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Checklist for Superorder Holometabola Heider, 1889 (Holometabolous Insects)
and region Upper Miocene (Miocene)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
102
243
280
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
101
242
279
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
91
216
229
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
68
182
196
Families
Genera
Species
 
Taxom
 


List of families for Upper Miocene
68 Families width 196 Species and 9 Subspecies
#PCBSFAEIGNHLTDMOKZ
Pamphiliidae Cameron, 1890Leaf-rolling sawflies2SpeciesTaxom
Carabidae Latreille, 1802Carabid beetles21SpeciesTaxom
Buprestidae Leach, 1815Flat-headed wood-borers5SpeciesTaxom
Scraptiidae Mulsant, 1856Antlike leaf beetles2SpeciesTaxom
Coccinellidae Latreille, 1807Coccinelles1SpeciesTaxom
Formicidae Latreille, 1809Ants32SpeciesTaxom
Scarabaeidae Latreille, 1802Dung beetles1SpeciesTaxom
Apidae Latreille, 1802Bumble bees8SpeciesTaxom
Pieridae Duponchel, 1835Brimstones3SpeciesTaxom
Erebidae Leach, 1815Erebid Moths3SpeciesTaxom
Staphylinidae Lameere, 1900Rove beetles5SpeciesTaxom
Bibionidae Fleming, 1821Bibionid flies7SpeciesTaxom
Bombyliidae Latreille, 1802Bee Flies4SpeciesTaxom
Ichneumonidae Latreille, 1802Ichneumon flies2SpeciesTaxom
Glaphyropteridae Pongracz, 19351SpeciesTaxom
Nymphalidae Rafinesque, 1815Admirals2SpeciesTaxom
Pteromalidae Dalman, 1820Pteromalid wasps2SpeciesTaxom
Noteridae C. G. Thomson, 1860Burrowing water beetle0SpeciesTaxom
Hydrophilidae Latreille, 1802Silver Water Beetles3SpeciesTaxom
Curculionidae Latreille, 1802Ambrosia beetles22SpeciesTaxom
Noctuidae Latreille, 1809Armyworms1SpeciesTaxom
Geometridae Leach, 1815Cankerworm moths3SpeciesTaxom
Lonchaeidae Loew, 1861Lance Flies0SpeciesTaxom
Chrysomelidae Latreille, 1802Casebearers4SpeciesTaxom
Tipulidae Latreille, 1802Crane Flies10SpeciesTaxom
Ptinidae Latreille, 1802Death-watch and Spider Beetles1SpeciesTaxom
Papilionidae Latreille, 1802Apollos1SpeciesTaxom
Dytiscidae Leach, 1815Diving beetles2SpeciesTaxom
Trichogrammatidae Haliday, 1851Trichogrammatid wasps1SpeciesTaxom
Melandryidae Leach, 1815False Darkling Beetle1SpeciesTaxom
Empididae Latreille, 1809Balloon flies1SpeciesTaxom
Latridiidae Erichson, 1842Minute brown scavenger beetle0SpeciesTaxom
Asilidae Latreille, 1802Assassin flies2SpeciesTaxom
Scydmaenidae Leach, 1815Ant-like stone beetles2SpeciesTaxom
Gelechiidae Stainton, 1854Gelechiid moths1SpeciesTaxom
Helophoridae Leach, 1815Water scavenger beetles1SpeciesTaxom
Otitidae Macquart, 1835Picture-winged flies1SpeciesTaxom
Crabronidae Latreille, 1802Bee Wolves1SpeciesTaxom
Lycaenidae Leach, 1815Blue butterflies1SpeciesTaxom
Keroplatidae Rondani, 1856Fungus gnats1SpeciesTaxom
Ptiliidae Heer, 1843Feather-wing beetles0SpeciesTaxom
Tephritidae Newman, 1834Fruit flies2SpeciesTaxom
Argidae Konow, 1890Argid flies1SpeciesTaxom
Lucanidae Latreille, 1804Stag Beetle1SpeciesTaxom
Gyrinidae Latreille, 1802Whirligig beetle1SpeciesTaxom
Byrrhidae Latreille, 1804Moss beetles2SpeciesTaxom
Meloidae Gyllenhal, 1810Blister Beetle1SpeciesTaxom
Mymarommatidae Debauche, 1948False fairy wasp1SpeciesTaxom
Zopheridae Solier, 1834Cylindrical bark beetles0SpeciesTaxom
Silphidae Latreille, 1807Burying And Carrion Beetles1SpeciesTaxom
Attelabidae Billberg, 1820Leaf Rolling Weevil2SpeciesTaxom
Scelionidae (Haliday, 1839)Scelionid wasps1SpeciesTaxom
Trogossitidae Latreille, 1802Bark gnawing beetle1SpeciesTaxom
Phryganeidae Leach, 1815Giant casemaker3SpeciesTaxom
Calliphoridae Brauer & Ber..., 1889Blow flies1SpeciesTaxom
Cerambycidae Latreille, 1802Long Horn beetles2SpeciesTaxom
Ptychopteridae Osten Sacken, 1862Phantom crane flies1SpeciesTaxom
Hepialidae Stephens, 1829Ghost Moths1SpeciesTaxom
Siricidae Billberg, 1820Horntail1SpeciesTaxom
Mycetophilidae Newman, 1834Fungus Gnat1SpeciesTaxom
Cecidomyiidae Newman, 1834Forest Midges1SpeciesTaxom
Ceratopogonidae Newman, 1834Biting midge1SpeciesTaxom
Sesiidae (Boisduval, 1828)Clear Wings1SpeciesTaxom
Therevidae (Newman, 1834)Stiletto flies0SpeciesTaxom
Pallopteridae Loew, 1862Flutter flies1SpeciesTaxom
Trachypachidae C. G. Thomson, 1857False ground beetle2SpeciesTaxom
Trichoceridae Rondani, 1841Winter Crane Flies1SpeciesTaxom
Scolebythidae Evans, 1963Scolebythid wasps4SpeciesTaxom


