Species list, Checklist of Subsection Calyptratae (Calyptrate muscoids) for Coto Brus (Brunca Region, Costa Rica)

Ants	Arachnids
Bees	Beetles
Booklice	Bugs
Bumblebees	Butterflies
Cicadas	Crane flies
Dragonflies	Earwigs
Flies	Isopods
Locusts	Mantises
Mayflies	Mosquitoes
Myriapods	Net-winged
Plant-parasitic	Scorpion flies
Termites	Wasps

Taxonomy & Distribution
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INSECTOIDGERENGDARK LIGHT

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Checklist for Subsection Calyptratae Robineau-Desvoidy, 1830 (Calyptrate muscoids)
and region Coto Brus (Brunca Region, Costa Rica)

Class	Insecta Linnaeus, 1758 Insects, True insects	300 3210 3826	Families Genera Species	Taxom
Subclass	Pterygota Lang, 1888 Winged insects	300 3210 3826	Families Genera Species	Taxom
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	292 3198 3811	Families Genera Species	Taxom
Superorder	Holometabola Heider, 1889 Holometabolous Insects	226 2911 3518	Families Genera Species	Taxom
Order	Diptera Linnaeus, 1758 Flies, Gnats, Hoverflies, Mosquitoes, True flies	62 525 381	Families Genera Species	Taxom
Suborder	Brachycera Macquart, 1834 Circular-seamed flies, Muscoid flies, Short-horned flies	50 481 334	Families Genera Species	Taxom
Infraorder	Muscomorpha Woodley, 1989	38 385 265	Families Genera Species	Taxom
Section	Schizophora Becher, 1882	35 302 117	Families Genera Species	Taxom
Subsection	Calyptratae Robineau-Desvoidy, 1830 Calyptrate muscoids	7 191 59	Families Genera Species	Taxom

List of families for Coto Brus
7 Families width 59 Species

Tachinidae Robineau-Des..., 1830	Bristle Flies	32	Species	Taxom
Sarcophagidae Macquart, 1834	Flesh flies	6	Species	Taxom
Calliphoridae Brauer & Ber..., 1889	Blow flies	12	Species	Taxom
Muscidae Latreille, 1802	Face flies	6	Species	Taxom
Richardiidae Snow, 1896	Picture-winged flies	2	Species	Taxom
Oestridae Leach, 1815	Bot flies	0	Species	Taxom
Hippoboscidae Samouelle, 1819	Keds	1	Species	Taxom

