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Checklist for Subfamily Aphodiinae Leach, 1815 (Aphodiine Dung Beetles)
and region Continents


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
2110
116133
1222976
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
2101
115931
1221959
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
1812
113780
1210168
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
929
84484
1007160
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
234
35522
419905
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
189
32170
363924
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
23
3594
50225
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
23
3594
50225
Families
Genera
Species
 
Taxom
 
FamilyScarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles
2963
42190
Genera
Species
 
Taxom
 
SubfamilyAphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles
378
4846
Genera
Species
 
Taxom
 


Genera for Continents
378 Genera with 4846 Species
#ABCDEFGHIJKLMNOPQRSTVXY
Acanthaphodius A.Schmidt, 19091SpeciesTaxom
Acanthobodilus Dellacasa, 19223SpeciesTaxom
Acrossidius Schmidt, 18926SpeciesTaxom
Acrossoides Schmidt, 18922SpeciesTaxom
Acrossus Mulsant, 196158SpeciesTaxom
Adebrattia Bordat & Howden, 19951SpeciesTaxom
Adeloparius Schmidt, 194214SpeciesTaxom
Aegialatis Kono, 19381SpeciesTaxom
Aegialia Latreille, 198659SpeciesTaxom
Afroataenius Petrovitz, 19691SpeciesTaxom
Afrodiapterna Dellacasa, 19842SpeciesTaxom
Afrodiastictus Pittino & Ma..., 19707SpeciesTaxom
Aganocrossus Reitter, 185727SpeciesTaxom
Agoliinus Schmidt, 192844SpeciesTaxom
Agolius Mulsant & Rey, 201412SpeciesTaxom
Agrilinellus Dellacasa, ..., 19117SpeciesTaxom
Agrilinus Mulsant & Rey, 196256SpeciesTaxom
Ahermodontus Baguena, 19303SpeciesTaxom
Aidophus Balthasar, 200113SpeciesTaxom
Airapus Stebnicka & ..., 199834SpeciesTaxom
Alloblackburneus Bordat, 189415SpeciesTaxom
Allobodilus Petrovitz, 19904SpeciesTaxom
Alocoderus Schmidt, 199021SpeciesTaxom
Amerilochus1SpeciesTaxom
Amerisaprus1SpeciesTaxom
Amidorus Mulsant & Rey, 190619SpeciesTaxom
Ammoecioides14SpeciesTaxom
Ammoecius Mulsant, 196022SpeciesTaxom
Anaetius Petrovitz, 20172SpeciesTaxom
Ankaratrotrox Paulian, 19735SpeciesTaxom
Annegialia Howden, 19711SpeciesTaxom
Anomalurobius Paulian, 19521SpeciesTaxom
Anomius Mulsant & Rey, 196215SpeciesTaxom
Aparammoecius Petrovitz, 198131SpeciesTaxom
Aphodaulacus Kozhantschikov, 187512SpeciesTaxom
Aphodiellus Schmidt, 18752SpeciesTaxom
Aphodiites Heer, 18651SpeciesTaxom
Aphodiopsis Paulian, 196615SpeciesTaxom
Aphodius Hellwig, 1958721SpeciesTaxom
Aphodobius Péringuey, 193717SpeciesTaxom
Aphodoharmogaster Endrodi, 18573SpeciesTaxom
Aphodopsammobius Endrodi, 18956SpeciesTaxom
Aphotaenius Cartwright, 19639SpeciesTaxom
Apsteiniella A.Schmidt, 19774SpeciesTaxom
Argeremazus1SpeciesTaxom
Arrowiellus Balthasar, 19331SpeciesTaxom
Arupaia1SpeciesTaxom
Aschnarhyparus Makhan, 19341SpeciesTaxom
Ataeniopsis Petrovitz, 200315SpeciesTaxom
Ataenius Harold, 2001372SpeciesTaxom
Aulonocnemis Klug, 189153SpeciesTaxom
Auperia Chevrolat, 19767SpeciesTaxom
Australagolius Mencl, 20081SpeciesTaxom
Australammoecius Petrovitz, 19236SpeciesTaxom
Australaphodius Balthasar, 19904SpeciesTaxom
Australoxenella Howden & Storey, 199611SpeciesTaxom
Ballucus Gordon and S..., 20074SpeciesTaxom
Basilewskyanus Endrodi, 19013SpeciesTaxom
Batesiana Erwin, 18872SpeciesTaxom
Biralus Mulsant & Rey, 17895SpeciesTaxom
Blackburneus Schmidt, 196471SpeciesTaxom
Bobiricola Endrodi, 19731SpeciesTaxom
Bodiloides4SpeciesTaxom
Bodilopsis Adám, 010SpeciesTaxom
Bodilus Mulsant & Rey, 197562SpeciesTaxom
Bordatius Pittino & Ma..., 19865SpeciesTaxom
Boucomontiellus Balthasar, 19132SpeciesTaxom
Brachiaphodius Koshantschikov, 200017SpeciesTaxom
Brindalus Landin, 19805SpeciesTaxom
Bruchaphodius Martínez, 19383SpeciesTaxom
Bruneixenus Howden & Storey, 19922SpeciesTaxom
Caelius Lewis, 18876SpeciesTaxom
Calamosternus Motschulsky, 199426SpeciesTaxom
Caligodorus Gordon and S..., 19313SpeciesTaxom
Calocolobopterus8SpeciesTaxom
Candezeollus3SpeciesTaxom
Carinaulus Tesar, 200021SpeciesTaxom
Cartwrightia Islas, 19673SpeciesTaxom
Cephalocyclus Dellacasa, ..., 186225SpeciesTaxom
Cesamexico1SpeciesTaxom
Chaetopisthes Westwood, 198220SpeciesTaxom
Cheleion Vårdal & For..., 20193SpeciesTaxom
Chilothorax Motschulsky, 198785SpeciesTaxom
Chiron MacLeay, 185517SpeciesTaxom
Chittius1SpeciesTaxom
Cinacanthus Schmidt, 200616SpeciesTaxom
Cnemargulus Semenov, 19034SpeciesTaxom
Cnematoplatys A.Schmidt, 20067SpeciesTaxom
Cnemisus Motschulsky, 18455SpeciesTaxom
Coelotrachelus Schmidt, 19117SpeciesTaxom
Colobopteridius Clement, 19581SpeciesTaxom
Colobopteroides Paulian, 03SpeciesTaxom
Colobopterus Mulsant, 08SpeciesTaxom
Coprimorphus Mulsant, 17891SpeciesTaxom
Coptochiroides Balthasar, 04SpeciesTaxom
Coptochirus Harold, 186812SpeciesTaxom
Corythoderus Klug, 19004SpeciesTaxom
Craterocyphus Schmidt, 196017SpeciesTaxom
Cryptaphodius2SpeciesTaxom
Cryptoscatomaseter Gordon and S..., 194630SpeciesTaxom
Daintreeola Howden & Storey, 20001SpeciesTaxom
Danielssonia Bordat & Howden, 19951SpeciesTaxom
Dellacasiellus Gordon and S..., 193213SpeciesTaxom
Dialytellus Brown, 19163SpeciesTaxom
Dialytes Harold, 19416SpeciesTaxom
Dialytodius Gordon and S..., 19714SpeciesTaxom
Diapterna Horn, 18486SpeciesTaxom
Diastictus Mulsant, 18055SpeciesTaxom
Diasticus Mulsant, 1842
0SpeciesTaxom
Didactylia Orbigny, 194228SpeciesTaxom
Dilortomaeus Bordat, 19085SpeciesTaxom
Doaphius Balthasar, 19419SpeciesTaxom
Drepanacanthoides Schmidt, 18702SpeciesTaxom
Drepanocanthoides Schmidt, 19956SpeciesTaxom
Drepanocanthus Péringuey, 07SpeciesTaxom
Dudleyellus11SpeciesTaxom
Emadiellus Schmidt, 19646SpeciesTaxom
Eocorythoderus Maruyama, 20121SpeciesTaxom
Eremazus Mulsant, 18516SpeciesTaxom
Erytodes Schmidt, 19161SpeciesTaxom
Erytus Mulsant & Rey, 196111SpeciesTaxom
Esymus Mulsant & Rey, 191622SpeciesTaxom
Eudolus Mulsant & Rey, 18922SpeciesTaxom
Euhemicyclium2SpeciesTaxom
Euheptaulacus Dellacasa, 18069SpeciesTaxom
Euorodalus Dellacasa, 184814SpeciesTaxom
Euparia Le Peletier ..., 193812SpeciesTaxom
Euparixia Brown, 012SpeciesTaxom
Euparixoides Hinton, 20055SpeciesTaxom
Euparotrix1SpeciesTaxom
Eupleurus Mulsant, 17584SpeciesTaxom
Exaphodius Endrodi, 19601SpeciesTaxom
Exoxyomus1SpeciesTaxom
Ferrerianus1SpeciesTaxom
Flaviellus Gordon and S..., 18789SpeciesTaxom
Flechtmanniella1SpeciesTaxom
Geomyphilus Gordon and S..., 200617SpeciesTaxom
Geopsammodius Gordon & Pit..., 200612SpeciesTaxom
Gilletianus Balthasar, 201542SpeciesTaxom
Goiginus Endrodi, 19672SpeciesTaxom
Gonaphodiellus Schmidt, 191613SpeciesTaxom
Gonaphodioides Dellacasa M...., 20189SpeciesTaxom
Gonaphodiopsis5SpeciesTaxom
Gonaphodius Reitter, 18771SpeciesTaxom
Gongrolophus Stebnicka & ..., 19961SpeciesTaxom
Gordonius1SpeciesTaxom
Grandinaphodius2SpeciesTaxom
Granulopsammodius Rakovic, 196111SpeciesTaxom
Guanyinaphodius1SpeciesTaxom
Hadrorhyparus Howden, 19951SpeciesTaxom
Harmodactylus Péringuey, 19902SpeciesTaxom
Harmogaster Harold, 199113SpeciesTaxom
Haroldaphodius3SpeciesTaxom
Haroldiataenius Chalumeau, 19289SpeciesTaxom
Haroldiellus Gordon and S..., 20223SpeciesTaxom
Hemicorythoderus Tangelder & ..., 19471SpeciesTaxom
Hemicyclium Schmidt, 19094SpeciesTaxom
Hemycorythoderus1SpeciesTaxom
Heptaulacus Mulsant, 180613SpeciesTaxom
Hornietus Stebnicka, 18871SpeciesTaxom
Iapurea Martínez, 1953
0SpeciesTaxom
Iarupea Martínez, 07SpeciesTaxom
Iguazua Stebnicka, 19973SpeciesTaxom
Ingogius Endrodi, 19761SpeciesTaxom
Irrasinus Gordon and S..., 18702SpeciesTaxom
Jalisco1SpeciesTaxom
Koreoxyomus1SpeciesTaxom
Koshantschikovius Schmidt, 201634SpeciesTaxom
Labarrus Mulsant and Rey, 196126SpeciesTaxom
Lechorodius Gordon and S..., 18232SpeciesTaxom
Leiopsammodius Rakovic, 198350SpeciesTaxom
Leptorhyparus Howden, 20214SpeciesTaxom
Limarus Mulsant & Rey, 18133SpeciesTaxom
Lindbergianus Petrovitz, 19611SpeciesTaxom
Lioglyptoxenus Pittino, 20061SpeciesTaxom
Liothorax Motschulsky, 016SpeciesTaxom
Lomanoxia Martínez, 19117SpeciesTaxom
Lomanoxoides Stebnicka, 19637SpeciesTaxom
Longaphodius Endrodi, 19801SpeciesTaxom
Loraphodius Reitter, 18355SpeciesTaxom
Loraspis Mulsant & Rey, 18701SpeciesTaxom
Lorditomaeus Péringuey, 196729SpeciesTaxom
Lunaphodius Balthasar, 18763SpeciesTaxom
Luxolinus Gordon and S..., 18871SpeciesTaxom
Macroretroides1SpeciesTaxom
Macroretrus Péringuey, 19404SpeciesTaxom
Macrotetrus Péringuey, 19081SpeciesTaxom
Maculaphodius Gordon and S..., 18871SpeciesTaxom
Manjarivolo Paulian, 20194SpeciesTaxom
Martineziana Chalumeau & ..., 19366SpeciesTaxom
Mecynodes Mulsant & Rey, 202112SpeciesTaxom
Megateloides Landin, 04SpeciesTaxom
Megatelus Reitter, 18713SpeciesTaxom
Melinopterus Mulsant, 197530SpeciesTaxom
Mendidaphodius Reitter, 196427SpeciesTaxom
Mendidius Harold, 199741SpeciesTaxom
Merogyrus Gordon and S..., 19071SpeciesTaxom
Mesontoplatys Motschulsky, 196227SpeciesTaxom
Messyrhus Rakovič & K..., 19971SpeciesTaxom
Micraegialia Brown, 18871SpeciesTaxom
Microtermitodius Pittino, 20061SpeciesTaxom
Microteuchestes Landin, 19741SpeciesTaxom
Monteitheolus Howden & Storey, 20001SpeciesTaxom