Genera for Upper Miocene
182 Genera with 196 Species
#ABCDEFGHIKLMNOPRSTUV
Acantholyda Costa, 17581SpeciesTaxom
Agonum Bonelli, 18441SpeciesTaxom
Agrilus Curtis, 19631SpeciesTaxom
Amara Bonelli, 1806Seed-Eating Ground Beetles1SpeciesTaxom
Anaspis Geoffroy, 18381SpeciesTaxom
Anatis Mulsant, 1932Giant Lady Beetles1SpeciesTaxom
Anthaxia Eschscholtz, 18701SpeciesTaxom
Aphaenogaster Mayr, 1960Collared Ants1SpeciesTaxom
Aphodius Hellwig, 19071SpeciesTaxom
Apis Linnaeus, 1960Honey Bee2SpeciesTaxom
Appias Hübner, 1862Albatrosses1SpeciesTaxom
Arctiites Rebel, 18981SpeciesTaxom
Arrhenopeplus Koch, 18281SpeciesTaxom
Bibio Geoffroy, 1883March flies3SpeciesTaxom
Blethisa Bonelli, 19831SpeciesTaxom
Bombus Latreille, 1991Bumble Bee3SpeciesTaxom
Bombylius Linnaeus, 1907Greater Bee Flies2SpeciesTaxom
Brachypimpla Strobl, 1902
0SpeciesTaxom
Buprestites Heer, 18371SpeciesTaxom
Callithomia Bates, 18601SpeciesTaxom
Campalita Motschulsky, 18601SpeciesTaxom
Camponotus Mayr, 1960Black carpenter ant3SpeciesTaxom
Canberrana Boucek, 19881SpeciesTaxom
Canthydrus Sharp, 1881
0SpeciesTaxom
Carabites Heer, 18921SpeciesTaxom
Carabus Linnaeus, 1998Ground beetle4SpeciesTaxom
Carebara Westwood, 18912SpeciesTaxom
Cataulacus Smith, 18912SpeciesTaxom
Catocala Schrank, 0Underwing moths1SpeciesTaxom
Cerdanyagrion Nel, Arillo ..., 19961SpeciesTaxom
Ceutorhynchus Germar, 1824
0SpeciesTaxom
Chalcophora Dejean, 19621SpeciesTaxom
Chrysodeixis Hübner, 18431SpeciesTaxom
Cleonolithus Bassi, 18411SpeciesTaxom
Coelostoma Brullé, 17751SpeciesTaxom
Colias Fabricius, 1891Clouded yellows1SpeciesTaxom
Crematogaster Lund, 1891Acrobat Ants1SpeciesTaxom
Ctenognophos Prout, 19661SpeciesTaxom
Curculionites Oppenheim, 18701SpeciesTaxom
Dasiops Rondani, 1856
0SpeciesTaxom
Diacheila Motschulsky, 18101SpeciesTaxom
Dicerca Eschscholtz, 20031SpeciesTaxom
Dicheirotrichus Jacquelin du..., 18271SpeciesTaxom
Dicladispa Gestro, 19881SpeciesTaxom
Dicranomyia Stephens, 19921SpeciesTaxom
Dicranoptycha Osten Sacken, 19921SpeciesTaxom
Dignomus Wollaston, 18071SpeciesTaxom
Dolichomitus Smith, 18291SpeciesTaxom
Donacia Fabricius, 18701SpeciesTaxom
Doritites Rebel, 18981SpeciesTaxom
Dyschirius Bonelli, 18131SpeciesTaxom
Dytiscus Linnaeus, 18621SpeciesTaxom
Ectatomma Smith, 18911SpeciesTaxom
Elaphropoda Lieftinck, 19661SpeciesTaxom
Elaphropus Motschoulsky, 19871SpeciesTaxom
Elaphrus Fabricius, 1758Marsh Ground Beetles1SpeciesTaxom
Emeria Skalski, 19881SpeciesTaxom
Emmesa Newman, 19001SpeciesTaxom
Empis Linnaeus, 18371SpeciesTaxom
Enicmus C. G. Thomson, 1859
