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| | | | Checklist of Tribe Onthophagini Lacordaire, 1856 for East Asia (Far East)
Class | Insecta Linnaeus, 1758 Insects, True insects | 1145 26237 148864 | Families Genera Species | Taxom | Subclass | Pterygota Lang, 1888 Winged insects | 1141 26214 148810 | Families Genera Species | Taxom | Infraclass | Neoptera van der Wulp, 1890 Modern Wing-folding insects | 1053 25800 147485 | Families Genera Species | Taxom | Superorder | Holometabola Heider, 1889 Holometabolous Insects | 627 19553 119028 | Families Genera Species | Taxom | Order | Coleoptera Linnaeus, 1758 Beetles, Coleopterans | 179 7286 50107 | Families Genera Species | Taxom | Suborder | Polyphaga Emery, 1886 Polyphags | 147 6426 41704 | Families Genera Species | Taxom | Infraorder | Scarabaeiformia Crowson, 1960
| 18 703 6141 | Families Genera Species | Taxom | Superfamily | Scarabaeoidea Latreille, 1802 Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 18 703 6141 | Families Genera Species | Taxom | | Family | Scarabaeidae Latreille, 1802 Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 515 4686 | Genera Species | Taxom | | Subfamily | Scarabaeinae Latreille, 1802 Dung Beetles, True dung beetles | 39 529 | Genera Species | Taxom | | Tribe | Onthophagini Lacordaire, 1856
| 9 326 | Genera Species | Taxom |
Genera for East Asia 9 Genera with 326 Species
Species list for East Asia 326 Species and 19 Subspecies | Caccobius (Caccobius) chujoi Paulian, 1942 | | | Caccobius (Caccophilus) scrofa (Balthasar, 1942) | ⇒ Cleptocaccobius scrofa Balthasar, 1942 | | Caccobius (Caccophilus) tortus Sharp, 1875 | | | Caccobius tortus matsudai Ochi & Araya, 1996 | | | Caccobius tortus tortus | | | Caccobius (Caccophilus) unicornis Fabricius, 1798 | | | Caccobius boucomonti Balthasar, 1935 | | | Caccobius brevis Waterhouse, 1875 | | | Caccobius castaneus (Klug, 1855) | ⇒ Caccocnemus castaneus Klug, 1855 | | Caccobius christophi Harold, 1879 | | | Caccobius demangei Boucomont, 1919 | | | Caccobius excavatus Bai, Zhang &..., 2007 | | | Caccobius flavolimbatus Balthasar, 1942 | | | Caccobius fukiensis Balthasar, 1942 | | | Caccobius gonoderus Fairmaire, 1888 | | | Caccobius himalayanus Jekel, 1872 | | | Caccobius hirayamai (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccobius imitans Balthasar, 1933 | | | Caccobius jessoensis Harold, 1867 | | | Caccobius jononis (Matsumura, 1936) | ⇒ Gibbonthophagus viduus Harold, 1875 | | Caccobius kelleri Olsoufieff, 1907 | | | Caccobius masumotoi Cambefort, 1990 | | | Caccobius nikkoensis Lewis, 1895 | | | Caccobius pullus Jekel, 1872 | | | Caccobius sappoensis Matsumura, 1936 | | | Caccobius sapporensis (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccobius schreberi Linnaeus, 1767 | | | Caccobius sordidus Harold, 1886 | | | Caccobius suzukii (Matsumura, 1936) | ⇒ Caccobius boucomonti Balthasar, 1935 | | Caccobius tsunoellus (Matsumara, 1938) | ⇒ Caccophilus tortus Sharp, 1875 | | Caccobius yamauchii (Matsumura, 1936) | ⇒ Caccobius unicornis Fabricius, 1798 | | Caccobius yubariensis (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Cassolus fukiensis Balthasar, 1960 | | | Cassolus gotoi Masumoto, 1986 | | | Cassolus nudus Sharp, 1875 | | | Cleptocaccobius scrofa Balthasar, 1942 | | | Cleptocaccobius simplex Boucomont, 1923 | | | Cleptocaccobius simplex simplex | | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Euonthophagus