Passalinae (Leach 1815) ⇒ Taxonomy and Distribution of Subfamily

Ants	Arachnids
Bees	Beetles
Booklice	Bugs
Bumblebees	Butterflies
Cicadas	Crane flies
Dragonflies	Earwigs
Flies	Isopods
Locusts	Mantises
Mayflies	Mosquitoes
Myriapods	Net-winged
Plant-parasitic	Scorpion flies
Termites	Wasps

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Angola Argentina Australia Bahamas Belize Bhutan Bolivia Brazil Cambodia Cameroon Canada Chile China Colombia Costa Rica Cuba Democratic Republic of the Congo Dominica Dominican Republic Ecuador El Salvador Equatorial Guinea Ethiopia Fiji France Gabon Georgia Germany Ghana Greece Grenada Guatemala Guinea Guyana Haiti Honduras India Indonesia Ivory Coast Jamaica Kenya Laos Liberia Madagascar Malawi Malaysia Mexico Micronesia Mozambique Myanmar Nepal Nicaragua Nigeria North Korea Pakistan Palau Panama Papua New Guinea Paraguay Peru Philippines Republic of the Congo Republic South Africa Russia Rwanda Saint Lucia Saint Vincent and the Grenadines São Tomé and Príncipe Sierra Leone Singapore Solomon Islands South Korea Spain Sri Lanka Suriname Sweden Taiwan Tanzania Thailand Togo Trinidad and Tobago Uganda United Kingdom Uruguay USA Vanuatu Venezuela Vietnam Zambia Zimbabwe

INSECTOIDGERENGDARK LIGHT

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Taxonomy of Subfamily Passalinae Leach, 1815

Class	Insecta Linnaeus, 1758 Insects, True insects	4179 190570 2444470	Families Genera Species
Subclass	Pterygota Lang, 1888 Winged insects	3816 189502 2439455	Families Genera Species
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	3399 186457 2415487	Families Genera Species
Superorder	Holometabola Heider, 1889 Holometabolous Insects	1891 142849 2048615	Families Genera Species
Order	Coleoptera Linnaeus, 1758 Beetles, Coleopterans	592 62779 813516	Families Genera Species
Suborder	Polyphaga Emery, 1886 Polyphags	274 54416 687671	Families Genera Species
Infraorder	Scarabaeiformia Crowson, 1960	32 5275 90050	Families Genera Species
Superfamily	Scarabaeoidea Latreille, 1802 Bess Beetles, Chafer, Dung Beetles, D´or Beetles	32 5275 90050	Families Genera Species
Family	Passalidae Leach, 1815 Bess beetle, Bess beetles, Betsy beetles, Passalid beetles	188 2026	Genera Species
Subfamily	Passalinae Leach, 1815	68 1570	Genera Species

Subnodes of Passalinae Leach, 1815

Tribe	Leptaulacini Kaup, 1871	3 187	Genera Species
Tribe	Macrolinini Kaup, 1871	17 282	Genera Species
Tribe	Passalini Leach, 1815	16 597	Genera Species
Tribe	Proculini Kaup, 1868	22 427	Genera Species
Tribe	Solenocyclini Kaup, 1871	9 74	Genera Species
Genus	Oileoides Gravely, 1918 ⇒ Popilius	3	Species