Genera for Coto Brus
191 Genera with 59 Species

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Acaulona van der Wulp, 1884		0	Species	Taxom
Acemya Robineau-Des..., 1849		1	Species	Taxom
Actinochaeta Brauer & von..., 1889		0	Species	Taxom
Actinodoria Townsend, 1927		0	Species	Taxom
Adejeania Townsend, 1913		0	Species	Taxom
Admontia Brauer & von..., 1889		0	Species	Taxom
Anepalpus Townsend, 1931		0	Species	Taxom
Angiometopa Brauer & Ber..., 1916		1	Species	Taxom
Angioneura Brauer & Ber..., 1859		1	Species	Taxom
Anisia van der Wulp, 1890		0	Species	Taxom
Atactosturmia Townsend, 1915		0	Species	Taxom
Athiplanomyia Séguy, 1937		1	Species	Taxom
Austrophorocera Townsend, 1916		0	Species	Taxom
Avibrissosturmia Townsend, 1927		0	Species	Taxom
Belvosia Robineau-Des..., 1830		0	Species	Taxom
Billaea Robineau-Des..., 1830		0	Species	Taxom
Bithoracochaeta Stein, 1911		0	Species	Taxom
Blaesoxipha Loew, 1861	Grasshopper Flesh Flies	0	Species	Taxom
Blepharipa Rondani, 0		1	Species	Taxom
Blondelia Robineau-Des..., 1863		0	Species	Taxom
Boettcheria Parker, 1914		0	Species	Taxom
Bombyliomyia Brauer and B..., 1889		0	Species	Taxom
Calliphora Robineau-Des..., 1830	Bluebottle	1	Species	Taxom
Calodexia van der Wulp, 1915		5	Species	Taxom
Calolydella Townsend, 1927		0	Species	Taxom
Campylocheta Rondani, 1859		0	Species	Taxom
Carcelia Robineau-Des..., 1830		0	Species	Taxom
Celatoria Coquillett, 1890		0	Species	Taxom
Chaetodoria Townsend, 1927		1	Species	Taxom
Chaetogaedia Brauer & von..., 1891		0	Species	Taxom
Chaetona van der Wulp, 1891		0	Species	Taxom
Chaetonodexodes Townsend, 1916		0	Species	Taxom
Chaetonopsis Townsend, 1915		0	Species	Taxom
Chaetostigmoptera Townsend, 1916		0	Species	Taxom
Chlorohystricia Townsend, 1927		0	Species	Taxom
Choeteprosopa Macquart, 1891		1	Species	Taxom
Cholomyia Bigot, 1884		0	Species	Taxom
Chrysoexorista Townsend, 1915		0	Species	Taxom
Chrysotachina Brauer & Ber..., 1891		0	Species	Taxom
Clythoxynops Walton, 1914		0	Species	Taxom
Cochliomyia Townsend, 1775	New World screwworm fly	1	Species	Taxom
Coenosia Meigen, 1940	Tiger Flies	1	Species	Taxom
Coilometopia Macquart, 1911		1	Species	Taxom
Compsomyiops Townsend, 1849		1	Species	Taxom
Copecrypta Walker, 1908		0	Species	Taxom
Crocinosoma Reinhard, 1947		0	Species	Taxom
Cryptomeigenia Brauer & von..., 1891		0	Species	Taxom
Cryptopalpus Rondani, 1850		0	Species	Taxom
Cuterebra Clark, 1815	Bot flies	0	Species	Taxom
Cyacyrtoneura Townsend, 1931		0	Species	Taxom
Cyrtoneuropsis Malloch, 1925		0	Species	Taxom
Cyrtophleba Rondani, 1856		0	Species	Taxom
Dasymorellia Malloch, 1923		0	Species	Taxom
Deopalpus Walker, 1908		0	Species	Taxom
Dexosarcophaga Townsend, 1917		0	Species	Taxom
Drino Robineau-Des..., 1836		1	Species	Taxom
Ebenia Macquart, 1846		0	Species	Taxom
Emdenimyia de Souza Lopes, 1946		0	Species	Taxom
Empheremyia Bischof, 1904		0	Species	Taxom
Epalpus Rondani, 1849		1	Species	Taxom
Eribella Mesnil, 1960		0	Species	Taxom
Erythromelana Townsend, 1919		0	Species	Taxom
Euantha van der Wulp, 1885		0	Species	Taxom
Eucelatoria Townsend, 1909		0	Species	Taxom
Euclytia Townsend, 1908		0	Species	Taxom
Eudejeania Townsend, 1912		0	Species	Taxom
Eudexia Brauer & von..., 1889		0	Species	Taxom
Euepalpus Townsend, 1908		1	Species	Taxom
Euhalidaya Walton, 1914		0	Species	Taxom
Eujuriniodes Townsend, 1935		0	Species	Taxom
Eulasiona Townsend, 1892		0	Species	Taxom
Eulasiopalpus Townsend, 1913		0	Species	Taxom
Eulobomyia Woodley & Ar..., 2008		1	Species	Taxom
Euthelyconychia Townsend, 1927		0	Species	Taxom
Eutrixa Coquillett, 1897		0	Species	Taxom
Exorista Meigen, 0		1	Species	Taxom
Gaediopsis Brauer & von..., 1891		0	Species	Taxom
Genea Vittad., 1831		0	Species	Taxom
Gnadochaeta Macquart, 1851		0	Species	Taxom
Graphomya Robineau-Des..., 1934		2	Species	Taxom
Gymnomma van der Wulp, 1888		1	Species	Taxom
Haematobia Lepeletier &..., 1758		1	Species	Taxom
Helicobia Coquillett, 1895		0	Species	Taxom
Hemilucilia Brauer, 1850		2	Species	Taxom
Hemisturmia Townsend, 1927		0	Species	Taxom
Henriquella Bonatto & Ma..., 1925		1	Species	Taxom
Homosaundersia Townsend, 1931		0	Species	Taxom
Houghia Coquillett, 1897		0	Species	Taxom
Hydrotaea Robineau-Des..., 1830		0	Species	Taxom
Hyphantrophaga Townsend, 1892		0	Species	Taxom
Hypotachina Robineau-Des..., 1830		0	Species	Taxom
Hystrichodexia Röder, 1886		0	Species	Taxom
Hystricia Macquart, 1942		2	Species	Taxom
Ichneumonops Townsend, 1908		0	Species	Taxom
Ischyrophaga Townsend, 1915		0	Species	Taxom