amyntas subviolaceus Ménétries, 1832 | | | Euonthophagus gibbosus Scriba, 1790 | | | Euonthophagus koshantschikoffi Reitter, 1891 | | | Euonthophagus schnabeli Splichal, 1910 | | | Haroldius hwangi Masumoto, Le..., 2005 | | | Haroldius jechaer Král & Hájek, 2012 | | | Haroldius perroti Paulian, 1939 | | | Haroldius turnai Kral, 2003 | | | Haroldius uenoi Masumoto & Yin, 1993 | | | Onthophagus (Altonthophagus) cervenkai Kabakov, 2008 | | | Onthophagus (Altonthophagus) inelegans Balthasar, 1935 | | | Onthophagus (Altonthophagus) kozlovi Kabakov, 1990 | | | Onthophagus (Altonthophagus) kukunorensis Kabakov, 1990 | | | Onthophagus (Altonthophagus) sibiricus Harold, 1877 | | | Onthophagus (Altonthophagus) turpidoides Kabakov, 2008 | | | Onthophagus (Colobonthophagus) aenescens Wiedemann, 1823 | | | Onthophagus (Colobonthophagus) dama Fabricius, 1798 | | | Onthophagus (Colobonthophagus) quadridentatus (Fabricius, 1798) | ⇒ Onthophagus fissicornis Stevens, 1809 | | Onthophagus (Colobonthophagus) tragus Fabricius, 1792 | | | Onthophagus (Euonthophagus) carbonarius Klug, 1855 | | | Onthophagus (Furconthophagus) furcatus (Fabricius, 1781) | ⇒ Onthophagus furcaticeps Masters, 1886 | | Onthophagus (Gibbonthophagus) apicetinctus (d´Orbigny, 1900) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus (Gibbonthophagus) atripennis Waterhouse, 1875 | | | Onthophagus (Gibbonthophagus) duporti Boucomont, 1914 | | | Onthophagus (Gibbonthophagus) gracilipes Boucomont, 1914 | | | Onthophagus (Gibbonthophagus) kentingensis Nomura, 1973 | | | Onthophagus (Gibbonthophagus) limbatus Herbst, 1789 | | | Onthophagus (Gibbonthophagus) luridipennis Boheman, 1858 | | | Onthophagus (Gibbonthophagus) taurinus White, 1844 | | | Onthophagus (Gibbonthophagus) viduus (Harold, 1875) | ⇒ Onthophagus orbus Boucomont, 1924 | | Onthophagus (Indachorius) arai Masumoto, 1989 | | | Onthophagus (Indachorius) chinensis Balthasar, 1952 | | | Onthophagus (Indachorius) hsui Masumoto, Ch..., 2004 | | | Onthophagus (Indachorius) koshunensis Balthasar, 1941 | | | Onthophagus (Indachorius) platypus Zhang, 1997 | | | Onthophagus (Indachorius) suginoi Ochi, 1984 | | | Onthophagus (Macronthophagus) cludtsi Ochi, 2003 | | | Onthophagus (Macronthophagus) diabolicus Harold, 1877 | | | Onthophagus (Macronthophagus) manipurensis Arrow, 1907 | | | Onthophagus (Macronthophagus) rubricollis Hope, 1831 | | | Onthophagus (Matashia) anguliceps (Boucomont, 1914) | ⇒ Onthophagus avocetta Arrow, 1933 | | Onthophagus (Matashia) yubarinus (Matsumura, 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus (Micronthophagus) konoi Matsumura, 1938 | | | Onthophagus (Micronthophagus) vigilans Boucomont, 1921 | | | Onthophagus (Micronthophagus) wangi Masumoto, Ch..., 2004 | | | Onthophagus (Onthophagiellus) popovi Kabakov, 1983 | | | Onthophagus (Onthophagiellus) taiyaruensis Masumoto, 1977 | | | Onthophagus (Onthophagiellus) unguiculatus Kabakov, 1994 | | | Onthophagus (Onthophagus) acuticollis (Gillet, 1927) | ⇒ Onthophagus amphioxus Arrow, 1933 | | Onthophagus (Onthophagus) acuticornis Endrödi, 1973 | | | Onthophagus (Onthophagus) anguicorius Boucomont, 1914 | | | Onthophagus (Onthophagus) dapcauensis Boucomont, 1921 | | | Onthophagus (Onthophagus) dubernardi Boucomont, 1914 | | | Onthophagus (Onthophagus) fissicornis Stevens, 1809 | | | Onthophagus (Onthophagus) formosanus Gillet, 1924 | | | Onthophagus (Onthophagus) gibbulus