Mothon Semenov & Me..., 19271SpeciesTaxom
Mozartius Nomura & Nakane, 011SpeciesTaxom
Myrhessus Balthasar, 20067SpeciesTaxom
Myrmecaphodius Martínez, 19521SpeciesTaxom
Mysarus Petrovitz, 19621SpeciesTaxom
Nanotermitodius Howden, 20032SpeciesTaxom
Napoa1SpeciesTaxom
Neagolius Kozhantschikov, 199734SpeciesTaxom
Neocalaphodius Bordat, 18012SpeciesTaxom
Neocolobopterus Landin, 19849SpeciesTaxom
Neodiapterna Dellacasa, 19972SpeciesTaxom
Neoemadiellus Clement, 200711SpeciesTaxom
Neoheptaulacus Paulian & Vi..., 18575SpeciesTaxom
Neopsammodius Rakovic, 19559SpeciesTaxom
Neorhyssemus4SpeciesTaxom
Neotrichaphodioides Dellacasa, D..., 20105SpeciesTaxom
Neotrichiorhyssemus Rakovič & K..., 196816SpeciesTaxom
Neotrichonotulus Dellacasa, ..., 19553SpeciesTaxom
Nettelislasia Martínez, 19451SpeciesTaxom
Nialaphodius Kolbe, 18013SpeciesTaxom
Nialosternus2SpeciesTaxom
Nialus Mulsant & Rey, 196915SpeciesTaxom
Nimbus Mulsant & Rey, 198418SpeciesTaxom
Nipponaphodius Nakane, 19511SpeciesTaxom
Nipponoagoliinus1SpeciesTaxom
Nobiellus4SpeciesTaxom
Nobius Mulsant & Rey, 193711SpeciesTaxom
Nolicus Petrovitz, 19622SpeciesTaxom
Notocaulus Quedenfeldt, 184816SpeciesTaxom
Obaphodius1SpeciesTaxom
Odochilus Harold, 199724SpeciesTaxom
Odontolochus Schmidt, 199615SpeciesTaxom
Odontolytes Koshantschikov, 200520SpeciesTaxom
Odontopsammodius Gordon and P..., 195513SpeciesTaxom
Orocanthus Endrodi, 19731SpeciesTaxom
Orodaliscoides Schmidt, 18594SpeciesTaxom
Orodaliscus Reitter, 19065SpeciesTaxom
Oromus Mulsant & Rey, 19204SpeciesTaxom
Oscarinus Gordon and S..., 193919SpeciesTaxom
Osmanius Branco & Baraud, 19883SpeciesTaxom
Otophorus Mulsant, 17581SpeciesTaxom
Oxyataenius1SpeciesTaxom
Oxycorythus Solsky, 18761SpeciesTaxom
Oxyomoides Dellacasa, D..., 19443SpeciesTaxom
Oxyomus Dejean, 199135SpeciesTaxom
Ozodius4SpeciesTaxom
Palnia Stebnicka, 20204SpeciesTaxom
Parabodilus3SpeciesTaxom
Paracoptochirus Balthasar, 19904SpeciesTaxom
Paracorythoderus Wasmann, 19824SpeciesTaxom
Paracrossidius Balthasar, 196311SpeciesTaxom
Paradeloparius Landin, 19014SpeciesTaxom
Paradidactylia Balthasar, 196137SpeciesTaxom
Parammoecius Seidlitz, 19608SpeciesTaxom
Paranimbus Schmidt, 19904SpeciesTaxom
Paraplesiataenius3SpeciesTaxom
Parapsammodius Verdú, Steb..., 18584SpeciesTaxom
Pararhyssemus Balthasar, 18871SpeciesTaxom
Parataenius Balthasar, 18686SpeciesTaxom
Paratrichaphodius3SpeciesTaxom
Pardalosus Gordon and S..., 19287SpeciesTaxom
Passalaphodius Martínez, 19531SpeciesTaxom
Passaliolla Balthasar, 18695SpeciesTaxom
Paulianellus Balthasar, 198612SpeciesTaxom
Petrovitzius Rakovič, 19861SpeciesTaxom
Phaeaphodius Reitter, 196714SpeciesTaxom
Phalacronothus Motschulsky, 191719SpeciesTaxom
Pharaphodius Reitter, 1941112SpeciesTaxom
Pholeoaphodius5SpeciesTaxom
Phycochus1SpeciesTaxom
Phycocus Broun, 18831SpeciesTaxom
Pittinius Rakovič & K..., 19972SpeciesTaxom
Plagiogonus Mulsant, 198031SpeciesTaxom
Planolinellus Dellacasa an..., 18253SpeciesTaxom
Planolinoides Dellacasa an..., 18575SpeciesTaxom
Planolinus Mulsant and Rey, 18257SpeciesTaxom
Platyderides Schmidt, 19958SpeciesTaxom
Platytomus Mulsant, 200630SpeciesTaxom
Pleuraphodius Schmidt, 199496SpeciesTaxom
Pleurophorus Mulsant, 198443SpeciesTaxom
Podotenoides Paulian, 19451SpeciesTaxom
Podotenus Schmidt, 199437SpeciesTaxom
Proctophanes Harold, 18468SpeciesTaxom
Psammodaphodius Endrodi, 19761SpeciesTaxom
Psammodius Fallen, 198360SpeciesTaxom
Psammoporus Thomson, 198623SpeciesTaxom
Psammorpha Stebnicka, 19941SpeciesTaxom
Pseudacrossus Reitter, 190828SpeciesTaxom
Pseudagolius Schmidt, 20074SpeciesTaxom
Pseudataenius Brown, 19384SpeciesTaxom
Pseuderytus1SpeciesTaxom
Pseudesymus Orbigny, 18451SpeciesTaxom
Pseudoahermodontus1SpeciesTaxom
Pseudocoelotrachelus2SpeciesTaxom
Pseudodrepanocanthus3SpeciesTaxom
Pseudogonaphodiellus M.Dellacasa ..., 20071SpeciesTaxom
Pseudoheptaulacus Dellacasa & ..., 20145SpeciesTaxom
Pseudomothon Pittino, 20172SpeciesTaxom
Pseudopharaphodius Bordat, 201519SpeciesTaxom
Pseudopodotenus1SpeciesTaxom
Pseudosaprosites Balthasar, 19671SpeciesTaxom
Pseudostereomera Bordat & Howden, 19951SpeciesTaxom
Pseudoteuchestes Dellacasa, 19551SpeciesTaxom
Pseudoxyomus Petrovitz, 197614SpeciesTaxom
Pterobius Maruyama, 20101SpeciesTaxom
Pubinus Mulsant & Rey, 17761SpeciesTaxom
Qingaphodius2SpeciesTaxom
Rakovicius Pittino, 19658SpeciesTaxom
Renaudius Balthasar, 19912SpeciesTaxom
Rhinocerotopsis Maruyama, 20091SpeciesTaxom
Rhodaphodius1SpeciesTaxom
Rhyparus Westwood, 1969103SpeciesTaxom
Rhysothorax Bedel, 17922SpeciesTaxom
Rhyssemodes Reitter, 198412SpeciesTaxom
Rhyssemorphus Clouët, 19905SpeciesTaxom
Rhyssemus Mulsant, 1956197SpeciesTaxom
Rugaphodius Gordon and S..., 19071SpeciesTaxom
Sahlbergianus1SpeciesTaxom
Saprolochus Stebnicka & ..., 20074SpeciesTaxom
Saprositellus Balthasar, 20035SpeciesTaxom
Saprosites Redtenbacher, 2019144SpeciesTaxom
Saprosties Redtenbacher, 1858
0SpeciesTaxom
Saprovisca2SpeciesTaxom
Saprus Blackburn, 19954SpeciesTaxom
Sariangus Rakovič & M..., 20111SpeciesTaxom
Scabrostomus Gordon and S..., 199110SpeciesTaxom
Schaefferellus Gordon and S..., 19071SpeciesTaxom
Selviria2SpeciesTaxom
Setodius Gordon and S..., 19293SpeciesTaxom
Setylaides8SpeciesTaxom
Siamaphodius1SpeciesTaxom
Sicardia Théry, 18962SpeciesTaxom
Sigorus Mulsant & Rey, 18022SpeciesTaxom
Silluvia Landin, 200814SpeciesTaxom
Simogonius5SpeciesTaxom
Sinaphodius7SpeciesTaxom
Sinodiapterna Dellacasa, 19917SpeciesTaxom
Sitiphus Fairmaire, 19583SpeciesTaxom
Skelleyanus M. Dellacasa..., 20111SpeciesTaxom
Stebnickiella1SpeciesTaxom
Stenotothorax Schmidt, 200633SpeciesTaxom
Stereomera Arrow, 19052SpeciesTaxom
Strigodius Gordon and S..., 19391SpeciesTaxom
Subrinus Mulsant & Rey, 19706SpeciesTaxom
Sugrames Reitter, 19285SpeciesTaxom
Sussorca Paulian, 19541SpeciesTaxom
Sybacodes Fairmaire, 08SpeciesTaxom
Sybax Boheman, 18573SpeciesTaxom
Symphodon Schmidt, 18742SpeciesTaxom
Tanyana Stebnicka, 20033SpeciesTaxom
Termitaxis Krikken, 19701SpeciesTaxom
Termitoderus Mateu, 19935SpeciesTaxom
Termitodiellus Nakane, 19879SpeciesTaxom
Termitodius Wasmann, 20226SpeciesTaxom
Termitopisthes Wasmann, 19117SpeciesTaxom
Termitotrox Reichensperger, 201515SpeciesTaxom
Tesarius Rakovič, 19047SpeciesTaxom
Tetraclipeoides Schmidt, 200716SpeciesTaxom
Teuchestes Mulsant, 198313SpeciesTaxom
Thinorycter Semenov-Tian..., 19755SpeciesTaxom
Trichaphodiellus Schmidt, 18401SpeciesTaxom
Trichaphodioides Paulian, 196720SpeciesTaxom
Trichaphodius Schmidt, 196647SpeciesTaxom
Trichiopsammobius Petrovitz, 20132SpeciesTaxom
Trichiorhyssemus Clouët, 200740SpeciesTaxom
Trichonotuloides Balthasar, 18875SpeciesTaxom
Trichonotulus Bedel, 197129SpeciesTaxom
Trigonoscelus Petrovitz, 20045SpeciesTaxom
Tristaphodius Balthasar, 19322SpeciesTaxom
Turanella Semenov, 18871SpeciesTaxom
Vladimirellus1SpeciesTaxom
Volinus Mulsant & Rey, 17981SpeciesTaxom
Xenoheptaulacus Hinton, 18962SpeciesTaxom
Xeropsamobeus Saylor, 187810SpeciesTaxom
Youngaphodius1SpeciesTaxom


Species list for Continents
4846 Species and 176 Subspecies
Acanthaphodius bruchi A.Schmidt, 1909
Acanthobodilus crimensis W.Koshantsch..., 1913
Acanthobodilus immundus Creutzer, 1799
Acanthobodilus languidulus Schmidt, 1922
Acrossidius brittoni Stebnicka & ..., 1995
Acrossidius delavayi Koshantschikov, 1916
Acrossidius kozlovi Koshantschikov, 1916
Acrossidius pseudotasmaniae Given, 1950Blackheaded pasture cockchafer
Acrossidius tasmaniae Hope, 1847Black-headed pasture cockchafer
Acrossidius yorkensis Blackburn, 1892
Acrossoides brittoni Stebnicka & ..., 1995
Acrossoides yorkensis Blackburn, 1892
Acrossus angustiarum Balthasar, 1967
Acrossus atratus Waterhouse, 1875
Acrossus belousovi Frolov, 2001
Acrossus bimaculatus Laxmann, 1770
Acrossus binaevulus Heyden, 1887
Acrossus bolognai Carpaneto & ..., 1989
Acrossus byki Minkina, 2018
Acrossus carpetanus Graells, 1846
Acrossus delavayi Koshantschikov, 1916
Acrossus depressus Kugelann, 1792
Acrossus devabhumi Mittal, 1993
Acrossus eberti Balthasar, 1965
Acrossus emodus Petrovitz, 1976
Acrossus formosanus Nomura & Nakane, 1951
Acrossus gagatinus Ménétriés, 1832
Acrossus haliki Balthasar, 1933
Acrossus histrio Balthasar, 1932
Acrossus hrubanti Balthasar, 1961
Acrossus humerospinosus Petrovitz, 1958
Acrossus igai Nakane, 1956
Acrossus igai pseudoigai Nakane & Mas..., 1967
Acrossus igai sakuraii Nakane & Mas..., 1967
Acrossus igai tsurugianus Nakane & Mas..., 1967
Acrossus impressiusculus Fairmaire, 1888
Acrossus japonicus Nomura & Nakane, 1951
Acrossus jedlickai Balthasar, 1932
Acrossus jeloneki Minkina, 2018
Acrossus jubingensis Balthasar, 1967
Acrossus klickai Balthasar, 1932
Acrossus koreanensis Kim, 1986
Acrossus kwanhsiensis Balthasar, 1945
Acrossus laticapitus Zhang, 1992
Acrossus laticollis Baudi, 1870
Acrossus longepilosus Schmidt, 1911
Acrossus luridus Fabricius, 1775
Acrossus melanodiscus Zhang, 1992
Acrossus milloti Paulian, 1945
Acrossus obenbergeri Balthasar, 1932
Acrossus opacipennis Schmidt, 1910
Acrossus planicollis Reitter, 1890
Acrossus pseudoobenbergeri Cervenka, 1995
Acrossus ritsemai Schmidt, 1909
Acrossus rubripennis Horn, 1870
Acrossus rufipes Linnaeus, 1758Night-Flying Dung Beetle
