Checklist of Superfamily Scarabaeoidea Latreille, 1802 (Bess Beetles)
for Ryukyu Islands (Okinawa Prefecture, Japan)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 348
2667
4080 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 348
2667
4080 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 333
2610
3989 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 220
2112
3277 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 73
984
1617 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 68
881
1437 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 8
71
117 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 8
71
117 | Families
Genera
Species |
Taxom
|
List of families for Ryukyu Islands
8 Families width 117 Species and 26 Subspecies
Genera for Ryukyu Islands
71 Genera with 117 Species
| Adoretus Laporte, 1965 | | 2 | Species | Taxom | | Aegus MacLeay, 1873 | | 9 | Species | Taxom | | Aganocrossus Reitter, 1862 | | 1 | Species | Taxom | | Allomyrina Arrow, 1771 | | 1 | Species | Taxom | | Amphicoma Latreille, 1942 | | 1 | Species | Taxom | | Anomala Samouelle, 1913 | Shining Leaf Chafer | 12 | Species | Taxom | | Anthypna Eschscholtz, 1818 | | 1 | Species | Taxom | | Aphodius Hellwig, 1878 | | 2 | Species | Taxom | | Apogonia Melichar, 1875 | | 3 | Species | Taxom | | Ataenius Harold, 1858 | | 1 | Species | Taxom | | Blitopertha Reitter, 1903 | | 1 | Species | Taxom | | Bolbelasmus Boucomont, 1977 | | 1 | Species | Taxom | | Bolbochromus Boucomont, 1984 | | 1 | Species | Taxom | | Chaetadoretus Ohaus, 1914 | | 1 | Species | Taxom | | Cheirotonus Hope, 1984 | | 2 | Species | Taxom | | Clinterocera Motschulsky, 1973 | | 1 | Species | Taxom | | Coenochilus Burmeister &..., 1874 | | 1 | Species | Taxom | | Copris Geoffroy, 0 | Mexican dung beetle | 2 | Species | Taxom | | Cosmiomorpha Saunders, 1899 | | 1 | Species | Taxom | | Dasylepida Moser, 1927 | | 1 | Species | Taxom | | Dorcus MacLeay, 1835 | | 1 | Species | Taxom | | Eophileurus Arrow, 1835 | | 1 | Species | Taxom | | Eurytrachellelus Didier, 1898 | | 1 | Species | Taxom | | Exomala Reitter, 1925 | | 1 | Species | Taxom | | Figulus MacLeay, 1986 | | 2 | Species | Taxom | | Gametis Burmeister, 1835 | | 2 | Species | Taxom | | Heterosoma Schaum, 1901 | | 1 | Species | Taxom | | Holotrichia Hope, 1837 | | 1 | Species | Taxom | | Hoplia Illiger, 1959 | Monkey Beetles | 1 | Species | Taxom | | Labarrus Mulsant and Rey, 1860 | | 1 | Species | Taxom | | Leiopsammodius Rakovic, 1981 | | 1 | Species | Taxom | | Lepadoretus Reitter, 1855 | | 1 | Species | Taxom | | Macrodorcas Motschulsky, 1857 | | 3 | Species | Taxom | | Macrodorcus Motschulsky, 1964 | | 2 | Species | Taxom | | Maladera Mulsant, 1963 | | 9 | Species | Taxom | | Melolontha Fabricius, 1952 | Billy witch | 2 | Species | Taxom | | Mimela Kirby, 1965 | | 3 | Species | Taxom | | Neolucanus Thomson, 1864 | | 8 | Species | Taxom | | Nigidius MacLeay, 1905 | | 1 | Species | Taxom | | Nipponovalgus Sawada, 1941 | | 1 | Species | Taxom | | Odochilus Harold, 1971 | | 1 | Species | Taxom | | Odontolabis Hope, 1842 | | 2 | Species | Taxom | | Onthophagus Latreille, 1930 | | 3 | Species | Taxom | | Oryctes Hellwig, 