Species list, Checklist of Superorder Holometabola (Holometabolous Insects) for Cacao Biological Station (Guanacaste National Park, Costa Rica)

Ants	Arachnids
Bees	Beetles
Booklice	Bugs
Bumblebees	Butterflies
Cicadas	Crane flies
Dragonflies	Earwigs
Flies	Isopods
Locusts	Mantises
Mayflies	Mosquitoes
Myriapods	Net-winged
Plant-parasitic	Scorpion flies
Termites	Wasps

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Guanacaste National Park

Cacao Biological Station

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INSECTOIDGERENGDARK LIGHT

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Checklist for Superorder Holometabola Heider, 1889 (Holometabolous Insects)
and region Cacao Biological Station (Guanacaste National Park, Costa Rica)

Class	Insecta Linnaeus, 1758 Insects, True insects	286 2832 3151	Families Genera Species	Taxom
Subclass	Pterygota Lang, 1888 Winged insects	286 2832 3151	Families Genera Species	Taxom
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	277 2821 3136	Families Genera Species	Taxom
Superorder	Holometabola Heider, 1889 Holometabolous Insects	208 2542 2843	Families Genera Species	Taxom

List of families for Cacao Biological Station
208 Families width 2843 Species and 101 Subspecies

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Agaonidae Walker, 1848	Fig wasp	1	Species	Taxom
Agromyzidae Fallen, 1823	Leaf miner flies	0	Species	Taxom
Aididae Schaus, 1906		1	Species	Taxom
Alucitidae Leach, 1815	Many Plumed Moths	1	Species	Taxom
Ampulicidae Shuckard, 1840	Ampulicid wasps	0	Species	Taxom
Andrenidae Webster, 1913	Andrenid bees	1	Species	Taxom
Anisopodidae Knab, 1912	Window Midges	0	Species	Taxom
Anthicidae Latreille, 1819	Ant beetles	1	Species	Taxom
Anthomyzidae Czerny, 1903	Anthomyzid flies	0	Species	Taxom
Anthribidae Billberg, 1820	Fungus Weevil	7	Species	Taxom
Apidae Latreille, 1802	Bumble bees	27	Species	Taxom
Arctiidae Leach, 1815	Footman moths	4	Species	Taxom
Argidae Konow, 1890	Argid flies	3	Species	Taxom
Asilidae Latreille, 1802	Assassin flies	15	Species	Taxom
Attelabidae Billberg, 1820	Leaf Rolling Weevil	15	Species	Taxom
Bethylidae Halliday, 1839	Bethylid wasps	1	Species	Taxom
Bibionidae Fleming, 1821	Bibionid flies	1	Species	Taxom
Bittacidae Handlirsch, 1906	Hanging scorpionflies	1	Species	Taxom
Bombycidae Latreille, 1802	Lappet moths	8	Species	Taxom
Bombyliidae Latreille, 1802	Bee Flies	5	Species	Taxom
Bostrichidae Latreille, 1802	Auger beetles	1	Species	Taxom
Braconidae Latreille, 1829	Braconid wasps	44	Species	Taxom
Brentidae Billberg, 1820	Brenthidae	27	Species	Taxom
Buprestidae Leach, 1815	Flat-headed wood-borers	14	Species	Taxom
Calamoceratidae Ulmer, 1905		2	Species	Taxom
Calliphoridae Brauer & Ber..., 1889	Blow flies	8	Species	Taxom
Cantharidae Imhoff, 1856	Cantharids	0	Species	Taxom
Carabidae Latreille, 1802	Carabid beetles	61	Species	Taxom