Acrossus semiopacus Reitter, 1887
Acrossus sichuanensis Zhang, 1992
Acrossus siculus Harold, 1862
Acrossus siculus buturensis Dellacasa, 1983
Acrossus siculus siculus
Acrossus superatratus Nomura & Nakane, 1951
Acrossus tiabi Masumoto, 1988
Acrossus tingitanus Reitter, 1892
Acrossus transmontanus Balthasar, 1932
Acrossus unifasciatus Nomura & Nakane, 1951
Acrossus unifasciatus kiiensis Nakane, 1960
Acrossus ustulatus Harold, 1862
Acrossus viturati Reitter, 1907
Acrossus yunlingensis Petrovitz, 1961
Adebrattia depressa Bordat & Howden, 1995
Adeloparius blattoides Landin, 1959
Adeloparius endroedii Landin, 1959
Adeloparius formosus Paulian, 1942
Adeloparius iridescens Péringuey, 1901
Adeloparius lemoulti Koshantschikov, 1916
Adeloparius luridipes Harold, 1861
Adeloparius nobilis Harold, 1874
Adeloparius overlaeti Boucomont, 1936
Adeloparius pulcherrimus Reiche, 1847
Adeloparius septemmaculatus Fabricius, 1781
Adeloparius trimaculatus Schmidt, 1909
Adeloparius vethi Lansberge, 1886
Adeloparius wahlbergi Boheman, 1857
Adeloparius wittei Paulian, 1942
Aegialatis stejnegeri Kono, 1938
Aegialia (Aegialia) amplipunctata Gordon & Car..., 1988Sharpnosed weever
Aegialia (Aegialia) arenaria Fabricius, 1787Dune sand scarab beetle
Aegialia (Aegialia) argentina (Martínez, Pe..., 1970)⇒ Mimaegialia argentina Martinez & P..., 1970
Aegialia (Aegialia) blanchardi Horn, 1887
Aegialia (Aegialia) carri Gordon & Car..., 1988Sharpnosed weever
Aegialia (Aegialia) cartwrighti Stebnicka, 1977
Aegialia (Aegialia) concinna Gordon & Car..., 1977Ciervo Aegialian Scarab
Aegialia (Aegialia) conferta Horn, 1871Scratchbush
Aegialia conferta conferta
Aegialia conferta nigrella Brown, 1931
Aegialia (Aegialia) convexa Fall, 1932
Aegialia (Aegialia) crassa LeConte, 1857
Aegialia (Aegialia) crescenta Gordon & Car..., 1977Crescent Dune aegialian scarab
Aegialia (Aegialia) hardyi Gordon & Car..., 1977Hardy's aegialian scarab
Aegialia (Aegialia) kelsoi Gordon & Car..., 1988Sharpnosed weever
Aegialia (Aegialia) knighti Gordon & Rust, 1997
Aegialia (Aegialia) latispina LeConte, 1878
Aegialia (Aegialia) magnifica Gordon & Car..., 1977Large aegialian scarab
Aegialia (Aegialia) mcclevei Gordon, 1990
Aegialia (Aegialia) nigrella Brown, 1931
Aegialia (Aegialia) opifex Horn, 1887
Aegialia (Aegialia) punctata Brown, 1931
Aegialia (Aegialia) spinosa Gordon & Car..., 1988Sharpnosed weever
Aegialia (Psammoporus) criddlei Brown, 1931
Aegialia (Psammoporus) cylindrica Eschscholtz, 1822
Aegialia (Psammoporus) insularis (Pittino, 2006)⇒ 
Aegialia (Psammoporus) lacustris LeConte, 1850
Aegialia (Psammoporus) nana Brown, 1931
Aegialia (Psammoporus) opaca Brown, 1931
Aegialia (Psammoporus) terminalis Brown, 1931
Aegialia (Rhysothorax) rufa (Fabricius, 1792)⇒ Caelius rufescens Horn, 1887
Aegialia abdita Nikritin, 1975
Aegialia arenarius Fabricius, 1787
Aegialia caelatus Laconte, 1857
Aegialia clypeata Say, 1824
Aegialia comis Lewis, 1895
Aegialia cylindricus Eschscholtz, 1822
Aegialia exaratus Mannerheim, 1853
Aegialia friebi (Balthasar, 1935)⇒ Psammoporus friebi Balthasar, 1935
Aegialia hybrida (Reitter, 1892)⇒ Aegialia comis Lewis, 1895
Aegialia kamtschaticus Motschulsky, 1860
Aegialia koreana Kim, 1979
Aegialia latipunctus Gredler, 1863
Aegialia matalini Gusakov, 2003
Aegialia mimica Pittino, 2006
Aegialia mimicus Pittino, 2006
Aegialia nakanei Masumoto, 1986
Aegialia nakanei horii Ochi & Kawahara, 2012
Aegialia nanus Brown, 1931
Aegialia nitida Waterhouse, 1875
Aegialia opacus Brown, 1931
Aegialia rufina (Silfverberg, 1977)⇒ Rhysothorax rufa Fabricius, 1792
Aegialia rupta Scudder, 1882
Aegialia sabuleti Panzer, 1797
Aegialia shimeki Lago & Freese, 2016
Aegialia sibirica Stebnicka, 1977
Aegialia sibiricus Stebnicka, 1977
Aegialia spissipes (LeConte, 1878)⇒ Rhysothorax rufa Fabricius, 1792
Aegialia tsukamotoi Masumoto, 1986
Afroataenius singularis Petrovitz, 1969
Afrodiapterna cambeforti Dellacasa, 1984
Afrodiapterna heleninae Dellacasa, 1984
Afrodiastictus apterus Pittino & Ma..., 1986
Afrodiastictus carinatus Paulian, 1954
Afrodiastictus ceratus Clement, 1969
Afrodiastictus endroedyyoungai Endrödi, 1973
Afrodiastictus insularis Petrovitz, 1969
Afrodiastictus mimicus Gordon & Car..., 1980
Afrodiastictus minutus Petrovitz, 1970
Aganocrossus albertisi Harold, 1877
Aganocrossus amoenus Boheman, 1857
Aganocrossus apicenotatus Müller, 1941
Aganocrossus bordatianus Masumoto & K..., 2003
Aganocrossus borneensis Petrovitz, 1962
Aganocrossus ciprianii Balthasar, 1939
Aganocrossus clementianus Endrödi, 1964
Aganocrossus fugitivus Péringuey, 1901
Aganocrossus hirtus Bordat, 1992
Aganocrossus hutteri Petrovitz, 1973
Aganocrossus ingloriosus Schmidt, 1916
Aganocrossus krikkeni Bordat, 1992
Aganocrossus meticulosus Paulian, 1942
Aganocrossus newmanae Bordat, 2008
Aganocrossus pallidicornis Walker, 1858
Aganocrossus paraciprianii Bordat, 2008
Aganocrossus periculosus Bordat, 1992
Aganocrossus plicatus Endrödi, 1964
Aganocrossus postpilosus Reitter, 1895
Aganocrossus pseudamoenus Müller, 1941
Aganocrossus pseudourostigma Balthasar, 1939
Aganocrossus pseudurostigma Balthasar, 1939
Aganocrossus rudolfpetrovitzi Bordat, 2013
Aganocrossus tacitus Bordat, 2008
Aganocrossus urostigma Harold, 1862
Aganocrossus vejdovskyi Balthasar, 1945
Aganocrossus vestitus Boheman, 1857
Agoliinus albertanus Brown, 1928
Agoliinus aleutus Eschscholtz, 1822
Agoliinus amurensis Iablokov-Khn..., 1972
Agoliinus anthracus Gordon & Ske..., 2007Lesser weever
Agoliinus aquilonarius Brown, 1928
Agoliinus ashworthi Gordon, 2006Greater weever
Agoliinus bidentatus Schmidt, 1906Spotted weever
Agoliinus canadensis Garnett, 1920Streaked weeverfish
Agoliinus congregatus Mannerheim, 1853
Agoliinus corruptor Brown, 1929
Agoliinus cruentatus LeConte, 1878
Agoliinus durrelli Rakovic & Mencl, 2011
Agoliinus explanatus LeConte, 1878
Agoliinus guttatus Eschscholtz, 1823
Agoliinus hatchi Hatch, 2007Lesser weever
Agoliinus idahoensis Gordon & Ske..., 2007Lesser weever
Agoliinus incommunis Fall, 1932
Agoliinus ingenursus Gordon & Ske..., 2007Lesser weever
Agoliinus isajevi Kabakov, 1996
Agoliinus kiuchii Masumoto, 1984
Agoliinus lapponum Gyllenhal, 1808Northern Dung Beetle
Agoliinus leopardus Horn, 1870
Agoliinus malkini Hatch, 1971
Agoliinus manitobensis Brown, 1928
Agoliinus morii Nakane, 1983
Agoliinus nemoralis Erichson, 1848
Agoliinus parastorkani Cervenka, 1995
Agoliinus piceatus Robinson, 1946Goby duckbill
Agoliinus piceus Gyllenhal, 1808
Agoliinus pittinoi Carpaneto, 1986
Agoliinus plutonicus Fall, 1907
Agoliinus poudreus Gordon & Ske..., 2007Lesser weever
Agoliinus praealtus Gordon & Ske..., 2007Lesser weever
Agoliinus pseudostorkani Stebnicka, 1982
Agoliinus ragusae Reitter, 1892
Agoliinus ragusai Reitter, 1892
Agoliinus satunini Olsoufieff, 1918
Agoliinus satyrus Reitter, 1892
Agoliinus setchan Masumoto, 1984
Agoliinus shibatai Nakane, 1983
Agoliinus sigmoideus Van Dyke, 1918
Agoliinus storkani Balthasar, 1932
Agoliinus tanakai Masumoto, 1981
Agoliinus wickhami Brown, 1928
Agolius abdominalis Bonelli, 1812
Agolius abdominalis abdominalis Bonelli, 1812
Agolius abdominalis balazuci Nicolas, 1971
Agolius abdominalis emilianus Dellacasa, 1983
Agolius abdominalis pecoudi Hoffmann, 1954
Agolius abdominalis tatrensis Dellacasa & ..., 2005
Agolius bonvouloiri Harold, 1860
Agolius bonvouloiri bonvouloiri
Agolius bonvouloiri cantabricus Pittino, 1981
Agolius limbolarius
Agolius moronensis Frolov, 2009
Agolius petri Frolov, 2014
Agrilinellus abbonai Dellacasa, D..., 2008
Agrilinellus antonioreyi Dellacasa, D..., 2008
Agrilinellus azteca Harold, 1863
Agrilinellus chiapasensis Galante, Ste..., 2003
Agrilinellus nuriae Dellacasa, D..., 2008Freckled pikeblenny
Agrilinellus oaxacaensis Dellacasa, D..., 2008
Agrilinellus ornatus Schmidt, 1911Banded pipefish
Agrilinus aquilus Schmidt, 1916
Agrilinus ater De Geer, 1774
Agrilinus ater ascendens Reiche, 1863
Agrilinus ater falsarius Reitter, 1892
Agrilinus ater lucasi Harold, 1859
Agrilinus azteca (Harold, 1863)⇒ Agrilinellus azteca Harold, 1863
Agrilinus bardus Balthasar, 1946
Agrilinus bellus (Schmidt, 1916)⇒ Agrilinus aquilus Schmidt, 1916
Agrilinus bollowi Balthasar, 1941
Agrilinus borealis
Agrilinus breviusculus Motschulsky, 1866
Agrilinus chiapasensis (Galante & St..., 2003)⇒ Agrilinellus chiapasensis Galante, Ste..., 2003
Agrilinus constans Duftschmid, 1805
Agrilinus convexus Erichson, 1848
Agrilinus hasegawai Nomura & Nakane, 1951
Agrilinus hasegawai aka Nakane, 1960
Agrilinus hasegawai akahane Nakane, 1960
Agrilinus hayachinensis Nomura & Nakane, 1951
Agrilinus ibericus Harold, 1874
Agrilinus ibericus basilicatus Fiori, 1907
Agrilinus ibericus ibericus
Agrilinus indutilis Harold, 1874
Agrilinus ishidai Masumoto & K..., 1987
Agrilinus jugurtha Balthasar, 1931
Agrilinus lansbergei Harold, 1874
Agrilinus lindbergi Petrovitz, 1959
Agrilinus lungaiensis Petrovitz, 1962
Agrilinus madara Nakane, 1960
Agrilinus matusiaki Minkina, 2022
Agrilinus monicae Stebnicka, 1981
Agrilinus monikae Kral, 2013
Agrilinus montisamator Balthasar, 1965
Agrilinus nemoralis Erichson, 1848
Agrilinus nikolajevi Berlov & Kal..., 1989