Distribution and Species lists of Passalinae Leach, 1815

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Angola
1 Genus  1 SpeciesArgentina
10 Genera  36 SpeciesAustralia
13 Genera  27 SpeciesBahamas
5 Genera  7 SpeciesBelize
6 Genera  8 SpeciesBhutan
5 Genera  12 SpeciesBolivia
6 Genera  51 SpeciesBrazil
12 Genera  151 SpeciesCambodia
4 Genera  9 SpeciesCameroon
3 Genera  13 SpeciesCanada
4 Genera  4 SpeciesChile
1 Genus  5 SpeciesChina
7 Genera  16 SpeciesColombia
15 Genera  168 SpeciesCosta Rica
17 Genera  91 SpeciesCuba
6 Genera  14 SpeciesDemocratic Republic of the Congo
3 Genera  17 SpeciesDominica
3 Genera  6 SpeciesDominican Republic
3 Genera  5 SpeciesEcuador
10 Genera  79 SpeciesEl Salvador
15 Genera  27 SpeciesEquatorial Guinea
3 Genera  9 SpeciesEthiopia
1 Genus  1 SpeciesFiji
1 Genus  1 SpeciesFrance
6 Genera  6 SpeciesGabon
2 Genera  5 SpeciesGeorgia
1 Genus  1 SpeciesGermany
1 Genus  0 SpeciesGhana
3 Genera  11 SpeciesGreece
1 Genus  1 SpeciesGrenada
2 Genera  4 SpeciesGuatemala
32 Genera  115 SpeciesGuinea
1 Genus  2 SpeciesGuyana
5 Genera  25 SpeciesHaiti
2 Genera  3 SpeciesHonduras
13 Genera  41 SpeciesIndia
8 Genera  30 SpeciesIndonesia
19 Genera  185 SpeciesIvory Coast
3 Genera  9 SpeciesJamaica
2 Genera  2 SpeciesKenya
2 Genera  3 SpeciesLaos
4 Genera  9 SpeciesLiberia
8 Genera  10 SpeciesMadagascar
7 Genera  12 SpeciesMalawi
1 Genus  1 SpeciesMalaysia
8 Genera  106 SpeciesMexico
22 Genera  131 SpeciesMicronesia
2 Genera  2 SpeciesMozambique
1 Genus  1 SpeciesMyanmar
7 Genera  26 SpeciesNepal
3 Genera  3 SpeciesNicaragua
13 Genera  35 SpeciesNigeria
2 Genera  2 SpeciesNorth Korea
1 Genus  1 SpeciesPakistan
1 Genus  1 SpeciesPalau
2 Genera  2 SpeciesPanama
13 Genera  43 SpeciesPapua New Guinea
11 Genera  80 SpeciesParaguay
6 Genera  30 SpeciesPeru
7 Genera  70 SpeciesPhilippines
8 Genera  30 SpeciesRepublic of the Congo
3 Genera  7 SpeciesRepublic South Africa
1 Genus  1 SpeciesRussia
2 Genera  3 SpeciesRwanda
3 Genera  6 SpeciesSaint Lucia
2 Genera  4 SpeciesSaint Vincent and the Grenadines
2 Genera  4 SpeciesSão Tomé and Príncipe
2 Genera  3 SpeciesSierra Leone
2 Genera  4 SpeciesSingapore
2 Genera  3 SpeciesSolomon Islands
3 Genera  3 SpeciesSouth Korea
1 Genus  1 SpeciesSpain
6 Genera  7 SpeciesSri Lanka
5 Genera  18 SpeciesSuriname
6 Genera  26 SpeciesSweden
4 Genera  4 SpeciesTaiwan
2 Genera  5 SpeciesTanzania
2 Genera  3 SpeciesThailand
7 Genera  23 SpeciesTogo
2 Genera  3 SpeciesTrinidad and Tobago
8 Genera  20 SpeciesUganda
2 Genera  9 SpeciesUnited Kingdom
4 Genera  9 SpeciesUruguay
1 Genus  1 SpeciesUSA
11 Genera  27 SpeciesVanuatu
1 Genus  2 SpeciesVenezuela
7 Genera  47 SpeciesVietnam
7 Genera  22 SpeciesZambia
1 Genus  0 SpeciesZimbabwe
1 Genus  1 Species

Distribution on islands

Admiralty Island
1 Genus  1 SpeciesAlor Archipelago
1 Genus  2 SpeciesAzores
1 Genus  1 SpeciesBacan islands
3 Genera  4 SpeciesBarat Daya Islands
1 Genus  1 SpeciesBasse-Terre Island
1 Genus  1 SpeciesCaroline Island
1 Genus  1 SpeciesIsla Chica
1 Genus  1 SpeciesIsla Grande
1 Genus  1 SpeciesIslands of Chile
1 Genus  1 SpeciesKai Islands
2 Genera  2 SpeciesLa Gomera
2 Genera  2 SpeciesLa Palma
2 Genera  2 SpeciesMacaronesia
5 Genera  5 SpeciesNias Island
1 Genus  1 SpeciesSouthern Line Islands
1 Genus  1 SpeciesTernate
3 Genera  6 Species

Links, sources and further information

Passalinae at itis.gov