Species list, Checklist of Infraorder Scarabaeiformia for Lower Pliocene (Pliocene)

Ants	Arachnids
Bees	Beetles
Booklice	Bugs
Bumblebees	Butterflies
Cicadas	Crane flies
Dragonflies	Earwigs
Flies	Isopods
Locusts	Mantises
Mayflies	Mosquitoes
Myriapods	Net-winged
Plant-parasitic	Scorpion flies
Termites	Wasps

Taxonomy & Distribution
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INSECTOIDGERENGDARK LIGHT

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Checklist for Infraorder Scarabaeiformia Crowson, 1960
and region Lower Pliocene (Pliocene)

Class	Insecta Linnaeus, 1758 Insects, True insects	12 20 15	Families Genera Species	Taxom
Subclass	Pterygota Lang, 1888 Winged insects	12 20 15	Families Genera Species	Taxom
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	12 20 15	Families Genera Species	Taxom
Superorder	Holometabola Heider, 1889 Holometabolous Insects	10 18 13	Families Genera Species	Taxom
Order	Coleoptera Linnaeus, 1758 Beetles, Coleopterans	9 17 12	Families Genera Species	Taxom
Suborder	Polyphaga Emery, 1886 Polyphags	7 15 10	Families Genera Species	Taxom
Infraorder	Scarabaeiformia Crowson, 1960	1 2 2	Family Genera Species	Taxom

List of families for Lower Pliocene
1 Family width 2 Species

Scarabaeidae Latreille, 1802

Dung beetles

Species

Taxom

Genera for Lower Pliocene
2 Genera with 2 Species

Calcitoryctes Krell, 2011		1	Species	Taxom
Copris Geoffroy, 1988	Mexican dung beetle	1	Species	Taxom

Species list for Lower Pliocene
2 Species

Calcitoryctes magnificus ♰ F. T. Krell, 2011
Copris (Copris) kartlinus ♰ O. N. Kabakov, 1988

Subnodes of Scarabaeiformia with occurrences in Lower Pliocene

Superfamily

Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles, Scarab Beetles, Stag Beetles

1
2
2

Family
Genera
Species

Taxom