Agrilinus obliviosus Reitter, 1892
Agrilinus piceus
Agrilinus pirinensis Balthasar, 1946
Agrilinus pseudolungaiensis Mencl & Rakovic, 2012
Agrilinus punctator Reitter, 1892
Agrilinus ritsukoae Kawai, 2004
Agrilinus rufoanalis Petrovitz, 1961
Agrilinus rufus (Moll, 1782)⇒ Bodilopsis rufa Moll, 1782
Agrilinus sallei Harold, 1863
Agrilinus shilenkovi Berlov, 1989
Agrilinus shukronajevi Nikolajev, 1998
Agrilinus sordidus (Fabricius, 1775)⇒ Bodilopsis sordida Fabricius, 1775
Agrilinus sordidus sordidus (Fabricius, 1775)⇒ Bodilopsis sordida sordida Fabricius, 1775
Agrilinus spinulosus Schmidt, 1910
Agrilinus striatus Schmidt, 1910
Agrilinus surdus Boucomont, 1929
Agrilinus tashigaonae Emberson & S..., 2001
Agrilinus tenax Balthasar, 1932
Agrilinus uliginosus
Agrilinus uniformis Waterhouse, 1875
Agrilinus virginalis Reitter, 1900
Agrilinus wassuensis Petrovitz, 1962
Ahermodontus ambrosi Pardo Alcaide, 1936
Ahermodontus bischoffi Vsetecka, 1939
Ahermodontus marini Baguena, 1930
Aidophus cabrali Petrovitz, 1973
Aidophus coheni Stebnicka & ..., 2005
Aidophus flaveolus Harold, 1867
Aidophus flechtmanni Dellacasa & ..., 2001
Aidophus gema Clavijo-Bust..., 2023
Aidophus impressus Petrovitz, 1970
Aidophus infuscatopennis Schmidt, 1909
Aidophus kolbei Schmidt, 1911
Aidophus notatus Harold, 1859
Aidophus panamensis Harold, 1859
Aidophus parcus Horn, 1887
Aidophus pellax Balthasar, 1960
Aidophus skelleyi Harpootlian ..., 2001
Airapus aegialiformis Minkina, 2020
Airapus bruxnerensis Stebnicka & ..., 1996
Airapus burrundieae Stebnicka & ..., 1996
Airapus cechovskyi Král, Mencl ..., 2019
Airapus cheesmani Paulian, 1939
Airapus gazelle Stebnicka, 1998
Airapus granulator Harold, 1877
Airapus halmaheraensis Minkina & Be..., 2023
Airapus henriettae Stebnicka & ..., 1996
Airapus hollandiae Stebnicka, 1998
Airapus idenburgii Paulian, 1937
Airapus interstitialis Fairmaire, 1883
Airapus jenisi Král, Mencl ..., 2019
Airapus lamingtonensis Minkina & Be..., 2023
Airapus lithuanicus Tamutis, Ale..., 2017
Airapus nondugli Stebnicka, 1998
Airapus novaebritanniae Balthasar, 1961
Airapus novaeirlandiae Balthasar, 1967
Airapus obscurus MacLeay, 1871
Airapus parvispinus Schmidt, 1920
Airapus popondettae Stebnicka, 1998
Airapus pygidialis Lea, 1923
Airapus rakovici Král & Lu, 2021
Airapus rugulopunctatus Krikken, 1970
Airapus sicardi Paulian, 1945
Airapus spinator Harold, 1877
Airapus sulawesianus Rakovi�, Me..., 2021
Airapus sumatrae Fairmaire, 1896
Airapus tyri Král, Mencl ..., 2019
Airapus uniseriatus Krikken, 1970
Airapus vangviengensis Minkina & Be..., 2023
Airapus wallaceanus Rakovi�, Me..., 2019
Airapus wampuensis Stebnicka, 1998
Airapus wauus Stebnicka, 1998
Alloblackburneus acutulus Bordat, 2009
Alloblackburneus aegrotus Horn, 1870
Alloblackburneus cavidomus Brown, 1927
Alloblackburneus cynomysi Brown, 1927
Alloblackburneus fordi Gordon, 1974
Alloblackburneus geomysi Cartwright, 1939
Alloblackburneus guadalajarae Dellacasa, D..., 2011
Alloblackburneus ibanezbernali Dellacasa, D..., 2011
Alloblackburneus lentus Horn, 1870
Alloblackburneus mashunensis Péringuey, 1901
Alloblackburneus poggianus Bordat, 2019
Alloblackburneus rubeolus Palisot de B..., 1809Surf blenny
Alloblackburneus saylori Hinton, 1934Sand stargazers
Alloblackburneus tenuistriatus Horn, 1887
Alloblackburneus troglodytes Hubbard, 1894
Allobodilus fascinatus Endrödi, 1980
Allobodilus notabilis Petrovitz, 1963
Allobodilus obibensis Endrödi, 1991
Allobodilus unicus Bordat, 1990
Alocoderus baleensis Bordat, 2005
Alocoderus ballerioi Bordat, 1997
Alocoderus carinifrons Reitter, 1892
Alocoderus decrepidus Petrovitz, 1975
Alocoderus digitalis Koshantschikov, 1894
Alocoderus elongatulus Fabricius, 1801
Alocoderus gilleti Schmidt, 1909
Alocoderus gilleti gilleti
Alocoderus gilleti katanganus Endrödi, 1964
Alocoderus holdereri Reitter, 1900
Alocoderus hydrochaeris Fabricius, 1798
Alocoderus imitator Koshanschikov, 1916
Alocoderus luangensis Emberson & S..., 2001
Alocoderus mineti Clement, 1981
Alocoderus semenowi Reitter, 1887
Alocoderus semienensis Bordat, 1990
Alocoderus strigimargo Reitter, 1892
Alocoderus teyrovskyi Balthasar, 1935
Alocoderus turbatus Baudi, 1870
Alocoderus volubilis Balthasar, 1961
Alocoderus wolfgangi Stebnicka, 1990
Amerilochus cinereus Skelley, 2007
Amerisaprus valdivia Stebnicka & ..., 2003
Amidorus alagoezi Olsoufieff, 1918
Amidorus cribrarius Brullé, 1832
Amidorus cribricollis Lucas, 1846
Amidorus dentellus Schmidt, 1908
Amidorus glyptus (Bates, 1887)⇒ Trichonotuloides glyptus Bates, 1887
Amidorus granulatus Boheman, 1857
Amidorus immaturus Mulsant, 1842
Amidorus koshantschikovi Jacobson, 1911
Amidorus moraguesi Baraud, 1978
Amidorus obscurus Fabricius, 1792
Amidorus obscurus latinus Dellacasa, 1983
Amidorus obscurus obscurus Fabricius, 1792
Amidorus porcus
Amidorus salebrosus Schmidt, 1911
Amidorus serrimargo Koshantschikov, 1913
Amidorus thermicola Sturm, 1800
Amidorus transaralicus Nikolajev, 1983
Amidorus umbrinus Koshantschikov, 1913
Amidorus zangi Schmidt, 1906
Ammoecioides adventicius Bordat, 1999
Ammoecioides catulus Balthasar, 1946
Ammoecioides davisianus Bordat, 1999
Ammoecioides facetus Bordat, 1999
Ammoecioides granosus Bordat, 2005
Ammoecioides insidiosus Bordat, 1999
Ammoecioides kovacsi Bordat, 1990
Ammoecioides mulanje Bordat, 1985
Ammoecioides nyika Bordat, 1999
Ammoecioides sparsepunctatus Petrovitz, 1964
Ammoecioides spectabilis Péringuey, 1901
Ammoecioides spectabilis spectabilis
Ammoecioides spectabilis tricarinulatus Schmidt, 1907
Ammoecioides uitsoekensis Bordat, 1999
Ammoecius amplicollis Peyerimhoff, 1949
Ammoecius brevis Erichson, 1848
Ammoecius dentatus Schmidt, 1908
Ammoecius dogueti Baraud, 1980
Ammoecius elevatus Olivier, 1789
Ammoecius eli Petrovitz, 1961
Ammoecius felscheanus Reitter, 1904
Ammoecius franzi Petrovitz, 1964
Ammoecius frigidus Brisout, 1866
Ammoecius incultus Petrovitz, 1961
Ammoecius lugubris Boheman, 1857
Ammoecius lusitanicus Erichson, 1848
Ammoecius meurguesae Clement, 1975
Ammoecius mimus Péringuey, 1901
Ammoecius muchei Petrovitz, 1963
Ammoecius naviauxi Baraud, 1971
Ammoecius numidicus Mulsant, 1851
Ammoecius pannonicus (Petrovitz, 1961)⇒ Loraspis frater Mulsant & Rey, 1870
Ammoecius rugifrons Aubé, 1850
Ammoecius satanas Carpaneto, 1976
Ammoecius terminatus Harold, 1869
Ammoecius yamato Nakane, 1960
Anaetius kuijteni Petrovitz, 1969
Anaetius vanharteni Kral, 2017
Ankaratrotrox centralis Paulian, 1954
Ankaratrotrox descarpentriesi Paulian, 1978
Ankaratrotrox griveaudi Paulian, 1973
Ankaratrotrox pauliani Cambefort, 1987
Ankaratrotrox peyrierasi Paulian, 1973
Annegialia ataeniformis Howden, 1971
Anomalurobius eburneus Paulian, 1952
Anomius annamariae Baraud, 1982
Anomius antii Gridelli, 1930
Anomius apollonius Petrovitz, 1971
Anomius baeticus Mulsant & Rey, 1869
Anomius castaneus Illiger, 1803
Anomius crovettii Dellacasa, 1983
Anomius hamricola Clement, 1928
Anomius lekefensis Baraud, 1981
Anomius neidae Petrovitz, 1971
Anomius permixtus Clément & Pe..., 1971
Anomius petrovitzi (Clement, 1976)⇒ Anomius lekefensis Baraud, 1981
Anomius peyerimhoffi Théry, 1925
Anomius saturnalis Ziani, 2011
Anomius segonzaci Bedel, 1904
Anomius theryi (Clément, 1962)⇒ Anomius permixtus Clément & Pe..., 1971
Aparammoecius annapurnae Stebnicka, 1994
Aparammoecius bagmatiensis Stebnicka, 1983
Aparammoecius balangensis Petrovitz, 1958
Aparammoecius chuangkuohungi Masumoto, Ki..., 2017
Aparammoecius gansuensis Stebnicka, 1997
Aparammoecius heinzi Pittino, 1997
Aparammoecius holzschuhi Pittino, 1997
Aparammoecius isaburoi Nakane, 1956
Aparammoecius ivani Stebnicka, 1989
Aparammoecius kanglae Stebnicka, 1990
Aparammoecius kaskiensis Ahrens & Ste..., 1997
Aparammoecius lamjungi Ahrens & Ste..., 1997
Aparammoecius leisteri Medvedev, 1968
Aparammoecius mahriensis Stebnicka, 1983
Aparammoecius mizo Nakane, 1967
Aparammoecius mudukensis Stebnicka, 1989
Aparammoecius muzaffarabadensis Pittino, 1997
Aparammoecius nomurai Stebnicka, 1986
Aparammoecius pallidiligonis Waterhouse, 1875
Aparammoecius phulcokiensis Stebnicka, 1986
Aparammoecius sabhae Stebnicka, 1990
Aparammoecius schawalleri Stebnicka, 1989
Aparammoecius shogranicus Pittino, 1997
Aparammoecius tarokensis Stebnicka, 1994
Aparammoecius topali Pittino, 1997
Aparammoecius waniaramensis Pittino, 1997
Aparammoecius yangricus Stebnicka, 1986
Aparammoecius yaralensis Stebnicka, 1986
Aparammoecius yenpingensis Stebnicka, 1986
Aparammoecius yushanus Masumoto, Ki..., 2017
Aparammoecius zojilae Stebnicka, 1981
Aphodaulacus assyricus Petrovitz, 1971
Aphodaulacus balachanicola Balthasar, 1973
Aphodaulacus grumi Frolov, 2002
Aphodaulacus ignobilis Reitter, 1887
Aphodaulacus kabaki Frolov, 2001
Aphodaulacus kizeritskyi Frolov, 2002
Aphodaulacus koltzei Reitter, 1892
Aphodaulacus nigrotessellatus Motschulsky, 1866
Aphodaulacus perblandus Balthasar, 1933
Aphodaulacus saraevi Nikolajev, 2004
Aphodaulacus turkestanicus Heyden, 1881
Aphodaulacus variabilis Waterhouse, 1875
Aphodiellus genialis Péringuey, 1901
Aphodiellus impunctatus Waterhouse, 1875
Aphodiites protogaeus Heer, 1865
