Species list, Checklist of Superorder Holometabola (Holometabolous Insects) for Lower Cretaceous (Cretaceous)

Ants	Arachnids
Bees	Beetles
Booklice	Bugs
Bumblebees	Butterflies
Cicadas	Crane flies
Dragonflies	Earwigs
Flies	Isopods
Locusts	Mantises
Mayflies	Mosquitoes
Myriapods	Net-winged
Plant-parasitic	Scorpion flies
Termites	Wasps

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INSECTOIDGERENGDARK LIGHT

⌕

Checklist for Superorder Holometabola Heider, 1889 (Holometabolous Insects)
and region Lower Cretaceous (Cretaceous)

Class	Insecta Linnaeus, 1758 Insects, True insects	566 2074 3587	Families Genera Species	Taxom
Subclass	Pterygota Lang, 1888 Winged insects	565 2073 3586	Families Genera Species	Taxom
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	506 1892 3348	Families Genera Species	Taxom
Superorder	Holometabola Heider, 1889 Holometabolous Insects	276 1132 2082	Families Genera Species	Taxom

List of families for Lower Cretaceous
276 Families width 2082 Species and 3 Subspecies

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Ademosynidae Ponomarenko, 1968		3	Species	Taxom
Agaonidae Walker, 1848	Fig wasp	1	Species	Taxom
Agyrtidae Thomson, 1859	Primitive carrion beetles	3	Species	Taxom
Alavarommatidae Ortega-Blanc..., 2011		1	Species	Taxom
Ampulicidae Shuckard, 1840	Ampulicid wasps	19	Species	Taxom
Anaxyelidae Martynov, 1925	Anaxyelids	9	Species	Taxom
Aneuretopsychidae Rasnitsyn & ..., 1990		5	Species	Taxom
Angarosphecidae Rasnitsyn, 1975		7	Species	Taxom
Anisopodidae Knab, 1912	Window Midges	8	Species	Taxom
Antefungivoridae Rohdendorf, 1938		5	Species	Taxom
Anthicidae Latreille, 1819	Ant beetles	1	Species	Taxom
Anthomyzidae Czerny, 1903	Anthomyzid flies	1	Species	Taxom
Anthribidae Billberg, 1820	Fungus Weevil	2	Species	Taxom
Apidae Latreille, 1802	Bumble bees	1	Species	Taxom
Archaeocynipidae Rasnitsyn & ..., 1988		3	Species	Taxom
Archaeoserphitidae Engel, 2015		1	Species	Taxom
Archisargidae Rohdendorf, 1962		5	Species	Taxom
Archizelmiridae Rohdendorf, 1962		2	Species	Taxom
Asilidae Latreille, 1802	Assassin flies	1	Species	Taxom
Asiochaoboridae Hong & Wang, 1990		4	Species	Taxom
Atelestidae Hennig, 1970		6	Species	Taxom
Athericidae Stuckenberg, 1973	Athericid flies	7	Species	Taxom
Attelabidae Billberg, 1820	Leaf Rolling Weevil	2	Species	Taxom
Aulacidae Shuckard, 1842	Aulacid wasps	29	Species	Taxom
Australochoristidae Krzemi?ski, ..., 2015		1	Species	Taxom
Axymyiidae Rohdendorf, 1946	Axymyiidae flies	3	Species	Taxom
Baissidae Rasnitsyn, 1975		1	Species	Taxom
Baissodidae Benton, 1993		8	Species	Taxom
Baissoferidae Sukatsheva, 1968		5	Species	Taxom
Belidae Schönherr, 1826	Belid Beetle	3	Species	Taxom
Bethylidae Halliday, 1839	Bethylid wasps	8	Species	Taxom
Bethylonymidae Rasnitsyn, 1975		2	Species	Taxom
Bibionidae Fleming, 1821	Bibionid flies	3	Species	Taxom
Bittacidae Handlirsch, 1906	Hanging scorpionflies	8	Species	Taxom
