Species list, Checklist of Infraorder Scarabaeiformia for Middle Eocene (Eocene)

Ants	Arachnids
Bees	Beetles
Booklice	Bugs
Bumblebees	Butterflies
Cicadas	Crane flies
Dragonflies	Earwigs
Flies	Isopods
Locusts	Mantises
Mayflies	Mosquitoes
Myriapods	Net-winged
Plant-parasitic	Scorpion flies
Termites	Wasps

Taxonomy & Distribution
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INSECTOIDGERENGDARK LIGHT

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Checklist for Infraorder Scarabaeiformia Crowson, 1960
and region Middle Eocene (Eocene)

Class	Insecta Linnaeus, 1758 Insects, True insects	106 278 448	Families Genera Species	Taxom
Subclass	Pterygota Lang, 1888 Winged insects	106 278 448	Families Genera Species	Taxom
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	94 264 433	Families Genera Species	Taxom
Superorder	Holometabola Heider, 1889 Holometabolous Insects	58 215 376	Families Genera Species	Taxom
Order	Coleoptera Linnaeus, 1758 Beetles, Coleopterans	22 107 209	Families Genera Species	Taxom
Suborder	Polyphaga Emery, 1886 Polyphags	20 97 195	Families Genera Species	Taxom
Infraorder	Scarabaeiformia Crowson, 1960	3 7 8	Families Genera Species	Taxom

List of families for Middle Eocene
3 Families width 8 Species

Scarabaeidae Latreille, 1802	Dung beetles	6	Species	Taxom
Geotrupidae Latreille, 1802	Dor beetles	1	Species	Taxom
Lucanidae Latreille, 1804	Stag Beetle	1	Species	Taxom

Genera for Middle Eocene
7 Genera with 8 Species

Eophyllocerus Haupt, 1950		2	Species	Taxom
Geotrupes Latreille, 1921	Dor Beetle	1	Species	Taxom
Gymnopleurus Illiger, 1921		1	Species	Taxom
Melolonthites Heer, 1929		1	Species	Taxom
Oryctoantiquus Ratcliffe & ..., 2005		1	Species	Taxom
Pelidnotites Cockerell, 1920		1	Species	Taxom
Protognathinus Chalumeau & ..., 2001		1	Species	Taxom

Species list for Middle Eocene
8 Species

Eophyllocerus glaucinus ♰ Haupt, 1950
Eophyllocerus scrobiculatus ♰ Haupt, 1950
Geotrupes messelensis ♰ F. Meunier, 1921
Gymnopleurus eocaenicus ♰ F. Meunier, 1921
Melolonthites collinsi ♰ H. F. Wickham, 1929
Oryctoantiquus borealis ♰ B. C. Ratcliffe, 2005
Pelidnotites atavus ♰ T. D. A. Coc..., 1920
Protognathinus spielbergi ♰ Chalumeau & ..., 2001

Subnodes of Scarabaeiformia with occurrences in Middle Eocene

Superfamily

Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles, Scarab Beetles, Stag Beetles

3
7
8

Families
Genera
Species

Taxom