Aphodiopsis camerunensis Petrovitz, 1969
Aphodiopsis cubita Petrovitz, 1966
Aphodiopsis evestigata Bordat, 2014
Aphodiopsis kumasianus Endrödi, 1973
Aphodiopsis latens Bordat, 2014
Aphodiopsis latipes Paulian, 1942
Aphodiopsis motoensis Endrödi, 1964
Aphodiopsis olseni Petrovitz, 1962
Aphodiopsis paulianellus Endrödi, 1964
Aphodiopsis problematicus Petrovitz, 1973
Aphodiopsis radians Endrödi, 1964
Aphodiopsis raucus Endrödi, 1964
Aphodiopsis subopaca Petrovitz, 1964
Aphodiopsis subopacus Petrovitz, 1964
Aphodiopsis temerarius Balthasar, 1966
Aphodius (Acanthobodilus) immundus (Creutzer, 1799)⇒ Pubinus tomentosus O.F.Muller, 1776
Aphodius (Acrossoides) yorkensis (Blackburn, 1892)⇒ Acrossidius yorkensis Blackburn, 1892
Aphodius (Acrossus) bimaculatus (Laxman, 1778)⇒ Teuchestes brachysomus Solsky, 1874
Aphodius (Acrossus) carpetanus Graells, 1847
Aphodius (Acrossus) depressus (Kugelann, 1792)⇒ Acrossus depressus Kugelann, 1792
Aphodius (Acrossus) formosanus Nomura & Nakane, 1951
Aphodius (Acrossus) laticollis (Baudi di Selve, 1870)⇒ Acrossus laticollis Baudi, 1870
Aphodius (Acrossus) luridus (Fabricius, 1775)⇒ Acrossus luridus Fabricius, 1775
Aphodius (Acrossus) rubripennis Horn, 1870
Aphodius (Acrossus) rufipes (Linnaeus, 1758)⇒ Acrossus rufipes Linnaeus, 1758
Aphodius (Acrossus) siculus Harold, 1862
Aphodius siculus buturensis Dellacasa, 1983
Aphodius siculus siculus Harold, 1862
Aphodius (Aganocrossus) urostigma Harold, 1862
Aphodius (Aganocrossus) vestitus Boheman, 1857
Aphodius (Agoliinus) canadensis (Garnett, 1920)⇒ Agoliinus canadensis Garnett, 1920
Aphodius (Agoliinus) congregatus Mannerheim, 1853
Aphodius (Agoliinus) corruptor Brown, 1929
Aphodius (Agoliinus) lapponum (Gyllenhal, 1806)⇒ Agoliinus lapponum Gyllenhal, 1808
Aphodius (Agoliinus) piceus (Gyllenhal, 1808)⇒ Agoliinus piceus Gyllenhal, 1808
Aphodius (Agoliinus) plutonicus (Fall, 1907)⇒ Aphodius aleutus Eschscholtz, 1822
Aphodius (Agoliinus) ragusae Reitter, 1892
Aphodius (Agoliinus) satyrus (Reitter, 1892)⇒ Agoliinus satyrus Reitter, 1892
Aphodius (Agoliinus) sigmoideus Van Dyke, 1918
Aphodius (Agolius) abdominalis (Bonelli, 1812)⇒ Agolius abdominalis Bonelli, 1812
Aphodius abdominalis abdominalis Bonelli, 1812
Aphodius abdominalis emilianus Dellacasa, 1983
Aphodius (Agolius) mixtus (Motschulsky, 1858)⇒ Agolius abdominalis Bonelli, 1812
Aphodius (Agrilinus) ater (DeGeer, 1774)⇒ Agrilinus ater De Geer, 1774
Aphodius (Agrilinus) constans (Duftschmid, 1805)⇒ Agrilinus constans Duftschmid, 1805
Aphodius (Agrilinus) convexus (Erichson, 1848)⇒ Agrilinus convexus Erichson, 1848
Aphodius (Agrilinus) ibericus Harold, 1874
Aphodius (Agrilinus) nemoralis (Erichson, 1848)⇒ Agoliinus nemoralis Erichson, 1848
Aphodius (Agrilinus) rufus Moll, 1782
Aphodius (Agrilinus) sordidus (Fabricius, 1775)⇒ Bodilopsis sordida Fabricius, 1775
Aphodius (Agrilinus) tenellus (Say, 1823)⇒ Planolinus fasciatus Olivier, 1789
Aphodius (Ahermodontus) marini Baguena, 1930
Aphodius (Alloblackburneus) aegrotus Horn, 1870
Aphodius (Alloblackburneus) cynomysi Brown, 1927
Aphodius (Alloblackburneus) lentus (Horn, 1870)⇒ Blackburneus lentus Horn, 1870
Aphodius (Alloblackburneus) rubeolus Beauvois, 1805
Aphodius (Alloblackburneus) tenuistriatus (Horn, 1887)⇒ Blackburneus tenuistriatus Horn, 1887
Aphodius (Alloblackburneus) troglodytes Hubbard, 1894Aphodius tortoise commensal scarab
Aphodius (Alocoderus) hydrochaeris
Aphodius (Amidorus) cribrarius Brullé, 1832
Aphodius (Amidorus) immaturus (Mulsant, 1842)⇒ Amidorus immaturus Mulsant, 1842
Aphodius (Amidorus) obscurus (Fabricius, 1792)⇒ Amidorus obscurus Fabricius, 1792
Aphodius obscurus latinus Dellacasa, 1983
Aphodius obscurus obscurus
Aphodius (Ammoecius) brevis (Erichson, 1848)⇒ Ammoecius brevis Erichson, 1848
Aphodius (Ammoecius) elevatus (Olivier, 1789)⇒ Ammoecius elevatus Olivier, 1789
Aphodius (Ammoecius) rugifrons (Horn, 1871)⇒ Aphodius fucosus Schmidt, 1922
Aphodius (Anomius) castaneus (Illiger, 1803)⇒ Anomius castaneus Illiger, 1803
Aphodius (Anomius) crovettii G. Dellacasa, 1983
Aphodius (Aparammoecius) balangensis Petrovitz, 1958
Aphodius (Aparammoecius) nishikawai (Masumoto, 1996)⇒ Aparammoecius tarokensis Stebnicka, 1994
Aphodius (Aparammoecius) tarokensis Stebnicka, 1994
Aphodius (Aphodiellus) impunctatus (Waterhouse, 1875)⇒ Melinopterus consputus Creutzer, 1799
Aphodius (Aphodius) aestivalis (Stephens, 1839)⇒ Rhodaphodius foetens Fabricius, 1787
Aphodius (Aphodius) coniugatus Panzer, 1795
Aphodius (Aphodius) conjugatus (Mulsant, 1842)⇒ Aphodius coniugatus Panzer, 1795
Aphodius (Aphodius) elegans Allibert, 1847
Aphodius (Aphodius) fimetarius Linnaeus, 1758Common dung beetle
Aphodius (Aphodius) foetens (Fabricius, 1787)⇒ Rhodaphodius foetens Fabricius, 1787
Aphodius (Aphodius) foetidus Herbst, 1783
Aphodius foetidus foetidus
Aphodius (Aphodius) pedellus (DeGeer, 1774)⇒ Aphodius fimetarius Linnaeus, 1758
Aphodius (Aphodius) pedrosi Wollaston, 1854
Aphodius (Aphodius) scybalarius (Fabricius, 1781)⇒ Bodilopsis rufa Moll, 1782
Aphodius (Aphodius) variabilis (C. O. Waterh..., 1875)⇒ Aphodaulacus nigrotessellatus Motschulsky, 1866
Aphodius (Aphodobius) misellus (Boheman, 1857)⇒ Trichaphodius misellus Boheman, 1857
Aphodius (Balthasarianus) babori Balthasar, 1938
Aphodius (Balthasarianus) pilifer Paulian, 1934
Aphodius (Basilewskyanus) elisabethae Endrodi, 1956
Aphodius (Biralus) menetriesi (Dejean, 1833)⇒ Biralus menetriesi Ménétriés, 1849
Aphodius (Biralus) satellitius (Herbst, 1789)⇒ Biralus satellitius Herbst, 1789
Aphodius (Blackburneus) cavidomus Brown, 1927
Aphodius (Blackburneus) stercorosus (Melsheimer, 1845)⇒ Blackburneus stercorosus Melsheimer, 1845
Aphodius (Bodilus) arvernicus (Hoffmann, 1928)⇒ Bodilopsis rufus Moll, 1782
Aphodius (Bodilus) beduinus Reitter, 1892
Aphodius (Bodilus) ghardimaouensis (Balthasar, 1929)⇒ Bodiloides ictericus ghardimaouensis Balthasar, 1929
Aphodius (Bodilus) gregarius Harold, 1871
Aphodius (Bodilus) hydrochaeris (Fabricius, 1798)⇒ Alocoderus hydrochaeris Fabricius, 1798
Aphodius (Bodilus) ictericus (Laicharting, 1781)⇒ Bodiloides ictericus Laicharting, 1781
Aphodius (Bodilus) longispina Küster, 1854
Aphodius (Bodilus) lugens (Creutzer, 1799)⇒ Bodilus lugens Creutzer, 1799
Aphodius (Bodilus) nigriventris Reitter, 1892
Aphodius (Bodilus) nitidulus (C. O. Waterh..., 1875)⇒ Bodiloides ictericus ictericus Laicharting, 1781
Aphodius (Bodilus) punctipennis Erichson, 1848
Aphodius (Bodilus) rufus (Moll, 1782)⇒ Bodilopsis rufa Moll, 1782
Aphodius (Bodilus) sordidus (Fabricius, 1775)⇒ Bodilopsis rufus Moll, 1782
Aphodius (Bodilus) wollastoni Harold, 1862
Aphodius (Calamosternus) granarius (Linnaeus, 1767)⇒ Calamosternus granarius Linnaeus, 1767
Aphodius (Calamosternus) hyxos Petrovitz, 1962
Aphodius hyxos algiricus Mariani & Pi..., 1983
Aphodius (Calamosternus) mayeri (Pilleri, 1953)⇒ Calamosternus mayeri Pilleri, 1953
Aphodius (Calamosternus) sublimbatus Motschulsky, 1860
Aphodius (Calamosternus) trucidatus (Chevrolat, 1863)⇒ Calamosternus trucidatus Harold, 1863
Aphodius (Calamosternus) unicolor (Lucas, 1846)⇒ Anomius castaneus Illiger, 1803
Aphodius (Calaphodius) moestus (Fabricius, 1801)⇒ Calamosternus granarius Linnaeus, 1767
Aphodius (Caligodorus) opacus (Motschulsky, 1874)⇒ Erytus opacior Koshantschikov, 1894
Aphodius (Caligodorus) vandykei Barret, 1931
Aphodius (Calocolobopterus) senegalensis (Klug, 1835)⇒ Calocolobopterus senegalensis Klug, 1835
Aphodius (Calomosternus) granarius Linnaeus, 1767
Aphodius (Carinaulus) pucholti Balthasar, 1961
Aphodius (Cephalocyclus) luteolus Horn, 1887
Aphodius (Chilothorax) cervorum (Fairmaire, 1871)⇒ Chilothorax cervorum Fairmaire, 1871
Aphodius (Chilothorax) clathratus Reitter, 1892
Aphodius (Chilothorax) conspurcatus (Linnaeus, 1758)⇒ Chilothorax conspurcatus Linnaeus, 1758
Aphodius (Chilothorax) distinctus O.F. Müller, 1776Maculated Dung Beetle
Aphodius (Chilothorax) exclamationis Motschulsky, 1849
Aphodius (Chilothorax) hieroglyphicus (Klug, 1845)⇒ Chilothorax hieroglyphicus Klug, 1845
Aphodius (Chilothorax) lineolatus (Illiger, 1803)⇒ Chilothorax lineolatus Illiger, 1803
Aphodius (Chilothorax) melanostictus (W.L.E. Schmidt, 1840)⇒ Chilothorax melanostictus Schmidt, 1840
Aphodius (Chilothorax) paykulli Bedel, 1907
Aphodius (Chilothorax) pictus (D. Koshantsc..., 1894)⇒ Chilothorax pictus Sturm, 1805
Aphodius (Chilothorax) sticticus (Panzer, 1798)⇒ Volinus sticticus Panzer, 1798
Aphodius (Chilothorax) tessulatus (Paykull, 1798)⇒ Chilothorax paykulli Bedel, 1907
Aphodius (Cinacanthus) militaris LeConte, 1858
Aphodius (Coelotrachelus) rudis LeConte, 1878
Aphodius (Colobopteroides) latus A.Schmidt, 1911
Aphodius (Colobopterus) erraticus (Linnaeus, 1758)⇒ Colobopterus erraticus Linnaeus, 1758
Aphodius (Coprimorphus) scrutator (Herbst, 1789)⇒ Coprimorphus scrutator Herbst, 1789
Aphodius (Coptochiroides) saoudi Paulian, 1980
Aphodius (Craterocyphus) magnificus A.Schmidt, 1920
Aphodius (Cryptoscatomaseter) acuminatus Cartwright, 1944
Aphodius (Cryptoscatomaseter) criddlei Brown, 1928
Aphodius (Cryptoscatomaseter) depressiusculus (Schmidt, 1907)⇒ Cryptoscatomaseter depressiusculus Schmidt, 1907