Carnidae Newman, 1834	Bird flies	1	Species	Taxom
Castniidae Boisduval, 1828	Castniid moths	2	Species	Taxom
Cecidomyiidae Newman, 1834	Forest Midges	2	Species	Taxom
Cerambycidae Latreille, 1802	Long Horn beetles	193	Species	Taxom
Ceratophyllidae Dampf, 1908	Bird Fleas	1	Species	Taxom
Ceratopogonidae Newman, 1834	Biting midge	4	Species	Taxom
Chalcididae Latreille, 1817	Chalcidid wasps	2	Species	Taxom
Chironomidae Erichson, 1842	Chironomids	3	Species	Taxom
Chloropidae Schoenherr, 1840	Chloropid flies	3	Species	Taxom
Choreutidae Stainton, 1854	Choreuitid Moths	1	Species	Taxom
Chrysididae Latreille, 1802	Chrysidid wasps	2	Species	Taxom
Chrysomelidae Latreille, 1802	Casebearers	99	Species	Taxom
Cleridae Latreille, 1802	Checkered Beetle	3	Species	Taxom
Clusiidae Schönherr, 1840	Clusiid flies	5	Species	Taxom
Coccinellidae Latreille, 1807	Coccinelles	8	Species	Taxom
Coleophoridae Hübner, 1825	Case moths	6	Species	Taxom
Colletidae Lepeletier, 1841	Colletid bees	4	Species	Taxom
Cossidae Leach, 1815	Borers	1	Species	Taxom
Crabronidae Latreille, 1802	Bee Wolves	0	Species	Taxom
Crambidae Latreille, 1810	Crambid Snout Moths	105	Species	Taxom
Ctenophthalmidae Rothschild, 1915		1	Species	Taxom
Cucujidae Latreille, 1802	Flat bark beetles	0	Species	Taxom
Culicidae Meigen, 1818	Mosquito	2	Species	Taxom
Curculionidae Latreille, 1802	Ambrosia beetles	129	Species	Taxom
Cynipidae Latreille, 1802	Cynipid gall wasps	1	Species	Taxom
Dalceridae Dyar, 1898	Dalcerid moths	4	Species	Taxom
Diapriidae Latreille, 1796	Diapriid wasps	1	Species	Taxom
Dolichopodidae Latreille, 1809	Long legged flies	9	Species	Taxom
Douglasiidae Heinemann & ..., 1876	Douglas Moths	1	Species	Taxom
Drosophilidae Rondani, 1856	Fruit flies	3	Species	Taxom
Dytiscidae Leach, 1815	Diving beetles	3	Species	Taxom
Elachistidae Bruand, 1851	Elachistid moths	27	Species	Taxom
Elateridae Leach, 1815	Click Beetle	24	Species	Taxom
Elmidae Curtis, 1830	Drive beetles	2	Species	Taxom
Empididae Latreille, 1809	Balloon flies	1	Species	Taxom
Encyrtidae Walker, 1837	Encyrtid wasps	2	Species	Taxom
Ephydridae Zetterstedt, 1837	Brine flies	2	Species	Taxom
Epipyropidae Dyar, 1903	Planthopper Parasite Moths	1	Species	Taxom
Erebidae Leach, 1815	Erebid Moths	295	Species	Taxom
Erotylidae Latreille, 1802	Pleasing Fungus Beetle	8	Species	Taxom
Eucharitidae Latreille, 1809	Eucharitid wasp	1	Species	Taxom
Eucnemidae Eschscholtz, 1829	Click Beetles	0	Species	Taxom
Eulophidae Westwood, 1829	Eulophid wasps	4	Species	Taxom
Eurytomidae Walker, 1832	Eurytomid wasps	0	Species	Taxom
Euteliidae Grote, 1882	Euteliid Moths	7	Species	Taxom
Evaniidae Latreille, 1802	Ensign wasp	0	Species	Taxom
Fanniidae Schnabl & Dz..., 1911	Latrine flies	1	Species	Taxom
Figitidae Thomson, 1862	Figitid wasps	1	Species	Taxom
Formicidae Latreille, 1809	Ants	47	Species	Taxom
Gasteruptiidae Ashmead, 1900	Carrot Wasps	0	Species	Taxom
