Checklist for Subfamily Aphodiinae Leach, 1815 (Aphodiine Dung Beetles)
and region East Asia (Far East)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 1146
26296
148959 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 1142
26273
148905 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 1054
25859
147577 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 627
19594
119086 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 179
7304
50129 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 147
6442
41727 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 18
706
6144 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 18
706
6144 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 518
4689 | Genera
Species |
Taxom
| | Subfamily | Aphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles | 127
674 | Genera
Species |
Taxom
|
Genera for East Asia
127 Genera with 674 Species #ABCDEGHKLMNOPQRSTV
| Acanthobodilus Dellacasa, 1922 | | 2 | Species | Taxom | | Acrossidius Schmidt, 1892 | | 4 | Species | Taxom | | Acrossus Mulsant, 1961 | | 38 | Species | Taxom | | Adeloparius Schmidt, 1942 | | 1 | Species | Taxom | | Aegialia Latreille, 1986 | | 9 | Species | Taxom | | Aganocrossus Reitter, 1862 | | 3 | Species | Taxom | | Agoliinus Schmidt, 1981 | | 12 | Species | Taxom | | Agolius Mulsant & Rey, 2014 | | 3 | Species | Taxom | | Agrilinus Mulsant & Rey, 1962 | | 26 | Species | Taxom | | Airapus Stebnicka & ..., 2021 | | 1 | Species | Taxom | | Alocoderus Schmidt, 1887 | | 3 | Species | Taxom | | Amidorus Mulsant & Rey, 1913 | | 4 | Species | Taxom | | Ammoecius Mulsant, 1960 | | 2 | Species | Taxom | | Aparammoecius Petrovitz, 2017 | | 10 | Species | Taxom | | Aphodaulacus Kozhantschikov, 1875 | | 8 | Species | Taxom | | Aphodiellus Schmidt, 1875 | | 1 | Species | Taxom | | Aphodiites Heer, 1865 | | 1 | Species | Taxom | | Aphodius Hellwig, 1951 | | 102 | Species | Taxom | | Aphodobius Péringuey, 1945 | | 1 | Species | Taxom | | Ataenius Harold, 1904 | | 10 | Species | Taxom | | Australaphodius Balthasar, 1892 | | 1 | Species | Taxom | | Bodiloides | | 1 | Species | Taxom | | Bodilopsis Adám, 0 | | 7 | Species | Taxom | | Bodilus Mulsant & Rey, 1975 | | 10 | Species | Taxom | | Boucomontiellus Balthasar, 1913 | | 2 | Species | Taxom | | Brachiaphodius Koshantschikov, 2000 | | 9 | Species | Taxom | | Caelius Lewis, 1887 | | 2 | Species | Taxom | | Calamosternus Motschulsky, 1994 | | 7 | Species | Taxom | | Carinaulus Tesar, 2000 | | 18 | Species | Taxom | | Chilothorax Motschulsky, 2001 | | 34 | Species | Taxom | | Cnemisus Motschulsky, 1903 | | 3 | Species | Taxom | | Colobopterus Mulsant, 1847 | | 4 | Species | Taxom | | Coprimorphus Mulsant, 1789 | | 1 | Species | Taxom | | Coptochiroides Balthasar, 0 | | 2 | Species | Taxom | | Craterocyphus Schmidt, 1847 | | 1 | Species | Taxom | | Cryptoscatomaseter Gordon and S..., 2001 | | 1 | Species | Taxom | | Dialytes Harold, 1845 | | 2 | Species | Taxom | | Diastictus Mulsant, 1805 | | 1 | Species | Taxom | Diasticus Mulsant, 1842 | | 0 | Species | Taxom | | Didactylia Orbigny, 1961 | | 1 | Species | Taxom | | Erytus Mulsant & Rey, 1892 | | 3 | Species | Taxom | | Esymus Mulsant & Rey, 1932 | | 4 | Species | Taxom | | Eudolus Mulsant & Rey, 1783 | | 1 | Species | Taxom | | Euheptaulacus Dellacasa, 1806 | | 4 | Species | Taxom | | Euorodalus Dellacasa, 1945 | | 2 | Species | Taxom | | Eupleurus Mulsant, 1758 | | 3 | Species | Taxom | | Gilletianus Balthasar, 2015 | | 11 | Species | Taxom | | Gonaphodius Reitter, 1877 | | 1 | Species | Taxom | | Granulopsammodius Rakovic, 1984 | | 4 | Species | Taxom | | Guanyinaphodius | | 1 | Species | Taxom | | Heptaulacus Mulsant, 1775 | | 2 | Species | Taxom | | Koreoxyomus | | 1 | Species | Taxom | | Koshantschikovius Schmidt, 1983 | | 1 | Species | Taxom | | Labarrus Mulsant and Rey, 1860 | | 6 | Species | Taxom | | Leiopsammodius Rakovic, 2012 | | 6 | Species | Taxom | | Limarus Mulsant & Rey, 1813 | | 2 | Species | Taxom | | Liothorax Motschulsky, 1767 | | 2 | Species | Taxom | | Loraphodius Reitter, 1878 | | 1 | Species | Taxom | | Loraspis Mulsant & Rey, 1870 | | 1 | Species | Taxom | | Melinopterus Mulsant, 1798 | | 7 | Species | Taxom | | Mendidius Harold, 1941 | | 8 | Species | Taxom | | Mesontoplatys Motschulsky, 1875 | | 1 | Species | Taxom | | Messyrhus RakoviÄ & K..., 1997 | | 1 | Species | Taxom | | Mozartius Nomura & Nakane, 0 | | 11 | Species | Taxom | | Myrhessus Balthasar, 2006 | | 5 | Species | Taxom | | Neagolius Kozhantschikov, 1997 | | 10 | Species | Taxom | | Neocalaphodius Bordat, 1801 | | 1 | Species | Taxom | | Neotrichiorhyssemus RakoviÄ & K..., 2013 | | 5 | Species | Taxom | | Nettelislasia Martínez, 1945 | | 1 | Species | Taxom | | Nialus Mulsant & Rey, 1805 | | 2 | Species | Taxom | | Nimbus Mulsant & Rey, 1823 | | 3 | Species | Taxom | | Nipponaphodius Nakane, 1951 | | 1 | Species | Taxom | | Nipponoagoliinus | | 1 | Species | Taxom | | Nobius Mulsant & Rey, 1799 | | 2 | Species | Taxom | | Odochilus Harold, 2016 | | 2 | Species | Taxom | | Orodaliscus Reitter, 1906 | | 2 | Species | Taxom | | Oromus Mulsant & Rey, 1763 | | 1 | Species | Taxom | | Otophorus Mulsant, 1758 | | 1 | Species | Taxom | | Oxyomus Dejean, 2014 | | 7 | Species | Taxom | | Palnia Stebnicka, 1999 | | 1 | Species | Taxom | | Paracrossidius Balthasar, 1963 | | 10 | Species | Taxom | | Paradidactylia Balthasar, 1941 | | 6 | Species | Taxom | | Parammoecius Seidlitz, 1960 | | 4 | Species | Taxom | | Paulianellus Balthasar, 2015 | | 5 | Species | Taxom | Petrovitzius RakoviÄ, 1979 | | 0 | Species | Taxom | | Phaeaphodius Reitter, 1866 | | 5 | Species | Taxom | | Phalacronothus Motschulsky, 1952 | | 5 | Species | Taxom | | Pharaphodius Reitter, 1875 | | 7 | Species | Taxom | | Plagiogonus Mulsant, 1951 | | 9 | Species | Taxom | | Planolinellus Dellacasa an..., 1825 | | 2 | Species | Taxom | | Planolinoides Dellacasa an..., 1827 | | 2 | Species | Taxom | | Planolinus Mulsant and Rey, 1951 | | 2 | Species | Taxom | | Platyderides Schmidt, 1963 | | 5 | Species | Taxom | | Platytomus Mulsant, 2006 | | 3 | Species | Taxom | | Pleuraphodius Schmidt, 1875 | | 1 | Species | Taxom | | Pleurophorus Mulsant, 1933 | | 5 | Species | Taxom | | Psammodius Fallen, 1980 | | 16 | Species | Taxom | | Psammoporus Thomson, 1986 | | 11 | Species | Taxom | | Pseudacrossus Reitter, 1922 | | 13 | Species | Taxom | | Pseudoahermodontus | | 1 | Species | Taxom | | Pseudoheptaulacus Dellacasa & ..., 1909 | | 1 | Species | Taxom | | Pubinus Mulsant & Rey, 1776 | | 1 | Species | Taxom | | Qingaphodius | | 2 | Species | Taxom | | Rakovicius Pittino, 2019 | | 7 | Species | Taxom | | Renaudius Balthasar, 1942 | | 1 | Species | Taxom | | Rhodaphodius | | 1 | Species | Taxom | | Rhyparus Westwood, 2019 | | 23 | Species | Taxom | | Rhyssemodes Reitter, 2017 | | 6 | Species | Taxom | | Rhyssemus Mulsant, 1951 | | 19 | Species | Taxom | | Saprosites Redtenbacher, 2019 | | 6 | Species | Taxom | | Scabrostomus Gordon and S..., 1991 | | 1 | Species | Taxom | | Setylaides | | 3 | Species | Taxom | | Sigorus Mulsant & Rey, 1792 | | 1 | Species | Taxom | | Silluvia Landin, 2008 | | 8 | Species | Taxom | | Sinaphodius | | 1 | Species | Taxom | | Sinodiapterna Dellacasa, 1991 | | 6 | Species | Taxom | | Stenotothorax Schmidt, 0 | | 2 | Species | Taxom | | Subrinus Mulsant & Rey, 1870 | | 1 | Species | Taxom | Sugrames Reitter, 1894 | | 0 | Species | Taxom | | Sybacodes Fairmaire, 0 | | 3 | Species | Taxom | | Termitodiellus Nakane, 2020 | | 1 | Species | Taxom | | Teuchestes Mulsant, 2006 | | 6 | Species | Taxom | | Trichiorhyssemus Clouët, 2007 | | 13 | Species | Taxom | | Trichonotulus Bedel, 1941 | | 4 | Species | Taxom | | Tristaphodius Balthasar, 1932 | | 2 | Species | Taxom | | Turanella Semenov, 1887 | | 1 | Species | Taxom | | Volinus Mulsant & Rey, 1798 | | 1 | Species | Taxom |
Species list for East Asia
674 Species and 37 Subspecies | Acanthobodilus immundus Creutzer, 1799 | | | Acanthobodilus languidulus Schmidt, 1922 | | | Acrossidius delavayi Koshantschikov, 1916 | | | Acrossidius kozlovi Koshantschikov, 1916 | | | Acrossidius tasmaniae Hope, 1847 | Black-headed pasture cockchafer | | Acrossidius yorkensis Blackburn, 1892 | | | Acrossus atratus Waterhouse, 1875 | | | Acrossus belousovi Frolov, 2001 | | | Acrossus binaevulus Heyden, 1887 | | | Acrossus byki Minkina, 2018 | | | Acrossus delavayi Koshantschikov, 1916 | | | Acrossus depressus Kugelann, 1792 | | | Acrossus formosanus Nomura & Nakane, 1951 | | | Acrossus haliki Balthasar, 1933 | | | Acrossus histrio Balthasar, 1932 | | | Acrossus hrubanti Balthasar, 1961 | | | Acrossus humerospinosus Petrovitz, 1958 | | | Acrossus igai Nakane, 1956 | | | Acrossus igai pseudoigai Nakane & Mas..., 1967 | | | Acrossus igai sakuraii Nakane & Mas..., 1967 | | | Acrossus igai tsurugianus Nakane & Mas..., 1967 | | | Acrossus impressiusculus Fairmaire, 1888 | | | Acrossus japonicus Nomura & Nakane, 1951 | | | Acrossus jedlickai Balthasar, 1932 | | | Acrossus jeloneki Minkina, 2018 | | | Acrossus klickai Balthasar, 1932 | | | Acrossus koreanensis Kim, 1986 | | | Acrossus kwanhsiensis Balthasar, 1945 | | | Acrossus laticapitus Zhang, 1992 | | | Acrossus luridus Fabricius, 1775 | | | Acrossus melanodiscus Zhang, 1992 | | | Acrossus obenbergeri Balthasar, 1932 | | | Acrossus opacipennis Schmidt, 1910 | | | Acrossus pseudoobenbergeri Cervenka, 1995 | | | Acrossus ritsemai Schmidt, 1909 | | | Acrossus rufipes Linnaeus, 1758 | Night-Flying Dung Beetle | | Acrossus semiopacus Reitter, 1887 | | | Acrossus sichuanensis Zhang, 1992 | | | Acrossus superatratus Nomura & Nakane, 1951 | | | Acrossus transmontanus Balthasar, 1932 | | | Acrossus unifasciatus Nomura & Nakane, 1951 | | | Acrossus unifasciatus kiiensis Nakane, 1960 | | | Acrossus viturati Reitter, 1907 | | | Acrossus yunlingensis Petrovitz, 1961 | | | Adeloparius formosus Paulian, 1942 | | | Aegialia abdita Nikritin, 1975 | | | Aegialia comis Lewis, 1895 | | | Aegialia friebi (Balthasar, 1935) | ⇒ Psammoporus friebi Balthasar, 1935 | | Aegialia kamtschaticus Motschulsky, 1860 | | | Aegialia koreana Kim, 1979 | | | Aegialia nakanei Masumoto, 1986 | | | Aegialia nakanei horii Ochi & Kawahara, 2012 | | | Aegialia nitida Waterhouse, 1875 | | | Aegialia tsukamotoi Masumoto, 1986 | | | Aganocrossus amoenus Boheman, 1857 | | | Aganocrossus postpilosus Reitter, 1895 | | | Aganocrossus urostigma Harold, 1862 | | | Agoliinus cruentatus LeConte, 1878 | | | Agoliinus kiuchii Masumoto, 1984 | | | Agoliinus lapponum Gyllenhal, 1808 | Northern Dung Beetle | | Agoliinus morii Nakane, 1983 | | | Agoliinus parastorkani Cervenka, 1995 | | | Agoliinus piceus Gyllenhal, 1808 | | | Agoliinus pseudostorkani Stebnicka, 1982 | | | Agoliinus satyrus Reitter, 1892 | | | Agoliinus setchan Masumoto, 1984 | | | Agoliinus shibatai Nakane, 1983 | | | Agoliinus storkani Balthasar, 1932 | | | Agoliinus tanakai Masumoto, 1981 | | | Agolius abdominalis Bonelli, 1812 | | | Agolius moronensis Frolov, 2009 | | | Agolius petri Frolov, 2014 | | | Agrilinus ater De Geer, 1774 | | | Agrilinus bardus Balthasar, 1946 | | | Agrilinus bollowi Balthasar, 1941 | | | Agrilinus breviusculus Motschulsky, 1866 | | | Agrilinus convexus Erichson, 1848 | | | Agrilinus hasegawai Nomura & Nakane, 1951 | | | Agrilinus hasegawai aka Nakane, 1960 | | | Agrilinus hasegawai akahane Nakane, 1960 | | | Agrilinus hayachinensis Nomura & Nakane, 1951 | | | Agrilinus ishidai Masumoto & K..., 1987 | | | Agrilinus lungaiensis Petrovitz, 1962 | | | Agrilinus madara Nakane, 1960 | | | Agrilinus matusiaki Minkina, 2022 | | | Agrilinus monikae Kral, 2013 | | | Agrilinus obliviosus Reitter, 1892 | | | Agrilinus pseudolungaiensis Mencl & Rakovic, 2012 | | | Agrilinus punctator Reitter, 1892 | | | Agrilinus ritsukoae Kawai, 2004 | | | Agrilinus sordidus (Fabricius, 1775) | ⇒ Bodilopsis sordida Fabricius, 1775 | | Agrilinus spinulosus Schmidt, 1910 | | | Agrilinus striatus Schmidt, 1910 | | | Agrilinus surdus Boucomont, 1929 | | | Agrilinus tenax Balthasar, 1932 | | | Agrilinus uniformis Waterhouse, 1875 | | | Agrilinus virginalis Reitter, 1900 | | | Agrilinus wassuensis Petrovitz, 1962 | | | Airapus rakovici Král & Lu, 2021 | | | Alocoderus elongatulus Fabricius, 1801 | | | Alocoderus holdereri Reitter, 1900 | | | Alocoderus semenowi Reitter, 1887 | | | Amidorus koshantschikovi Jacobson, 1911 | | | Amidorus obscurus Fabricius, 1792 | | | Amidorus serrimargo Koshantschikov, 1913 | | | Amidorus umbrinus Koshantschikov, 1913 | | | Ammoecius brevis Erichson, 1848 | | | Ammoecius yamato Nakane, 1960 | | | Aparammoecius balangensis Petrovitz, 1958 | | | Aparammoecius chuangkuohungi Masumoto, Ki..., 2017 | | | Aparammoecius gansuensis Stebnicka, 1997 | | | Aparammoecius isaburoi Nakane, 1956 | | | Aparammoecius mizo Nakane, 1967 | | | Aparammoecius nomurai Stebnicka, 1986 | | | Aparammoecius pallidiligonis Waterhouse, 1875 | | | Aparammoecius tarokensis Stebnicka, 1994 | | | Aparammoecius yenpingensis Stebnicka, 1986 | | | Aparammoecius yushanus Masumoto, Ki..., 2017 | | | Aphodaulacus grumi Frolov, 2002 | | | Aphodaulacus ignobilis Reitter, 1887 | | | Aphodaulacus kabaki Frolov, 2001 | | | Aphodaulacus koltzei Reitter, 1892 | | | Aphodaulacus nigrotessellatus Motschulsky, 1866 | | | Aphodaulacus perblandus Balthasar, 1933 | | | Aphodaulacus turkestanicus Heyden, 1881 | | | Aphodaulacus variabilis Waterhouse, 1875 | | | Aphodiellus impunctatus Waterhouse, 1875 | | | Aphodiites protogaeus ♰ Heer, 1865 | | | Aphodius (Acanthobodilus) immundus (Creutzer, 1799) | ⇒ Pubinus tomentosus O.F.Muller, 1776 | | Aphodius (Acrossoides) yorkensis (Blackburn, 1892) | ⇒ Acrossidius yorkensis Blackburn, 1892 | | Aphodius (Acrossus) formosanus Nomura & Nakane, 1951 | | | Aphodius (Acrossus) rufipes (Linnaeus, 1758) | ⇒ Acrossus rufipes Linnaeus, 1758 | | Aphodius (Aganocrossus) urostigma Harold, 1862 | | | Aphodius (Ahermodontus) marini Baguena, 1930 | | | Aphodius (Aparammoecius) balangensis Petrovitz, 1958 | | | Aphodius (Aparammoecius) nishikawai (Masumoto, 1996) | ⇒ Aparammoecius tarokensis Stebnicka, 1994 | | Aphodius (Aparammoecius) tarokensis Stebnicka, 1994 | | | Aphodius (Aphodiellus) impunctatus (Waterhouse, 1875) | ⇒ Melinopterus consputus Creutzer, 1799 | | Aphodius (Aphodius) elegans Allibert, 1847 | | | Aphodius (Aphodius) fimetarius Linnaeus, 1758 | Common dung beetle | | Aphodius (Aphodius) foetidus Herbst, 1783 | | | Aphodius (Aphodius) pedellus (DeGeer, 1774) | ⇒ Aphodius fimetarius Linnaeus, 1758 | | Aphodius (Aphodius) variabilis (C. O. Waterh..., 1875) | ⇒ Aphodaulacus nigrotessellatus Motschulsky, 1866 | | Aphodius (Balthasarianus) babori Balthasar, 1938 | | | Aphodius (Balthasarianus) pilifer Paulian, 1934 | | | Aphodius (Bodilus) gregarius Harold, 1871 | | | Aphodius (Bodilus) punctipennis Erichson, 1848 | | | Aphodius (Bodilus) sordidus (Fabricius, 1775) | ⇒ Bodilopsis rufus Moll, 1782 | | Aphodius (Calamosternus) granarius (Linnaeus, 1767) | ⇒ Calamosternus granarius Linnaeus, 1767 | | Aphodius (Calamosternus) sublimbatus Motschulsky, 1860 | | | Aphodius (Calamosternus) trucidatus (Chevrolat, 1863) | ⇒ Calamosternus trucidatus Harold, 1863 | | Aphodius (Calaphodius) moestus (Fabricius, 1801) | ⇒ Calamosternus granarius Linnaeus, 1767 | | Aphodius (Carinaulus) pucholti Balthasar, 1961 | | | Aphodius (Colobopteroides) latus A.Schmidt, 1911 | | | Aphodius (Colobopterus) erraticus (Linnaeus, 1758) | ⇒ Colobopterus erraticus Linnaeus, 1758 | | Aphodius (Diapterna) hamatus Say, 1824 | | | Aphodius (Euorodalus) pusillus (Herbst, 1789) | ⇒ Esymus pusillus Herbst, 1789 | | Aphodius (Eupleurus) subterraneus (Linnaeus, 1758) | ⇒ Eupleurus subterraneus Linnaeus, 1758 | | Aphodius (Gonaphodius) scoparius Harold, 1877 | | | Aphodius (Loboparius) kotoensis Miwa, 1930 | | | Aphodius (Melinopterus) prodromus (Braham, 1790) | ⇒ Melinopterus prodromus Brahm, 1790 | | Aphodius (Nialus) insularis (Petrovitz, 1961) | ⇒ Labarrus hoabinhensis Balthasar, 1946 | | Aphodius (Nialus) plagiatus (Linnaeus, 1758) | ⇒ Liothorax plagiatus Linnaeus, 1767 | | Aphodius (Nialus) sturmi (Harold, 1870) | ⇒ Subrinus sturmi Harold, 1870 | | Aphodius (Oromus) alpinus (Scopoli, 1763) | ⇒ Euheptaulacus carinatus carinatus Germar, 1824 | | Aphodius (Otophorus) haemorrhoidalis (Linnaeus, 1758) | ⇒ Otophorus haemorrhoidalis Linnaeus, 1758 | | Aphodius (Paremadus) isaburoi Nakane, 1967 | | | Aphodius (Paulianellus) maderi Balthasar, 1938 | | | Aphodius (Phaeaphodius) rectus Motschulsky, 1866 | | | Aphodius (Phalacronothus) pseudoavunculus Masumoto, 1977 | | | Aphodius (Pharaphodius) marginellus (Fabricius, 1781) | ⇒ Ataenius luteomargo Chapin, 1940 | | Aphodius (Pharaphodius) priscus Motschulsky, 1858 | | | Aphodius (Plagiogonus) arenarius Olivier, 1789 | | | Aphodius (Plagiogonus) putridus (Herbst, 1789) | ⇒ Planolinoides borealis Gyllenhal, 1827 | | Aphodius (Planolinellus) vittatus Mulsant, 1842 | | | Aphodius (Planolinoides) borealis (Gyllenhal, 1827) | ⇒ Planolinoides borealis Gyllenhal, 1827 | | Aphodius (Pleuraphodius) lewisi C. O. Waterh..., 1875 | | | Aphodius (Sinodiapterna) troitzkyi Jacobson, 1897 | | | Aphodius (Sinodiapterna) zeni Ochi, 1991 | | | Aphodius (Teuchestes) analis (Gistel, 1857) | ⇒ Rhodaphodius foetens Fabricius, 1787 | | Aphodius (Trichaphodius) reichei Harold, 1859 | | | Aphodius (Trichonotulus) scrofa (Fabricius, 1787) | ⇒ Trichonotulus scrofa Fabricius, 1787 | | Aphodius anteactus ♰ Krell, 2000 | | | Aphodius apicalis (Schilsky, 1888) | ⇒ Calamosternus granarius Linnaeus, 1767 | | Aphodius armatus (Roth, 1851) | ⇒ Craterocyphus rhinocerus Reiche, 1847 | | Aphodius australasiae (Boheman, 1858) | ⇒ Ataenius castanicolor Motschulsky, 1858 | | Aphodius bifasciatus (Dalla Torre, 1879) | ⇒ Chilothorax melanostictus Schmidt, 1840 | | Aphodius bonariensis Klug, 1829 | | | Aphodius calichromus Balthasar, 1932 | | | Aphodius caminarius (Faldermann, 1835) | ⇒ Pseudacrossus tenebricosus Schmidt, 1922 | | Aphodius cancelliventris (Motschulsky, 1858) | ⇒ Ataenius australasiae Boheman, 1858 | | Aphodius castanicolor (Motschulsky, 1858) | ⇒ Ataenius castanicolor Motschulsky, 1858 | | Aphodius catulus (Balthasar, 1946) | ⇒ Australaphodius frenchi Blackburn, 1892 | | Aphodius chokaiensis (Nomura & Nakane, 1951) | ⇒ Gilletianus comatus Schmidt, 1920 | | Aphodius corallifer Koshantschikov, 1913 | | | Aphodius cruentatus (Dejean, 1833) | ⇒ Planolinellus vittatus Say, 1825 | | Aphodius ealaensis (Endrödi, 1956) | ⇒ Adeloparius formosus Paulian, 1942 | | Aphodius edai (Balthasar, 1956) | ⇒ Phaeaphodius rectus Motschulsky, 1866 | | Aphodius epotus (Dellacasa, 1983) | ⇒ Chilothorax conspurcatus Linnaeus, 1758 | | Aphodius fasciger Harold, 1881 | | | Aphodius gestroi (Harold, 1877) | ⇒ Ataenius australasiae Boheman, 1858 | | Aphodius guangdongensis Maté, 2008 | | | Aphodius haroldianus (Balthasar, 1932) | ⇒ Colobopterus quadratus Reiche, 1847 | | Aphodius hibernalis Nakane & Tsu..., 1956 | | | Aphodius igai Nakane, 1956 | | | Aphodius illigeri (Dallatorre, 1879) | ⇒ Bodilopsis rufus Moll, 1782 | | Aphodius informis (Mulsant, 1842) | ⇒ Acrossus luridus Fabricius, 1775 | | Aphodius irakensis (Baraud, 1977) | ⇒ Erytus opacior Koshantschikov, 1894 | | Aphodius irregularis (Westwood, 1839) | ⇒ Chilothorax paykulli Bedel, 1907 | | Aphodius jengi Masumoto, Ki..., 2014 | | | Aphodius lapponicus (Villa, 1833) | ⇒ Oromus alpinus Scopoli, 1763 | | Aphodius madari (Tesar, 1945) | ⇒ Nimbus makowskyi Koshantschikov, 1891 | | Aphodius mara (Krikken & Kaas, 1984) | ⇒ Didactylia szunyoghyi Endrödi, 1961 | | Aphodius marginatus (LeConte, 1878) | ⇒ Cryptoscatomaseter depressiusculus Schmidt, 1907 | | Aphodius miniatulus (Balthasar, 1931) | ⇒ Agoliinus lapponum Gyllenhal, 1808 | | Aphodius mizo Nakane, 1967 | | | Aphodius multiplex Reitter, 1897 | | | Aphodius nasutus (Dellacasa, 1983) | ⇒ Chilothorax melanostictus Schmidt, 1840 | | Aphodius nigerrimus (Dallatorre, 1879) | ⇒ Oromus alpinus Scopoli, 1763 | | Aphodius nigrotessellatus (Nakane, 1972) | ⇒ Chilothorax punctatus Waterhouse, 1875 | | Aphodius obsoleteguttatus (Waterhouse, 1875) | ⇒ Aphodaulacus nigrotessellatus Motschulsky, 1866 | | Aphodius plasoni Kaufel, 1914 | | | Aphodius postpilosus Reitter, 1895 | | | Aphodius przewalskyi Reitter, 1887 | | | Aphodius reginae Kral, 1997 | | | Aphodius rudii Endrodi, 1968 | | | Aphodius rugostriatus (Reitter, 1892) | ⇒ Pharaphodius rugosostriatus Waterhouse, 1875 | | Aphodius snowi (Brown, 1929) | ⇒ Agoliinus cruentatus LeConte, 1878 | | Aphodius subcostatus H. J. Kolbe, 1886 | | | Aphodius unifasciatus Nomura & Nakane, 1951 | | | Aphodobius amplipennis Balthasar, 1945 | | | Ataenius australasiae Boheman, 1858 | | | Ataenius castanicolor Motschulsky, 1858 | | | Ataenius fukiensis (Balthasar, 1942) | ⇒ Ataenius australasiae Boheman, 1858 | | Ataenius luteomargo Chapin, 1940 | | | Ataenius malayianus (Petrovitz, 1961) | ⇒ Ataenius australasiae Boheman, 1858 | | Ataenius okinawensis (Nakane, 1960) | ⇒ Ataenius australasiae Boheman, 1858 | | Ataenius peregrinator Harold, 1877 | Birthworts | | Ataenius picinus Harold, 1861 | Antarctic blennies | | Ataenius striatocrenatus Fairmaire, 1889 | | | Ataenius tweedensis (Blackburn, 1904) | ⇒ Ataenius castanicolor Motschulsky, 1858 | | Australaphodius frenchi Blackburn, 1892 | Little pasture cockchafer | | Bodiloides ictericus Laicharting, 1781 | | | Bodilopsis aquilus Schmidt, 1916 | | | Bodilopsis rufa Moll, 1782 | | | Bodilopsis rufus Moll, 1782 | | | Bodilopsis sordida Fabricius, 1775 | | | Bodilopsis sordidus Fabricius, 1775 | | | Bodilopsis sordidus changajicus Endrödi, 1965 | | | Bodilopsis sordidus sordidus | | | Bodilus caspius Menestries, 1832 | | | Bodilus crassus Schmidt, 1916 | | | Bodilus gregarius Harold, 1871 | | | Bodilus kaznakovi Frolov, 2001 | | | Bodilus longeciliatus Reitter, 1887 | | | Bodilus nigriventris (Reitter, 1892) | ⇒ Bodilus lugens Creutzer, 1799 | | Bodilus punctipennis Erichson, 