Checklist for Superorder Holometabola Heider, 1889 (Holometabolous Insects)
and region Middle Pennsylvanian (Upper Carboniferous)
List of families for Middle Pennsylvanian
49 Families width 68 Species #ECLABSNFPTDOHIRXM
| Elateridae Leach, 1815 | Click Beetle | 1 | Species | Taxom | | Culicidae Meigen, 1818 | Mosquito | 1 | Species | Taxom | | Leiodidae Fleming, 1821 | Round fungus beetles | 1 | Species | Taxom | | Archaeolepiidae Whalley, 1985 | | 1 | Species | Taxom | | Belidae Schönherr, 1826 | Belid Beetle | 1 | Species | Taxom | | Stratiomyidae Latreille, 1802 | Soldier flies | 1 | Species | Taxom | | Curculionidae Latreille, 1802 | Ambrosia beetles | 6 | Species | Taxom | | Notodontidae Stephens, 1829 | Bag Shelter Moths | 8 | Species | Taxom | | Formicidae Latreille, 1809 | Ants | 2 | Species | Taxom | | Ptinidae Latreille, 1802 | Death-watch and Spider Beetles | 1 | Species | Taxom | | Tineidae Latreille, 1810 | & Allies | 1 | Species | Taxom | | Latridiidae Erichson, 1842 | Minute brown scavenger beetle | 1 | Species | Taxom | | Eremochaetidae Ussatchov, 1968 | | 1 | Species | Taxom | | Drosophilidae Rondani, 1856 | Fruit flies | 1 | Species | Taxom | | Protopleciidae Rohdendorf, 1946 | | 1 | Species | Taxom | | Tabanidae Latreille, 1802 | Breeze flies | 1 | Species | Taxom | | Oedemeridae Latreille, 1810 | False Blister Beetle | 1 | Species | Taxom | | Eucnemidae Eschscholtz, 1829 | Click Beetles | 1 | Species | Taxom | | Ceratopogonidae Newman, 1834 | Biting midge | 1 | Species | Taxom | | Staphylinidae Lameere, 1900 | Rove beetles | 1 | Species | Taxom | | Limoniidae Speiser, 1909 | Craneflies | 1 | Species | Taxom | | Helotidae Reitter, 1876 | | 1 | Species | Taxom | | Nemonychidae Bedel, 1882 | Nemonychid weevils | 1 | Species | Taxom | | Ichneumonidae Latreille, 1802 | Ichneumon flies | 1 | Species | Taxom | | Rhagionidae Latreille, 1802 | Snipe flies | 1 | Species | Taxom | | Cantharidae Imhoff, 1856 | Cantharids | 1 | Species | Taxom | | Noctuidae Latreille, 1809 | Armyworms | 1 | Species | Taxom | | Hydrophilidae Latreille, 1802 | Silver Water Beetles | 2 | Species | Taxom | | Tenebrionidae Latreille, 1802 | Darkling Beetle | 1 | Species | Taxom | | Scarabaeidae Latreille, 1802 | Dung beetles | 1 | Species | Taxom | | Cylindrotomidae Mulsant, 1842 | Cylindrotomid Crane Flies | 1 | Species | Taxom | | Ithyceridae Schönherr, 1826 | New York weevil | 1 | Species | Taxom | | Dryophthoridae Schönherr, 1825 | Palm weevils | 2 | Species | Taxom | | Palaeomanteidae Handlirsch, 1906 | | 1 | Species | Taxom | | Athericidae Stuckenberg, 1973 | Athericid flies | 1 | Species | Taxom | | Trogossitidae Latreille, 1802 | Bark gnawing beetle | 1 | Species | Taxom | | Sepulcidae Rasnitsyn, 