Gelechiidae Stainton, 1854	Gelechiid moths	21	Species	Taxom
Geometridae Leach, 1815	Cankerworm moths	184	Species	Taxom
Geotrupidae Latreille, 1802	Dor beetles	2	Species	Taxom
Gracillariidae Stainton, 1854	Blotch leaf miner moths	1	Species	Taxom
Gyrinidae Latreille, 1802	Whirligig beetle	0	Species	Taxom
Halictidae Thomson, 1869	Flower bees	7	Species	Taxom
Hedylidae Guenée, 1857	American Butterfly Moths	4	Species	Taxom
Heleomyzidae (Westwood, 1840)	Heleomyzid flies	0	Species	Taxom
Heliozelidae Heinemann & ..., 1876	Leaf miners	1	Species	Taxom
Hesperiidae Latreille, 1809	Skipper	49	Species	Taxom
Hippoboscidae Samouelle, 1819	Keds	1	Species	Taxom
Histeridae Gyllenhal, 1808	Clown beetle	12	Species	Taxom
Hybosoridae Erichson, 1847	Pill scarab beetles	10	Species	Taxom
Hybotidae Meigen, 1820	Dance flies	1	Species	Taxom
Hydraenidae Mulsant, 1844	Cascade Beetles	1	Species	Taxom
Hydrobiosidae Ulmer, 1905		2	Species	Taxom
Hydrophilidae Latreille, 1802	Silver Water Beetles	3	Species	Taxom
Hydropsychidae Curtis, 1835	Net Spinning caddisflies	2	Species	Taxom
Ichneumonidae Latreille, 1802	Ichneumon flies	208	Species	Taxom
Ischnopsyllidae Wahlgren, 1907	Bat fleas	1	Species	Taxom
Keroplatidae Rondani, 1856	Fungus gnats	1	Species	Taxom
Lampyridae Latreille, 1817	Fireflies	2	Species	Taxom
Lasiocampidae Harris, 1841	Eggar Moths	20	Species	Taxom
Lauxaniidae Macquart, 1835	Beach flies	1	Species	Taxom
Lecithoceridae Le Marchand, 1947	Long-horned moths	1	Species	Taxom
Leiodidae Fleming, 1821	Round fungus beetles	5	Species	Taxom
Lepidostomatidae Ulmer, 1903	Bizarre Caddisflies	1	Species	Taxom
Leptoceridae Leach, 1815	Long Horned caddisflies	1	Species	Taxom
Leptopsyllidae Rothschild &..., 1915	Bird fleas	1	Species	Taxom
Limacodidae Duponchel, 1845	Cup moths	13	Species	Taxom
Limnichidae Erichson, 1846	Minute marsh loving beetle	0	Species	Taxom
Liopteridae Ashmead, 1895	Liopterid wasps	0	Species	Taxom
Lonchaeidae Loew, 1861	Lance Flies	0	Species	Taxom
Lycaenidae Leach, 1815	Blue butterflies	13	Species	Taxom
Lycidae Laporte, 1836	Lycid beetles	0	Species	Taxom
Megachilidae Schenk, 1860	And Resin Bees	0	Species	Taxom
Megalodontesidae Konow, 1897		1	Species	Taxom
Megalopygidae Herrich-Schä..., 1855	Crinkled Flannel Moths	5	Species	Taxom
Meloidae Gyllenhal, 1810	Blister Beetle	0	Species	Taxom
Melyridae Leach, 1815	Malachite Beetles	0	Species	Taxom
Micropezidae Illiger, 1798	Stilt Flies	1	Species	Taxom
Milichiidae Schiner, 1862	Filth flies	0	Species	Taxom
Mimallonidae Burmeister, 1878	Mimallonid Moths	6	Species	Taxom
Monomachidae Ashmead, 1902	Monomachid wasps	0	Species	Taxom
Mordellidae Latreille, 1802	Pintail beetles	1	Species	Taxom
Muscidae Latreille, 1802	Face flies	3	Species	Taxom
Mutillidae André, 1907	Cow ant	3	Species	Taxom
Mycetophagidae Leach, 1815	Hairy fungus beetle	1	Species	Taxom
Mycetophilidae Newman, 1834	Fungus Gnat	0	Species	Taxom