Blephariceridae Loew, 1861	Mountain midges	1	Species	Taxom
Boholdoyidae Kovalev, 1985		1	Species	Taxom
Bolitophilidae Malloch, 1917		1	Species	Taxom
Bombyliidae Latreille, 1802	Bee Flies	2	Species	Taxom
Boreidae Latreille, 1816	Snow fleas	1	Species	Taxom
Bostrichidae Latreille, 1802	Auger beetles	1	Species	Taxom
Brachycentridae Ulmer, 1903	Humpless Casemaker Caddisflies	1	Species	Taxom
Braconidae Latreille, 1829	Braconid wasps	5	Species	Taxom
Brentidae Billberg, 1820	Brenthidae	4	Species	Taxom
Buprestidae Leach, 1815	Flat-headed wood-borers	51	Species	Taxom
Byrrhidae Latreille, 1804	Moss beetles	6	Species	Taxom
Calamoceratidae Ulmer, 1905		1	Species	Taxom
Carabidae Latreille, 1802	Carabid beetles	49	Species	Taxom
Caridae Thompson, 1992		26	Species	Taxom
Cecidomyiidae Newman, 1834	Forest Midges	6	Species	Taxom
Cephidae Newman, 1834	Cephids	2	Species	Taxom
Cerambycidae Latreille, 1802	Long Horn beetles	9	Species	Taxom
Ceratopogonidae Newman, 1834	Biting midge	47	Species	Taxom
Cerophytidae Latreille, 1834	Cerophytid beetles	4	Species	Taxom
Cerylonidae Billberg, 1820	Cerylid Kingfishers	1	Species	Taxom
Chaoboridae Newman, 1834	Phantom midge	16	Species	Taxom
Chelonariidae Blanchard, 1845	Chelonarid beetles	1	Species	Taxom
Chimeromyiidae Grimaldi & C..., 2009		7	Species	Taxom
Chironomidae Erichson, 1842	Chironomids	42	Species	Taxom
Choristidae Esben-Petersen, 1915		1	Species	Taxom
Chrysididae Latreille, 1802	Chrysidid wasps	1	Species	Taxom
Chrysomelidae Latreille, 1802	Casebearers	7	Species	Taxom
Cimbicidae Kirby, 1837	Cimbicid sawflies	1	Species	Taxom
Cimbrophlebiidae Willmann, 1977	Hangingfly	3	Species	Taxom
Clambidae Fischer, 1821	Fringe-winged beetles	1	Species	Taxom
Cleridae Latreille, 1802	Checkered Beetle	1	Species	Taxom
Coccinellidae Latreille, 1807	Coccinelles	1	Species	Taxom
Coelocatiniidae Ponomarenko, 1968		2	Species	Taxom
Coleocatiniidae Ponomarenko ..., 2015		1	Species	Taxom
Coptoclavidae Ponomarenko, 1961	Extinct family	18	Species	Taxom
Crabronidae Latreille, 1802	Bee Wolves	5	Species	Taxom
Cryptophagidae Kirby, 1937	Mould beetles	4	Species	Taxom
Culicidae Meigen, 1818	Mosquito	2	Species	Taxom
Cupedidae Laporte, 1836	Cupedids	43	Species	Taxom
Curculionidae Latreille, 1802	Ambrosia beetles	16	Species	Taxom
Cynipidae Latreille, 1802	Cynipid gall wasps	3	Species	Taxom
Dascillidae Guérin-Ménev..., 1843	Orchid beetles	1	Species	Taxom
Dermestidae Hope, 1840	Carpet beetles	2	Species	Taxom
Diapriidae Latreille, 1796	Diapriid wasps	8	Species	Taxom
Dolichopodidae Latreille, 1809	Long legged flies	2	Species	Taxom
Dryinidae Haliday, 1840	Dryinid wasps	9	Species	Taxom
Dysoneuridae Sukatsheva, 1968		4	Species	Taxom
Dytiscidae Leach, 1815	Diving beetles	14	Species	Taxom
Elateridae Leach, 1815	Click Beetle	30	Species	Taxom
Elliidae Krzeminska, ..., 1993		3	Species	Taxom
Elodophthalmidae Kirejtshuk &..., 2008		2	Species	Taxom
Embolemidae Förster, 1856	Embolemid wasps	1	Species	Taxom
