Taxonomy of Genus Leptaulax Kaup, 1868
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 4179
190570
2444470 | Families
Genera
Species |
Subclass
| Pterygota
Lang, 1888
Winged insects | 3816
189502
2439455 | Families
Genera
Species |
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 3399
186457
2415487 | Families
Genera
Species |
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 1891
142849
2048615 | Families
Genera
Species |
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 592
62779
813516 | Families
Genera
Species |
Suborder
| Polyphaga
Emery, 1886
Polyphags | 274
54416
687671 | Families
Genera
Species |
Infraorder
| Scarabaeiformia
Crowson, 1960
| 32
5275
90050 | Families
Genera
Species |
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 32
5275
90050 | Families
Genera
Species | | Family | Passalidae
Leach, 1815
Bess beetle, Bess beetles, Betsy beetles, Passalid beetles | 188
2026 | Genera
Species | | Subfamily | Passalinae
Leach, 1815
| 68
1570 | Genera
Species | | Tribe | Leptaulacini
Kaup, 1871
| 3
187 | Genera
Species | | Genus | Leptaulax
Kaup, 1868
| 169 | Species |
Subnodes of Leptaulax Kaup, 1868 | Species | Leptaulax abdominibarbatus Kuwert, 1898
| | | | Species | Leptaulax abdominisculptus Kuwert, 1891
| 1 | Subspecies | | Species | Leptaulax acutangularis Iwase, 1998
| | | | Species | Leptaulax analis Zang, 1906
| | | | Species | Leptaulax anaulax Zang, 1905
| | | | Species | Leptaulax andamanarum Zang, 1905 ⇒ Leptaulax roepstorfi
| | | | Species | Leptaulax angustifrons Kuwert, 1898 ⇒ Leptaulax cyclotaenius
| | | | Species | Leptaulax anibarbis Kuwert, 1898
| | | | Species | Leptaulax anipunctatus Zang, 1905
| | | | Species | Leptaulax anna Zang, 1905
| | | | Species | Leptaulax apicalis Iwase, 1996
| | | | Species | Leptaulax arayai Johki & Kon, 2003
| | | | Species | Leptaulax arrowi Hincks, 1933
| | | | Species | Leptaulax aurivillii Kuwert, 1891 ⇒ Leptaulax differentispina
| | | | Species | Leptaulax barbicauda Zang, 1905
| | | | Species | Leptaulax batchianae Kuwert, 1891
| | | | Species | Leptaulax batschianae Kuwert, 1891
| | | | Species | Leptaulax beccarii Kuwert, 1891
| | | | Species | Leptaulax bicolor Fabricius, 1801 ⇒ Leptaulax dentatus
| 3 | Subspecies | | Species | Leptaulax biroi Endrodi, 1953
| | | | Species | Leptaulax bornensis Kuwert, 1891
| | | | Species | Leptaulax calcuttae Kuwert, 1891
| | | | Species | Leptaulax castilloae Kon, Johki & Araya, 2005
| | | | Species | Leptaulax cazicus Kuwert, 1891
| | | | Species | Leptaulax celebensis Kuwert, 1891
| 2 | Subspecies | | Species | Leptaulax cicatricosus Kuwert, 1891
| | | | Species | Leptaulax circulus Iwase, 1996
| | | | Species | Leptaulax colombinus Kuwert, 1891
| | | | Species | Leptaulax columbinus Kuwert, 1891
| | | | Species | Leptaulax consequens Kuwert, 1898
| | | | Species | Leptaulax crockerensis Iwase, 1998
| | | | Species | Leptaulax cyclotaenius Kuwert, 1891
| | | | Species | Leptaulax darjeilingi Kuwert, 1891 ⇒ Leptaulax dentatus
| | | | Species | Leptaulax darjelingi Kuwert, 1891
| | | | Species | Leptaulax datar Iwase, 1996
| | | | Species | Leptaulax dentatus Fabricius, 1792