Mycteridae Blanchard, 1845	Mycterid beetles	0	Species	Taxom
Neriidae Hendel, 1916	Banana stalk flies	0	Species	Taxom
Nitidulidae Latreille, 1802	Nitidules	0	Species	Taxom
Noctuidae Latreille, 1809	Armyworms	136	Species	Taxom
Nolidae Hampson, 1894	Hatted Caterpillars	14	Species	Taxom
Nosodendridae Erichson, 1846	Nosodendrid beetles	0	Species	Taxom
Notodontidae Stephens, 1829	Bag Shelter Moths	100	Species	Taxom
Nymphalidae Rafinesque, 1815	Admirals	70	Species	Taxom
Odontoceridae Wallengren, 1891	Mortarjoint Casemaker Caddisflies	0	Species	Taxom
Oecophoridae Bruand, 1851	Concealer Moths	6	Species	Taxom
Oestridae Leach, 1815	Bot flies	1	Species	Taxom
Otitidae (Macquart, 1835)	Picture-winged flies	5	Species	Taxom
Pantophthalmidae Williston, 1906	Timber flies	3	Species	Taxom
Papilionidae Latreille, 1802	Apollos	12	Species	Taxom
Passalidae Leach, 1815	Bess beetle	11	Species	Taxom
Pelecinidae Haliday, 1839	Ancestral And Modern Pelecinid Wasps	1	Species	Taxom
Philopotamidae Stephens, 1829	Finger Net caddisflies	3	Species	Taxom
Phoridae Curtis, 1833	Coffin Flies	10	Species	Taxom
Pieridae Duponchel, 1835	Brimstones	24	Species	Taxom
Pipunculidae Walker, 1834	Big headed flies	1	Species	Taxom
Platygastridae Haliday, 1833	Platygastrid wasps	1	Species	Taxom
Polycentropodidae Ulmer, 1903	Tube Maker Caddisflies	1	Species	Taxom
Pompilidae Latreille, 1805	Pompilid wasps	6	Species	Taxom
Proctotrupidae Latreille, 1802	Proctotrupid wasps	0	Species	Taxom
Prodoxidae Riley, 1881	Yucca Moths	1	Species	Taxom
Psilidae Macquart, 1835	Carrot flies	0	Species	Taxom
Psychodidae Newman, 1834	Drain flies	1	Species	Taxom
Pteromalidae Dalman, 1820	Pteromalid wasps	5	Species	Taxom
Pterophoridae Zeller, 1841	Plume Moths	1	Species	Taxom
Ptilodactylidae Laporte, 1836	Ptilodactylid beetles	0	Species	Taxom
Ptinidae Latreille, 1802	Death-watch and Spider Beetles	1	Species	Taxom
Pygiopsyllidae Wagner, 1939		1	Species	Taxom
Pyralidae Latreille, 1802	Flour Moths	39	Species	Taxom
Rhinophoridae Robineau-Des..., 1863	Woodlouse flies	3	Species	Taxom
Rhopalopsyllidae Oudemans, 1909	Marsupial fleas	1	Species	Taxom
Rhynchitidae Gistel, 1848	Leaf-rolling weevils	1	Species	Taxom
Richardidae Snow, 1896	Richardid flies	1	Species	Taxom
Riodinidae Grote, 1895	Metalmark Butterflies	12	Species	Taxom
Rutelidae (MacLeay, 1819)		2	Species	Taxom
Salpingidae Leach, 1815	Narrow-waisted bark beetle	0	Species	Taxom
Sarcophagidae Macquart, 1834	Flesh flies	4	Species	Taxom
Saturniidae Boisduval, 1837	And Giant Silk Moths	32	Species	Taxom
Scarabaeidae Latreille, 1802	Dung beetles	137	Species	Taxom
Scelionidae Haliday, 1839	Scelionid wasps	1	Species	Taxom
Sciaridae Billberg, 1820	Black fungus gnats	0	Species	Taxom
Scoliidae Leach, 1815	Mammoth wasps	0	Species	Taxom
Scydmaenidae (Leach, 1815)	Ant-like stone beetles	1	Species	Taxom
Sematuridae Guenée, 1858	American Swallowtail Moths	2	Species	Taxom