Empididae Latreille, 1809	Balloon flies	9	Species	Taxom
Encyrtidae Walker, 1837	Encyrtid wasps	1	Species	Taxom
Englathaumatidae Novokshonov ..., 2016		2	Species	Taxom
Eoichneumonidae (Jell & Duncan, 1986)		15	Species	Taxom
Eolepidopterigidae Rasnitsyn, 1983		2	Species	Taxom
Eomeropidae Cockerell, 1909		3	Species	Taxom
Ephialtitidae Handlirsch, 1906		10	Species	Taxom
Erebidae Leach, 1815	Erebid Moths	1	Species	Taxom
Eremazidae Iablokoff-Kh..., 1977		2	Species	Taxom
Eremochaetidae Ussatchov, 1968		17	Species	Taxom
Eucinetidae Lacordaire, 1857	Plate-thigh beetle	4	Species	Taxom
Eucnemidae Eschscholtz, 1829	Click Beetles	1	Species	Taxom
Evaniidae Latreille, 1802	Ensign wasp	18	Species	Taxom
Falsiformicidae Rasnitsyn, 1975		1	Species	Taxom
Formicidae Latreille, 1809	Ants	7	Species	Taxom
Fungivoritidae Rohdendorf, 1957		1	Species	Taxom
Gasteruptiidae Ashmead, 1900	Carrot Wasps	1	Species	Taxom
Geotrupidae Latreille, 1802	Dor beetles	5	Species	Taxom
Glaphyridae MacLeay, 1819	Bumble bee scarab beetle	23	Species	Taxom
Glaresidae Kolbe, 1905	Enigmatic scarab beetle	6	Species	Taxom
Glossosomatidae Wallengren, 1891	Little Black Caddisflies	1	Species	Taxom
Gyrinidae Latreille, 1802	Whirligig beetle	4	Species	Taxom
Haliplidae Aubé, 1836	Crawling Water Beetle	3	Species	Taxom
Helophoridae (Leach, 1815)	Water scavenger beetles	1	Species	Taxom
Heloridae Förster, 1856	Helorid wasps	11	Species	Taxom
Hesperiidae Latreille, 1809	Skipper	2	Species	Taxom
Heteroceridae MacLeay, 1825	Mud beetles	2	Species	Taxom
Hilarimorphidae Hendel, 1936	Hilarimorphid flies	1	Species	Taxom
Histeridae Gyllenhal, 1808	Clown beetle	1	Species	Taxom
Huaxiasciaritidae (Hong, 2002)		2	Species	Taxom
Hybosoridae Erichson, 1847	Pill scarab beetles	11	Species	Taxom
Hydraenidae Mulsant, 1844	Cascade Beetles	1	Species	Taxom
Hydrophilidae Latreille, 1802	Silver Water Beetles	35	Species	Taxom
Hydroscaphidae LeConte, 1874	Skiff beetle	1	Species	Taxom
Ichneumonidae Latreille, 1802	Ichneumon flies	35	Species	Taxom
Ichneumonomimidae Rasnitsyn, 1975		2	Species	Taxom
Ironomyiidae McAlpine & M..., 1966	Ironic flies	17	Species	Taxom
Ithyceridae (Schönherr, 1826)	New York weevil	3	Species	Taxom
Kateretidae Kirkby, 1837	Pollen Beetles	1	Species	Taxom
Keroplatidae Rondani, 1856	Fungus gnats	2	Species	Taxom
Khutelchalcididae Rasnitsyn & al., 2004		1	Species	Taxom
Lasiosynidae Kirejtshuk, ..., 2010		14	Species	Taxom
Latridiidae Erichson, 1842	Minute brown scavenger beetle	3	Species	Taxom
Leiodidae Fleming, 1821	Round fungus beetles	1	Species	Taxom
Lepidostomatidae Ulmer, 1903	Bizarre Caddisflies	2	Species	Taxom
Leptoceridae Leach, 1815	Long Horned caddisflies	2	Species	Taxom
Liadytidae Ponomarenko, 1977		4	Species	Taxom
Limnephilidae Kolenati, 1848	Northern caddisflies	71	Species	Taxom
Limoniidae (Speiser, 1909)	Craneflies	12	Species	Taxom
Lithoscarabaeidae Nikolajev, 1992		3	Species	Taxom
Lonchopteridae Curtis, 1839	Pointed-wing flies	2	Species	Taxom