0SpeciesTaxom
Eremisca Hull, 19641SpeciesTaxom
Euconnus Thomson, 19321SpeciesTaxom
Eurylychnus Bates, 01SpeciesTaxom
Exoteleia Wallengren, 19181SpeciesTaxom
Formica Linnaeus, 1935And Field Ants3SpeciesTaxom
Georissus Latreille, 1809
Minute mud-loving beetles0SpeciesTaxom
Geranomyia Haliday, 19861SpeciesTaxom
Gonioctena Motschulsky, 1870Broad-shouldered leaf beetles1SpeciesTaxom
Gonomyia Meigen, 19921SpeciesTaxom
Helophorus Fabricius, 18081SpeciesTaxom
Helopium Westwood, 18541SpeciesTaxom
Hemipenthes Loew, 19072SpeciesTaxom
Herina Robineau-Des..., 18261SpeciesTaxom
Hydaticus Leach, 18671SpeciesTaxom
Hydroporus Clairville, 1806
0SpeciesTaxom
Hypera Germar, 02SpeciesTaxom
Hyphantria Harris, 19121SpeciesTaxom
Hypopomyrmex Emery, 18911SpeciesTaxom
Incertogaster Boudinot, 18911SpeciesTaxom
Isochnus C.G. Thomson, 1865
0SpeciesTaxom
Kalissus LeConte, 18741SpeciesTaxom
Larrophanes Handlirsch, 19071SpeciesTaxom
Lasius Fabricius, 1870Citronella ants3SpeciesTaxom
Lathrobium Gravenhorst, 1802
0SpeciesTaxom
Ledouxnebria Deuve, 19981SpeciesTaxom
Leistus Frölich, 1799
0SpeciesTaxom
Lepidophorus W. Kirby, 18371SpeciesTaxom
Lepropus Schonherr, 1823
0SpeciesTaxom
Leptomyrmula Emery, 18911SpeciesTaxom
Leptotyphlus Fauvel, 19841SpeciesTaxom
Limonia Meigen, 19921SpeciesTaxom
Lixus Fabricius, 19931SpeciesTaxom
Lonchomyrmex Mayr, 18701SpeciesTaxom
Lycaenites Rebel, 18991SpeciesTaxom
Macrocera Meigen, 19721SpeciesTaxom
Martintella Artigas, 19011SpeciesTaxom
Megarhynchaenus Kohring, 19891SpeciesTaxom
Meliponorytes Tosi, 18961SpeciesTaxom
Mesochorus Gravenhorst, 19991SpeciesTaxom
Micridium Motschulsky, 1869
0SpeciesTaxom
Microlynx Kohring & Sc..., 19891SpeciesTaxom
Micropeplus Latreille, 19702SpeciesTaxom
Mioacinia Gentilini & ..., 20061SpeciesTaxom
Mioarge Nel, 20041SpeciesTaxom
Miocenidorcus Riou, 19991SpeciesTaxom
Miodineutes Hatch, 19271SpeciesTaxom
Morychus Erichson, 17751SpeciesTaxom
Mylabris Fabricius, 19351SpeciesTaxom
Mymaromella Girault, 19901SpeciesTaxom
Namunaria Reitter, 1882
0SpeciesTaxom
Napeogenes Bates, 18991SpeciesTaxom
Nebria Latreille, 1802
Gazelle Beetles0SpeciesTaxom
Necrodes Leach, 19271SpeciesTaxom
Notaris Germar, 18821SpeciesTaxom
Notiophilus Dumeril, 17581SpeciesTaxom
Oecophylla Smith, 1891Green ants2SpeciesTaxom
Olophrum Erichson, 1839
0SpeciesTaxom
Oreda White, 19161SpeciesTaxom
Paleauletobius Legalov, 18471SpeciesTaxom
Paridris Kieffer, 20001SpeciesTaxom
Patrobus Dejean, 18281SpeciesTaxom
Peltis Müller, 19351SpeciesTaxom
Pheidole Westwood, 1849Big-headed Ants1SpeciesTaxom