1848 | | | Bodilus rudii Endrödi, 1968 | | | Bodilus sordescens Harold, 1869 | | | Bodilus zunbajanicus Stebnicka, 1975 | | | Boucomontiellus pelliculosus Balthasar, 1932 | | | Boucomontiellus umbrinus Koshantschikov, 1913 | | | Brachiaphodius babori Balthasar, 1938 | | | Brachiaphodius eccoptus Bates, 1889 | | | Brachiaphodius jendeki Kral, 2000 | | | Brachiaphodius pilifer Paulian, 1934 | | | Brachiaphodius pilosus Harold, 1874 | | | Brachiaphodius ruupbai Masumoto, 1991 | | | Brachiaphodius taiwanicus Petrovitz, 1976 | | | Brachiaphodius weibao Kral, 2000 | | | Brachiaphodius yuan Kral, 2000 | | | Caelius denticollis (Lewis, 1895) | ⇒ Silluvia elongata Landin, 1949 | | Caelius rufescens Horn, 1887 | | | Calamosternus boreosinicus Cervenka, 1994 | | | Calamosternus granarius Linnaeus, 1767 | Granular Small Dung Beetle | | Calamosternus prusai Tesar, 1945 | | | Calamosternus sauteri Petrovitz, 1961 | | | Calamosternus trucidatus Harold, 1863 | | | Calamosternus uniplagiatus Waterhouse, 1875 | | | Calamosternus zmoskev Cervenka, 1994 | | | Carinaulus armandidavidi Cervenka, 2000 | | | Carinaulus coreensis Kim, 1986 | | | Carinaulus dundarum Cervenka, 2000 | | | Carinaulus gansuicus Cervenka, 2000 | | | Carinaulus imitabilis Cervenka, 2000 | | | Carinaulus inexpectatus Balthasar, 1935 | | | Carinaulus jakli Cervenka, 2000 | | | Carinaulus krali Cervenka, 2000 | | | Carinaulus meghalayensis Cervenka, 2000 | | | Carinaulus milaninikodymi Cervenka, 2000 | | | Carinaulus nanxiensis Masumoto, Ki..., 2018 | | | Carinaulus pseudovseteckai Cervenka, 2003 | | | Carinaulus pseudozhanglaensis Cervenka, 2003 | | | Carinaulus pucholti Balthasar, 1961 | | | Carinaulus tryznai Cervenka, 2000 | | | Carinaulus vseteckai Tesar, 1945 | | | Carinaulus wrzecionkoi Cervenka, 2000 | | | Carinaulus zhanglaensis Cervenka, 2000 | | | Chilothorax alexis Frolov, 2001 | | | Chilothorax altaicus Nikolajev, 1984 | | | Chilothorax bistriga Reitter, 1900 | | | Chilothorax chandmanicus Frolov, 2001 | | | Chilothorax comma Reitter, 1892 | | | Chilothorax conspurcatus Linnaeus, 1758 | | | Chilothorax distinctus (Müller, 1776) | ⇒ Aphodius distinctus O.F. Müller, 1776 | | Chilothorax dzongensis Petrovitz, 1976 | | | Chilothorax flavimargo Reitter, 1901 | | | Chilothorax glebi Frolov, 2006 | | | Chilothorax grafi Reitter, 1901 | | | Chilothorax hozuensis Ochi, Tsukam..., 2015 | | | Chilothorax huangyuanensis Cervenka, 1994 | | | Chilothorax jacobsoni Koshantschikov, 1911 | | | Chilothorax kardonensis Huang & Du, 1999 | | | Chilothorax kerzhneri Nikolajev, 1984 | | | Chilothorax kukunorensis Semenov, 1898 | | | Chilothorax lieni Masumoto, Ki..., 2014 | | | Chilothorax melanostictus Schmidt, 1840 | | | Chilothorax melanostictus melanostictus W. Schmidt, 1840 | | | Chilothorax mongolaltaicus Nikolajev, 1984 | | | Chilothorax nigrivittis impubis Koshantschikov, 1913 | | | Chilothorax nigrivittis kongurensis Ochi, Tsukam..., 2015 | | | Chilothorax nigrivittis nigrivittis | | | Chilothorax ohishii Masumoto, 1975 | | | Chilothorax okadai Nakane, 1951 | | | Chilothorax okadai kiritachiensis Ochi, Kawaha..., 2012 | | | Chilothorax okadai okadai | | | Chilothorax paykulli Bedel, 1907 | | | Chilothorax potanini Frolov, 2001 | | | Chilothorax punctatus Waterhouse, 1875 | | | Chilothorax shansianus Nakane & Shi..., 1957 | | | Chilothorax sinicus Cervenka, 1994 | | | Chilothorax tanhensis Frolov, 2001 | | | Cnemisus kaszabi (Endrödi, 1964) | ⇒ Cnemisus kaznakovi Semenov, 1903 | | Cnemisus kaznakovi Semenov, 1903 | | | Cnemisus kozlovi Semenov, 1903 | | | Colobopterus erraticus Linnaeus, 1758 | Erratic small dung beetle | | Colobopterus indagator Mannerheim, 1849 | | | Colobopterus propraetor Balthasar, 1932 | | | Colobopterus quadratus Reiche, 1847 | | | Coprimorphus scrutator Herbst, 1789 | | | Coptochiroides subcostatus Kolbe, 1886 | | | Coptochiroides subcostatus subcostatus | | | Craterocyphus rhinocerus Reiche, 1847 | | | Cryptoscatomaseter gambrinus Skelley & Go..., 2001 | Warteye stargazer | | Dialytes foveatus (Schmidt, 1909) | ⇒ Setylaides foveatus Schmidt, 1909 | | Dialytes truncatus Melsheimer, 1845 | | | Diastictus vulneratus Sturm, 1805 | | | Didactylia szunyoghyi Endrödi, 1961 | | | Erytus aequalis Schmidt, 1907 | | | Erytus opacior Koshantschikov, 1894 | | | Erytus pruinosus Reitter, 1892 | | | Esymus merdarius Fabricius, 1775 | | | Esymus pusillus Herbst, 1789 | | | Esymus pusillus pusillus Herbst, 1789 | | | Esymus pusillus roubali Balthasar, 1932 | | | Eudolus quadriguttatus Herbst, 1783 | | | Euheptaulacus carinatus Germar, 1824 | | | Euheptaulacus carinatus carinatus Germar, 1824 | | | Euheptaulacus sus Herbst, 1783 | | | Euheptaulacus villosus Gyllenhal, 1806 | | | Euorodalus paracoenosus Balthasar & ..., 1960 | | | Euorodalus stribrnyi Tesar, 1945 | | | Eupleurus antiquus Faldermann, 1835 | | | Eupleurus subterraneus Linnaeus, 1758 | Subterranean small dung beetle | | Eupleurus subterraneus subterraneus Linnaeus, 1758 | | | Gilletianus atsushii Ochi, 1986 | | | Gilletianus comatus Schmidt, 1920 | | | Gilletianus commatoides Balthasar, 1961 | | | Gilletianus costatulus Schmidt, 1908 | | | Gilletianus fukiensis Balthasar, 1952 | | | Gilletianus kratochvili Balthasar, 1941 | | | Gilletianus nigrovirgatus Schmidt, 1911 | | | Gilletianus proclivis Balthasar, 1933 | | | Gilletianus reichei Harold, 1859 | | | Gilletianus segmentaroides Schmidt, 1909 | | | Gilletianus yunnanensis Král, Rakovi..., 2015 | | | Gonaphodius scoparius