenusMartynopsocus Karny, 1930
GenusMepachycera Enderlein, 1925
GenusNescus Navas, 1925
GenusOnychotroctes Badonnel, 1969
GenusOxypsocus Tillyard, 1923
GenusPelmatocoria Enderlein, 1910
GenusPermembia Tillyard, 1928
GenusPtenocorium Enderlein, 1931
GenusPteroxaniella Karny, 1932
GenusPtilopsocus Enderlein, 1900
GenusRhyopsocopsis Pearman, 1929
GenusRosega Navas, 1931
GenusScoliopsyllopsis Enderlein, 1912
GenusSemnopsocus Laing, 1925
GenusStenepipsocus Badonnel, 1946
GenusStenotroctes Enderlein, 1905
GenusSyngonosoma Kolbe, 1883
GenusTeratopsocus Reuter, 1894
GenusTitella Navas, 1912
GenusTrigonosceliscus Enderlein, 1910
GenusTrocticus Bertkau, 1882
GenusTropusia Hagen, 1883
GenusUdamolepis Enderlein, 1912
GenusUndensis Vishnyakova, 1981
Synonyms

Copeognatha (Enderlein, 1903)
Psocina (Burmeister, 1839)
Psocoptera (Shipley, 1904)
English
bark lice,
Barklice,
book lice,
Booklice,
Lice,
Parasitic Lice,
true lice


Distribution and Species lists for Psocodea Yoshizawa & Johnson, 2006
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Z

