AntsAntsArachnidsArachnids
BeesBeesBeetlesBeetles
BookliceBookliceBugsBugs
BumblebeesBumblebeesButterfliesButterflies
CicadasCicadasCrane fliesCrane flies
DragonfliesDragonfliesEarwigsEarwigs
FliesFliesIsopodsIsopods
LocustsLocustsMantisesMantises
MayfliesMayfliesMosquitoesMosquitoes
MyriapodsMyriapodsNet-wingedNet-winged
Plant-parasiticPlant-parasiticScorpion fliesScorpion flies
TermitesTermitesWaspsWasps
INSECTOIDGERENGDARK LIGHT
Home Arthropoda Imprint Contact
TAXFAMGENSPESUB
⌕
Checklist for Superfamily Coccoidea Handlirsch, 1903 (Coccids)
and region Puerto Rico (Greater Antilles)


Class
 		Insecta
Linnaeus, 1758
Insects, True insects		352
2671
5038		Families
Genera
Species		 
Taxom
 

Subclass
 		Pterygota
Lang, 1888
Winged insects		351
2670
5037		Families
Genera
Species		 
Taxom
 

Infraclass
 		Neoptera
van der Wulp, 1890
Modern Wing-folding insects		343
2630
4959		Families
Genera
Species		 
Taxom
 

Superorder
 		Paraneoptera
Martynov, 1923
86
537
1084		Families
Genera
Species		 
Taxom
 

Order
 		Hemiptera
Linnaeus, 1758
Aphids, Bugs, Cicadas, Hemipterans, Hoppers, True bugs		69
490
1016		Families
Genera
Species		 
Taxom
 

Suborder
 		Sternorrhyncha
Amyot & Serville, 1843
Aphids, Coccids, Jumping Plant Lice, Mealybugs, Psyllids		20
128
240		Families
Genera
Species		 
Taxom
 

Superfamily
 		Coccoidea
Handlirsch, 1903
Coccids, Mealy Bugs, Mealybugs, Scale Insects, Scales		13
89
171		Families
Genera
Species		 
Taxom
 


List of families for Puerto Rico
13 Families width 171 Species and 2 Subspecies
Diaspididae Maskell, 1878Armored scale insects90SpeciesTaxom
Aclerdidae Ferris, 1937Aclerdid scales2SpeciesTaxom
Pseudococcidae Heymons, 1915Mealy Bugs30SpeciesTaxom
Asterolecaniidae Cockerell, 1896Asterolecaniids3SpeciesTaxom
Coccidae Stephens, 1829Soft scale33SpeciesTaxom
Cerococcidae Balachowsky, 1942Cerococcids1SpeciesTaxom
Conchaspididae Green, 1896False armored scales1SpeciesTaxom
Monophlebidae Signoret, 1875Giant scale3SpeciesTaxom
Eriococcidae Cockerell, 1899Eriococcid scales1SpeciesTaxom
Ortheziidae Green, 1896Ensign coccids2SpeciesTaxom
Lecanodiaspididae Borchsenius, 1965False pit scales2SpeciesTaxom
Putoidae Beardsley, 1969Giant mealybugs1SpeciesTaxom
Rhizoecidae Williams, 19692SpeciesTaxom


