Species list, Checklist of Infraorder Scarabaeiformia for Main (Columbia County, Pennsylvania, USA)

Ants	Arachnids
Bees	Beetles
Booklice	Bugs
Bumblebees	Butterflies
Cicadas	Crane flies
Dragonflies	Earwigs
Flies	Isopods
Locusts	Mantises
Mayflies	Mosquitoes
Myriapods	Net-winged
Plant-parasitic	Scorpion flies
Termites	Wasps

Taxonomy & Distribution
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INSECTOIDGERENGDARK LIGHT

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Checklist for Infraorder Scarabaeiformia Crowson, 1960
and region Main (Columbia County, Pennsylvania, USA)

Class	Insecta Linnaeus, 1758 Insects, True insects	77 163 169	Families Genera Species	Taxom
Subclass	Pterygota Lang, 1888 Winged insects	77 163 169	Families Genera Species	Taxom
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	73 158 165	Families Genera Species	Taxom
Superorder	Holometabola Heider, 1889 Holometabolous Insects	59 142 149	Families Genera Species	Taxom
Order	Coleoptera Linnaeus, 1758 Beetles, Coleopterans	16 55 63	Families Genera Species	Taxom
Suborder	Polyphaga Emery, 1886 Polyphags	15 44 45	Families Genera Species	Taxom
Infraorder	Scarabaeiformia Crowson, 1960	3 6 6	Families Genera Species	Taxom

List of families for Main (Pennsylvania)
3 Families width 6 Species

Scarabaeidae Latreille, 1802	Dung beetles	3	Species	Taxom
Geotrupidae Latreille, 1802	Dor beetles	1	Species	Taxom
Lucanidae Latreille, 1804	Stag Beetle	2	Species	Taxom

Genera for Main (Pennsylvania)
6 Genera with 6 Species

Anomala Samouelle, 1941	Shining Leaf Chafer	1	Species	Taxom
Anoplotrupes Jekel, 1791		1	Species	Taxom
Calamosternus Motschulsky, 1767		1	Species	Taxom
Dorcus MacLeay, 1758		1	Species	Taxom
Lucanus Scopoli, 1758		1	Species	Taxom
Phyllophaga Harris, 1950	June beetles	1	Species	Taxom

Species list for Main (Pennsylvania)
6 Species

Anomala adscita Robinson, 1941
Anoplotrupes stercorosus Scriba, 1791	Dor Beetle
Calamosternus granarius Linnaeus, 1767	Granular Small Dung Beetle
Dorcus parallelipipedus Linnaeus, 1758	Lesser stag beetle
Lucanus (Lucanus) cervus Linnaeus, 1758	Stag beetle
Phyllophaga (Phyllophaga) panorpa Sanderson, 1950

Subnodes of Scarabaeiformia with occurrences in Main (Pennsylvania)

Superfamily

Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles, Scarab Beetles, Stag Beetles

3
6
6

Families
Genera
Species

Taxom