1758 | Coconut rhinoceros beetles | 2 | Species | Taxom | | Oxycetonia Arrow, 1910 | | 1 | Species | Taxom | | Pachyserica Brenske, 1959 | | 1 | Species | Taxom | | Paraserica Reitter, 1866 | | 1 | Species | Taxom | | Parastasia Westwood, 1895 | | 2 | Species | Taxom | | Pedinotrichia | | 1 | Species | Taxom | | Pentodon Hope, 1902 | | 1 | Species | Taxom | | Phaeochrous Laporte de C..., 1961 | | 2 | Species | Taxom | | Platytomus Mulsant, 2006 | | 1 | Species | Taxom | | Polybathes Kirby, 1827 | | 1 | Species | Taxom | | Polyphylla Harris, 1890 | Lined June Beetles | 1 | Species | Taxom | | Popillia Leach, 1913 | | 4 | Species | Taxom | | Proagopertha Reitter, 1965 | | 1 | Species | Taxom | | Prosopocoilus Westwood, 1976 | | 7 | Species | Taxom | | Protaetia Burmeister, 1879 | | 9 | Species | Taxom | | Psalidoremus Motschoulsky, 1985 | | 2 | Species | Taxom | | Psammodius Fallen, 1973 | | 1 | Species | Taxom | | Rhomborhina Hope, 1964 | | 1 | Species | Taxom | | Rhyparus Westwood, 1905 | | 3 | Species | Taxom | | Serrognathus Motschulsky, 1985 | | 6 | Species | Taxom | | Smaragdesthes Kraatz, 1836 | | 1 | Species | Taxom | | Sophrops Fairmaire, 1989 | | 2 | Species | Taxom | | Subrinus Mulsant & Rey, 1870 | | 1 | Species | Taxom | | Trichiorhyssemus Clouët, 2001 | | 1 | Species | Taxom | | Tricholontha | | 1 | Species | Taxom | | Trox Fabricius, 1986 | Hide Beetles | 1 | Species | Taxom | | Trypoxylus Arrow, 1771 | | 1 | Species | Taxom | | Xylonychus Blanchard, 1924 | | 1 | Species | Taxom |
Species list for Ryukyu Islands
117 Species and 26 Subspecies | Adoretus (Adoretus) falciungulatus Nomura, 1965 | | | Aegus eschscholtzii eschscholtzii abei Ichikaw..., 1976 | | | Aegus eschscholtzii nakanei Ichikawa & I..., 1976 | | | Aegus (Aegus) laevicollis Saunders, 1854 | | | Aegus laevicollis laevicollis abei Ichikaw..., 1976 | | | Aegus laevicollis nakanei (Ichikawa & I..., 1976) | ⇒ Aegus eschscholtzi nakanei Ichikawa & I..., 1976 | | Aegus eschscholtzi nakanei Ichikawa & I..., 1976 | | | Aegus ishigakiensis Nomura, 1960 | | | Aegus nayani Schenk, 2008 | | | Aegus subnitidus (C. O. Waterh..., 1873) | ⇒ Aegus eschscholtzi subnitidus Waterhouse, 1872 | | Aganocrossus urostigma Harold, 1862 | | | Allomyrina dichotoma Linnaeus, 1771 | Japanese horned beetle | | Amphicoma splendens Yawata, 1942 | | | Anomala albopilosa Hope, 1839 | Green chafer | | Anomala albopilosa yashiroi Sawada, 1950 | | | Anomala cpustulata Matsumura, 1940 | | | Anomala cuprea (Hope, 1839) | ⇒ Anomala corruscans Chevrolat, 1841 | | Anomala esakii Sawada, 1950 | | | Anomala limbifera Ohaus, 1915 | | | Anomala miyakona Nomura, 1977 | | | Anomala niijimae Ohaus, 1925 | | | Anomala rufocuprea (Motschulsky, 1860) | ⇒ Anomala lucens Ballion, 1871 | | Anomala shirakii Nomura, 1959 | | | Anomala triangularis Schönfeldt, 1890 | | | Anomala xanthopleura Arrow, 1913 | | | Aphodius marginatus (LeConte, 1878) | ⇒ Cryptoscatomaseter depressiusculus Schmidt, 1907 | | Apogonia bicarinata Lewis, 1896 | | | Apogonia cupreoviridis Kolbe, 1886 | | | Apogonia major Waterhouse, 1875 | | | Ataenius australasiae Boheman, 1858 | | | Bolbelasmus nativus Krikken, 1977 | | | Bolbochromus