Genera for Puerto Rico
89 Genera with 171 Species
#ABCDEFGHIKLMNOPRSTUV
Abgrallaspis Balachowsky, 18691SpeciesTaxom
Aclerda Signoret, 19322SpeciesTaxom
Acutaspis Ferris, 19632SpeciesTaxom
Antonina Signoret, 18971SpeciesTaxom
Antoninoides Ferris, 19031SpeciesTaxom
Aonidiella Berlese and ..., 19066SpeciesTaxom
Aonidomytilus Leonardi, 18931SpeciesTaxom
Aspidaspis Ferris, 19091SpeciesTaxom
Aspidiella Leonardi, 18932SpeciesTaxom
Aspidiotus Bouche, 18333SpeciesTaxom
Asterolecanium Targioni-Toz..., 19412SpeciesTaxom
Aulacaspis Cockerell, 19774SpeciesTaxom
Bambusaspis Cockerell, 18691SpeciesTaxom
Brevennia Goux, 19431SpeciesTaxom
Cardiococcus Cockerell, 19031SpeciesTaxom
Cerococcus Comstock, 19661SpeciesTaxom
Ceroplastes Gray, 1758Wax Scales10SpeciesTaxom
Chaetococcus Maskell, 18931SpeciesTaxom
Chorizococcus Mckenzie, 19921SpeciesTaxom
Chrysomphalus Ashmead, 18892SpeciesTaxom
Clavaspis Macgillivray, 19091SpeciesTaxom
Coccus Linnaeus, 18893SpeciesTaxom
Conchaspis Cockerell, 18931SpeciesTaxom
Crenulaspidiotus Macgillivray, 19102SpeciesTaxom
Crypticerya Cockerell, 18901SpeciesTaxom
Cryptostigma Ferris, 19202SpeciesTaxom
Dactylaspis Ferris, 18931SpeciesTaxom
Diaspidiotus Berlese, 18961SpeciesTaxom
Diaspis Costa, 18333SpeciesTaxom
Distichlicoccus Ferris, 19921SpeciesTaxom
Duplaspidiotus Macgillivray, 18992SpeciesTaxom
Dysmicoccus Ferris, 19006SpeciesTaxom
Eriococcus Targioni Toz..., 18791SpeciesTaxom
Eucalymnatus Cockerell, 18731SpeciesTaxom
Ferrisia Fullaway, 18933SpeciesTaxom
Fiorinia Targioni-Toz..., 18671SpeciesTaxom
Froggattiella Leonardi, 19051SpeciesTaxom
Furcaspis Lindinger, 18931SpeciesTaxom
Genaparlatoria McGillivray, 19051SpeciesTaxom
Geococcus Green, 19331SpeciesTaxom
Gymnaspis Newstead, 18981SpeciesTaxom
Haliaspis Takagi, 19782SpeciesTaxom
Hemiberlesia Leonardi, 18814SpeciesTaxom
Hemigymnaspis Lindinger, 19341SpeciesTaxom
Howardia Berlese and ..., 18831SpeciesTaxom
Icerya Signoret, 1878cottony cushion scales2SpeciesTaxom
Inglisia Maskell, 18941SpeciesTaxom
Insignorthezia Kozár, 18871SpeciesTaxom
Ischnaspis Douglas, 18821SpeciesTaxom
Kilifia De Lotto, 18731SpeciesTaxom
Kuwanaspis Macgillivray, 19221SpeciesTaxom
Lecanodiaspis Targioni Toz..., 19721SpeciesTaxom
Lepidosaphes Shimer, 19378SpeciesTaxom
Lopholeucaspis Balachowsky, 18991SpeciesTaxom
Maconellicoccus Ezzat, 19081SpeciesTaxom
Melanaspis Cockerell, 19445SpeciesTaxom
Milviscutulus Williams & W..., 18891SpeciesTaxom
Morganella Cockerell, 18891SpeciesTaxom
Mycetaspis Cockerell, 18831SpeciesTaxom
Nipaecoccus Sulc, 18931SpeciesTaxom
Oceanaspidiotus Takagi, 18832SpeciesTaxom
Odonaspis Leonardi, 18972SpeciesTaxom
Palinaspis Ferris, 19231SpeciesTaxom
Parasaissetia Takahashi, 18611SpeciesTaxom
Parlatoria Targioni-Toz..., 18537SpeciesTaxom
Phenacoccus Cockerell, 18985SpeciesTaxom
Pinnaspis Cockerell, 18994SpeciesTaxom
Planococcus Ferris, 19052SpeciesTaxom
Praelongorthezia Kozár, 18911SpeciesTaxom
Protopulvinaria Cockerell, 18942SpeciesTaxom
Pseudaonidia Cockerell, 18961SpeciesTaxom
Pseudaulacaspis Macgillivray, 19613SpeciesTaxom
Pseudischnaspis Macgillivray, 18931SpeciesTaxom
Pseudococcus Westwood, 1996Spike Primrose3SpeciesTaxom
Pseudokermes Cockerell, 18941SpeciesTaxom
Pseudoparlatoria Cockerell, 18921SpeciesTaxom
Pulvinaria Targioni-Toz..., 1893Cottony Scales5SpeciesTaxom
Puto Signoret, 1950Giant mealybugs1SpeciesTaxom
Radionaspis Ferris, 19081SpeciesTaxom
Rhizoecus Künckel, 19712SpeciesTaxom
Rutherfordia Macgillivray, 18941SpeciesTaxom
Saccharicoccus Ferris, 18951SpeciesTaxom
Saissetia Deplanche, 19693SpeciesTaxom
Selenaspidus Cockerell, 18891SpeciesTaxom
Trionymus Berg, 19261SpeciesTaxom
Unaspis Macgillivray, 18831SpeciesTaxom
Vinculaspis Ferris, 19411SpeciesTaxom
Vinsonia Signoret, 18711SpeciesTaxom
Virgulicoccus Lambdin & Ko..., 19111SpeciesTaxom