Aphodius (Cryptoscatomaseter) haldemani Horn, 1887
Aphodius (Cryptoscatomaseter) oklahomensis (Brown, 1928)⇒ Cryptoscatomaseter oklahomensis Brown, 1928
Aphodius (Cryptoscatomaseter) rossi (Cartwright, 1944)⇒ Cryptoscatomaseter rossi Cartwright, 1944
Aphodius (Cryptoscatomaseter) umbricollis Fall, 1907
Aphodius (Dellacasiellus) claudus (Fall, 1932)⇒ Dellacasiellus claudus Fall, 1932
Aphodius (Dellacasiellus) concavus (Horn, 1887)⇒ Dellacasiellus concavus Say, 1823
Aphodius (Dellacasiellus) fucosus Schmidt, 1922
Aphodius (Dellacasiellus) kirni Cartwright, 1944
Aphodius (Dellacasiellus) laevigatus (Haldeman, 1848)⇒ Aphodiellus impunctatus Waterhouse, 1875
Aphodius (Dellacasiellus) ruficlarus (Fall, 1932)⇒ Dellacasiellus ruficlarus Fall, 1932
Aphodius (Dialytodius) decipiens (Horn, 1887)⇒ Dialytodius decipiens Horn, 1887
Aphodius (Diapterna) hamatus Say, 1824
Aphodius hamatus garmani (Brown, 1929)⇒ Diapterna hamata Say, 1824
Aphodius (Drepanocanthoides) walshi G. Horn, 1870
Aphodius (Drepanocanthoides) walshii Horn, 1870
Aphodius (Erytus) aequalis (A. SCHMIDT, 1907)⇒ Erytus aequalis Schmidt, 1907
Aphodius (Erytus) cognatus (Fairmaire, 1860)⇒ Alocoderus hydrochaeris Fabricius, 1798
Aphodius (Esymus) merdarius (Fabricius, 1775)⇒ Esymus merdarius Fabricius, 1775
Aphodius (Esymus) pusillus Herbst, 1789
Aphodius pusillus pusillus
Aphodius (Eudolus) quadriguttatus (Herbst, 1783)⇒ Eudolus quadriguttatus Herbst, 1783
Aphodius (Euorodalus) coenosus (Panzer, 1798)⇒ Euorodalus coenosus Panzer, 1798
Aphodius (Euorodalus) paracoenosus (Balthasar & ..., 1960)⇒ Euorodalus paracoenosus Balthasar & ..., 1960
Aphodius (Euorodalus) pusillus (Herbst, 1789)⇒ Esymus pusillus Herbst, 1789
Aphodius (Euorodalus) tersus (Rosenhauer, 1856)⇒ Euorodalus tersus Erichson, 1848
Aphodius (Eupleurus) subterraneus (Linnaeus, 1758)⇒ Eupleurus subterraneus Linnaeus, 1758
Aphodius subterraneus subterraneus
Aphodius (Flaviellus) perfimbriatus (Gordon, 1977)⇒ Flaviellus perfimbriatus Gordon, 1977
Aphodius (Geomyphilus) insolitus (Brown, 1928)⇒ Geomyphilus insolitus Brown, 1928
Aphodius (Geomyphilus) rubiginosus (Horn, 1870)⇒ Geomyphilus rubiginosus Horn, 1870
Aphodius (Gonaphodiellus) bimaculosus (Schmidt, 1909)⇒ Gonaphodiellus bimaculosus Schmidt, 1909
Aphodius (Gonaphodiellus) xalapensis (Galante & St..., 2003)⇒ Gonaphodiellus bimaculosus Schmidt, 1909
Aphodius (Gonaphodius) scoparius Harold, 1877
Aphodius (Haroldiellus) sallei (Harold, 1863)⇒ Haroldiellus sallei Harold, 1863
Aphodius (Hemicyclium) unicornutus A.Schmidt, 1909
Aphodius (Hornosus) larreae Horn, 1887
Aphodius (Irrasinus) stupidus (Horn, 1870)⇒ Irrasinus stupidus Horn, 1870
Aphodius (Labarrus) lividus (Olivier, 1789)⇒ Labarrus lividus Olivier, 1789
Aphodius (Labarrus) pseudolividus (Balthasar, 1941)⇒ Labarrus pseudolividus Balthasar, 1941
Aphodius (Lechorodius) lutulentus (Haldeman, 1843)⇒ Lechorodius lutulentus Haldeman, 1843
Aphodius (Lechorodius) terminalis (Say, 1823)⇒ Lechorodius terminalis Say, 1823
Aphodius (Limarus) maculatus (Sturm, 1800)⇒ Limarus maculatus Sturm, 1800
Aphodius (Limarus) zenkeri (Germar, 1813)⇒ Limarus zenkeri Germar, 1813
Aphodius (Lindbergianus) solitarius Petrovitz, 1961
Aphodius (Liothorax) alternatus Horn, 1870
Aphodius (Liothorax) niger (Panzer, 1797)⇒ Planolinellus vittatus Say, 1825
Aphodius (Liothorax) plagiatus (Linnaeus, 1767)⇒ Liothorax plagiatus Linnaeus, 1767
Aphodius (Lithorax) linearis (Reiche & Saulcy, 1856)⇒ Mendidaphodius linearis Reiche & Saulcy, 1856
Aphodius (Loboparius) kotoensis Miwa, 1930
Aphodius (Loraphodius) suarius (Faldermann, 1835)⇒ Loraphodius suarius Faldermann, 1835
Aphodius (Lorapsis) frater (Mulsant & Rey, 1869)⇒ Loraspis frater Mulsant & Rey, 1870
Aphodius (Loraspis) frater Mulsant & Rey, 1870
Aphodius (Lunaphodius) lunifer Solsky, 1876
Aphodius (Luxolinus) luxatus Horn, 1887
Aphodius (Maculaphodius) conspersus Horn, 1887
Aphodius (Mecynodes) striatulus (Waltl, 1835)⇒ Mecynodes striatulus Waltl, 1835
Aphodius (Megateloides) dimidiatus (Roth, 1851)⇒ Megateloides dimidiatus Roth, 1851
Aphodius (Megatelus) contractus Klug, 1845
Aphodius contractus scolytoides Lucas, 1846
Aphodius (Melinopterus) consentaneus (Dejean, 1833)⇒ Coptochirus excisus Harold, 1868
Aphodius (Melinopterus) consputus (Creutzer, 1799)⇒ Melinopterus consputus Creutzer, 1799
Aphodius (Melinopterus) femoralis (Sahlberg, 1900)⇒ Teuchestes fossor Linnaeus, 1758
Aphodius (Melinopterus) prodromus (Braham, 1790)⇒ Melinopterus prodromus Brahm, 1790
Aphodius (Melinopterus) pubescens (Sturm, 1800)⇒ Melinopterus pubescens Sturm, 1800
Aphodius (Melinopterus) punctatosulcatus (Sturm, 1805)⇒ Melinopterus punctatosulcatus punctatosulcatus Sturm, 1805
Aphodius (Melinopterus) reyi (Reitter, 1892)⇒ Melinopterus reyi Reitter, 1892
Aphodius (Melinopterus) sabulicola (Thomson, 1868)⇒ Melinopterus punctatosulcatus hirtipes Fischer von ..., 1844
Aphodius (Melinopterus) sphacelatus (Panzer, 1798)⇒ Melinopterus sphacelatus Panzer, 1798
Aphodius (Melinopterus) stolzi (Reitter, 1906)⇒ Melinopterus stolzi Reitter, 1906
Aphodius (Melinopterus) tingens (Reitter, 1892)⇒ Melinopterus tingens Reitter, 1892
Aphodius (Mendidius) aculeatus Robinson, 1940
Aphodius (Merogyrus) rotundiceps (Fall, 1907)⇒ Merogyrus rotundiceps Fall, 1907
Aphodius (Microteuchestes) brusewitzi Landin, 1974
Aphodius (Neagolius) amblyodon (K. Daniel, 1900)⇒ Neagolius amblyodon Daniel, 1900
Aphodius (Neagolius) heydeni Harold, 1871
Aphodius (Neagolius) liguricus (J. Daniel, 1902)⇒ Neagolius liguricus Daniel, 1902
Aphodius (Neagolius) limbolarius Reitter, 1892
Aphodius limbolarius danielorum A. Semenov, 1902
Aphodius limbolarius limbolarius Reitter, 1892
Aphodius (Neagolius) montanus (Schmidt, 1840)⇒ Neagolius montanus Erichson, 1848
Aphodius (Neagolius) montivagus (Erichson, 1848)⇒ Neagolius montivagus Erichson, 1848
Aphodius (Neagolius) penninus J. Daniel, 1902
Aphodius (Neagolius) pollicatus Erichson, 1848
Aphodius (Neagolius) praecox Erichson, 1848
Aphodius (Neagolius) schlumbergeri (Seidlitz, 1891)⇒ Neagolius schlumbergeri Seidlitz, 1891
Aphodius schlumbergeri consobrinus
Aphodius schlumbergeri samniticus J. Daniel, 1902
Aphodius (Neocolobopterus) abyssinicus Harold, 1861
Aphodius (Neodiapterna) erichsoni Harold, 1861
Aphodius (Nialus) insularis (Petrovitz, 1961)⇒ Labarrus hoabinhensis Balthasar, 1946
Aphodius (Nialus) kraatzi Harold, 1868
Aphodius (Nialus) lividus (Olivier, 1789)⇒ Acrossus luridus Fabricius, 1775
Aphodius (Nialus) nigrita (Fabricius, 1801)⇒ Nialaphodius nigrita Fabricius, 1801
Aphodius (Nialus) plagiatus (Linnaeus, 1758)⇒ Liothorax plagiatus Linnaeus, 1767
Aphodius (Nialus) sturmi (Harold, 1870)⇒ Subrinus sturmi Harold, 1870
Aphodius (Nialus) varians (Duftschmid, 1805)⇒ Nialus varians Duftschmid, 1805
Aphodius (Nimbus) affinis (Lucas, 1846)⇒ Agrilinus ater De Geer, 1774
Aphodius (Nimbus) contaminatus (Herbst, 1783)⇒ Nimbus contaminatus Herbst, 1783
Aphodius (Nimbus) franzinii (Pittino, 1978)⇒ Nimbus franzinii Pittino, 1978
Aphodius (Nimbus) johnsoni Baraud, 1976
Aphodius (Nimbus) obliteratus (Panzer, 1823)⇒ Nimbus obliteratus Panzer, 1823
Aphodius (Nobius) bonnairei (Reitter, 1892)⇒ Nobiellus bonnairei Reitter, 1892
Aphodius (Nobius) gresseri Semenov, 1899
Aphodius (Nobius) serotinus Creutzer, 1799
Aphodius (Oromus) alpinus (Scopoli, 1763)⇒ Euheptaulacus carinatus carinatus Germar, 1824
Aphodius (Oscarinus) abusus (Fall, 1907)⇒ Mendidius crassulus Horn, 1870
Aphodius (Oscarinus) brimleyi (Cartwright, 1939)⇒ Oscarinus brimleyi Cartwright, 1939
Aphodius (Oscarinus) crassuloides (Fall, 1907)⇒ Oscarinus crassuloides Fall, 1907
Aphodius (Oscarinus) rusicola (Melsheimer, 1845)⇒ Oscarinus rusicola Melsheimer, 1845
Aphodius (Oscarinus) texensis (Cartwright, 1972)⇒ Oscarinus texensis Cartwright, 1972
Aphodius (Otophorus) haemorrhoidalis (Linnaeus, 1758)⇒ Otophorus haemorrhoidalis Linnaeus, 1758
Aphodius (Paradeloparius) circumdatus (Klug, 1855)⇒ Otophorus haemorrhoidalis Linnaeus, 1758
Aphodius (Paradidactylia) venloides Endrodi, 1960
Aphodius (Parammoecius) corvinus (Haldeman, 1848)⇒ Parammoecius corvinus Erichson, 1848
Aphodius (Parammoecius) gibbus (Germar, 1817)⇒ Parammoecius gibbus Germar, 1817
Aphodius (Parammoecius) pyrenaeus (Jacquelin du..., 1860)⇒ Parammoecius pyrenaeus Jacquelin du..., 1863
Aphodius (Paranimbus) peruanus Erichson, 1834
Aphodius (Pardalosus) pardalis LeConte, 1857
Aphodius (Pardalosus) pumilio (Schmidt, 1907)⇒ Pardalosus pumilio Schmidt, 1907
Aphodius (Pardalosus) serval (Say, 1835)⇒ Pardalosus serval Say, 1835
Aphodius (Pardalosus) slevini (Van Dyke, 1928)⇒ Pardalosus slevini Van Dyke, 1928
Aphodius (Paremadus) isaburoi Nakane, 1967
Aphodius (Paulianellus) maderi Balthasar, 1938
Aphodius (Phaeaphodius) rectus Motschulsky, 1866
Aphodius (Phalacronothus) biguttatus Dallatorre, 1879
Aphodius (Phalacronothus) pseudoavunculus Masumoto, 1977
Aphodius (Phalacronothus) quadrimaculatus Linné, 1761
Aphodius quadrimaculatus quadrimaculatus
Aphodius (Phalocronothus) biguttatus (Dallatorre, 1879)⇒ Phalacronothus biguttatus Germar, 1824