Harold, 1877 | | | Granulopsammodius mongol Endrödi, 1969 | | | Granulopsammodius plicatuloides Pittino, 1984 | | | Granulopsammodius sinicus Rakovic, 1986 | | | Granulopsammodius tienshanicus Pittino, 1984 | | | Guanyinaphodius kishimotoi Masumoto & K..., 2001 | | | Heptaulacus carinatus (Gebler, 1833) | ⇒ Euheptaulacus carinatus Germar, 1824 | | Heptaulacus testudinarius Fabricius, 1775 | | | Koreoxyomus koreanus Kim, 1996 | | | Koshantschikovius chovdgolensis Endrödi, 1983 | | | Labarrus hoabinhensis Balthasar, 1946 | | | Labarrus lividus Olivier, 1789 | Scarab beetle | | Labarrus lividus lividus | | | Labarrus nabeleki Balthasar, 1936 | | | Labarrus rigidus Balthasar, 1936 | | | Labarrus sublimbatus Motschulsky, 1860 | | | Leiopsammodius chifengleei Kiuchi & Tsai, 2018 | | | Leiopsammodius gestroi Clouët, 1900 | | | Leiopsammodius indicus Harold, 1877 | | | Leiopsammodius japonicus Harold, 1878 | | | Leiopsammodius laevicollis Klug, 1845 | | | Leiopsammodius nomurai Masumoto, 2012 | | | Limarus maculatus Sturm, 1800 | | | Limarus zenkeri Germar, 1813 | | | Liothorax niger Illiger, 1798 | Beaulieu dung beetle | | Liothorax plagiatus Linnaeus, 1767 | | | Loraphodius kashmirensis Sharp, 1878 | | | Loraspis frater Mulsant & Rey, 1870 | | | Melinopterus consputus Creutzer, 1799 | | | Melinopterus prodromus Brahm, 1790 | Spring small dung beetle | | Melinopterus punctatosulcatus Sturm, 1805 | | | Melinopterus punctatosulcatus hirtipes Fischer von ..., 1844 | | | Melinopterus reyi Reitter, 1892 | | | Melinopterus scuticollis Semenov, 1898 | | | Melinopterus sphacelatus Panzer, 1798 | | | Mendidius bidens Solsky, 1876 | | | Mendidius bispinifrons Reitter, 1889 | | | Mendidius bivittatus Schmidt, 1922 | | | Mendidius fimbriolatus Mannerheim, 1849 | | | Mendidius gobiensis Endrödi, 1964 | | | Mendidius multiplex Reitter, 1897 | | | Mendidius nelsinae Medvedev, 1968 | | | Mendidius pseudobidens Balthasar, 1941 | | | Mesontoplatys arabicus Harold, 1875 | | | Messyrhus brachypterus Rakovic & Kral, 1997 | | | Mozartius jugosus Lewis, 1895 | | | Mozartius jugosus horii Ochi, Kawaha..., 2006 | | | Mozartius jugosus jugosus | | | Mozartius jugosus shingoi Ochi, Kawaha..., 2006 | | | Mozartius kyushuensis Ochi & Kawah..., 2002 | | | Mozartius testaceus Nomura & Nakane, 1951 | | | Mozartius testaceus shikokuensis Masumoto, 1984 | | | Mozartius testaceus testaceus | | | Mozartius uenoi Masumoto, 1984 | | | Mozartius uenoi hadai Kawai, 2003 | | | Mozartius uenoi uenoi | | | Myrhessus kubotai Ochi, Kawaha..., 2012 | | | Myrhessus nishinoi Ochi, Kawaha..., 2012 | | | Myrhessus samurai Balthasar, 1941 | | | Myrhessus sunadai Ochi, Kawaha..., 2019 | | | Myrhessus yorikoae Ochi, Kawaha..., 2006 | | | Neagolius bilimecki Seidlitz, 1891 | | | Neagolius falcispinis Koshantschikov, 1912 | | | Neagolius haroldi Koshantschikov, 1894 | | | Neagolius hengduanensis Kral, 1997 | | | Neagolius montanus Erichson, 1848 | | | Neagolius montisjuldi Pittino, 1988 | | | Neagolius moronensis Frolov, 2009 | | | Neagolius petri Frolov, 2014 | | | Neagolius takin Kral, 1995 | | | Neagolius zangbo Kral, 1997 | | | Neocalaphodius moestus Fabricius, 1801 | | | Neotrichiorhyssemus boninensis Nakane, 1960 | | | Neotrichiorhyssemus esakii Nomura, 1943 | | | Neotrichiorhyssemus hauseri Balthasar, 1933 | | | Neotrichiorhyssemus hirsutus Clouët, 1901 | | | Neotrichiorhyssemus kentingensis Masumoto & al., 2013 | | | Nettelislasia euparinoides Islas, 1945 | | | Nialus nefandus Reitter, 1907 | | | Nialus varians Duftschmid, 1805 | | | Nimbus contaminatus Herbst, 1783 | | | Nimbus makowskyi Koshantschikov, 1891 | | | Nimbus obliteratus Panzer, 1823 | | | Nipponaphodius gotoi Nomura & Nakane, 1951 | | | Nipponoagoliinus yasutakai Ochi & Kawahara, 2001 | | | Nobius bosaki Tesar, 1945 | | | Nobius serotinus Panzer, 1799 | | | Odochilus convexus Nomura, 1971 | | | Odochilus taiwanus Masumoto, La..., 2016 | | | Orodaliscus rotundangulus Reitter, 1900 | | | Orodaliscus zangi Schmidt, 1906 | | | Oromus alpinus Scopoli, 1763 | | | Otophorus haemorrhoidalis Linnaeus, 1758 | Bloody small dung beetle | | Oxyomus alligator Ho, Hu & Fik..., 2022 | | | Oxyomus cameratus Schmidt, 1908 | | | Oxyomus ishidai Nakane, 1977 | | | Oxyomus masumotoi Nomura, 1973 | | | Oxyomus nanxiensis Masumoto, Ki..., 2018 | | | Oxyomus sylvestris Scopoli, 1763 | Forest small dung beetle | | Oxyomus taipingensis Masumoto, Ki..., 2014 | | | Palnia cambeforti Kral, 1999 | | | Paracrossidius impressiusculus Fairmaire, 1888 | | | Paracrossidius instigator Balthasar, 1932 | | | Paracrossidius kozlovi Koshantschikov, 1916 | | | Paracrossidius laticapitus Zhang, 1992 | | | Paracrossidius melanodiscus Zhang, 1992 | | | Paracrossidius planipennis Petrovitz, 1961 | | | Paracrossidius radekcervenkai Cervenka, 2017 | | | Paracrossidius sichuanensis Zhang, 1992 | | | Paracrossidius transmontanus Balthasar, 1932 | | | Paracrossidius truculentus Balthasar, 1963 | | | Paradidactylia avuncula Balthasar, 1946 | | | Paradidactylia avunculus Balthasar, 1946 | | | Paradidactylia carinulata Motschulsky, 1863 | | | Paradidactylia carinulatus Motschulsky, 1863 | | | Paradidactylia semiangulus Balthasar, 1952 | | | Paradidactylia shantungensis Balthasar, 1941 | | | Parammoecius corvinus Erichson, 1848 | | | Parammoecius gibbus Germar, 1817 | | | Parammoecius pyrenaeus Jacquelin du..., 1863 | | | Parammoecius yamato (Nakane, 1960) | ⇒ Ammoecius yamato Nakane, 1960 | | Paulianellus andrei Minkina & Král, 2018 | | | Paulianellus asahinai Nakane, 1951 | | | Paulianellus maderi Balthasar, 1938 | | | Paulianellus neomaderi Mencl, Král ..., 2015 | | | Paulianellus taiwanensis Mencl, Král ..., 2015 | | | Phaeaphodius costalis Gebler, 1848 | | | Phaeaphodius dauricus Harold, 1863 | | | Phaeaphodius kiulungensis Balthasar, 1932 | | | Phaeaphodius plutenkoi Kral, 2002 | | | Phaeaphodius rectus Motschulsky, 1866 | | | Phalacronothus biguttatus Germar, 1824 | | | Phalacronothus botulus Balthasar, 1945 | | | Phalacronothus pseudoavunculus Masumoto, 1977 | | | Phalacronothus quadrimaculatus Linnaeus, 1761 | | | Phalacronothus semiangulus Balthasar, 1952 | | | Pharaphodius attritus (Balthasar, 1933) | ⇒ Pharaphodius marginellus Fabricius, 1781 | | Pharaphodius crenatus Harold, 1862 | | | Pharaphodius marginellus Fabricius, 1781 | | | Pharaphodius orientalis Harold, 1862 | | | Pharaphodius priscus Motschulsky, 1858 | | | Pharaphodius putearius Reitter, 1895 | | | Pharaphodius rugosostriatus Waterhouse, 1875 | | | Plagiogonus arenarius Olivier, 1789 | | | Plagiogonus bortkevitschi Koshantschikov, 1916 | | | Plagiogonus burgaltaicus Csiki, 1901 | | | Plagiogonus chan Endrödi, 1967 | | | Plagiogonus culminarius Reitter, 1900 | | | Plagiogonus inflatus Balthasar, 1938 | | | Plagiogonus putridus (Geoffroy, 1785) | ⇒ Plagiogonus arenarius Olivier, 1789 | | Plagiogonus quadriclypealis Cervenka, 1995 | | | Plagiogonus shirahatai Nakane, 1951 | | | Planolinellus grzegorzi Minkina, 2017 | | | Planolinellus vittatus Say, 1825 | | | Planolinoides aenictus Cooper & Gordon, 1987 | Benttooths | | Planolinoides borealis Gyllenhal, 1827 | | | Planolinus fasciatus Olivier, 1789 | | | Planolinus pratensis Nomura & Nakane, 1951 | | | Platyderides arrowi Schmidt, 1910 | | | Platyderides jengi Masumoto, Ki..., 2014 | | | Platyderides klapperichi Balthasar, 1942 | | | Platyderides planipennis Petrovitz, 1961 | | | Platyderides truculentus Balthasar, 1963 | | | Platytomus mongolicus Medvedev, 1974 | | | Platytomus variolosus Kolenati, 1846 | | | Platytomus yadai Ochi, Kawaha..., 2006 | | | Pleuraphodius lewisi Waterhouse, 1875 | | | Pleurophorus apicipennis Reitter, 1892 | | | Pleurophorus caesus Creutzer, 1796 | Cut small dung beetle | | Pleurophorus formosanus Pittino & Ma..., 1986 | | | Pleurophorus pannonicus Petrovitz, 1961 | | | Pleurophorus tonkinensis Balthasar, 1933 | | | Psammodius asper (Fabricius, 1775) | ⇒ Psammodius sulcicollis Illiger, 1802 | | Psammodius convexus Waterhouse, 1875 | | | Psammodius coreanus (Kim, 1980) | ⇒ Rakovicius coreanus Kim, 1980 | | Psammodius flavolittoralis Kim, 1980 | | | Psammodius generosus (Reitter, 1892) | ⇒ Psammodius nocturnus Reitter, 1892 | | Psammodius hangangnensis Kim, 1980 | | | Psammodius inscitus (Walker, 1858) | ⇒ Rhyssemus inscitus Walker, 1858 | | Psammodius japonicus (Harold, 1878) | ⇒ Leiopsammodius japonicus Harold, 1878 | | Psammodius kobayashii Nomura, 1973 | | | Psammodius kondoi (Masumoto, 1984) | ⇒ Psammodius kobayashii Nomura, 1973 | | Psammodius laevicollis (Klug, 1845) | ⇒ Leiopsammodius laevicollis Klug, 1845 | | Psammodius maruyamai Ochi, Kawaha..., 2011 | | | Psammodius nocturnus Reitter, 1892 | | | Psammodius pierottii Pittino, 1979 | Mud blenny | | Psammodius sculpticollis (Fairmaire, 1897) | ⇒ Leiopsammodius indicus Harold, 1877 | | Psammodius sungshinarum Kim, 1980 | | | Psammoporus abdita Nikritin, 1975 | | | Psammoporus comis Lewis, 1895 | | | Psammoporus friebi Balthasar, 1935 | | | Psammoporus hybridus Reitter, 1892 | | | Psammoporus kamtschaticus Motschulsky, 1860 | | | Psammoporus latipunctus Gredler, 1866 | | | Psammoporus mimica Pittino, 2006 | | | Psammoporus nakanei (Masumoto, 1986) | ⇒ Aegialia kamtschaticus Motschulsky, 1860 | | Psammoporus nakanei horii Ochi & Kawahara, 2012 | | | Psammoporus nakanei nakanei | | | Psammoporus tsukamotoi Masumoto, 1986 | | | Pseudacrossus absconditus Balthasar, 1932 | | | Pseudacrossus grebenschikovi Balthasar, 1961 | | | Pseudacrossus grebenscikovi Balthasar, 1961 | | | Pseudacrossus grombczewskyi Koshantschikov, 1891 | | | Pseudacrossus juxtus Petrovitz, 1972 | | | Pseudacrossus kalabi Kral, 1997 | | | Pseudacrossus nasutus Reitter, 1887 | | | Pseudacrossus przewalskyi Reitter, 1887 | | | Pseudacrossus qinghaiensis Kral, 1997 | | | Pseudacrossus serrimargo Koshantschikov, 1913 | | | Pseudacrossus smetanai Kral, 2011 | | | Pseudacrossus subsericeus Ballion, 1878 | | | Pseudacrossus tenebricosus Schmidt, 1922 | | | Pseudoahermodontus benesi Dellacasa & ..., 1997 | | | Pseudoheptaulacus iniquus Schmidt, 1909 | | | Pubinus tomentosus O.F.Muller, 1776 | | | Qingaphodius nikodymi Kral, 1997 | | | Qingaphodius ochotonarum Kral, 1997 | | | Rakovicius ainu Lewis, 1895 | | | Rakovicius coreanus Kim, 1980 | | | Rakovicius kawaii (Pittino, 2006) | ⇒ Rakovicius subopacus Nomura, 1973 | | Rakovicius kyotoensis Ochi, Kawaha..., 2019 | | | Rakovicius lanae Masumoto, 2012 | | | Rakovicius subopacus Nomura, 1973 | | | Rakovicius takashii Ochi, Kawaha..., 2019 | | | Renaudius tesari Paulian, 1942 | | | Rhodaphodius foetens Fabricius, 1787 | | | Rhyparus amamianus (Nakane, 1956) | ⇒ Rhyparus helophoroides Fairmaire, 1893 | | Rhyparus australiae (Lea, 1923) | ⇒ Rhyparus helophoroides Fairmaire, 1893 | | Rhyparus azumai Nakane, 1956 | | | Rhyparus azumai loebli Paulian, 1983 | | | Rhyparus barclayi Masumoto, Oc..., 2019 | | | Rhyparus chinensis Balthasar, 1952 | | | Rhyparus helephoroides (Fairmaire, 1893) | ⇒ Rhyparus helophoroides