1968 | | 1 | Species | Taxom | | Phryganeidae Leach, 1815 | Giant casemaker | 3 | Species | Taxom | | Pseudopolycentropodidae Handlirsch, 1925 | | 1 | Species | Taxom | | Ptiliidae Heer, 1843 | Feather-wing beetles | 1 | Species | Taxom | | Xyelydidae Rasnitsyn, 1968 | | 1 | Species | Taxom | | Micropterigidae Herrich-Schä..., 1855 | Mandibulate Archaic Moths | 1 | Species | Taxom | | Tipulidae Latreille, 1802 | Crane Flies | 2 | Species | Taxom | | Platystomatidae Schiner, 1862 | Picture-winged flies | 1 | Species | Taxom | | Ademosynidae Ponomarenko, 1968 | | 1 | Species | Taxom | | Srokalarvidae Haug & al., 2016 | | 1 | Species | Taxom | | Tenthredinidae Latreille, 1804 | Common sawflies | 1 | Species | Taxom | | Anthribidae Billberg, 1820 | Fungus Weevil | 1 | Species | Taxom | | Cleridae Latreille, 1802 | Checkered Beetle | 1 | Species | Taxom |
Genera for Middle Pennsylvanian
53 Genera with 68 Species #ABCDEGHKLMOPRST
| Adiaphtharsia Handlirsch, 1906 | | 1 | Species | Taxom | | Aedoeophasma Scudder, 1885 | | 1 | Species | Taxom | | Agogoblattina Handlirsch, 1885 | | 1 | Species | Taxom | | Archaeologus Handlirsch, 1906 | | 1 | Species | Taxom | | Archimulacris Dawson, 1868 | | 1 | Species | Taxom | | Asyncritus Handlirsch, 1911 | | 1 | Species | Taxom | | Balduria Meunier, 1922 | | 1 | Species | Taxom | | Breyeria Janse, 1964 | | 8 | Species | Taxom | | Bruayaphis Nel & al., 2012 | | 1 | Species | Taxom | | Cacurgellus Pruvost, 1919 | | 1 | Species | Taxom | | Chrestotes Scudder, 1868 | | 1 | Species | Taxom | | Diexodus Handlirsch, 1911 | | 1 | Species | Taxom | | Distasis Handlirsch, 1904 | | 1 | Species | Taxom | | Drybrookia Bolton, 1924 | | 1 | Species | Taxom | | Dytiscomorphus Fritsch, 1907 | | 1 | Species | Taxom | | Eopanorpella Schmidt, 1962 | | 1 | Species | Taxom | | Eumecoptera Handlirsch, 1877 | | 1 | Species | Taxom | | Eurytaenia Handlirsch, 1906 | | 1 | Species | Taxom | | Eurythmopteryx Handlirsch, 1906 | | 1 | Species | Taxom | | Gegenemene Handlirsch, 1883 | | 1 | Species | Taxom | | Gyrophlebia Handlirsch, 1906 | | 1 | Species | Taxom | | Haplophlebium Scudder, 1867 | | 1 | Species | Taxom | | Hemeristia Dana, 1864 | | 1 | Species | Taxom | | Hypermegethes Handlirsch, 1906 | | 2 | Species | Taxom | | Kustia Handlirsch, 1919 | | 1 | Species | Taxom | | Leptoblattina Woodward, 1887 | | 1 | Species | Taxom | | Litoneura Scudder, 1854 | | 1 | Species | Taxom | | Mammia Handlirsch, 1906 | | 1 | Species | Taxom | | Mecynoptera Schaus, 1904 | | 1 | Species | Taxom | | Megathentomum Scudder, 1906 | | 2 | Species | Taxom | | Melanoblattula Cockerell, 1927 | | 1 | Species | Taxom | | Metacheliphlebia Handlirsch, 1885 | | 1 | Species | Taxom | | Oryctomastax Pruvost, 1919 | | 1 | Species | Taxom | | Palaeocarria Cockerell, 1917 | | 1 | Species | Taxom | | Palaeodictyopteron Handlirsch, 1906 | | 2 | Species | Taxom | | Palaeomastax Handlirsch, 1904 | | 1 | Species | Taxom | | Palaiotaptus Handlirsch, 1906 | | 1 | Species | Taxom | | Paracridites Handlirsch, 1873 | | 1 | Species | Taxom | | Parapaolia Handlirsch, 1885 | | 1 | Species | Taxom | | Phryganea Linnaeus, 1901 | | 3 | Species | Taxom | | Polyernus Scudder, 1885 | | 1 | Species | Taxom | | Polyetes Handlirsch, 1934 | | 2 | Species | Taxom | | Pseudopolyernus Handlirsch, 1885 | | 1 | Species | Taxom | | Ptenodera Bolton, 1922 | | 1 | Species | Taxom | | Rectineura Bolton, 1934 | | 1 | Species | Taxom | | Sabitaptus Pruvost, 1930 | | 1 | Species | Taxom | | Scepasma Handlirsch, 1992 | | 2 | Species | Taxom | | Schuchertiella Handlirsch, 1911 | | 1 | Species | Taxom | | Sherborniella Handlirsch, 1934 | | 1 | Species | Taxom | | Srokalarva Haug & al., 2016 | | 1 | Species | Taxom | | Synaptoneura Konow, 1919 | | 1 | Species | Taxom | | Sypharoptera Handlirsch, 1911 | | 1 | Species | Taxom | | Titanodictya Handlirsch, 1885 | | 1 | Species | Taxom |
Species list for Middle Pennsylvanian
68 Species | Adiaphtharsia ferrea ♰ A. Handlirsch, 1906 | | | Aedoeophasma anglica Scudder, 1885 | | | Agogoblattina occidua ♰ S. H. Scudder, 1885 | | | Archaeologus falcatus ♰ Handlirsch, 1906 | | | Archimulacris acadica ♰ S. H. Scudder, 1868 | | | Asyncritus reticulatus ♰ A. Handlirsch, 1911 | | | Balduria archaica Meunier, 1922 | | | Breyeria bistrata ♰ C. Brauckmann, 1995 | | | Breyeria borinensis ♰ A. P. de Borre, 1875 | | | Breyeria brauckmanni ♰ I. D. Pinto, 1992 | | | Breyeria brevis ♰ C. Brauckmann, 2002 | | | Breyeria delruei ♰ D. Laurentiaux, 1949 | | | Breyeria rappi ♰ F. M. Carpenter, 1967 | | | Breyeria solida ♰ C. Brauckmann, 2000 | | | Breyeria vrankeni ♰ F. Laurentia..., 1964 | | | Bruayaphis oudardi ♰ A. Nel, 2012 | | | Cacurgellus barryi ♰ P. Pruvost, 1919 | | | Chrestotes lapidea ♰ S. H. Scudder, 1868 | | | Diexodus debilis ♰ A. Handlirsch, 1911 | | | Distasis rhipiphora ♰ A. Handlirsch, 1904 | | | Drybrookia cubitalis ♰ H. Bolton, 1924 | | | Dytiscomorphus larvalis ♰ A. Fritsch, 1907 | | | Eopanorpella ernsti ♰ W. Schmidt, 1962 | | | Eumecoptera laxa Goldenberg, 1877 | | | Eurytaenia virginiana ♰ A. Handlirsch, 1906 | | | Eurythmopteryx antiqua ♰ A. Handlirsch, 1906 | | | Gegenemene sinuosa ♰ M. Kliver, 1883 | | | Gyrophlebia longicollis A. Handlirsch, 1906 | | | Haplophlebium barnesii ♰ S. H. Scudder, 1867 | | | Hemeristia occidentalis ♰ J. D. Dana, 1864 | | | Hypermegethes northumbriae ♰ H. Bolton, 1916 | | | Hypermegethes schucherti ♰ A. Handlirsch, 1906 | | | Kustia bohemica ♰ A. Handlirsch, 1919 | | | Leptoblattina