Species list for Puerto Rico
171 Species and 2 Subspecies
Abgrallaspis cyanophylli Signoret, 1869Armored scale
Aclerda sacchari (Teague, 1925)⇒ Aclerda takahashii Kuwana, 1932
Aclerda takahashii Kuwana, 1932Flat grass scale
Acutaspis aliena Newstead, 1901Alien Scale
Acutaspis morrisonorum Kosztarab, 1963Round conifer scale
Antonina graminis (Maskell, 1897)⇒ Antoninoides parrotti Cockerell in..., 1903
Antoninoides parrotti Cockerell in..., 1903Felted grass coccid
Aonidiella aurantii (Maskell, 1879)⇒ Aonidiella citrina Coquillett, 1891
Aonidiella citrina Coquillett, 1891California red scale
Aonidiella comperei Mckenzie, 1937Compere scale
Aonidiella inornata Mckenzie, 1938Inornate scale
Aonidiella orientalis Newstead, 1894Oriental scale
Aonidiella taxus Leonardi, 1906Armored scale
Aonidomytilus albus Cockerell, 1893Cassava scale
Aspidaspis arctostaphyli Cockerell & ..., 1909Arctostaphylis scale
Aspidiella hartii Cockerell, 1895Ubi scale
Aspidiella sacchari Cockerell, 1893Brown sugarcane scale
Aspidiotus destructor Cockerell, 1903Bourbon scale
Aspidiotus excisus Green, 1896Cyanotis scale
Aspidiotus nerii Bouche, 1833Aucuba scale
Asterolecanium ingae Russell, 1941Inga pit scale
Asterolecanium viennae Russell, 1941
Aulacaspis rosae (Bouche, 1833)⇒ Aulacaspis rosarum Borchsenius, 1958
Aulacaspis rosarum Borchsenius, 1958Armoured scale
Aulacaspis tubercularis Newstead, 1906Cinnamomum Scale
Aulacaspis yasumatsui Takagi, 1977Asian cycad scale
Bambusaspis bambusae Boisduval, 1869Bamboo fringed scale
Brevennia rehi Lindinger, 1943Rice Mealybug
Cardiococcus umbonatus Cockerell, 1903
Cerococcus deklei Kosztarab & ..., 1966Grenade scale
Ceroplastes ceriferus (Fabricius, 1798)⇒ Ceroplastes destructor Newstead, 1917
Ceroplastes cirripediformis Comstock, 1881Barnacle scale
Ceroplastes cistudiformis Cockerell, 1893Soft scale
Ceroplastes destructor Newstead, 1917Citrus waxy scale
Ceroplastes dugesii Lichtenstein, 1885Duges wax scale
Ceroplastes floridensis Comstock, 1881Cochonilha-cerosa
Ceroplastes japonicus Green, 1921Fig wax scale
Ceroplastes pseudoceriferus Green, 1935Horned wax scale
Ceroplastes rubens Maskell, 1893Pink Wax Scale
Ceroplastes rusci (Linnaeus, 1758)⇒ Ceroplastes japonicus Green, 1921
Chaetococcus bambusae Maskell, 1893Bamboo Mealybug
Chorizococcus nakaharai Williams & G..., 1992
Chrysomphalus aonidum Linnaeus, 1758Circular black scale
Chrysomphalus dictyospermi Morgan, 1889Armored scale
Clavaspis herculeana Cockerell & ..., 1909Armored scale
Coccus longulus Douglas, 1887Long Brown Scale
Coccus moestus De Lotto, 1959Moestus soft scale
Coccus viridis Green, 1889Coffee green scale
Conchaspis angraeci Cockerell, 1893Angraecum scale
Crenulaspidiotus dicentron Miller & Dav..., 1981
Crenulaspidiotus portoricensis Lindinger, 1910
Crypticerya rosae Riley, 1890Rose Icerya
Cryptostigma ingae (Ferris, 1922)⇒ Cryptostigma inquilina Newstead, 1920
Cryptostigma inquilina Newstead, 1920
Dactylaspis crotonis Cockerell, 1893Croton Mussel Scale
Diaspidiotus forbesi Johnson, 1896Cherry Aspidiotus
