Species list, Checklist of Infraorder Scarabaeiformia for Lower Miocene (Miocene)

Ants	Arachnids
Bees	Beetles
Booklice	Bugs
Bumblebees	Butterflies
Cicadas	Crane flies
Dragonflies	Earwigs
Flies	Isopods
Locusts	Mantises
Mayflies	Mosquitoes
Myriapods	Net-winged
Plant-parasitic	Scorpion flies
Termites	Wasps

Taxonomy & Distribution
Browser for Arthropods

Region search

Countries

#ABCDEFGHIJKLMNOPQRSTUVYZ

Afghanistan Albania Algeria Andorra Angola Antigua and Barbuda Argentina Armenia Australia Austria Azerbaijan Bahamas Bahrain Bangladesh Barbados Belarus Belgium Belize Benin Bhutan Bolivia Bosnia and Herzegovina Botswana Brazil Brunei Bulgaria Burkina Faso Burundi Cambodia Cameroon Canada Cape Verde Central African Republic Chad Chile China Colombia Comoros Costa Rica Croatia Cuba Cyprus Czech Republic Democratic Republic of the Congo Denmark Djibouti Dominica Dominican Republic East Timor Ecuador Egypt El Salvador Equatorial Guinea Eritrea Estonia Ethiopia Fiji Finland France Gabon Gambia Georgia Germany Ghana Greece Grenada Guatemala Guinea Guinea-Bissau Guyana Haiti Honduras Hungary Iceland India Indonesia Iran Iraq Ireland Israel Italy Ivory Coast Jamaica Japan Jordan Kazakhstan Kenya Kiribati Kosovo Kuwait Kyrgyzstan Laos Latvia Lebanon Lesotho Liberia Libya Liechtenstein Lithuania Luxembourg Madagascar Malawi Malaysia Maldives Mali Malta Marshall Islands Mauritania Mauritius Mexico Micronesia Moldova Monaco Mongolia Montenegro Morocco Mozambique Myanmar Namibia Nauru Nepal Netherlands New Zealand Nicaragua Niger Nigeria Niue North Korea Norway Oman Pakistan Palau Palestine Panama Papua New Guinea Paraguay Peru Philippines Poland Portugal Qatar Republic of the Congo Republic South Africa Romania Russia Rwanda Saint Kitts and Nevis Saint Lucia Saint Vincent and the Grenadines Samoa San Marino São Tomé and Príncipe Saudi Arabia Senegal Serbia Seychelles Sierra Leone Singapore Slovakia Slovenia Solomon Islands Somalia South Korea South Sudan Spain Sri Lanka Sudan Suriname Sweden Switzerland Syria Taiwan Tajikistan Tanzania Thailand Togo Tonga Transnistria Trinidad and Tobago Tunisia Turkey Turkmenistan Tuvalu Uganda Ukraine United Arab Emirates United Kingdom Uruguay USA Uzbekistan Vanuatu Venezuela Vietnam Yemen Zambia Zimbabwe

INSECTOIDGERENGDARK LIGHT

⌕

Checklist for Infraorder Scarabaeiformia Crowson, 1960
and region Lower Miocene (Miocene)

Class	Insecta Linnaeus, 1758 Insects, True insects	165 413 592	Families Genera Species	Taxom
Subclass	Pterygota Lang, 1888 Winged insects	163 411 590	Families Genera Species	Taxom
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	154 390 570	Families Genera Species	Taxom
Superorder	Holometabola Heider, 1889 Holometabolous Insects	96 287 470	Families Genera Species	Taxom
Order	Coleoptera Linnaeus, 1758 Beetles, Coleopterans	26 77 84	Families Genera Species	Taxom
Suborder	Polyphaga Emery, 1886 Polyphags	21 58 62	Families Genera Species	Taxom
Infraorder	Scarabaeiformia Crowson, 1960	2 8 6	Families Genera Species	Taxom

List of families for Lower Miocene
2 Families width 6 Species

Scarabaeidae Latreille, 1802	Dung beetles	6	Species	Taxom
Geotrupidae (Latreille, 1802)	Dor beetles	0	Species	Taxom

Genera for Lower Miocene
8 Genera with 6 Species

Anachalcos Hope, 1976		1	Species	Taxom
Aphodius Hellwig, 1798		0	Species	Taxom
Copris Geoffroy, 1976	Mexican dung beetle	1	Species	Taxom
Geotrupes Latreille, 1796	Dor Beetle	0	Species	Taxom
Heliocopris Hope, 1968		1	Species	Taxom
Melolontha Fabricius, 1878	Billy witch	1	Species	Taxom
Melolonthites Heer, 1926		1	Species	Taxom
Metacatharsius Paulian, 1976		1	Species	Taxom

Species list for Lower Miocene
6 Species

Anachalcos mfwangani ♰ R. Paulian, 1976
Copris leakeyorum ♰ R. Paulian, 1976
Heliocopris antiquus ♰ I. Fujiyama, 1968
Melolontha solitaria ♰ O. Novák, 1878
Melolonthites interemtus ♰ Cockerell, 1926
Metacatharsius rusingae ♰ R. Paulian, 1976

Subnodes of Scarabaeiformia with occurrences in Lower Miocene

Superfamily

Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles, Scarab Beetles, Stag Beetles

2
8
6

Families
Genera
Species

Taxom