| 1 | Subspecies | | Species | Leptaulax dentifrons Iwase, 1996
| | | | Species | Leptaulax differentispina Kuwert, 1891
| 1 | Subspecies | | Species | Leptaulax dindigalensis Kuwert, 1898
| | | | Species | Leptaulax divaricatus Kuwert, 1898
| | | | Species | Leptaulax drescheri Doesburg, 1956
| | | | Species | Leptaulax elegans Iwase, 1996
| | | | Species | Leptaulax eschscholtzi Kaup, 1868
| | | | Species | Leptaulax evidens Kuwert, 1898 ⇒ Leptaulax differentispina
| | | | Species | Leptaulax exterris Kuwert, 1898
| | | | Species | Leptaulax formosanus Doesburg, 1992
| | | | Species | Leptaulax fruhstorferi Zang, 1905
| | | | Species | Leptaulax fujiokae Iwase, 1995
| | | | Species | Leptaulax geminatus Kuwert, 1898
| | | | Species | Leptaulax geminus Kuwert, 1898
| | | | Species | Leptaulax glaber Kirsche, 1877
| 1 | Subspecies | | Species | Leptaulax glabricollis Kuwert, 1898
| | | | Species | Leptaulax glabriventris Gravely, 1914
| | | | Species | Leptaulax hamidi Kon, Araya & Johki, 2021
| | | | Species | Leptaulax hansemani Kuwert, 1898
| | | | Species | Leptaulax hansemanni Kuwert, 1898
| | | | Species | Leptaulax hidakai Johki, Araya & Kon, 2003
| | | | Species | Leptaulax himalayae Kuwert, 1898 ⇒ Leptaulax cyclotaenius
| | | | Species | Leptaulax hirsutus Hincks, 1956
| | | | Species | Leptaulax humerosus Kuwert, 1891 ⇒ Leptaulax dentatus
| | | | Species | Leptaulax immarginalis Iwase, 1996
| 2 | Subspecies | | Species | Leptaulax incipiens Kuwert, 1891
| | | | Species | Leptaulax indicus Kuwert, 1891
| 1 | Subspecies | | Species | Leptaulax innocuus Dejean, 1837
| | | | Species | Leptaulax insignis Kuwert, 1898
| | | | Species | Leptaulax intermedius Iwase, 1996
| | | | Species | Leptaulax interponendus Kuwert, 1898 ⇒ Leptaulax dentatus
| | | | Species | Leptaulax iwasei Johki, Araya & Kon, 2007
| | | | Species | Leptaulax jaechi Iwase, 1998
| | | | Species | Leptaulax jenisi Iwase, 1998
| | | | Species | Leptaulax joliveti Endrodi, 1971
| | | | Species | Leptaulax karoensis Iwase, 1996
| | | | Species | Leptaulax kecil Iwase, 1996
| | | | Species | Leptaulax klugi Kaup, 1868
| | | | Species | Leptaulax klugii Kaup, 1868
| | | | Species | Leptaulax koreanus Nomura, Kon, Johki & Lee, 1993
| | | | Species | Leptaulax labricollis Kuwert, 1898
| | | | Species | Leptaulax labyrinthus Iwase, 1998
| | | | Species | Leptaulax laevis Iwase, 1998
| | | | Species | Leptaulax loebli Kon, Johki & Araya, 2003
| | | | Species | Leptaulax luzonicus Iwase, 1996
| | | | Species | Leptaulax macassariensis Kuwert, 1898 ⇒ Leptaulax differentispina
| | | | Species | Leptaulax malaccae Kuwert, 1891
| | | | Species | Leptaulax malitiosus Kuwert, 1898
| | | | Species | Leptaulax manillae Kuwert, 1891
| | | | Species | Leptaulax manis Iwase, 1996
| | | | Species | Leptaulax masatakai Johki, Araya & Kon, 2003
| | | | Species | Leptaulax masayukii Iwase, 1997
| | | | Species | Leptaulax masumotoi Johki, Araya & Kon, 2011
| | | | Species | Leptaulax matsumotoi Kon, Johki & Araya, 2003
| 3 | Subspecies | | Species | Leptaulax medius Kuwert, 1891
| 1 | Subspecies | | Species | Leptaulax mindanaoensis Iwase, 1996