Diaspis boisduvalii Signoret, 1869Armored scale
Diaspis bromeliae (Kerner, 1778)⇒ Diaspis boisduvalii Signoret, 1869
Diaspis echinocacti Bouche, 1833Cactus scale
Distichlicoccus digitariae Williams & G..., 1992
Duplaspidiotus fossor Newstead, 1914
Duplaspidiotus tesseratus Grandpre & C..., 1899Armoured scale
Dysmicoccus bispinosus (Beardsley, 1965)⇒ Dysmicoccus texensis Tinsley, 1900
Dysmicoccus boninsis Kuwana, 1909Citrophilus mealybug
Dysmicoccus brevipes Cockerell, 1893Chanchito Blanca De La Pina
Dysmicoccus grassii Leonardi, 1913Mealybug
Dysmicoccus neobrevipes Beardsley, 1959Annona Mealybug
Dysmicoccus texensis Tinsley, 1900
Eriococcus araucariae araucariae Maskell, 1879Araucaria mealybug
Eucalymnatus tessellatus Signoret, 1873Cochonilha-reticulata
Ferrisia malvastra McDaniel, 1962Grey mealybug
Ferrisia pitcairnia Kaydan & Gullan, 2012
Ferrisia virgata (Cockerell, 1893)⇒ Ferrisia malvastra McDaniel, 1962
Fiorinia fioriniae Targioni Toz..., 1867Avocado Scale
Froggattiella penicillata Green, 1905Avocado scale
Furcaspis biformis Cockerell, 1893Armored scale
Genaparlatoria pseudaspidiotus Lindinger, 1905Armored scale
Geococcus coffeae Green, 1933Brown Scale
Gymnaspis aechmeae Newstead, 1898Armored scale
Haliaspis arecibo Howell, 1978Arecibo scale
Haliaspis nakaharai Howell, 1978Nakahara scale
Hemiberlesia lataniae Signoret, 1869Lantana scale
Hemiberlesia musae Takagi & Yam..., 1974
Hemiberlesia palmae Cockerell, 1893Scale insect
Hemiberlesia rapax Comstock, 1881Greedy scale
Hemigymnaspis eugeniae Lindinger, 1934
Howardia biclavis Comstock, 1883Armored scale
Icerya montserratensis Riley & Howard, 1890
Icerya purchasi purchasi Maskell, 1878Cottony Cushion Scale
Inglisia vitrea (Cockerell, 1894)⇒ Pseudokermes vitreus Cockerell, 1894
Insignorthezia insignis Browne, 1887Croton bug
Ischnaspis longirostris Signoret, 1882Black Line Scale
Kilifia acuminata Signoret, 1873Acuminate scale
Kuwanaspis linearis Green, 1922Bamboo long scale
Lecanodiaspis ingae Howell & Kos..., 1972
Lepidosaphes beckii Newman, 1869Citrus mussel scale
Lepidosaphes conchiformis (Gmelin, 1790)⇒ Lepidosaphes ulmi Linnaeus, 1758
Lepidosaphes gloverii Packard, 1869Citrus Long Scale
Lepidosaphes lasianthi (Green, 1900)⇒ Lepidosaphes tokionis Kuwana, 1902
Lepidosaphes rubrovittata Cockerell, 1905Guava scale
Lepidosaphes tokionis Kuwana, 1902Armored scale
Lepidosaphes ulmi Linnaeus, 1758Apple mussel scale
Lepidosaphes vermiculus Mamet, 1937Elongate coconut scale
Lopholeucaspis cockerelli Grandpre & C..., 1899Armored scale
Maconellicoccus hirsutus Green, 1908Grape
Melanaspis bromiliae Leonardi, 1899Ananas Scale
Melanaspis coccolobae Ferris, 1943
Melanaspis nigropunctata Cockerell, 1896
Melanaspis odontoglossi Cockerell, 1893Armoured scale
Melanaspis tenax Mckenzie, 1944
Milviscutulus mangiferae Green, 1889Mango shield scale
Morganella longispina Morgan, 1889Champaca scale
Mycetaspis personata Comstock, 1883Masked scale
Nipaecoccus nipae Maskell, 1893Avocado mealybug
Oceanaspidiotus araucariae Adachi & Ful..., 1953Armored scale