Italispidea Townsend, 1927		0	Species	Taxom
Jurinia Robineau-Des..., 1830		0	Species	Taxom
Juriniopsis Townsend, 1916		0	Species	Taxom
Lepidodexia Brauer & Ber..., 1989		1	Species	Taxom
Leschenaultia Robineau-Des..., 1830		1	Species	Taxom
Leskia Robineau-Des..., 1830		0	Species	Taxom
Lespesia Robineau-Des..., 1863		0	Species	Taxom
Limnophora Robineau-Des..., 1830		0	Species	Taxom
Linnaemya Robineau-Des..., 1830		0	Species	Taxom
Lixophaga Townsend, 1908		0	Species	Taxom
Lucilia Robineau-Des..., 1836	Greenbottle Flies	2	Species	Taxom
Lydina Robineau-Des..., 1830		0	Species	Taxom
Lydinolydella Townsend, 1927		0	Species	Taxom
Lypha Robineau-Des..., 1830		0	Species	Taxom
Macromya Robineau-Des..., 1830		0	Species	Taxom
Masiphya Brauer & Ber..., 1891		0	Species	Taxom
Mauromyia Coquillett, 1897		0	Species	Taxom
Mesembrinella Giglio-Tos, 1922		2	Species	Taxom
Minthoplagia Townsend, 1890		1	Species	Taxom
Morellia Robineau-Des..., 1830		0	Species	Taxom
Mydaea Robineau-Des..., 1835		1	Species	Taxom
Myiopharus Brauer & Ber..., 1889		1	Species	Taxom
Myospila Rondani, 1856		0	Species	Taxom
Myothyriopsis Townsend, 1919		0	Species	Taxom
Mystacella Reinhard, 1967		0	Species	Taxom
Nemorilla Rondani, 1856		0	Species	Taxom
Neobrachelia Townsend, 1931		0	Species	Taxom
Neodexiopsis Malloch, 1920		0	Species	Taxom
Neomintho Brauer & Ber..., 1891		0	Species	Taxom
Neomuscina Townsend, 1919		0	Species	Taxom
Neosolieria Townsend, 1927		0	Species	Taxom
Nephochaetopteryx Townsend, 1934		0	Species	Taxom
Notochaeta Aldrich, 1916		0	Species	Taxom
Oebalia Robineau-Des..., 1989		1	Species	Taxom
Oestrophasia Brauer & Ber..., 1889		0	Species	Taxom
Opsidia Coquillett, 1926		1	Species	Taxom
Ormia Robineau-Des..., 1830		0	Species	Taxom
Ornithoctona Speiser, 1817		1	Species	Taxom
Oxynops Townsend, 1912		0	Species	Taxom
Oxysarcodexia Townsend, 1917		0	Species	Taxom
Oxyvinia Dodge, 1966		0	Species	Taxom
Panzeria Robineau-Des..., 1830		0	Species	Taxom
Paracraspedothrix Villeneuve, 1919		0	Species	Taxom
Paradejeania Brauer & von..., 1867	Spiny Tachina Fly	1	Species	Taxom
Paradidyma Brauer & Ber..., 1891		0	Species	Taxom
Paralucilia Brauer & Ber..., 1877		1	Species	Taxom
Parepalpus Coquillett, 1902		0	Species	Taxom
Patelloa Townsend, 1916		0	Species	Taxom
Peckia Robineau-Des..., 1830		0	Species	Taxom
Phaonia Robineau-Des..., 1830		0	Species	Taxom
Phasia Robineau-Des..., 1830		0	Species	Taxom
Phryxe Robineau-Des..., 1810		1	Species	Taxom
Phyllophilopsis Townsend, 1915		0	Species	Taxom
Phytomyptera Rondani, 1844		0	Species	Taxom
Platyphasia Skuse, 1890		0	Species	Taxom
Polietina Schnabl & Dz..., 1911		0	Species	Taxom
Polistiopsis Townsend, 1915		0	Species	Taxom
Prorhynchops Brauer & von..., 1891		0	Species	Taxom
Prosenoides Brauer & von..., 1891		0	Species	Taxom
Protodejeania Townsend, 1915		0	Species	Taxom
Pseudobombyliomyia Townsend, 1931		1	Species	Taxom
Pseudodexia Brauer & von..., 1830		1	Species	Taxom
Pseudoptilolepis Snyder, 1949		0	Species	Taxom
Ptilodexia Brauer & von..., 1889		0	Species	Taxom
Quadratosoma Townsend, 1914		0	Species	Taxom
Ravinia Robineau-Des..., 1968		1	Species	Taxom
Rhachoepalpus Townsend, 1908		1	Species	Taxom
Sarcophaga Meigen, 1908	Common Flesh Flies	1	Species	Taxom
Scotiptera Macquart, 1835		0	Species	Taxom
Setellida Hendel, 1911		1	Species	Taxom
Siphosturmia Coquillett, 1897		0	Species	Taxom
Sitellitergus Reinhard, 1964		0	Species	Taxom
Spathidexia Townsend, 1912		0	Species	Taxom
Sphaerina Wulp, 1890		0	Species	Taxom
Spilogona Schnabl, 1911		0	Species	Taxom
Strongygaster Macquart, 1834		0	Species	Taxom
Telothyria van der Wulp, 1890		0	Species	Taxom
Thelairodes van der Wulp, 1889		1	Species	Taxom
Thelairodoria Townsend, 1927		0	Species	Taxom
Thelyoxynops Townsend, 1927		0	Species	Taxom
Trafoia Brauer & von..., 1893		0	Species	Taxom
Tricharaea Thomson, 1869		0	Species	Taxom
Trichodura Macquart, 1843		0	Species	Taxom
Trichopyrrhosia Townsend, 1927		0	Species	Taxom
Triodontopyga Townsend, 1927		0	Species	Taxom
Tropidopsiomorpha Townsend, 1927		0	Species	Taxom
Uramya Robineau-Des..., 1830		0	Species	Taxom
Vanderwulpia Townsend, 1891		0	Species	Taxom
Voria Robineau-Des..., 1830		0	Species	Taxom
Winthemia Robineau-Des..., 1890		4	Species	Taxom
Xanthodexia van der Wulp, 1891		0	Species	Taxom
Xanthoepalpus Townsend, 1914		0	Species	Taxom
Xanthophyto Townsend, 1916		0	Species	Taxom
Xanthozona Hampson, 1830		1	Species	Taxom
Xenophasia Townsend, 1934		1	Species	Taxom
Zizyphomyia Townsend, 1916		0	Species	Taxom