exilis ♰ H. Woodward, 1887 | | | Litoneura anthracophila ♰ F. Goldenberg, 1854 | | | Mammia alutacea ♰ A. Handlirsch, 1906 | | | Mecynoptera splendida ♰ A. Handlirsch, 1904 | | | Megathentomum pustulatum ♰ S. H. Scudder, 1868 | | | Megathentomum scudderi Handlirsch, 1906 | | | Melanoblattula nigrescens ♰ T. D. A. Coc..., 1927 | | | Metacheliphlebia elongata ♰ S. H. Scudder, 1885 | | | Oryctomastax duboisi Pruvost, 1919 | | | Palaeocarria ornata ♰ T. D. A. Coc..., 1917 | | | Palaeodictyopteron latipenne Handlirsch, 1906 | | | Palaeodictyopteron mazonum Handlirsch, 1906 | | | Palaeomastax carbonis ♰ A. Handlirsch, 1904 | | | Palaiotaptus mazonus ♰ A. Handlirsch, 1906 | | | Paracridites formosus ♰ F. Goldenberg, 1873 | | | Parapaolia superba ♰ S. H. Scudder, 1885 | | | Phryganea arenacea ♰ A Fritsch, 1901 | | | Phryganea kolbi ♰ A Fritsch, 1895 | | | Phryganea solitaria ♰ A Fritsch, 1901 | | | Polyernus complanatus ♰ S. H. Scudder, 1885 | | | Polyetes furcifer ♰ A. Handlirsch, 1906 | | | Polyetes problematica ♰ H. Bolton, 1934 | | | Pseudopolyernus laminarum ♰ S. H. Scudder, 1885 | | | Ptenodera dubius ♰ H. Bolton, 1922 | | | Rectineura lineata ♰ H. Bolton, 1934 | | | Sabitaptus lagagei ♰ P. Pruvost, 1930 | | | Scepasma gigas ♰ A. Handlirsch, 1911 | | | Scepasma mediomatricorum ♰ Brauckmann &..., 1992 | | | Schuchertiella gracilis ♰ A. Handlirsch, 1911 | | | Sherborniella hemingwayi ♰ H. Bolton, 1934 | | | Srokalarva berthei Haug & al., 2016 | | | Synaptoneura champyi ♰ P. Pruvost, 1919 | | | Sypharoptera pneuma Handlirsch, 1911 | | | Titanodictya jucunda ♰ S. H. Scudder, 1885 | |
Subnodes of Holometabola with occurrences in Middle Pennsylvanian | Order | Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 22
25
29 | Families
Genera
Species |
Taxom
| | Order | Diptera
Linnaeus, 1758
Flies, Gnats, hoverflies, Mosquitoes, True flies, Two-Winged Flies | 13
13
14 | Families
Genera
Species |
Taxom
| | Order | Hymenoptera
Linnaeus, 1758
Ants, Bees, Sawflies, Wasps | 5
6
6 | Families
Genera
Species |
Taxom
| | Order | Lepidoptera
Linnaeus, 1758
Butterflies, Lepidopterans, Moths | 5
5
12 | Families
Genera
Species |
Taxom
| | Order | Mecoptera
Hyatt & Arms, 1891
Hanging flies, Hangingflies, Scorpion flies, Scorpionflies, snow scorpionflies, Snowflies | 1
2
2 | Family
Genera
Species |
Taxom
| | Order | Miomoptera ♰
Martynov, 1928
| 1
1
1 | Family
Genus
Species |
Taxom
| | Order | Trichoptera
Kirby, 1813
Caddis-Flies, Caddisflies, Caddisfly, Cased Caddis, Cased Caddis-Flies, Caseless Caddis, Caseless Caddis-Flies, Rail-flies, Sedge-flies | 2
2
4 | Families
Genera
Species |
Taxom
|
Links, sources and further information Holometabola at itis.gov
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