Aphodius (Phalocronothus) quadrimaculatus (Linnaeus, 1761)⇒ Teuchestes brachysomus Solsky, 1874
Aphodius (Pharaphodius) marginellus (Fabricius, 1781)⇒ Ataenius luteomargo Chapin, 1940
Aphodius (Pharaphodius) priscus Motschulsky, 1858
Aphodius (Plagiogonus) arenarius Olivier, 1789
Aphodius (Plagiogonus) putridus (Herbst, 1789)⇒ Planolinoides borealis Gyllenhal, 1827
Aphodius (Planolinellus) vittatus Mulsant, 1842
Aphodius (Planolinoides) borealis (Gyllenhal, 1827)⇒ Planolinoides borealis Gyllenhal, 1827
Aphodius (Planolinoides) neotomae Fall & Cocke..., 1907
Aphodius (Planolinoides) pectoralis (Guillebeau, 1888)⇒ Planolinoides pectoralis LeConte, 1857
Aphodius (Planolinus) borealis (Gyllenhal, 1827)⇒ Planolinoides borealis Gyllenhal, 1827
Aphodius (Planolinus) fasciatus (Olivier, 1789)⇒ Limarus maculatus Sturm, 1800
Aphodius (Planolinus) uliginosus (Hardy, 1847)⇒ Planolinus fasciatus Olivier, 1789
Aphodius (Pleuraphodius) lewisi C. O. Waterh..., 1875
Aphodius (Pleuraphodius) mutilus A.Schmidt, 1911
Aphodius (Podotenoides) jeannelianus Balthasar, 1945
Aphodius (Podotenus) gracilipes (Harold, 1867)⇒ Podotenus gracilipes Harold, 1867
Aphodius (Podotenus) suberosus (Blackburn, 1892)⇒ Podotenus suberosus Blackburn, 1904
Aphodius (Pseudacrossus) cribricollis Lucas, 1849
Aphodius (Pseudacrossus) grombczewskyi D.Koshant., 1912
Aphodius (Pseudacrossus) suffertus A. SCHMIDT, 1916
Aphodius suffertus ampliatus Reitter, 1892
Aphodius suffertus defiorei Fiori, 1907
Aphodius (Pseudacrossus) thermicola (Sturm, 1800)⇒ Amidorus thermicola Sturm, 1800
Aphodius (Pseudagolius) bicolor (Dallatorre, 1879)⇒ Aphodius fimetarius Linnaeus, 1758
Aphodius (Pseudagolius) coloradensis G. Horn, 1870
Aphodius (Pseudocalamosternus) rigidus Balthasar, 1936
Aphodius (Pseudopharaphodius) anthrax Gerstaecker, 1871
Aphodius (Pseudoteuchestes) schirparakensis Petrovitz, 1955
Aphodius (Pubinus) tomentosus (O.F.Muller, 1776)⇒ Pubinus tomentosus O.F.Muller, 1776
Aphodius (Rugaphodius) rugatus (Schmidt, 1907)⇒ Rugaphodius rugatus Schmidt, 1907
Aphodius (Scabrostomus) peculiosus Schmidt, 1922
Aphodius (Scabrostomus) sepultus (Cartwright, 1944)⇒ Scabrostomus sepultus Cartwright, 1944
Aphodius (Schaefferellus) arizonensis (Schaeffer, 1907)⇒ Schaefferellus arizonensis Schaeffer, 1907
Aphodius (Sigorius) porcus Fabricius, 1..., 1869
Aphodius (Sigorus) porcus (Fabricius, 1792)⇒ Sigorus porcus Fabricius, 1792
Aphodius (Sinodiapterna) troitzkyi Jacobson, 1897
Aphodius (Sinodiapterna) zeni Ochi, 1991
Aphodius (Sitiphus) brevitarsis (Reitter, 1894)⇒ Australaphodius frenchi Blackburn, 1892
Aphodius (Stenotothorax) badipes Melsheimer, 1844
Aphodius (Stenotothorax) oblongus (Say, 1823)⇒ Aphodius badipes Melsheimer, 1844
Aphodius (Stenotothorax) ovipennis Horn, 1870
Aphodius (Stenotothorax) sparsus (Barret, 1932)⇒ Stenotothorax sparsus LeConte, 1878
Aphodius (Strigodius) robinsoni (Cartwright, 1939)⇒ Strigodius robinsoni Cartwright, 1939
Aphodius (Subrinus) sturmi (Harold, 1870)⇒ Subrinus sturmi Harold, 1870
Aphodius (Subrinus) vitellinus (Klug, 1845)⇒ Subrinus vitellinus Klug, 1845
Aphodius (Sussorca) lusingae Paulian, 1954
Aphodius (Symphodon) anomalus Harold, 1874
Aphodius (Tetraclipeoides) acutissimus (Gordon, 1976)⇒ Tetraclipeoides acutissimus Gordon, 1976
Aphodius (Tetraclipeoides) aemulus (Horn, 1887)⇒ Tetraclipeoides aemulus Horn, 1887
Aphodius (Tetraclipeoides) anomaliceps (Brown, 1929)⇒ Tetraclipeoides anomaliceps Brown, 1929
Aphodius (Tetraclipeoides) denticulatus Haldeman, 1848
Aphodius (Tetraclipeoides) dentiger (LeConte, 1858)⇒ Tetraclipeoides dentiger LeConte, 1858
Aphodius (Tetraclipeoides) dentigerulus Brown, 1929
Aphodius (Tetraclipeoides) formidatus Gordon, 1976
Aphodius (Tetraclipeoides) moquinus Fall, 1907
Aphodius (Tetraclipeoides) testaceiventris Fall, 1932
Aphodius (Teuchestes) analis (Gistel, 1857)⇒ Rhodaphodius foetens Fabricius, 1787
Aphodius (Teuchestes) dejeani Harold, 1862
Aphodius (Teuchestes) fossor (Linnaeus, 1758)⇒ Teuchestes fossor Linnaeus, 1758
Aphodius (Trichaphodiellus) brasiliensis Laporte de C..., 1840
Aphodius (Trichaphodioides) calcaratus Boheman, 1857
Aphodius (Trichaphodius) humilis Roth, 1851
Aphodius (Trichaphodius) reichei Harold, 1859
Aphodius (Trichonotulus) scrofa (Fabricius, 1787)⇒ Trichonotulus scrofa Fabricius, 1787
Aphodius (Volinus) sticticus Panzer, 1798
Aphodius abeillei Sietti, 1903
Aphodius aboriginalis H. F. Wickham, 1912
Aphodius acerbus Horn, 1887
Aphodius aenictus Cooper & Gordon, 1987
Aphodius aggregatus (Gordon, 1977)⇒ Flaviellus aggregatus Gordon, 1977
Aphodius albertanus (Brown, 1928)⇒ Agoliinus albertanus Brown, 1928
Aphodius albertisi Harold, 1877
Aphodius aleutus Eschscholtz, 1822
Aphodius ambiguus (Boheman, 1858)⇒ Australaphodius frenchi Blackburn, 1892
Aphodius ambiquus
Aphodius angulatus (Schmidt, 1907)⇒ Orodaliscus angulatulus Schmidt, 1916
Aphodius angustatus (Mulsant, 1842)⇒ Melinopterus prodromus Brahm, 1790
Aphodius angustus (Philippi & P..., 1864)⇒ Agoliinus hatchi Hatch, 2007
Aphodius annamariae Baraud, 1982
Aphodius anomalipus Péringuey, 1901
Aphodius anteactus Krell, 2000
Aphodius anthracinus (Sturm, 1843)⇒ Parammoecius gibbus Germar, 1817
Aphodius apicalis (Schilsky, 1888)⇒ Calamosternus granarius Linnaeus, 1767
Aphodius aquilonarius (Brown, 1928)⇒ Agoliinus aquilonarius Brown, 1928
Aphodius ardoei (Landin, 1954)⇒ Agoliinus nemoralis Erichson, 1848
Aphodius armaticeps Péringuey, 1901
Aphodius armatus (Roth, 1851)⇒ Craterocyphus rhinocerus Reiche, 1847
Aphodius asellus Schmidt, 1907
Aphodius asphaltinus Kolenati, 1846
Aphodius ataeniodes (Hinton, 1938)⇒ Gonaphodioides chapini Hinton, 1934
Aphodius atwateri (Cartwright, 1944)⇒ Cryptoscatomaseter oklahomensis Brown, 1928
Aphodius australasiae (Boheman, 1858)⇒ Ataenius castanicolor Motschulsky, 1858
Aphodius badius (Sturm, 1826)⇒ Calamosternus unicolor Olivier, 1789
Aphodius baeticus Mulsant & Rey, 1870
Aphodius baraudi (Villarreal, 1975)⇒ Euorodalus boiteli Théry, 1918
Aphodius bayeri (Endrödi, 1956)⇒ Nialaphodius calidus Harold, 1871
Aphodius beloni Mulsant & Go..., 1879
Aphodius biauriculatus (Clement, 1962)⇒ Anomius permixtus Clément & Pe..., 1971
Aphodius bicoloratus (Schmidt, 1908)⇒ Aphodius fimetarius Linnaeus, 1758
Aphodius bidens (LeConte, 1878)⇒ Agoliinus bidentatus Schmidt, 1906
Aphodius bidentatus (Schmidt, 1906)⇒ Agoliinus bidentatus Schmidt, 1906
Aphodius bidentellus (A. SCHMIDT, 1910)⇒ Mendidius fimbriolatus Mannerheim, 1849
Aphodius bidentulus (Fairmaire, 1892)⇒ Trichonotulus pilosellus Schmidt, 1907
Aphodius bifasciatus (Dalla Torre, 1879)⇒ Chilothorax melanostictus Schmidt, 1840
Aphodius blaisdelli (Fall, 1905)⇒ Stenotothorax sparsus LeConte, 1878
Aphodius bonariensis Klug, 1829
Aphodius bonvouloiri (Harold, 1860)⇒ Agolius bonvouloiri Harold, 1860
Aphodius boryslavicus A. M. Lomnicki, 1894
Aphodius bosniaskii A. Handlirsch, 1907
Aphodius brancsiki Reitter, 1899
Aphodius brevicollis (LeConte, 1878)⇒ Cryptoscatomaseter brevicollis LeConte, 1878
Aphodius brevipennis (O. Heer, 1862)⇒ Aphodius foetidus Herbst, 1783
Aphodius browni (Hinton, 1934)⇒ Cryptoscatomaseter browni Hinton, 1934
Aphodius brunneus (Schilsky, 1888)⇒ Erytus cognatus Fairmaire, 1860
Aphodius bryanti (Brown, 1929)⇒ Setodius bryanti Brown, 1929
Aphodius buettikeri (R. Paulian, 1980)⇒ Erytus pruinosus Reitter, 1892
Aphodius cadaverinus (Mannerheim, 1843)⇒ Aphodius rugoclypeus Hinton, 1934
Aphodius calgaryensis (Balthasar, 1970)⇒ Agoliinus corruptor Brown, 1929
Aphodius calichromus Balthasar, 1932
Aphodius caminarius (Faldermann, 1835)⇒ Pseudacrossus tenebricosus Schmidt, 1922
Aphodius campestris (Blatchley, 1912)⇒ Alloblackburneus aegrotus Horn, 1870
Aphodius cancelliventris (Motschulsky, 1858)⇒ Ataenius australasiae Boheman, 1858
Aphodius capicola (Harold, 1862)⇒ Acrossus rufipes Linnaeus, 1758
Aphodius capitulatus (Clouët, 1898)⇒ Didactylia capitulata Clouët, 1898
Aphodius captivus (Cartwright, 1944)⇒ Cryptoscatomaseter captivus Cartwright, 1944
Aphodius cardinalis (Reitter, 1892)⇒ Aphodius fimetarius Linnaeus, 1758
Aphodius carinulatus (Motschulsky, 1864)⇒ Ammoecioides spectabilis tricarinulatus Schmidt, 1907
Aphodius carri (Brown, 1929)⇒ Dialytodius carri Brown, 1929
Aphodius caseyi Saylor, 1940
Aphodius caspicus
Aphodius caspius Ménétriés, 1832
Aphodius castanicolor (Motschulsky, 1858)⇒ Ataenius castanicolor Motschulsky, 1858
Aphodius catulus (Balthasar, 1946)⇒ Australaphodius frenchi Blackburn, 1892
Aphodius chapini (Hinton, 1934)⇒ Gonaphodiellus chapini Hinton, 1934
Aphodius charauxi L. Piton, 1940
Aphodius charmionus (Bates, 1887)⇒ Blackburneus charmionus Bates, 1887
Aphodius chokaiensis (Nomura & Nakane, 1951)⇒ Gilletianus comatus Schmidt, 1920
Aphodius circumcinctus W. L. E. Sch..., 1840
Aphodius citellorum Semenov & S...., 1929
Aphodius clypeatus Fischer von ..., 1821
Aphodius columbiensis (Wickham, 1913)⇒ Caligodorus opacus LeConte, 1872
Aphodius comanchi (Robinson, 1940)⇒ Pseudagolius bicolor Say, 1823
Aphodius communis (Broun, 1880)⇒ Saprosites communis Broun, 1880
Aphodius conigatus Panzer, 1795
Aphodius connexus (Klug, 1855)⇒ Trichaphodius copulatus Schmidt, 1916
Aphodius consociatus Horn, 1887