Lucanidae Latreille, 1804	Stag Beetle	12	Species	Taxom
Lygistorrhinidae Hendel, 1936	Long beaked fungus gnat	1	Species	Taxom
Lymexylidae Fleming, 1821	Ship-timber beetle	1	Species	Taxom
Magnocoleidae Hong, 1998		1	Species	Taxom
Maimetshidae (Rasnitsyn, 1975)		7	Species	Taxom
Megachilidae Schenk, 1860	And Resin Bees	2	Species	Taxom
Megalodontesidae Konow, 1897		5	Species	Taxom
Megalyridae Schletterer, 1889	Megalyrid wasps	13	Species	Taxom
Melandryidae Leach, 1815	False Darkling Beetle	2	Species	Taxom
Meloidae Gyllenhal, 1810	Blister Beetle	1	Species	Taxom
Mesopsychidae Tillyard, 1917		6	Species	Taxom
Mesosciophilidae Rohdendorf, 1946		10	Species	Taxom
Mesoserphidae Kozlov, 1970		7	Species	Taxom
Micromalthidae Barber, 1913	Micromalthid beetles	1	Species	Taxom
Micropterigidae Herrich-Schä..., 1855	Mandibulate Archaic Moths	4	Species	Taxom
Monotomidae Laporte, 1840	Minute Clubbed Beetles	5	Species	Taxom
Mordellidae Latreille, 1802	Pintail beetles	7	Species	Taxom
Mycetophilidae Newman, 1834	Fungus Gnat	56	Species	Taxom
Mydidae Bezzi, 1903	Mydas flies	1	Species	Taxom
Mymaridae Haliday, 1833	Fairflies	3	Species	Taxom
Mymarommatidae Debauche, 1948	False fairy wasp	1	Species	Taxom
Nannochoristidae Tillyard, 1917		1	Species	Taxom
Necrotauliidae Handlirsch, 1906		13	Species	Taxom
Nemestrinidae Macquart, 1834	Tangle-veined flies	8	Species	Taxom
Nemonychidae Bedel, 1882	Nemonychid weevils	47	Species	Taxom
Ningxiapsychidae Hong & Li, 2004		1	Species	Taxom
Nitidulidae Latreille, 1802	Nitidules	5	Species	Taxom
Nymphalidae Rafinesque, 1815	Admirals	1	Species	Taxom
Ochodaeidae Mulsant & Rey, 1870	Sand-loving scarab beetle	3	Species	Taxom
Oedemeridae Latreille, 1810	False Blister Beetle	2	Species	Taxom
Ommatidae Sharp & Muir, 1912		81	Species	Taxom
Opetiidae Rondani, 1856	Flat-footed flies	4	Species	Taxom
Orthophlebiidae Handlirsch, 1906		14	Species	Taxom
Pachyneuridae Schiner, 1864	Dark-winged fungus gnats	1	Species	Taxom
Pamphiliidae Cameron, 1890	Leaf-rolling sawflies	1	Species	Taxom
Papilionidae Latreille, 1802	Apollos	1	Species	Taxom
Parandrexidae Kirejtshuk, 1994		1	Species	Taxom
Paraxymyiidae Rohdendorf, 1946	Paxylommatid wasps	1	Species	Taxom
Paroryssidae Martynov, 1925		2	Species	Taxom
Pediciidae Osten Sacken, 1860	Hairy-Eyed Crane Flies	3	Species	Taxom
Pelecinidae Haliday, 1839	Ancestral And Modern Pelecinid Wasps	42	Species	Taxom
Peradeniidae Naumann & Ma..., 1985		1	Species	Taxom
Perissommatidae Colless, 1962		2	Species	Taxom
Permochoristidae Tillyard, 1918		2	Species	Taxom
Permocupedidae Martynov, 1933		1	Species	Taxom
Phoridae Curtis, 1833	Coffin Flies	3	Species	Taxom
Phryganeidae (Leach, 1815)	Giant casemaker	4	Species	Taxom
Pieridae Duponchel, 1835	Brimstones	1	Species	Taxom
Platypezidae Fallen, 1815	Flat footed flies	11	Species	Taxom
Pleciofungivoridae Rohdendorf, 1946		7	Species	Taxom
Pleciomimidae Rodendorf, 1949		3	Species	Taxom