ryukyuensis Masumoto, 1984 | | | Chaetadoretus formosanus Ohaus, 1914 | | | Cheirotonus jambar Kurosawa, 1984 | Yanbaru Long-armed Scarab Beetle | | Clinterocera ishikawai Kurosawa, 1973 | | | Coenochilus striatus Westwood, 1874 | | | Copris brachypterus Nomura, 1964 | | | Copris brachypterus brachypterus | | | Cosmiomorpha similis Fairmaire, 1899 | | | Dasylepida ishigakiensis Niijima & Ki..., 1927 | | | Dorcus titanus (Boisduval, 1835) | ⇒ Serrognathus typhon Boileau, 1905 | | Eophileurus chinensis Faldermann, 1835 | | | Eurytrachellelus costatus Boileau, 1898 | | | Exomala okiwaensis Ohaus, 1925 | | | Figulus (Figulus) punctatus (Waterhouse, 1873) | ⇒ Figulus daitoensis Fujita & Ich..., 1986 | | Figulus daitoensis Fujita & Ich..., 1986 | | | Gametis forticula Janson, 1881 | | | Gametis jucunda Faldermann, 1835 | Scarab | | Heterosoma exasperata Fairmaire, 1901 | | | Hoplia shirakii Nomura, 1959 | | | Labarrus sublimbatus Motschulsky, 1860 | | | Lepadoretus sinicus Burmeister, 1855 | | | Macrodorcas recta okinawana Nomura, 1964 | | | Macrodorcas recta recta Motschulsky, 1857 | | | Macrodorcus okinawanus Nomura, 1964 | | | Maladera amamiana Nomura, 1959 | | | Maladera imasakai Miyake & Yamaya, 1995 | | | Maladera nitidipes Moser, 1915 | | | Maladera okinawaensis Kobayashi, 1978 | | | Maladera opima Nomura, 1967 | | | Maladera oshimana Nomura, 1962 | | | Maladera shimogana Ahrens & Fab..., 2016 | | | Maladera yaeyamana Nomura, 1963 | | | Melolontha frater Arrow, 1913 | | | Melolontha masafumii Nomura, 1952 | | | Mimela ignicauda Bates, 1866 | | | Mimela sakishimana Nomura, 1973 | | | Mimela yonaguniensis Nomura, 1965 | | | Neolucanus insularis Miwa, 1929 | | | Neolucanus insulicola Y.Kurosawa, 1976 | | | Neolucanus insulicola insulicola hamaii Mizunuma, 1994 | | | Neolucanus insulicola okinawanus Sakaino, 1984 | | | Neolucanus protogenetivus Kurosawa, 1976 | | | Neolucanus protogenetivus okinawanus Sakaino, 1984 | | | Neolucanus saundersi Parry, 1864 | | | Nigidius lewisi Boileau, 1905 | | | Nipponovalgus yonakuniensis (Sawada, 1941) | ⇒ Dasyvalgus yonakuniensis Sawada, 1941 | | Odochilus convexus Nomura, 1971 | | | Odontolabis cuvera (Hope, 1842) | ⇒ Odontolabis delessertii Guérin-Ménev..., 1843 | | Onthophagus (Gibbonthophagus) viduus (Harold, 1875) | ⇒ Onthophagus orbus Boucomont, 1924 | | Onthophagus (Indachorius) suginoi Ochi, 1984 | | | Onthophagus malthinus Gillet, 1930 | | | Oryctes hisamatsui Nagai, 2002 | | | Oryctes rhinoceros Linnaeus, 1758 | Asiatic rhinoceros beetle | | Pachyserica yanoi Nomura, 1959 | | | Paraserica grisea Motschulsky, 1866 | | | Parastasia ferrieri Nonfried, 1895 | | | Pedinotrichia parallela Motschulsky, 1854 | | | Pentodon idiota memnon Jakovlev, 1902 | | | Phaeochrous emarginatus Laporte, 1840 | | | Phaeochrous tokaraensis Nomura, 1961 | | | Platytomus yadai Ochi, Kawaha..., 2006 | | | Polyphylla schoenfeldti Brenske, 1890 | | | Popillia comma Niijima & Ma..., 1923 | | | Popillia dilutipennis Fairmaire, 1888 | | | Popillia insularis Lewis, 1895 | | | Popillia lewisi Arrow, 1913 | | | Proagopertha ohbayashii Nomura, 1965 | | | Prosopocoilus (Cyclotropus) occipitalis Hope & Westwood, 1845 | | | Prosopocoilus