Afghanistan
3 Families  3 Genera  3 Species
Albania
3 Families  4 Genera  5 Species
Algeria
11 Families  14 Genera  27 Species
Angola
25 Families  76 Genera  243 Species
Argentina
19 Families  39 Genera  81 Species
Armenia
7 Families  7 Genera  12 Species
Australia
31 Families  133 Genera  472 Species
Austria
15 Families  37 Genera  83 Species
Bahamas
5 Families  7 Genera  14 Species
Bahrain
1 Family  1 Genus  1 Species
Bangladesh
7 Families  8 Genera  8 Species
Barbados
2 Families  2 Genera  2 Species
Belarus
4 Families  4 Genera  8 Species
Belgium
23 Families  48 Genera  111 Species
Belize
6 Families  8 Genera  19 Species
Benin
9 Families  11 Genera  14 Species
Bhutan
9 Families  18 Genera  24 Species
Bolivia
11 Families  25 Genera  33 Species
Bosnia and Herzegovina
15 Families  35 Genera  77 Species
Brazil
29 Families  100 Genera  411 Species
Bulgaria
13 Families  23 Genera  33 Species
Burkina Faso
2 Families  2 Genera  2 Species
Cambodia
1 Family  1 Genus  1 Species
Cameroon
10 Families  13 Genera  18 Species
Canada
18 Families  44 Genera  112 Species
Cape Verde
8 Families  10 Genera  18 Species
Chile
20 Families  48 Genera  103 Species
China
29 Families  126 Genera  753 Species
Colombia
23 Families  74 Genera  261 Species
Comoros
2 Families  3 Genera  3 Species
Costa Rica
17 Families  29 Genera  46 Species
Croatia
15 Families  35 Genera  84 Species
Cuba
20 Families  40 Genera  79 Species
Cyprus
14 Families  24 Genera  51 Species
Czech Republic
16 Families  35 Genera  81 Species
Democratic Republic of the Congo
1 Family  1 Genus  1 Species
Denmark
20 Families  41 Genera  85 Species
Djibouti
2 Families  2 Genera  2 Species
Dominica
4 Families  4 Genera  4 Species
Dominican Republic
8 Families  11 Genera  14 Species
Ecuador
18 Families  39 Genera  67 Species
Egypt
8 Families  10 Genera  15 Species
El Salvador
2 Families  2 Genera  2 Species
Equatorial Guinea
13 Families  22 Genera  34 Species
Estonia
18 Families  35 Genera  60 Species
Ethiopia
1 Family  1 Genus  1 Species
Fiji
9 Families  22 Genera  74 Species
Finland
16 Families  37 Genera  88 Species
France
28 Families  77 Genera  169 Species
Gabon
4 Families  5 Genera  6 Species
Georgia
7 Families  10 Genera  13 Species
Germany
23 Families  55 Genera  128 Species
Ghana
5 Families  6 Genera  5 Species
Greece
20 Families  38 Genera  80 Species
Guatemala
18 Families  34 Genera  78 Species
Guinea
6 Families  8 Genera  9 Species
Guinea-Bissau
3 Families  3 Genera  3 Species
Guyana
9 Families  9 Genera  12 Species
Haiti
6 Families  8 Genera  10 Species
Honduras
4 Families  8 Genera  17 Species
Hungary
15 Families  32 Genera  73 Species
Iceland
19 Families  34 Genera  48 Species
India
22 Families  65 Genera  136 Species
Indonesia
24 Families  82 Genera  263 Species
Iran
4 Families  4 Genera  4 Species
Ireland
18 Families  42 Genera  92 Species
Israel
16 Families  27 Genera  47 Species
Italy
19 Families  50 Genera  133 Species
Ivory Coast
13 Families  25 Genera  42 Species
Jamaica
21 Families  40 Genera  87 Species
Japan
21 Families  60 Genera  124 Species
Jordan
1 Family  1 Genus  2 Species
Kazakhstan
3 Families  3 Genera  5 Species
Kenya
14 Families  34 Genera  75 Species
Kyrgyzstan
4 Families  4 Genera  5 Species
Laos
1 Families  2 Genera  4 Species
Latvia
14 Families  19 Genera  26 Species
Lebanon
6 Families  10 Genera  12 Species
Lithuania
12 Families  16 Genera  22 Species
Luxembourg
17 Families  33 Genera  86 Species
Madagascar
19 Families  46 Genera  210 Species
Malawi
1 Family  1 Genus  1 Species
Malaysia
23 Families  55 Genera  181 Species
Malta
3 Families  3 Genera  7 Species
Mauritius
6 Families  7 Genera  7 Species
Mexico
32 Families  110 Genera  440 Species
Micronesia
14 Families  30 Genera  95 Species
Monaco
2 Families  2 Genera  2 Species
Mongolia
11 Families  25 Genera  45 Species
Montenegro
1 Family  1 Genus  2 Species
Morocco
16 Families  32 Genera  68 Species
Mozambique
9 Families  15 Genera  24 Species
Myanmar
8 Families  14 Genera  17 Species
Namibia
2 Families  2 Genera  2 Species
Nepal
12 Families  23 Genera  50 Species
Netherlands
18 Families  41 Genera  93 Species
New Zealand
15 Families  41 Genera  84 Species
Nicaragua
13 Families  21 Genera  20 Species
Niger
1 Family  1 Genus  1 Species
Nigeria
16 Families  33 Genera  50 Species
Norway
17 Families  87 Genera  388 Species
Oman
3 Families  3 Genera  4 Species
Pakistan
7 Families  7 Genera  8 Species
Palau
1 Family  1 Genus  1 Species
Palestine
1 Family  1 Genus  2 Species
Panama
25 Families  61 Genera  83 Species
Papua New Guinea
10 Families  28 Genera  59 Species
Paraguay
10 Families  19 Genera  24 Species
Peru
18 Families  46 Genera  109 Species
Philippines
16 Families  35 Genera  86 Species
Poland
19 Families  45 Genera  96 Species
Portugal
18 Families  36 Genera  89 Species
Republic South Africa
19 Families  50 Genera  83 Species
Romania
15 Families  34 Genera  71 Species
Russia
25 Families  71 Genera  153 Species
Saint Lucia
2 Families  2 Genera  2 Species
Saint Vincent and the Grenadines
4 Families  5 Genera  10 Species
Samoa
9 Families  14 Genera  24 Species
São Tomé and Príncipe
1 Family  1 Genus  1 Species
Saudi Arabia
6 Families  7 Genera  7 Species
Senegal
3 Families  6 Genera  16 Species
Serbia
4 Families  4 Genera  7 Species
Seychelles
11 Families  25 Genera  62 Species
Sierra Leone
4 Families  4 Genera  4 Species
Singapore
16 Families  43 Genera  72 Species
Slovakia
14 Families  30 Genera  68 Species
Slovenia
15 Families  35 Genera  83 Species
Solomon Islands
6 Families  10 Genera  16 Species
South Korea
4 Families  7 Genera  6 Species
Spain
24 Families  57 Genera  151 Species
Sri Lanka
18 Families  45 Genera  81 Species
Sudan
2 Families  2 Genera  1 Species
Suriname
7 Families  12 Genera  18 Species
Sweden
20 Families  49 Genera  113 Species
Switzerland
17 Families  37 Genera  105 Species
Syria
2 Families  2 Genera  2 Species
Taiwan
24 Families  79 Genera  221 Species
Tanzania
21 Families  54 Genera  100 Species
Thailand
16 Families  22 Genera  30 Species
Togo
12 Families  18 Genera  24 Species
Tonga
8 Families  13 Genera  22 Species
Trinidad and Tobago
13 Families  20 Genera  41 Species
Tunisia
11 Families  16 Genera  29 Species
Turkey
11 Families  17 Genera  23 Species
Turkmenistan
3 Families  3 Genera  6 Species
Uganda
5 Families  10 Genera  14 Species
Ukraine
7 Families  8 Genera  9 Species
United Arab Emirates
7 Families  9 Genera  11 Species
United Kingdom
22 Families  60 Genera  131 Species
Uruguay
4 Families  6 Genera  7 Species
USA
33 Families  119 Genera  441 Species
Vanuatu
5 Families  5 Genera  10 Species
Venezuela
24 Families  57 Genera  110 Species
Vietnam
10 Families  15 Genera  26 Species
Yemen
4 Families  5 Genera  6 Species
Zambia
4 Families  6 Genera  6 Species
Zimbabwe
12 Families  17 Genera  24 Species


Distribution on islands