Pallas, 1781 | | | Onthophagus (Onthophagus) hajimei Masumoto, 1984 | | | Onthophagus (Onthophagus) hastifer Lansberge, 1885 | | | Onthophagus (Onthophagus) hayashii Masumoto, 1991 | | | Onthophagus (Onthophagus) illyricus Scopoli, 1763 | | | Onthophagus (Onthophagus) koryoensis Kim, 1985 | | | Onthophagus (Onthophagus) kuatunensis Balthasar, 1942 | | | Onthophagus (Onthophagus) lutosopictus Fairmaire, 1897 | | | Onthophagus (Onthophagus) masumotoi Ochi, 1985 | | | Onthophagus (Onthophagus) mendicus Gillet, 1924 | | | Onthophagus (Onthophagus) mulleri van Lansberge, 1883 | | | Onthophagus (Onthophagus) orientalis (Harold, 1868) | ⇒ Onthophagus fissinasus Fairmaire, 1895 | | Onthophagus (Onthophagus) proletarius Harold, 1875 | | | Onthophagus (Onthophagus) pseudoarmatus Balthasar, 1944 | | | Onthophagus (Onthophagus) punctator Reitter, 1893 | | | Onthophagus (Onthophagus) roubali Balthasar, 1935 | | | Onthophagus (Onthophagus) rugulosus Harold, 1886 | | | Onthophagus (Onthophagus) sakainoi Masumoto, 1991 | | | Onthophagus (Onthophagus) sauteri Gillet, 1924 | | | Onthophagus (Onthophagus) setchan Masumoto, 1984 | | | Onthophagus (Onthophagus) sobrius Balthasar, 1959 | | | Onthophagus (Onthophagus) striatulus Palisot de B..., 1809 | Surf blenny | | Onthophagus (Onthophagus) tatsienluensis Balthasar, 1942 | | | Onthophagus (Onthophagus) taurus Schreber, 1759 | Bull Headed Dung Beetle | | Onthophagus (Onthophagus) terminatus Eschscholtz, 1822 | | | Onthophagus (Onthophagus) tritinctus Boucomont, 1914 | | | Onthophagus (Onthophagus) trituber Wiedemann, 1823 | | | Onthophagus trituber jacobsoni Kabakov, 2006 | | | Onthophagus (Onthophagus) yangi Masumoto, Ts..., 2006 | | | Onthophagus (Palaeonthophagus) baraudi Nicolas, 1964 | | | Onthophagus (Palaeonthophagus) clitellifer Reitter, 1894 | | | Onthophagus (Palaeonthophagus) coenobita Herbst, 1783 | | | Onthophagus (Palaeonthophagus) finschi Harold, 1877 | | | Onthophagus (Palaeonthophagus) fissicornis Steven, 1809 | | | Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790) | ⇒ Onthophagus similis Scriba, 1790 | | Onthophagus (Palaeonthophagus) gibbulus Pallas, 1781 | | | Onthophagus gibbulus gibbulus | | | Onthophagus (Palaeonthophagus) gorodinskii Kabakov, 2008 | | | Onthophagus (Palaeonthophagus) grossepunctatus Reitter, 1905 | | | Onthophagus (Palaeonthophagus) haroldi Ballion, 1870 | | | Onthophagus (Palaeonthophagus) joannae Goljan, 1953 | | | Onthophagus (Palaeonthophagus) laticornis Gebler, 1823 | | | Onthophagus (Palaeonthophagus) lemur Fabricius, 1781 | | | Onthophagus (Palaeonthophagus) marginalis Gebler, 1817 | | | Onthophagus marginalis marginalis | | | Onthophagus marginalis przewalski Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) medius (Kugelann, 1792) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus (Palaeonthophagus) muelleri (Lansberge, 1883) | ⇒ Onthophagus statzi Krell, 1990 | | Onthophagus (Palaeonthophagus) nikolajevi Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) novaki Boucomont, 1927 | | | Onthophagus (Palaeonthophagus) nuchicornis Linnaeus, 1758 | Mottled dung beetle | | Onthophagus (Palaeonthophagus) olsoufieffi Boucomont, 1924 | | | Onthophagus (Palaeonthophagus) opacicollis Reitter, 1893 | | | Onthophagus (Palaeonthophagus) ovatus Linnaeus, 1767 | | | Onthophagus (Palaeonthophagus) pygargus Motschulsky, 1845 | | | Onthophagus (Palaeonthophagus) rachelis Martín Piera, 