Philonthus Stephens, 1829
0SpeciesTaxom
Phryganea Linnaeus, 18833SpeciesTaxom
Phytophtirites Pampaloni, 19021SpeciesTaxom
Pilaria Sintenis, 19921SpeciesTaxom
Pinacopteryx Wallengren, 18191SpeciesTaxom
Plagiolepis Mayr, 18911SpeciesTaxom
Plateumaris C.G. Thomson, 2001reed leaf beetle1SpeciesTaxom
Platypodidarum Kohring & Sc..., 19891SpeciesTaxom
Platypus Herbst, 18381SpeciesTaxom
Plecia Wiedemann, 1935love bug4SpeciesTaxom
Podomyrma Smith, 18911SpeciesTaxom
Pollenia Robineau-Des..., 1930Attic flies1SpeciesTaxom
Polyrhachis Smith, 20091SpeciesTaxom
Ponera Latreille, 18911SpeciesTaxom
Prionus Geoffroy, 1999Giant root borers1SpeciesTaxom
Probittacomorpha Freiwald & W..., 20121SpeciesTaxom
Problongos Mérit & Mérit, 20081SpeciesTaxom
Protainophthalmus Zherikhin, 18561SpeciesTaxom
Protohepialus Pierce, 19451SpeciesTaxom
Proximoxeris Riou, 19911SpeciesTaxom
Pseudolimnophila Alexander, 19831SpeciesTaxom
Pseudoschoenionta Breuning, 18941SpeciesTaxom
Psiloptera Dejean, 19351SpeciesTaxom
Pterostichus Bonelli, 1851Woodland Ground Beetle3SpeciesTaxom
Rhynchaenus Clairville &..., 18271SpeciesTaxom
Rhyssomatus Schönherr, 19651SpeciesTaxom
Rhytideres Schönherr, 18561SpeciesTaxom
Scardamia Guenée, 18921SpeciesTaxom
Sciara Meigen, 19611SpeciesTaxom
Scolytus Geoffroy, 2000True fir bark beetles2SpeciesTaxom
Scraptia Latreille, 18381SpeciesTaxom
Sequoiomyia Möhn, 19601SpeciesTaxom
Serafimea Marinescu & ..., 19991SpeciesTaxom
Sicelomyrmex Wheeler, 18911SpeciesTaxom
Sicilomyrmex Wheeler, 18911SpeciesTaxom
Simplocaria Stephens, 19031SpeciesTaxom
Sitophilus Schönherr, 2000Granary Weevils1SpeciesTaxom
Solenopsis Westwood, 2017Fire ants1SpeciesTaxom
Sphenophorus Schoenherr, 1935Billbug Weevils1SpeciesTaxom
Stilobezzia Kieffer, 19591SpeciesTaxom
Symplecta Meigen, 19861SpeciesTaxom
Tachinus Gravenhorst, 1802
0SpeciesTaxom
Tapholyda Rasnitsyn, 19331SpeciesTaxom
Tapinoma Förster, 18911SpeciesTaxom
Technomyrmex Mayr, 18911SpeciesTaxom
Teleosphecia Le Cerf, 19161SpeciesTaxom
Tephritis Latreille, 20061SpeciesTaxom
Thanatophilus Leach, 1815
0SpeciesTaxom
Thereva Latreille, 1796
Stiletto flies0SpeciesTaxom
Tipula Linnaeus, 2012Cranefly2SpeciesTaxom
Tournotaris Alonso-Zaraz..., 17871SpeciesTaxom
Toxeuma Walker, 18481SpeciesTaxom
Toxoneura Macquart, 17801SpeciesTaxom
Trachypachus Motschulsky, 18272SpeciesTaxom
Trichocera Meigen, 1986Winter gnat1SpeciesTaxom
Trigona Jurine, 1896Vulture bees1SpeciesTaxom
Ulmeriella Meunier, 19834SpeciesTaxom
Veraphis Casey, 1897
0SpeciesTaxom
Vitavitus Kissinger, 19731SpeciesTaxom


