Species list, Checklist of Infraorder Staphyliniformia for Middle Pennsylvanian (Upper Carboniferous)

Ants	Arachnids
Bees	Beetles
Booklice	Bugs
Bumblebees	Butterflies
Cicadas	Crane flies
Dragonflies	Earwigs
Flies	Isopods
Locusts	Mantises
Mayflies	Mosquitoes
Myriapods	Net-winged
Plant-parasitic	Scorpion flies
Termites	Wasps

Taxonomy & Distribution
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INSECTOIDGERENGDARK LIGHT

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Checklist for Infraorder Staphyliniformia Lameere, 1900
and region Middle Pennsylvanian (Upper Carboniferous)

Class	Insecta Linnaeus, 1758 Insects, True insects	168 324 534	Families Genera Species	Taxom
Subclass	Pterygota Lang, 1888 Winged insects	166 322 533	Families Genera Species	Taxom
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	133 247 432	Families Genera Species	Taxom
Superorder	Holometabola Heider, 1889 Holometabolous Insects	49 54 68	Families Genera Species	Taxom
Order	Coleoptera Linnaeus, 1758 Beetles, Coleopterans	22 25 29	Families Genera Species	Taxom
Suborder	Polyphaga Emery, 1886 Polyphags	21 24 28	Families Genera Species	Taxom
Infraorder	Staphyliniformia Lameere, 1900	4 4 5	Families Genera Species	Taxom

List of families for Middle Pennsylvanian
4 Families width 5 Species

Leiodidae Fleming, 1821	Round fungus beetles	1	Species	Taxom
Staphylinidae Lameere, 1900	Rove beetles	1	Species	Taxom
Hydrophilidae Latreille, 1802	Silver Water Beetles	2	Species	Taxom
Ptiliidae Heer, 1843	Feather-wing beetles	1	Species	Taxom

Genera for Middle Pennsylvanian
4 Genera with 5 Species

Agogoblattina Handlirsch, 1885	1	Species	Taxom
Gyrophlebia Handlirsch, 1906	1	Species	Taxom
Megathentomum Scudder, 1906	2	Species	Taxom
Ptenodera Bolton, 1922	1	Species	Taxom

Species list for Middle Pennsylvanian
5 Species

Agogoblattina occidua ♰ S. H. Scudder, 1885
Gyrophlebia longicollis A. Handlirsch, 1906
Megathentomum pustulatum ♰ S. H. Scudder, 1868
Megathentomum scudderi Handlirsch, 1906
Ptenodera dubius ♰ H. Bolton, 1922

Subnodes of Staphyliniformia with occurrences in Middle Pennsylvanian

Superfamily	Hydrophiloidea Latreille, 1802 Histerids, Hydriphiloids, Hydrophilids, Water Scavenger and Clown Beetles	1 1 2	Family Genus Species	Taxom
Superfamily	Staphylinoidea Latreille, 1802 Carrion Beetles, Fungus Beetles, Rove Beetles, Staphylinid Beetles	3 3 3	Families Genera Species	Taxom

Links, sources and further information

Staphyliniformia at itis.gov