1985 | | | Onthophagus (Palaeonthophagus) ruficapillus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) scabriusculus Harold, 1873 | | | Onthophagus (Palaeonthophagus) semicornis Panzer, 1798 | | | Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767 | | | Onthophagus (Palaeonthophagus) verticicornis (Laicharting, 1781) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus (Palaeonthophagus) vitulus (Fabricius, 1776) | ⇒ Onthophagus immundus Boheman, 1858 | | Onthophagus (Paleonthophagus) andalusicus Waltl, 1835 | | | Onthophagus (Paraphanaeomorphus) argyropygus Gillet, 1927 | | | Onthophagus (Paraphanaeomorphus) carinensis Boucomont, 1914 | | | Onthophagus (Paraphanaeomorphus) vaulogeri Boucomont, 1923 | | | Onthophagus (Parascatonomus) kiuchianus Masumoto, Ya..., 2004 | | | Onthophagus (Parascatonomus) koichii (Ochi & Araya, 1994) | ⇒ Caccobius jessoensis Harold, 1867 | | Onthophagus (Parascatonomus) miyakei Ochi & Araya, 1992 | | | Onthophagus (Parascatonomus) murasakianus Nomura, 1976 | | | Onthophagus (Parascatonomus) tricornis (Wiedemann, 1823) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus (Phanaeomorphus) ater Waterhouse, 1875 | | | Onthophagus (Phanaeomorphus) cernyi (Balthasar, 1935) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus (Phanaeomorphus) lenzi Harold, 1875 | | | Onthophagus potanini hiurai Ochi, 1984 | | | Onthophagus (Serrophorus) laevis (Harold, 1880) | ⇒ Onthophagus laevissimus d´Orbigny, 1905 | | Onthophagus laevis asiaticus Boucomont, 1919 | | | Onthophagus laevis laevis | | | Onthophagus laevis lampromelas Fairmaire, 1891 | | | Onthophagus (Serrophorus) oblongus Zhang, 1997 | | | Onthophagus (Serrophorus) rectecornutus Lansberge, 1883 | | | Onthophagus (Serrophorus) sagittarius Fabricius, 1775 | Sagittarius dung beetle | | Onthophagus (Strandius) changshouensis Zhang, 1997 | | | Onthophagus (Strandius) japonicus Harold, 1875 | | | Onthophagus (Strandius) kuraruanus (Matsumura, 1938) | ⇒ Onthophagus lenzi Harold, 1875 | | Onthophagus (Strandius) potanini R. C. Brusca..., 1990 | | | Onthophagus potanini hiurai Ochi, 1984 | | | Onthophagus potanini potanini | | | Onthophagus (Strandius) taiwanus Nomura, 1973 | | | Onthophagus acrisius Balthasar, 1946 | | | Onthophagus aequiperus Matsumura & ..., 1937 | | | Onthophagus agilis (Matsumura, 1938) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus akirai (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus amamiensis Nomura, 1965 | | | Onthophagus amamiensis amamiensis | | | Onthophagus amamiensis matsumurai Ochi, Kawaha..., 2019 | | | Onthophagus amenthus Matsumura, 1937 | | | Onthophagus amphioxus Arrow, 1933 | | | Onthophagus amycoides Kabakov, 2014 | | | Onthophagus armatus Blanchard, 1853 | | | Onthophagus avocetta Arrow, 1933 | | | Onthophagus balthasari Vsetecka, 1939 | | | Onthophagus basicruentatus Goidanich, 1926 | | | Onthophagus bivertex Heyden, 1887 | | | Onthophagus boucomontianus Balthasar, 1935 | | | Onthophagus brutus Arrow, 1931 | | | Onthophagus campestris Masumoto, Ki..., 2018 | | | Onthophagus capitosus Harold, 1867 | | | Onthophagus carpophilus Pereira & Ha..., 1961 | | | Onthophagus chibanus Matsumura & ..., 1937 | | | Onthophagus chokakurianus Matsumura, 1938 | | | Onthophagus chosensis Matsumura, 1937 | | | Onthophagus chuzenjianus (Matsumura & ..., 1937) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus collaris (Kabakov, 1983) | ⇒ Proagoderus loricatus Klug, 1855 | | Onthophagus convexicollis Boheman, 1858 | | | Onthophagus coracinoides Kabakov, 1983 | | | Onthophagus coracinus Boucomont, 1914 | | | Onthophagus crassicollis Boucomont, 1913 | | | Onthophagus cupreiceps Arrow, 1907 | | | Onthophagus curvispina Reitter, 1893 | | | Onthophagus deflexicollis Lansberge, 1883 | | | Onthophagus dinoderus d´Orbigny, 1913 | | | Onthophagus dissentaneus Balthasar, 1964 | | | Onthophagus dorsofasciatus Fairmaire, 1893 | | | Onthophagus egurianus Matsumura, 1938 | | | Onthophagus euplynes Bates, 1888 | | | Onthophagus excavatus (Redtenbacher, 1848) | ⇒ Liatongus phanaeoides Westwood, 1839 | | Onthophagus fissinasus Fairmaire, 1895 | | | Onthophagus fodiens Waterhouse, 1875 | | | Onthophagus frontalis Raffray, 1877 | | | Onthophagus gagates Hope, 1831 | | | Onthophagus ginyunensis Vsetecka, 1942 | | | Onthophagus heyrovskyi Vsetecka, 1943 | | | Onthophagus hikosanus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus ibonus (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus incollaris Kabakov, 2008 | | | Onthophagus itoi Nomura, 1976 | | | Onthophagus jeannelianus shokhini Kabakov, 2014 | | | Onthophagus jedensis Matsumura, 1937 | | | Onthophagus jiupengensis Masumoto, La..., 2017 | | | Onthophagus kandai (Matsumura, 1937) | ⇒ Onthophagus olsoufieffi Boucomont, 1924 | | Onthophagus kawarinus (Matsumura & ..., 1937) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus klapperichi Balthasar, 1952 | | | Onthophagus kleinei Balthasar, 1935 | | | Onthophagus kogatanus (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus koma Matsumura, 1937 | | | Onthophagus komabellus Matsumura & ..., 1937 | | | Onthophagus komareki Balthasar, 1935 | | | Onthophagus kozunonis (Matsumura, 1937) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus kulti Balthasar, 1952 | | | Onthophagus kuluensis Bates, 1891 | | | Onthophagus laevigatus Fabricius, 1798 | | | Onthophagus laevissimus d´Orbigny, 1905 | | | Onthophagus lenzii Harold, 1874 | | | Onthophagus liui Masumoto, Oc..., 2014 | | | Onthophagus loricatus Klug, 1855 | | | Onthophagus lunatus Harold, 1868 | | | Onthophagus magnini Paulian, 1933 | | | Onthophagus malthinus Gillet, 1930 | | | Onthophagus matsuokai (Matsumura & ..., 1937) | ⇒ Onthophagus sauteri Gillet, 1924 | | Onthophagus micellus Matsumura, 1937 | | | Onthophagus midorianus (Matsumura, 1938) | ⇒ Parascatonomus anguicorius Boucomont, 1914 | | Onthophagus minoi Matsumura & ..., 1937 | | | Onthophagus minokuchianus (Matsumura, 1934) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus misujianus Matsumura & ..., 1937 | | | Onthophagus miyabei (Matsumura, 1937) | ⇒ Onthophagus punctator Reitter, 1893 | | Onthophagus miyazakianus Matsumura & ..., 1937 | | | Onthophagus mushanus Matsumura, 1938 | | | Onthophagus mushensis Matsumura, 1938 | | | Onthophagus nagasawai Matsumura, 1938 | | | Onthophagus nakatomii (Matsumura, 1937) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus necessarius Reitter, 1893 | | | Onthophagus neofurcatus Goidanich, 1926 | | | Onthophagus nitidulus Klug, 1845 | | | Onthophagus nitidus Waterhouse, 1875 | | | Onthophagus obliviosus Balthasar, 1935 | | | Onthophagus ocellatopunctatus Waterhouse, 1875 | | | Onthophagus ohbayashii Nomura, 1939 | | | Onthophagus oishii Matsumura & ..., 1937 | | | Onthophagus okushiranus Matsumura & ..., 1937 | | | Onthophagus okushirianus Matsumura, 