Species list for Upper Miocene
196 Species and 9 Subspecies
Acantholyda (Acantholyda) erythrocephala Linnaeus, 1758Pine false webworm
Agonum quinquepuncatum Motschulsky, 1844
Agrilus syrphus Descarpentri..., 1963
Amara (Celia) quenseli Schönherr, 1806
Anaspis antica Guérin-Ménev..., 1838
Anatis gortanii Fiori, 1932
Anthaxia buschi Assmann, 1870
Aphaenogaster pannonica Bachmayer, 1960
Aphodius bosniaskii A. Handlirsch, 1907
Apis (Apis) melisuga A. Handlirsch, 1907
Apis catanensis Roussy, 1960
Appias paulina galathea Felder, 1862
Arctiites deletus Rebel, 1898
Arrhenopeplus tesserula (Curtis, 1828)⇒ Arrhenoplus tesserula Curtis, 1828
Bibio boulayi B. Riou, 1999
Bibio joursacensis Theobald, 1935
Bibio kochii Staub, 1883
Blethisa inexpectata (Goulet & Sme..., 1983)⇒ Blethisa multipunctata aurata Fischer von ..., 1828
Bombus cerdanyensis Dehon & al., 2014
Bombus coironensis B. Riou, 1999
Bombus vetustus A. P. Rasnitsyn, 1991
Bombylius fossilis A. Handlirsch, 1907
Bombylius tertiarius A. Handlirsch, 1907
Buprestites carbonum Germar, 1837
Callithomia lenea xantho C. & R. Felder, 1860
Campalita deplanatum O. Heer, 1860
Camponotus crozei B. Riou, 1999
Camponotus obesus L. Piton, 1935
Camponotus ullrichi F. Bachmayer, 1960
Canberrana hypera Boucek, 1988
Carabites capellinii Cecconi, 1892
Carabus (Diocarabus) truncaticollis Eschscholtz, 1833
Carabus (Tachypus) cancellatus Illiger, 1798Dwarf sand verbena
Carabus foveolatus (L. Piton, 1935)⇒ Carabus cribratus cribratus Quensel, 1806
Carabus neli T. Deuve, 1998
Carebara schossnicensis Assmann, 1870
Carebara sophiae Emery, 1891
Cataulacus planiceps Emery, 1891
Cataulacus silvestrii Emery, 1891
Catocala actaea actaea
Cerdanyagrion miocenicus A. Nel, 1996
Chalcophora espanoli Fernández de..., 1962
Chrysodeixis (Chrysodeixis) eriosoma Doubleday, 1843Cabbage semi-looper
Cleonolithus antiquus C. A. Bassi, 1841
Coelostoma (Coelostoma) orbiculare Fabricius, 1775
Colias cocandica maja (Grum-Grshima..., 1891)⇒ Colias cocandica Erschoff, 1874
Crematogaster praecursor Emery, 1891
Ctenognophos eolaria anax Wiltshire, 1966
Curculionites silesiacus A. Assmann, 1870
Diacheila arctica Gyllenhal, 1810
Dicerca reticulatoides Bellamy, 2003
Dicheirotrichus (Trichocellus) cognatus Gyllenhal, 1827
Dicladispa muratensis A. Nel, 1988
Dicranomyia sergio W. Krzemi..., 1992
Dicranoptycha anna W. Krzemi..., 1992
Dignomus (Dignomus) dilophus Illiger, 1807
Dolichomitus messor messor Gravenhorst, 1829
Donacia letzneri A. Assmann, 1870
Doritites bosniaskii Rebel, 1898
Dyschirius dyschirius (Bonelli, 1813)⇒ Dyschirius bonellii Putzeys, 1846
Dytiscus avunculus C.Heyden, 1862
Ectatomma gracile Emery, 1891
Elaphropoda bembidion Lieftinck, 1966Cascade cave amphipod
Elaphropus (Tachyura) leptothorax Baehr, 1987
Elaphrus (Trichelaphrus) riparius Linnaeus, 1758
Emeria simetitia Skalski, 1988
Emmesa colorata Scudder, 1900
Empis carbonum E. F. Germar, 1837
Eremisca dipogon Lehr, 1964