occipitalis occipitalis Anoa Kriesche, 1920 | | | Prosopocoilus (Psalidoremus) dissimilis Boileau, 1898 | | | Prosopocoilus dissimilis dissimilis dissimilis B..., 1898 | | | Prosopocoilus dissimilis okinawanus Nomura, 1962 | | | Prosopocoilus (Psalidoremus) motschulskii Waterhouse, 1869 | | | Prosopocoilus pseudodissimilis Y. Kurosawa, 1976 | | | Protaetia lewisi lewisi | | | Protaetia (Calopotosia) orientalis Gory & Perch..., 1833 | Asian flower beetle | | Protaetia (Heteroprotaetia) fusca Herbst, 1790 | Asian Mango Flower Beetle | | Protaetia (Liocola) formosana Moser, 1910 | | | Protaetia (Liocola) miyakoensis Niijima & Ki..., 1923 | | | Protaetia (Pseudocalopotosia) ishigakia Kurosawa, 1959 | | | Protaetia (Pyropotosia) pryeri Janson, 1888 | Midway emerald beetle | | Protaetia (Tomentoprotaetia) culta Waterhouse, 1879 | | | Psalidoremus dissimilis Boileau, 1898 | | | Psalidoremus dissimilis kumejimaensis Ichikawa & F..., 1985 | | | Psammodius kobayashii Nomura, 1973 | | | Rhomborhina hamai Nomura, 1964 | | | Rhyparus azumai Nakane, 1956 | | | Rhyparus helophoroides Fairmaire, 1893 | | | Rhyparus peninsularis Arrow, 1905 | | | Serrognathus titanus daitoensis Fujita & Ich..., 1986 | | | Serrognathus titanus elegans Boileau, 1899 | | | Serrognathus titanus okinawanus Kriesche, 1922 | | | Serrognathus titanus okinoerabuensis Fujita & Ich..., 1985 | | | Serrognathus titanus takaraensis Fujita & Ich..., 1985 | | | Smaragdesthes africana smaragdina Chevrolat, 1836 | | | Sophrops kawadai okinawaensis Nomura, 1977 | | | Sophrops takatoshii Itoh, 1989 | | | Subrinus sturmi Harold, 1870 | | | Trichiorhyssemus kitayamai Ochi & Kawah..., 2001 | | | Tricholontha papagena Nomura, 1952 | | | Trox (Trox) aphanocephalus (Scholtz, 1986) | ⇒ Omorgus aphanocephalus Scholtz, 1986 | | Trypoxylus (Allomyrina) dichotomus (Linnaeus, 1771) | ⇒ Allomyrina dichotoma Linnaeus, 1771 | | Xylonychus eucalypti pentaphyllus Arrow, 1924 | |
Subnodes of Scarabaeoidea with occurrences in Ryukyu Islands | Family | Geotrupidae
Latreille, 1802
Dor beetles, Dung beetles, Earth-Boring Dung Beetle, Earth-boring dung beetles, Earth-boring scarab beetles | 2
2 | Genera
Species |
Taxom
| | Family | Glaphyridae
MacLeay, 1819
bumble bee scarab beetle, Bumble bee scarab beetles, The bumble bee scarab beetles | 2
1 | Genera
Species |
Taxom
| | Family | Hybosoridae
Erichson, 1847
Pill scarab beetles, scavenger scarab beetle, Scavenger scarab beetles, Scavenging scarab beetles | 1
2 | Genus
Species |
Taxom
| | Family | Lucanidae
Latreille, 1804
Stag Beetle, Stag beetles | 12
24 | Genera
Species |
Taxom
| | Family | Melolonthidae
Leach, 1817
Chafers, June beetles, May beetles | 1
1 | Genus
Species |
Taxom
| | Family | Rutelidae
MacLeay, 1819
| 1
1 | Genus
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Monkey beetles, Rain beetles, Scarab Beetle, Scarab beetles, Scarabs, Tumble bugs | 51
85 | Genera
Species |
Taxom
| | Family | Trogidae
MacLeay, 1819
Gnawer beetles, Hide, hide beetle, Hide beetles, Keratin Beetles, Skin and hide beetles, Skin beetles | 1
1 | Genus
Species |
Taxom
|
Links, sources and further information
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