Species list, Checklist of Infraorder Scarabaeiformia for Upper Pliocene (Pliocene)

Ants	Arachnids
Bees	Beetles
Booklice	Bugs
Bumblebees	Butterflies
Cicadas	Crane flies
Dragonflies	Earwigs
Flies	Isopods
Locusts	Mantises
Mayflies	Mosquitoes
Myriapods	Net-winged
Plant-parasitic	Scorpion flies
Termites	Wasps

Taxonomy & Distribution
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INSECTOIDGERENGDARK LIGHT

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Checklist for Infraorder Scarabaeiformia Crowson, 1960
and region Upper Pliocene (Pliocene)

Class	Insecta Linnaeus, 1758 Insects, True insects	87 164 186	Families Genera Species	Taxom
Subclass	Pterygota Lang, 1888 Winged insects	87 164 186	Families Genera Species	Taxom
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	82 157 180	Families Genera Species	Taxom
Superorder	Holometabola Heider, 1889 Holometabolous Insects	57 117 140	Families Genera Species	Taxom
Order	Coleoptera Linnaeus, 1758 Beetles, Coleopterans	21 69 71	Families Genera Species	Taxom
Suborder	Polyphaga Emery, 1886 Polyphags	18 56 58	Families Genera Species	Taxom
Infraorder	Scarabaeiformia Crowson, 1960	1 3 3	Family Genera Species	Taxom

List of families for Upper Pliocene
1 Family width 3 Species

Scarabaeidae Latreille, 1802

Dung beetles

Species

Taxom

Genera for Upper Pliocene
3 Genera with 3 Species

Aphodius Hellwig, 1758		1	Species	Taxom
Melolonthites Heer, 1935		1	Species	Taxom
Oryctes Hellwig, 1758	Coconut rhinoceros beetles	1	Species	Taxom

Species list for Upper Pliocene
3 Species

Aphodius (Acrossus) rufipes (Linnaeus, 1758)	⇒ Acrossus rufipes Linnaeus, 1758
Melolonthites laterosinuatus Piton, 1935
Oryctes (Oryctes) nasicornis Linnaeus, 1758	European rhinoceros beetle

Subnodes of Scarabaeiformia with occurrences in Upper Pliocene

Superfamily

Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles, Scarab Beetles, Stag Beetles

1
3
3

Family
Genera
Species

Taxom