Euconnus (Tetramelus) siculus Fiori, 1932
Eurylychnus dyschiriodes
Exoteleia oribatella Rebel, 1918
Formica auxillacensis L. Piton, 1935
Formica cantalica L. Piton, 1935
Formica maculipennis N. Theobald, 1935
Geranomyia messiniana G. Gentilini, 1986
Gonioctena primordialis A. Assmann, 1870
Gonomyia andrea W. Krzemi..., 1992
Helophorus (Kyphohelophorus) tuberculatus Gyllenhal, 1808
Helopium agabus J. O. Westwood, 1854
Hemipenthes gabbroensis Handlirsch, 1907
Hemipenthes tertiarius Handlirsch, 1907
Herina lugubris Meigen, 1826
Hydaticus laevipennis (C. G. Thomson, 1867)⇒ Hydaticus aruspex Clark, 1864
Hypera (Hypera) nigrirostris Fabricius, 1775Black-beaked Green Weevil
Hypera nigrirostres
Hyphantria penthetria Dyar, 1912
Hypopomyrmex bombiccii C. Emery, 1891
Incertogaster praecursor Emery, 1891
Kalissus nitidus LeConte, 1874
Larrophanes ophthalmicus A. Handlirsch, 1907
Lasius chambonensis Piton & Théo..., 1935
Lasius crispus Wilson, 1955
Lasius oblongus Assmann, 1870Little-file macoma
Ledouxnebria brisaci T. Deuve, 1998
Lepidophorus lineaticollis Kirby, 1837
Leptomyrmula maravignae C. Emery, 1891
Leptotyphlus omessae mutilus Orousset, 1984
Limonia luca W. Krzemi..., 1992
Lixus (Eulixus) balazuci J. F. Voisin, 1993
Lonchomyrmex nigritus A. Assmann, 1870Little-file macoma
Lycaenites gabbroensis Rebel, 1899
Macrocera nephrotoma Matile, 1972
Martintella lestes Williston, 1901
Megarhynchaenus schalowae Kohring, 1989
Meliponorytes succini A. Tosi, 1896
Mesochorus flexus Schwenke, 1999
Microlynx franzbesucheti Kohring & Sc..., 1989
Micropeplus hoogendorni J. V. Matthews, 1970
Micropeplus hopkinsi J. V. Matthews, 1970
Mioacinia adriatica Gentilini & ..., 2006
Mioarge azari A. Nel, 2004
Miocenidorcus andancensis B. Riou, 1999
Miodineutes heeri Hatch, 1927
Morychus aeneus Fabricius, 1775
Mylabris punctatissima N. Theobald, 1935
Mymaromella duerrenfeldi Schlüter & K..., 1990
Napeogenes tolosa amara Godman, 1899Tolosa Tigerwing
Necrodes tricostata Hatch, 1927
Notaris eversmanni Faust, 1882
Notiophilus aquaticus Linnaeus, 1758
Oecophylla grandimandibula B. Riou, 1999
Oecophylla sicula C. Emery, 1891
Oreda hylastes Heller, 1916
Paleauletobius silenus Heer, 1847
Paridris stenus (Kononova & P..., 2000)⇒ Paridris stena
Patrobus (Patrobus) septentrionis Dejean, 1828
Peltis minuscula L. Piton, 1935
Pheidole pallidula Nylander, 1849Cumberland leafshell
Phryganea antiqua O Heer, 1849
Phryganea kryshtofovichi T. D. A. Coc..., 1923
Phryganea parschlugiana O Heer, 1883
Phytophtirites disodilis L. Pampaloni, 1902
Pilaria castellariana W. Krzemi..., 1992
Pinacopteryx eriphia eriphia Godart, , 1819Zebra White
Plagiolepis labilis Emery, 1891
Plateumaris kinugasana Hayashi, 2001
Platypodidarum ferrarae Kohring & Sc..., 1989
Platypus maravignae F. E. Guérin..., 1838
Plecia baglii G. Gentilini, 1991
Plecia castellaroi G. Gentilini, 1991
Plecia pisaurensis G. Gentilini, 1991
Plecia vergnei Piton, 1935