Plectrotarsidae Mosely, 1953		1	Species	Taxom
Pleocomidae LeConte, 1861	Rain beetle	3	Species	Taxom
Polycentropodidae Ulmer, 1903	Tube Maker Caddisflies	2	Species	Taxom
Pompilidae Latreille, 1805	Pompilid wasps	9	Species	Taxom
Praeaulacidae Rasnitsyn, 1972		13	Species	Taxom
Praeichneumonidae Rasnitsyn, 1983		5	Species	Taxom
Praesiricidae Rasnitsyn, 1968		14	Species	Taxom
Procramptonomyiidae Kovalev, 1985		2	Species	Taxom
Proctotrupidae Latreille, 1802	Proctotrupid wasps	15	Species	Taxom
Protapioceridae Ren, 1998		4	Species	Taxom
Protendipedidae Rohdendorf, 1962		1	Species	Taxom
Protomphralidae Rohdendorf, 1957		2	Species	Taxom
Protopleciidae Rohdendorf, 1946		5	Species	Taxom
Protorhyphidae Handlirsch, 1906		1	Species	Taxom
Pseudopulicidae Gao, Shih & Ren, 2012		3	Species	Taxom
Pseudosiricidae Handlirsch, 1908		1	Species	Taxom
Psychodidae Newman, 1834	Drain flies	20	Species	Taxom
Ptinidae Latreille, 1802	Death-watch and Spider Beetles	1	Species	Taxom
Ptismidae Kirejtshuk &..., 2016		1	Species	Taxom
Ptychopteridae Osten Sacken, 1862	Phantom crane flies	19	Species	Taxom
Pyrochroidae Latreille, 1807	Cardinal beetles	1	Species	Taxom
Radiophronidae Ortega-Blanc..., 2010		2	Species	Taxom
Rhagionemestriidae Ussatchov, 1968		4	Species	Taxom
Rhagionidae Latreille, 1802	Snipe flies	23	Species	Taxom
Rhipiphoridae Gemminger & ..., 1870	Hump back beetles	1	Species	Taxom
Rhopalosomatidae Ashmead, 1896	Rhopalosomatid wasps	1	Species	Taxom
Roeslerianidae Martins-Neto, 1997		1	Species	Taxom
Roproniidae Bradley, 1905	Roproniid wasps	5	Species	Taxom
Sapygidae Latreille, 1810	Club-horned wasps	1	Species	Taxom
Saurophthiridae Ponomarenko, 1986		2	Species	Taxom
Scarabaeidae Latreille, 1802	Dung beetles	56	Species	Taxom
Scatopsidae Newman, 1834	Dung midges	1	Species	Taxom
Scelionidae Haliday, 1839	Scelionid wasps	15	Species	Taxom
Schizophoridae Ponomarenko, 1968		3	Species	Taxom
Schizopodidae (LeConte, 1859)	False Jewel Beetles	1	Species	Taxom
Sclerogibbidae Ashmead, 1902	Sclerogibbid wasp	1	Species	Taxom
Scolebythidae Evans, 1963	Scolebythid wasps	6	Species	Taxom
Scoliidae Leach, 1815	Mammoth wasps	15	Species	Taxom
Scraptiidae Mulsant, 1856	Antlike leaf beetles	2	Species	Taxom
Scydmaenidae (Leach, 1815)	Ant-like stone beetles	1	Species	Taxom
Septiventeridae Bai & al., 2013		1	Species	Taxom
Sepulcidae Rasnitsyn, 1968		29	Species	Taxom
Serendipidae Evenhuis, 1994		3	Species	Taxom
Serphitidae Brues, 1937		4	Species	Taxom
Sesiidae Boisduval, 1828	Clear Wings	1	Species	Taxom
Silphidae Latreille, 1807	Burying And Carrion Beetles	0	Species	Taxom
Silvanidae Kirby, 1837	Flat bark beetles	1	Species	Taxom
Simuliidae Newman, 1834	Black flies	4	Species	Taxom
Sinosiricidae Hong You-Chong, 1975		1	Species	Taxom
Siricidae Billberg, 1820	Horntail	7	Species	Taxom
Sonidae Pritykina, 1986		1	Species	Taxom
Spathiopterygidae Engel & Orte..., 2013		3	Species	Taxom
Sphecidae Latreille, 1802	Digger & Sand wasps	1	Species	Taxom