Species list for Coto Brus
59 Species

Acemya masurius Walker, 1849
Angiometopa ravinia Parker, 1916
Angioneura cyrtoneurina Zetterstedt, 1859	Dark Least Blowfly
Athiplanomyia mydaea (Seguy, 1937)	⇒ Ochromusca trifaria Bigot, 1878
Blepharipa albicauda
Calliphora peruviana (Robineau-Des..., 1830)	⇒ Neta chilensis Walker, 1836
Calodexia flavipes Bigot, 1889
Calodexia fumosa Townsend, 1917
Calodexia major Curran, 1934
Calodexia majuscula Wulp, 1891
Calodexia similis Townsend, 1915
Chaetodoria conica Townsend, 1927
Choeteprosopa hedemanni Brauer & Ber..., 1891
Cochliomyia macellaria Fabricius, 1775	Secondary screw worm fly
Coenosia hippelates Emden, 1940
Coilometopia longicornis Hendel, 1911
Compsomyiops verena Walker, 1849
Drino piceiventris Walker, 1836
Epalpus alligans Walker, 1849
Euepalpus flavicauda Townsend, 1908
Eulobomyia neotropica Woodley & Ar..., 2008
Exorista heterusia
Graphomya amazonensis Couri, 2005
Graphomya auriceps Malloch, 1934
Gymnomma nitidiventris Wulp, 1888
Haematobia irritans Linnaeus, 1758	Buffalo fly
Hemilucilia segmentaria Fabricius, 1805
Hemilucilia semidiaphana (Rondani, 1850)	⇒ Hemilucilia segmentaria Fabricius, 1805
Henriquella spicata Aldrich, 1925
Hystricia amoena Macquart, 1843
Hystricia laxa Curran, 1942
Lepidodexia woodorum Pape, 1989
Leschenaultia leucophrys Wiedemann, 1830
Lucilia eximia (Wiedemann, 1819)	⇒ Neomyia lauta Wiedemann, 1830
Lucilia purpurescens Walker, 1836
Mesembrinella pictipennis Aldrich, 1922
Mesembrinella umbrosa Aldrich, 1922
Minthoplagia setifrons Wulp, 1890
Mydaea orthonevra Macquart, 1835
Myiopharus metopia Brauer & Ber..., 1889	Cadelle
Oebalia costarica Pape, 1989
Opsidia metopioides Allen, 1926
Ornithoctona erythrocephala Leach, 1817
Paradejeania rutilioides Jaennicke, 1867	Spiny Tachina Fly
Paralucilia fulvinota Bigot, 1877
Phryxe vulgaris Fallen, 1810
Pseudobombyliomyia linellii Townsend, 1931
Pseudodexia eques Wiedemann, 1830
Ravinia postnoda Dodge, 1968
Rhachoepalpus olivaceus Townsend, 1908	Curtiss' milkwort
Sarcophaga (Heteronychia) metopina Villeneuve, 1908
Setellida caerulescens Hendel, 1911
Thelairodes vittigerus Bigot, 1889
Winthemia peruviana Townsend, 1928
Winthemia pinguis Fabricius, 1805
Winthemia tessellata Wulp, 1890
Winthemia tricolor Wulp, 1890
Xanthozona melanopyga Wiedemann, 1830
Xenophasia xanthomelanodes Townsend, 1934

Subnodes of Calyptratae with occurrences in Coto Brus

Superfamily	Hippoboscoidea Samouelle, 1819	1 1 1	Family Genus Species	Taxom
Superfamily	Muscoidea Latreille, 1802	2 21 8	Families Genera Species	Taxom
Superfamily	Oestroidea Leach, 1815	4 169 50	Families Genera Species	Taxom