Podomyrma mayri Emery, 1891
Pollenia ibalia Seguy, 1930
Polyrhachis annosus Wappler & al., 2009
Ponera leptocephala Emery, 1891
Prionus rhodanensis B. Riou, 1999
Probittacomorpha brisaci W. Krzemi..., 2012
Problongos baudiliensis X. Mérit, 2008
Protainophthalmus asperulus O. Heer, 1856
Protohepialus comstocki Pierce, 1945
Proximoxeris tripoliensis A. Nel, 1991
Pseudolimnophila siciliana W. Krzemi..., 1983
Pseudoschoenionta libellula Jordan, 1894
Psiloptera martyi L. Piton, 1935
Pterostichus (Melanius) corvinus Dejean, 1828Bynoe's goby
Pterostichus (Steroperis) costatus Ménétriés, 1851
Pterostichus (Steroperis) vermiculosus Ménétriés, 1851
Rhynchaenus velaris (Gyllenhal, 1827)⇒ Pelenomus velaris Gyllenhal, 1827
Rhyssomatus miocenae Pierce, 1965
Rhytideres sexsulcatus O. Heer, 1856
Scardamia chrysolina Meyrick, 1892
Sciara schremmeri F. Bachmayer, 1961
Scolytus duplicatus A. V. Petrov, 2000
Scolytus tshernjaki A. V. Petrov, 2000
Scraptia ovata F. E. Guérin..., 1838
Sequoiomyia krauseli E. Möhn, 1960
Serafimea bergeri F. Marinescu, 1999
Sicelomyrmex corniger Emery, 1891
Sicilomyrmex corniger C. Emery, 1891
Simplocaria basalis Sahlberg, 1903
Sitophilus punctatissimus V. V. Zherikhin, 2000
Solenopsis atavina Perfilieva, ..., 2017
Sphenophorus elegans N. Theobald, 1935
Stilobezzia (Acanthohelea) acrotrichis Tokunaga, 1959
Symplecta (Psiloconopa) savchenkoi G. Gentilini, 1986
Tapholyda caplani T. D. A. Coc..., 1933
Tapinoma minutissimum Emery, 1891
Technomyrmex deletus Emery, 1891
Teleosphecia bibio (Le Cerf, 1916)⇒ Teleosphecia unicolor Walker, 1864
Tephritis sophus Gentilini & ..., 2006
Tipula maclurei S. H. Scudder, 1894
Tipula paleopannonia M. S. Engel, 2012
Tournotaris bimaculatus Fabricius, 1787
Toxeuma acilius Walker, 1848
Toxoneura muliebris Harris, 1780
Trachypachus holmbergi Mannerheim, 1853Bigtooth aspen
Trachypachus zetterstedti Gyllenhal, 1827
Trichocera fujiyamai G. Gentilini, 1986
Trigona sicula A. Tosi, 1896
Ulmeriella martynovi F. Zeuner, 1938
Ulmeriella pliocenica A. Nel, 1987
Ulmeriella shizukuishiensis I. Fujiyama, 1983
Ulmeriella uemurai I. Fujiyama, 1983
Vitavitus thulius D. G. Kissinger, 1973

Subnodes of Holometabola with occurrences in Upper Miocene
OrderColeoptera
Linnaeus, 1758
Beetles, Coleopterans
28
93
85
Families
Genera
Species

Taxom

OrderDiptera
Linnaeus, 1758
Flies, Gnats, hoverflies, Mosquitoes, True flies, Two-Winged Flies
17
29
35
Families
Genera
Species

Taxom

OrderHymenoptera
Linnaeus, 1758
Ants, Bees, Sawflies, Wasps
12
42
56
Families
Genera
Species

Taxom

OrderLepidoptera
Linnaeus, 1758
Butterflies, Lepidopterans, Moths
10
17
17
Families
Genera
Species

Taxom

OrderTrichoptera
Kirby, 1813
Caddis-Flies, Caddisflies, Caddisfly, Cased Caddis, Cased Caddis-Flies, Caseless Caddis, Caseless Caddis-Flies, Rail-flies, Sedge-flies
1
1
3
Family
Genus
Species

Taxom



Links, sources and further information
Holometabola at itis.gov