Sphindidae Jacquelin du..., 1860	Cryptic slime mold beetle	5	Species	Taxom
Staphylinidae Lameere, 1900	Rove beetles	83	Species	Taxom
Stigmaphronidae Kozlov, 1975	Stigmaphronid wasps	11	Species	Taxom
Stratiomyidae Latreille, 1802	Soldier flies	4	Species	Taxom
Synteliidae Lewis, 1882		6	Species	Taxom
Syrphidae Latreille, 1802	Drone flies	2	Species	Taxom
Tabanidae Latreille, 1802	Breeze flies	8	Species	Taxom
Tachinidae (Robineau-Des..., 1830)	Bristle Flies	0	Species	Taxom
Taldycupedidae Rohdendorf, 1961		3	Species	Taxom
Tanyderidae Alexander, 1919	Primitive crane flies	2	Species	Taxom
Tenebrionidae Latreille, 1802	Darkling Beetle	4	Species	Taxom
Tenthredinidae Latreille, 1804	Common sawflies	3	Species	Taxom
Tethepomyiidae Grimaldi & A..., 2008		1	Species	Taxom
Tetratomidae Billberg, 1820	Polypore fungus beetle	1	Species	Taxom
Throscidae Laporte, 1840	False metallic wood-boring beetles	2	Species	Taxom
Thynnidae Shuckard, 1841	Thynnid Wasps	1	Species	Taxom
Tipulidae Latreille, 1802	Crane Flies	18	Species	Taxom
Trachypachidae C. G. Thomson, 1857	False ground beetle	23	Species	Taxom
Trichoceridae Rondani, 1841	Winter Crane Flies	9	Species	Taxom
Trigonalidae (Cresson, 1887)	Trigonalid sawflies	3	Species	Taxom
Trigonalyidae		1	Species	Taxom
Trogidae (MacLeay, 1819)	Gnawer beetles	10	Species	Taxom
Trogossitidae Latreille, 1802	Bark gnawing beetle	13	Species	Taxom
Ulyanidae (Zherikhin, 1993)		1	Species	Taxom
Undopterigidae (Kozlov, 1988)		1	Species	Taxom
Vermileonidae Williston, 1886	Wormlion	2	Species	Taxom
Vespidae Leach, 1815	And Allies	8	Species	Taxom
Vitimotauliidae Sukatsheva, 1968		25	Species	Taxom
Volitorididae Hong, 1982		1	Species	Taxom
Xyelidae Newman, 1835	Sawfly	32	Species	Taxom
Xyelotomidae Rasnitsyn, 1968		11	Species	Taxom
Xyelydidae Rasnitsyn, 1968		8	Species	Taxom
Xylomyidae Verrall, 1901	Wood soldier flies	1	Species	Taxom
Zhangsolvidae Nagatomi & Yang, 1998		5	Species	Taxom
Zygaenidae Latreille, 1809	Burnet Forester	1	Species	Taxom

Subnodes of Holometabola with occurrences in Lower Cretaceous

Order	Coleoptera Linnaeus, 1758 Beetles, Coleopterans	90 455 834	Families Genera Species	Taxom
Order	Diptera Linnaeus, 1758 Flies, Gnats, hoverflies, Mosquitoes, True flies, Two-Winged Flies	70 302 491	Families Genera Species	Taxom
Order	Hymenoptera Linnaeus, 1758 Ants, Bees, Sawflies, Wasps	73 288 538	Families Genera Species	Taxom
Order	Lepidoptera Linnaeus, 1758 Butterflies, Lepidopterans, Moths	10 14 15	Families Genera Species	Taxom
Order	Mecoptera Hyatt & Arms, 1891 Hanging flies, Hangingflies, Scorpion flies, Scorpionflies, snow scorpionflies, Snowflies	15 36 54	Families Genera Species	Taxom
Order	Trichoptera Kirby, 1813 Caddis-Flies, Caddisflies, Caddisfly, Cased Caddis, Cased Caddis-Flies, Caseless Caddis, Caseless Caddis-Flies, Rail-flies, Sedge-flies	18 37 150	Families Genera Species	Taxom

Links, sources and further information

Holometabola at itis.gov