1937 | | | Onthophagus oniellus (Matsumura, 1937) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus orbus Boucomont, 1924 | | | Onthophagus oshimanus Nakane, 1960 | | | Onthophagus pacificus Lansberge, 1885 | | | Onthophagus parvulus (Fabricius, 1798) | ⇒ Onthophagus frontalis Raffray, 1877 | | Onthophagus phanaeoides (Westwood, 1839) | ⇒ Proagoderus lanista Castelnau, 1840 | | Onthophagus poeophagus Kabakov, 1979 | | | Onthophagus procurvus Balthasar, 1935 | | | Onthophagus productus Arrow, 1907 | | | Onthophagus pseudobrutus Kabakov, 1983 | | | Onthophagus pseudoconvexicollis Bai & Yang, 2016 | | | Onthophagus pseudojaponicus Balthasar, 1941 | | | Onthophagus pseudojaponicus fukiensis Balthasar, 1941 | | | Onthophagus pseudojaponicus pseudojaponicus | | | Onthophagus pumilus Kabakov, 2014 | | | Onthophagus pupillatus Kolbe, 1886 | | | Onthophagus purpurascens Boucomont, 1914 | | | Onthophagus putealis (Matsumura, 1938) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus schaefernai Balthasar, 1935 | | | Onthophagus scrutator Harold, 1877 | | | Onthophagus senescens Péringuey, 1908 | | | Onthophagus senex Boucomont, 1914 | | | Onthophagus seniculus (Fabricius, 1781) | ⇒ Onthophagus senescens Péringuey, 1908 | | Onthophagus shibatai Nakane, 1960 | | | Onthophagus shigeo (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus shigeoi Matsumura, 1937 | | | Onthophagus shinanensis (Matsumura, 1934) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus shirakii Nakane, 1960 | | | Onthophagus shoyozanus (Matsumura, 1937) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus shurianus Matsumura & ..., 1937 | | | Onthophagus simplicifrons Reitter, 1893 | | | Onthophagus sinicus Zhang & Wang, 1997 | | | Onthophagus smetanai Balthasar, 1952 | | | Onthophagus sobosanus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus solivagus Harold, 1886 | | | Onthophagus sonani Miwa, 1930 | | | Onthophagus spinifex Fabricius, 1781 | | | Onthophagus spitsbergeniensis Krell, 2010 | | | Onthophagus statzi ♰ Krell, 1990 | | | Onthophagus sternax Balthasar, 1959 | | | Onthophagus strandi Balthasar, 1935 | | | Onthophagus sulci Balthasar, 1935 | | | Onthophagus susterai Balthasar, 1952 | | | Onthophagus sutleinensis Splichal, 1910 | | | Onthophagus sycophanta (Fairmaire, 1887) | ⇒ Euonthophagus gibbosus Scriba, 1790 | | Onthophagus tabidus Balthasar, 1935 | | | Onthophagus takabayashii Matsumura, 1937 | | | Onthophagus tibetanus Arrow, 1907 | | | Onthophagus tragoides Boucomont, 1914 | | | Onthophagus tricolor Boucomont, 1914 | | | Onthophagus turfanicus Balthasar, 1971 | | | Onthophagus turpidus Reitter, 1887 | | | Onthophagus uedanus (Matsumura, 1937) | ⇒ Onthophagus olsoufieffi Boucomont, 1924 | | Onthophagus uniformis Heyden, 1886 | | | Onthophagus ushiodai (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus yakuinsulanus Nakane, 1984 | | | Onthophagus yanoi Matsumura, 1938 | | | Onthophagus yaoi Masumoto, Oc..., 2014 | | | Onthophagus yugianus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus yumigatanus Matsumura, 1937 | | | Onthophagus zavreli Balthasar, 1935 | | | Onthophagus zimmermanni Balthasar, 1959 | | | Proagoderus amplexus Sharp, 1875 | | | Proagoderus lanista Castelnau, 1840 | | | Proagoderus loricatus Klug, 1855 | | | Proagoderus panoplus Bates, 1888 | | | Proagoderus yunnanus Boucomont, 1912 | |
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