Oceanaspidiotus spinosus Comstock, 1883Spined scale insect
Odonaspis ruthae Kotinsky, 1915Bermuda grass scale
Odonaspis saccharicaulis Zehntner, 1897Armored scale
Palinaspis quohogiformis Merrill, 1923Quohog-shaped scale
Parasaissetia nigra Nietner, 1861Black coffee scale
Parlatoria cinerea Hadden, 1909Apple parlatoria
Parlatoria crotonis Douglas, 1887Croton parlatoria scale
Parlatoria oleae Colvée, 1880Olive parlatoria scale
Parlatoria pergandii (Comstock, 1881)⇒ Parlatoria oleae Colvée, 1880
Parlatoria proteus Curtis, 1843Cattleya scale
Parlatoria pseudaspidiotus (Lindinger, 1905)⇒ Genaparlatoria pseudaspidiotus Lindinger, 1905
Parlatoria ziziphi Lucas, 1853Black parlatoria scale
Phenacoccus colemani Ehrhorn, 1906Coleman mealybug
Phenacoccus madeirensis Green, 1923Madeira mealybug
Phenacoccus parvus Morrison, 1924Lantana Mealybug
Phenacoccus solani Ferris, 1918Soft scale
Phenacoccus solenopsis Tinsley, 1898Cotton mealybug
Pinnaspis buxi Bouche, 1851Bamboo Mussel-scale
Pinnaspis dracaenae Cooley, 1899
Pinnaspis dracarnae
Pinnaspis strachani Cooley, 1899Cotton white scale
Planococcus citri (Risso, 1813)⇒ Planococcus lilacinus Cockerell, 1905
Planococcus lilacinus Cockerell, 1905Cacao mealybug
Praelongorthezia praelonga Douglas, 1891Citrus orthezia
Protopulvinaria longivalvata Green, 1909Protopulvinarialongivalvata
Protopulvinaria pyriformis Cockerell, 1894Mealy shield scale
Pseudaonidia trilobitiformis Green, 1896Armoured scale
Pseudaulacaspis eugeniae Maskell, 1892Oleander scale
Pseudaulacaspis pentagona (Targioni-Toz..., 1886)⇒ Pseudaulacaspis simplex Takagi, 1961
Pseudaulacaspis simplex Takagi, 1961Mulberry scale
Pseudischnaspis bowreyi Cockerell, 1893Bowrey scale
Pseudococcus jackbeardsleyi Gimpel & Miller, 1996Banana mealybug
Pseudococcus longispinus Targioni-Toz..., 1867Cotonet Longispino
Pseudococcus puertoricensis Gimpel & Miller, 1996Puerto Rican mealybug
Pseudokermes vitreus Cockerell, 1894Glassy Scale
Pseudoparlatoria ostreata Cockerell, 1892Acalypha scale
Pulvinaria elongata (Newstead, 1917)⇒ Pulvinaria tenuivalvata Newstead, 1911
Pulvinaria iceryi Signoret, 1869Cottony grass scale
Pulvinaria psidii Maskell, 1893Green Shield Scale
Pulvinaria tenuivalvata Newstead, 1911Cottony grass scale
Pulvinaria urbicola Cockerell, 1893Capsicum mealy scale
Puto barberi Ferris, 1950Barber's giant mealybug
Radionaspis indica Marlatt, 1908Mango scale
Rhizoecus americanus Hambleton, 1946Mealybug
Rhizoecus hibisci Kawai & Takagi, 1971Root mealybug
Rutherfordia major Cockerell, 1894Armored scale
Saccharicoccus sacchari Cockerell, 1895Pink sugar-cane mealybug
Saissetia coffeae Walker, 1852Brown coffee scale
Saissetia miranda Cockerell & ..., 1899Mexican black scale
Saissetia neglecta De Lotto, 1969Caribbean black scale
Selenaspidus articulatus Morgan, 1889Armored scale
Trionymus radicicola Morrison, 1926Grass-root mealybug
Unaspis citri Comstock, 1883Armored scale
Vinculaspis virgata Ferris, 1941Fasciola scale
Vinsonia stellifera Westwood, 1871Stellate Scale
Virgulicoccus americanus Leonardi, 1911