Fairmaire, 1893 | | Rhyparus helophoroides Fairmaire, 1893 | | | Rhyparus huaphanensis Ochi, Kon & ..., 2019 | | | Rhyparus kitanoi Miyake, 1982 | | | Rhyparus kitanoi kitanoi | | | Rhyparus kitanoi taiwanus Ochi, 2001 | | | Rhyparus klapperichorum Paulian, 1983 | | | Rhyparus loebli (Paulian, 1983) | ⇒ Rhyparus azumai loebli Paulian, 1983 | | Rhyparus myslenickorum Minkina, 2019 | | | Rhyparus orousseti (Paulian, 1981) | ⇒ Rhyparus helophoroides Fairmaire, 1893 | | Rhyparus peninsularis Arrow, 1905 | | | Rhyparus punctithoracicus Ochi, Kon & Bai, 2019 | | | Rhyparus risbeci (Paulian, 1934) | ⇒ Rhyparus helophoroides Fairmaire, 1893 | | Rhyparus schoolmeestersi Ochi, Kon & ..., 2018 | | | Rhyparus semikitanoi Ochi, Kon & ..., 2018 | | | Rhyparus wushensis Ochi & Masumoto, 2018 | | | Rhyparus yangi Ochi, Kon & Bai, 2019 | | | Rhyssemodes bouvieri Clouët, 1901 | | | Rhyssemodes malyi Rakovic, 1982 | | | Rhyssemodes ningxia RakoviÄ�, Kr..., 2017 | | | Rhyssemodes orientalis Mulsant & Go..., 1874 | | | Rhyssemodes sindicus Pittino, 1984 | | | Rhyssemodes taklamakan RakoviÄ�, Kr..., 2017 | | | Rhyssemus australis (Petrovitz, 1963) | ⇒ Rhyssemus inscitus Walker, 1858 | | Rhyssemus chinensis Pittino, 1996 | | | Rhyssemus feae Clouët, 1901 | | | Rhyssemus germanus Linnaeus, 1767 | Germany small dung beetle | | Rhyssemus histrio Balthasar, 1961 | | | Rhyssemus inscitus Walker, 1858 | | | Rhyssemus insignicollis (Lea, 1923) | ⇒ Rhyssemus inscitus Walker, 1858 | | Rhyssemus koreanus Stebnicka, 1980 | | | Rhyssemus limbolarius Petrovitz, 1963 | | | Rhyssemus malasiacus (Lansberg, 1886) | ⇒ Rhyssemus inscitus Walker, 1858 | | Rhyssemus mongol (Endrödi, 1969) | ⇒ Granulopsammodius mongol Endrödi, 1969 | | Rhyssemus mussardi (Petrovitz, 1975) | ⇒ Rhyssemus inscitus Walker, 1858 | | Rhyssemus nanshanchicus Masumoto, 1977 | | | Rhyssemus papuanus (Petrovitz, 1965) | ⇒ Rhyssemus inscitus Walker, 1858 | | Rhyssemus samurai (Balthasar, 1941) | ⇒ Myrhessus samurai Balthasar, 1941 | | Rhyssemus tarsalis (Waterhouse, 1876) | ⇒ Rhyssemus inscitus Walker, 1858 | | Rhyssemus thailandicus Pittino, 1996 | | | Rhyssemus tonkineus (Balthasar, 1945) | ⇒ Rhyssemus feae Clouët, 1901 | | Rhyssemus tuberculatus (Nomura & Nakane, 1951) | ⇒ Myrhessus samurai Balthasar, 1941 | | Saprosites formosensis (Nomura, 1939) | ⇒ Saprosites japonicus Waterhouse, 1875 | | Saprosites japonicus Waterhouse, 1875 | | | Saprosites longethorax Paulian, 1944 | | | Saprosites narae Lewis, 1895 | | | Saprosites yanoi Nomura, 1939 | | | Saprosites yasutakai Ochi & Kawahara, 2019 | | | Scabrostomus tanytarsus Skelley & Wo..., 1991 | Masked stargazer | | Setylaides foveatus Schmidt, 1909 | | | Setylaides nakanoi Ochi & Kawahara, 2019 | | | Setylaides tsujii Kakizoe, Kaw..., 2021 | | | Sigorus porcus Fabricius, 1792 | Mahogany Ridgeback scarab | | Silluvia elongata Landin, 1949 | | | Silluvia gansuensis Frolov, 2008 | | | Silluvia igori Frolov, 2008 | | | Silluvia kabaki Frolov, 2002 | | | Silluvia shashi Stebnicka, 1977 | | | Silluvia sinica Stebnicka, 1977 | | | Silluvia wassuensis Petrovitz, 1963 | | | Silluvia yunnanica Frolov, 2008 | | | Sinaphodius yunnanus Cervenka, 1994 | | | Sinodiapterna hammondi Dellacasa, 1986 | | | Sinodiapterna songrini Stebnicka & ..., 1992 | | | Sinodiapterna troitzkyi Jacobson, 1897 | | | Sinodiapterna walamiensis Masumoto, Wa..., 2017 | | | Sinodiapterna yushana Li & Wang, 2017 | | | Sinodiapterna zeni Ochi, 1991 | | | Stenotothorax hibernalis Nakane & Tsu..., 1956 | | | Stenotothorax hibernalis hibernalis | | | Subrinus sturmi Harold, 1870 | | | Sybacodes simplicicollis Fairmaire, 1896 | | | Sybacodes simplicicollis aureopilosus Schmidt, 1909 | | | Sybacodes simplicicollis simplicicollis | | | Termitodiellus hainanensis Jiang & Wang, 2020 | | | Teuchestes analis Fabricius, 1787 | | | Teuchestes brachysomus Solsky, 1874 | | | Teuchestes fossor Linnaeus, 1758 | Digger small dung beetle | | Teuchestes guangdong Rakovic & Mencl, 2012 | | | Teuchestes sinofraternus Dellacasa & ..., 1983 | | | Teuchestes uenoi Ochi, Kawaha..., 2006 | | | Trichiorhyssemus asperulus Waterhouse, 1875 | | | Trichiorhyssemus boninensis (Nakane, 1960) | ⇒ Neotrichiorhyssemus esakii Nomura, 1943 | | Trichiorhyssemus fokiensis Petrovitz, 1968 | | | Trichiorhyssemus hatakei Ochi, Kawaha..., 2019 | | | Trichiorhyssemus hauseri (Balthasar, 1933) | ⇒ Neotrichiorhyssemus hauseri Balthasar, 1933 | | Trichiorhyssemus immaturus Minkina, 2022 | | | Trichiorhyssemus kitayamai Ochi & Kawah..., 2001 | | | Trichiorhyssemus lasionotus Clouët, 1901 | | | Trichiorhyssemus microtrichius Ochi, Kawaha..., 2011 | | | Trichiorhyssemus nakoshii Ochi, Kawaha..., 2021 | | | Trichiorhyssemus nishinoi Ochi, Kawaha..., 2020 | | | Trichiorhyssemus taiwanus Ochi, Masumo..., 2015 | | | Trichiorhyssemus yumikoae Pittino & Kawai, 2007 | | | Trichonotulus amdoensis Král, Rakovi..., 2015 | | | Trichonotulus dzamosanicus Stebnicka, 1973 | | | Trichonotulus scrofa Fabricius, 1787 | Sow small dung beetle | | Trichonotulus shantungensis Balthasar, 1941 | | | Tristaphodius asperulus Cervenka, 1994 | | | Tristaphodius mortuorum Balthasar, 1932 | | | Turanella latevittis Reitter, 1887 | | | Volinus sticticus Panzer, 1798 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Subnodes of Aphodiinae with occurrences in East Asia
Links, sources and further information Aphodiinae at artfakta.se
Aphodiinae at dyntaxa.se
Aphodiinae at itis.gov
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