| | | | Species | Leptaulax mixtus Kuwert, 1898
| | | | Species | Leptaulax negrosensis Iwase, 1996
| | | | Species | Leptaulax niae Kuwert, 1898
| | | | Species | Leptaulax nietneri Zang, 1905
| | | | Species | Leptaulax niisatoi Kon, Johki & Pham, 2017
| | | | Species | Leptaulax novaeguinea Kuwert, 1891
| | | | Species | Leptaulax novaeguineae Kuwert, 1891
| 1 | Subspecies | | Species | Leptaulax obtusidens Kuwert, 1891
| | | | Species | Leptaulax pacholatkoi Iwase, 1998
| 2 | Subspecies | | Species | Leptaulax palawanicus Zang, 1905
| | | | Species | Leptaulax papua Kuwert, 1898 ⇒ Leptaulax anibarbis
| | | | Species | Leptaulax papuanus Zang, 1906
| | | | Species | Leptaulax parvus Iwase, 1995
| 2 | Subspecies | | Species | Leptaulax planicollis Kuwert, 1898
| | | | Species | Leptaulax planus Illiger, 1800 ⇒ Leptaulax roepstorffi
| | | | Species | Leptaulax pulchellus Arrow, 1907
| | | | Species | Leptaulax punctatus Iwase, 1995
| | | | Species | Leptaulax punctipectus Kaup, 1868
| | | | Species | Leptaulax ribbei Kuwert, 1891
| | | | Species | Leptaulax riekoae Iwase, 1998
| 2 | Subspecies | | Species | Leptaulax roepstorffi Kuwert, 1898
| | | | Species | Leptaulax roepstorfi Kuwert, 1898
| | | | Species | Leptaulax rufidorsum Doesburg, 1956
| | | | Species | Leptaulax rugulosus Zang, 1905
| | | | Species | Leptaulax sabahensis Iwase, 1996
| | | | Species | Leptaulax saigonicus Kuwert, 1891
| | | | Species | Leptaulax sakaii Iwase, 1996
| | | | Species | Leptaulax sambawae Gravely, 1918
| | | | Species | Leptaulax sarawakus Iwase, 1996
| 2 | Subspecies | | Species | Leptaulax sawaii Johki, Araya & Kon, 2015
| | | | Species | Leptaulax schillhammeri Iwase, 1998
| | | | Species | Leptaulax schoedli Iwase, 1998
| | | | Species | Leptaulax separadus Kuwert, 1891
| | | | Species | Leptaulax separandus Kuwert, 1891
| 2 | Subspecies | | Species | Leptaulax sequens Kuwert, 1898
| | | | Species | Leptaulax shunichii Kon, Johki & Araya, 2021
| | | | Species | Leptaulax similis Iwase, 1996
| | | | Species | Leptaulax strbai Iwase, 1998
| | | | Species | Leptaulax submedius Kuwert, 1898
| | | | Species | Leptaulax subsequens Kuwert, 1891
| | | | Species | Leptaulax sumatrae Kuwert, 1898
| | | | Species | Leptaulax tamijii Kon, Araya & Johki, 2021
| | | | Species | Leptaulax tenesserimensis Kuwert, 1898
| | | | Species | Leptaulax thoreyi Kaup, 1868
| | | | Species | Leptaulax timoriensis Percheron, 1841
| | | | Species | Leptaulax tonkinensis Zang, 1905
| | | | Species | Leptaulax trajae Iwase, 1998
| | | | Species | Leptaulax uenoi Iwase, 1995 ⇒ Leptaulax arrowi
| | | | Species | Leptaulax ursulus Schaufuss, 1885
| | | | Species | Leptaulax vicinus Percheron, 1844
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Synonyms
Distribution of Leptaulax Kaup, 1868 Australia, Bhutan, Cambodia, China, Colombia, Guatemala, India, Indonesia, Laos, Malaysia, Mexico, Myanmar, Nepal, Papua New Guinea, Philippines, South Korea, Sri Lanka, Taiwan, Thailand, Vietnam
Distribution on islands Alor Archipelago
Links, sources and further information
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