Subnodes of Coccoidea with occurrences in Puerto Rico
FamilyAclerdidae
Ferris, 1937
Aclerdid scales, flat grass scales		1
2		Genus
Species
Taxom

FamilyAsterolecaniidae
Cockerell, 1896
Asterolecaniids, Pit scales		2
3		Genera
Species
Taxom

FamilyCerococcidae
Balachowsky, 1942
Cerococcids, Ornate Pit Scale Insects, Ornate pit scales		1
1		Genus
Species
Taxom

FamilyCoccidae
Stephens, 1829
soft scale, Soft Scale Insects, Soft scales, Tortoise scales, Wax scales		14
33		Genera
Species
Taxom

FamilyConchaspididae
Green, 1896
False armored scales, False armoured scales		1
1		Genus
Species
Taxom

FamilyDiaspididae
Maskell, 1878
Armored scale insects, Armored scales, armoured scale insect, Armoured Scales, Diaspidids		45
90		Genera
Species
Taxom

FamilyEriococcidae
Cockerell, 1899
Eriococcid scales, Eriococcids, felt scale, Felt scales, Felted scales, Scales		1
1		Genus
Species
Taxom

FamilyLecanodiaspididae
Borchsenius, 1965
False pit scales, Lecanodiaspidids		2
2		Genera
Species
Taxom

FamilyMonophlebidae
Signoret, 1875
giant scale, Giant Scale Insects, Giant scales, Monophlebids		2
3		Genera
Species
Taxom

FamilyOrtheziidae
Green, 1896
Ensign coccids, ensign scale insect, Ensign scales, Ortheziids		2
2		Genera
Species
Taxom

FamilyPseudococcidae
Heymons, 1915
Mealy Bugs, mealybug, Mealybugs		16
30		Genera
Species
Taxom

FamilyPutoidae ♰
Beardsley, 1969
Giant mealybugs, Putoids		1
1		Genus
Species
Taxom

FamilyRhizoecidae
Williams, 1969
1
2		Genus
Species
Taxom



Links, sources and further information
Coccoidea at artfakta.se
Coccoidea at dyntaxa.se
Coccoidea at itis.gov