Sesiidae Boisduval, 1828	Clear Wings	6	Species	Taxom
Silphidae Latreille, 1807	Burying And Carrion Beetles	1	Species	Taxom
Silvanidae Kirby, 1837	Flat bark beetles	1	Species	Taxom
Simuliidae Newman, 1834	Black flies	1	Species	Taxom
Sphaeroceridae Macquart, 1835	Dung flies	6	Species	Taxom
Sphecidae Latreille, 1802	Digger & Sand wasps	9	Species	Taxom
Sphingidae Latreille, 1802	Hawk Moths	53	Species	Taxom
Staphylinidae Lameere, 1900	Rove beetles	60	Species	Taxom
Stratiomyidae Latreille, 1802	Soldier flies	15	Species	Taxom
Syrphidae Latreille, 1802	Drone flies	59	Species	Taxom
Tabanidae Latreille, 1802	Breeze flies	11	Species	Taxom
Tachinidae (Robineau-Des..., 1830)	Bristle Flies	21	Species	Taxom
Tanypezidae Rondani, 1856	Stretched-foot flies	1	Species	Taxom
Tenebrionidae Latreille, 1802	Darkling Beetle	22	Species	Taxom
Tenthredinidae Latreille, 1804	Common sawflies	3	Species	Taxom
Tephritidae Newman, 1834	Fruit flies	12	Species	Taxom
Therevidae Newman, 1834	Stiletto flies	0	Species	Taxom
Throscidae Laporte, 1840	False metallic wood-boring beetles	0	Species	Taxom
Thyrididae Herrich-Schä..., 1846	Picture Winged Leaf Moths	2	Species	Taxom
Tineidae Latreille, 1810	& Allies	1	Species	Taxom
Tiphiidae Leach, 1815	Flower wasps	0	Species	Taxom
Tipulidae Latreille, 1802	Crane Flies	4	Species	Taxom
Tortricidae Latreille, 1803	Bell Moths	18	Species	Taxom
Torymidae Walker, 1833	Apple seed chalcids	0	Species	Taxom
Trigonalidae (Cresson, 1887)	Trigonalid sawflies	1	Species	Taxom
Trogidae (MacLeay, 1819)	Gnawer beetles	1	Species	Taxom
Ulidiidae Macquart, 1835	Picture-winged Flies	1	Species	Taxom
Uraniidae Blanchard, 1845	Swallowtail Moths	1	Species	Taxom
Urodidae Kyrki, 1984	False Burnet Moths	0	Species	Taxom
Vespidae Leach, 1815	And Allies	20	Species	Taxom
Xiphydriidae Leach, 1819	Wood wasps	0	Species	Taxom
Yponomeutidae Stephens, 1829	Ermine Moths	1	Species	Taxom
Zopheridae Solier, 1834	Cylindrical bark beetles	0	Species	Taxom

Subnodes of Holometabola with occurrences in Cacao Biological Station

Order	Coleoptera Linnaeus, 1758 Beetles, Coleopterans	51 924 870	Families Genera Species	Taxom
Order	Diptera Linnaeus, 1758 Flies, Gnats, hoverflies, Mosquitoes, True flies, Two-Winged Flies	52 376 233	Families Genera Species	Taxom
Order	Hymenoptera Linnaeus, 1758 Ants, Bees, Sawflies, Wasps	42 391 408	Families Genera Species	Taxom
Order	Lepidoptera Linnaeus, 1758 Butterflies, Lepidopterans, Moths	48 834 1313	Families Genera Species	Taxom
Order	Mecoptera Hyatt & Arms, 1891 Hanging flies, Hangingflies, Scorpion flies, Scorpionflies, snow scorpionflies, Snowflies	7 7 7	Families Genera Species	Taxom
Order	Trichoptera Kirby, 1813 Caddis-Flies, Caddisflies, Caddisfly, Cased Caddis, Cased Caddis-Flies, Caseless Caddis, Caseless Caddis-Flies, Rail-flies, Sedge-flies	8 10 12	Families Genera Species	Taxom

Links, sources and further information

Holometabola at itis.gov