Species list, Checklist of Infraorder Scarabaeiformia for Madison (Clarion County, Pennsylvania, USA)

Ants	Arachnids
Bees	Beetles
Booklice	Bugs
Bumblebees	Butterflies
Cicadas	Crane flies
Dragonflies	Earwigs
Flies	Isopods
Locusts	Mantises
Mayflies	Mosquitoes
Myriapods	Net-winged
Plant-parasitic	Scorpion flies
Termites	Wasps

Taxonomy & Distribution
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INSECTOIDGERENGDARK LIGHT

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Checklist for Infraorder Scarabaeiformia Crowson, 1960
and region Madison (Clarion County, Pennsylvania, USA)

Class	Insecta Linnaeus, 1758 Insects, True insects	113 365 465	Families Genera Species	Taxom
Subclass	Pterygota Lang, 1888 Winged insects	113 365 465	Families Genera Species	Taxom
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	105 347 442	Families Genera Species	Taxom
Superorder	Holometabola Heider, 1889 Holometabolous Insects	80 297 381	Families Genera Species	Taxom
Order	Coleoptera Linnaeus, 1758 Beetles, Coleopterans	22 68 77	Families Genera Species	Taxom
Suborder	Polyphaga Emery, 1886 Polyphags	19 61 67	Families Genera Species	Taxom
Infraorder	Scarabaeiformia Crowson, 1960	1 6 6	Family Genera Species	Taxom

List of families for Madison (Clarion County)
1 Family width 6 Species

Scarabaeidae Latreille, 1802

Dung beetles

Species

Taxom

Genera for Madison (Clarion County)
6 Genera with 6 Species

Aphodius Hellwig, 1774		1	Species	Taxom
Blackburneus Schmidt, 1845		1	Species	Taxom
Cotalpa Burmeister, 1758		1	Species	Taxom
Cotinis Burmeister, 1758	Green June Beetles	1	Species	Taxom
Melinopterus Mulsant, 1790		1	Species	Taxom
Phyllophaga Harris, 1856	June beetles	1	Species	Taxom

Species list for Madison (Clarion County)
6 Species

Aphodius (Aphodius) pedellus (DeGeer, 1774)	⇒ Aphodius fimetarius Linnaeus, 1758
Blackburneus stercorosus Melsheimer, 1845
Cotalpa lanigera Linnaeus, 1758	Goldsmith beetle
Cotinis (Cotinis) nitida Linnaeus, 1758	Common Green June Beetle
Melinopterus prodromus Brahm, 1790	Spring small dung beetle
Phyllophaga (Phyllophaga) nitida LeConte, 1856

Subnodes of Scarabaeiformia with occurrences in Madison (Clarion County)

Superfamily

Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles, Scarab Beetles, Stag Beetles

1
6
6

Family
Genera
Species

Taxom