Aphodius conspicatus (Brullé, 1839)⇒ Chilothorax hieroglyphicus Klug, 1845
Aphodius constrictus Robinson, 1946
Aphodius coquilletti (Linell, 1896)⇒ Cinacanthus coquilletti Linell, 1896
Aphodius corallifer Koshantschikov, 1913
Aphodius corniculatus (Boucomont, 1933)⇒ Neoheptaulacus puberulus Boheman, 1857
Aphodius costatulus (Endrödi, 1964)⇒ Pleuraphodius burorum Endrödi, 1980
Aphodius costulatus (Fairmaire, 1849)⇒ Nobius costatellus Schmidt, 1922
Aphodius crassulus (Horn, 1870)⇒ Oscarinus crassulus Horn, 1870
Aphodius crassus (A. SCHMIDT, 1916)⇒ Adeloparius overlaeti Boucomont, 1936
Aphodius crenatus (Dejean, 1833)⇒ Pharaphodius crenatus Harold, 1862
Aphodius crenicollis Fall, 1932
Aphodius cressoni (Robinson, 1946)⇒ Dellacasiellus concavus Say, 1823
Aphodius cretaceous G. V. Nikolajev, 2008
Aphodius cribratulus Schmidt, 1922
Aphodius cribratus (LeConte, 1878)⇒ Calamosternus granarius Linnaeus, 1767
Aphodius crudus (A. SCHMIDT, 1916)⇒ Otophorus haemorrhoidalis Linnaeus, 1758
Aphodius cruentatus (Dejean, 1833)⇒ Planolinellus vittatus Say, 1825
Aphodius crux Wiedemann, 1823
Aphodius cuniculus (Chevrolat, 1864)⇒ Nialaphodius nigrita Fabricius, 1801
Aphodius curtulus Harold, 1866
Aphodius curtus (Haldeman, 1848)⇒ Oscarinus rusicola Melsheimer, 1845
Aphodius cylindricus (Dejean, 1833)⇒ Mendidaphodius linearis Reiche & Saulcy, 1856
Aphodius dalmatinus (Schmidt, 1840)⇒ Amidorus cribrarius Brullé, 1832
Aphodius davisi (Fall, 1930)⇒ Pardalosus slevini Van Dyke, 1928
Aphodius davisianus
Aphodius decipens
Aphodius deplanatus (Roth, 1851)⇒ Chilothorax flammulatus Harold, 1876
Aphodius dialytoides (Fall & Cocke..., 1907)⇒ Dialytellus dialytoides Fall & Cocke..., 1907
Aphodius diecki (Harold, 1870)⇒ Phalacronothus quadrimaculatus Linnaeus, 1761
Aphodius dilaticollis Saylor, 1935
Aphodius discoidalis (Gistel, 1857)⇒ Melinopterus prodromus Brahm, 1790
Aphodius divisus A. SCHMIDT, 1908
Aphodius dixoni (Oke, 1925)⇒ Podotenus callabonnensis Blackburn, 1895
Aphodius dubiosus Péringuey, 1901
Aphodius duplex (Bates, 1887)⇒ Planolinoides duplex LeConte, 1878
Aphodius dyspistus (Skelley & Wo..., 1991)⇒ Scabrostomus dyspistus Skelley & Wo..., 1991
Aphodius ealaensis (Endrödi, 1956)⇒ Adeloparius formosus Paulian, 1942
Aphodius edai (Balthasar, 1956)⇒ Phaeaphodius rectus Motschulsky, 1866
Aphodius edmontonus (Brown, 1929)⇒ Setodius edmontonus Brown, 1929
Aphodius elongatulus (Fabricius, 1801)⇒ Alocoderus elongatulus Fabricius, 1801
Aphodius epotus (Dellacasa, 1983)⇒ Chilothorax conspurcatus Linnaeus, 1758
Aphodius equestris (Panzer, 1798)⇒ Volinus sticticus Panzer, 1798
Aphodius essigi (Saylor, 1935)⇒ Geomyphilus essigi Saylor, 1935
Aphodius euprosopus (Bates, 1887)⇒ Agrilinellus azteca Harold, 1863
Aphodius evanescens (Bouskell, 1901)⇒ Planolinoides borealis Gyllenhal, 1827
Aphodius excavatus (Robinson, 1940)⇒ Cryptoscatomaseter excavatus Robinson, 1940
Aphodius excultus (Péringuey, 1908)⇒ Dilorditomaeus excultus Péringuey, 1908
Aphodius explanatus (LeConte, 1878)⇒ Agoliinus explanatus LeConte, 1878
Aphodius faldermanni (Sperk, 1835)⇒ Bodilus lugens Creutzer, 1799
Aphodius fasciger Harold, 1881
Aphodius fenyesi (Van Dyke, 1947)⇒ Coelotrachelus rudis LeConte, 1878
Aphodius fimbripes (Brown, 1928)⇒ Tetraclipeoides fimbripes Brown, 1928
Aphodius fimitarius C. Linnaeus, 1758
Aphodius floridanus (Robinson, 1947)⇒ Oscarinus floridanus Robinson, 1947
Aphodius florissantensis H. F. Wickham, 1911
Aphodius forcipatus (Harold, 1868)⇒ Mecynodes leucopterus Klug, 1845
Aphodius fordi (Gordon, 1974)⇒ Blackburneus fordi Gordon, 1974
Aphodius fortunus (Fall, 1932)⇒ Tetraclipeoides aemulus Horn, 1887
Aphodius frenchi (Blackburn, 1892)⇒ Australaphodius frenchi Blackburn, 1892
Aphodius fugitivus Péringuey, 1901
Aphodius fuliginosus (Harold, 1863)⇒ Cephalocyclus fuliginosus Harold, 1863
Aphodius fulvescens (Dejean, 1833)⇒ Trichonotulus fulvescens Harold, 1874
Aphodius fumosus Mulsant, 1842
Aphodius geminatus (MacLeay, 1871)⇒ Demarziella geminata MacLeay, 1871
Aphodius genialis Péringuey, 1901
Aphodius gentilis Horn, 1887
Aphodius geomysi (Cartwright, 1939)⇒ Blackburneus geomysi Cartwright, 1939
Aphodius gestroi (Harold, 1877)⇒ Ataenius australasiae Boheman, 1858
Aphodius goffi (Cartwright, 1939)⇒ Dellacasiellus laevigatus Haldeman, 1848
Aphodius gordoni (Ratcliffe, 1988)⇒ Flaviellus gordoni Ratcliffe, 1988
Aphodius gracilis Boheman, 1857
Aphodius granarioides H. F. Wickham, 1913
Aphodius granosus
Aphodius gravis Fall, 1927
Aphodius gregori (Balthasar, 1941)⇒ Brachiaphodius pilosus Harold, 1874
Aphodius grombczewskii
Aphodius guangdongensis Maté, 2008
Aphodius guillebeaui (Reitter, 1891)⇒ Melinopterus guillebeaui Reitter, 1892
Aphodius guttatus Eschscholtz, 1823
Aphodius haagi (Becker, 1867)⇒ Liothorax kraatzi Harold, 1868
Aphodius haroldi D. Koshantsc..., 1894
Aphodius haroldianus (Balthasar, 1932)⇒ Colobopterus quadratus Reiche, 1847
Aphodius haywardi (Brown, 1928)⇒ Agoliinus canadensis Garnett, 1920
Aphodius helvolus G. Statz, 1952
Aphodius henryi Gordon, 1976
Aphodius hepaticolor (Quedenfeldt, 1884)⇒ Neoheptaulacus hepaticolor Quedenfeldt, 1884
Aphodius hibernalis Nakane & Tsu..., 1956
Aphodius hirsutus Brown, 1928
Aphodius hirtipes (Fischer von ..., 1844)⇒ Melinopterus punctatosulcatus Sturm, 1805
Aphodius holdereri Reitter, 1900
Aphodius hubbelli Skelley & Wo..., 1991
Aphodius humeralis (LeConte, 1878)⇒ Dialytellus humeralis LeConte, 1868
Aphodius hyperboreus LeConte, 1850
Aphodius hypocoprus (Gistel, 1857)⇒ Chilothorax distinctus distinctus O. F. Müller, 1776
Aphodius igai Nakane, 1956
Aphodius illigeri (Dallatorre, 1879)⇒ Bodilopsis rufus Moll, 1782
Aphodius incommunis (Fall, 1932)⇒ Agoliinus incommunis Fall, 1932
Aphodius indistinctus (Mulsant, 1842)⇒ Nimbus contaminatus Herbst, 1783
Aphodius informis (Mulsant, 1842)⇒ Acrossus luridus Fabricius, 1775
Aphodius infuscatopennis (Schmidt, 1909)⇒ Aidophus infuscatopennis Schmidt, 1909
Aphodius innexus (Say, 1835)⇒ Liothorax innexus Say, 1835
Aphodius inquinatus (Creutzer, 1799)⇒ Aphodius distinctus O.F. Müller, 1776
Aphodius insidiosus
Aphodius intercalaris Péringuey, 1901
Aphodius inundatus H. F. Wickham, 1914
Aphodius inutilis (Horn, 1887)⇒ Calamosternus granarius Linnaeus, 1767
Aphodius inyoensis (Van Dyke, 1947)⇒ Stenotothorax oviformis Robinson, 1946
Aphodius iowensis (Wickham, 1913)⇒ Cryptoscatomaseter iowensis Wickham, 1913
Aphodius irakensis (Baraud, 1977)⇒ Erytus opacior Koshantschikov, 1894
Aphodius irregularis (Westwood, 1839)⇒ Chilothorax paykulli Bedel, 1907
Aphodius jeanneli (Paulian, 1942)⇒ Aphodius jeannelianus Balthasar, 1945
Aphodius jengi Masumoto, Ki..., 2014
Aphodius kansanus (Wickham, 1905)⇒ Aidophus parcus Horn, 1887
Aphodius khuntianus (Balthasar, 1940)⇒ Aganocrossus pallidicornis Walker, 1858
Aphodius klugi (Schmidt, 1910)⇒ Erytus cognatus Fairmaire, 1860
Aphodius knausi (Fall, 1905)⇒ Aidophus parcus Horn, 1887
Aphodius koshantschikovi (Jacobson, 1911)⇒ Amidorus koshantschikovi Jacobson, 1911
Aphodius krantzi von Heyden &..., 1866
Aphodius laminicola H. F. Wickham, 1910
Aphodius lanei Saylor, 1940
Aphodius lanuginosus Péringuey, 1901
Aphodius lapponicus (Villa, 1833)⇒ Oromus alpinus Scopoli, 1763
Aphodius leptotarsis (Brown, 1928)⇒ Geomyphilus leptotarsis Brown, 1928
Aphodius leucopterus Klug, 1845
Aphodius limbatus (Germar, 1824)⇒ Rhodaphodius foetens Fabricius, 1787
Aphodius lineatosulcatus (Harold, 1859)⇒ Oxyomus lineatosulcatus Harold, 1859
Aphodius linsleyi Van Dyke, 1933
Aphodius longitarsis (Fall, 1907)⇒ Paranimbus longitarsis Harold, 1860
Aphodius lucidus (Dejean, 1862)⇒ Pharaphodius marginellus Fabricius, 1781
Aphodius lunatus Mulsant, 1842
Aphodius macgregori (Islas, 1955)⇒ Geomyphilus macgregori Islas, 1955
Aphodius maculicollis (Montrouzier, 1860)⇒ Labarrus lividus Olivier, 1789
Aphodius madari (Tesar, 1945)⇒ Nimbus makowskyi Koshantschikov, 1891
Aphodius magnificans Robinson, 1940
Aphodius magnificiens
Aphodius magnopuctatus
Aphodius magnopunctatus (Hinton, 1934)⇒ Agrilinellus ornatus Schmidt, 1911
Aphodius malkini Hatch, 1972
Aphodius manitobensis (Brown, 1928)⇒ Agoliinus manitobensis Brown, 1928
Aphodius mara (Krikken & Kaas, 1984)⇒ Didactylia szunyoghyi Endrödi, 1961
Aphodius marginalis (Stephens, 1830)⇒ Melinopterus sphacelatus Panzer, 1798
Aphodius marginatus (LeConte, 1878)⇒ Cryptoscatomaseter depressiusculus Schmidt, 1907
Aphodius martini Van Dyke, 1928
Aphodius mashunensis Péringuey, 1901
Aphodius maurus (Gebler, 1833)⇒ Agoliinus lapponum Gyllenhal, 1808
Aphodius mcdonaldi (Robinson, 1946)⇒ Oscarinus rusicola Melsheimer, 1845
Aphodius mediaevus H. F. Wickham, 1914
Aphodius metallicus (Haldeman, 1848)⇒ Calamosternus granarius Linnaeus, 1767
Aphodius mexicanus (Harold, 1862)⇒ Cephalocyclus mexicanus Harold, 1862
Aphodius meyeri Heer, 1847
Aphodius micros Walker, 1871
Aphodius mimus Péringuey, 1901
Aphodius minatorius Péringuey, 1908
Aphodius miniatorius
Aphodius miniatulus (Balthasar, 1931)⇒ Agoliinus lapponum Gyllenhal, 1808
Aphodius mirus (Schmidt, 1916)⇒ Craterocyphus rhinocerus mirus Schmidt, 1916
Aphodius mizo Nakane, 1967
Aphodius moclipsi (Hatch, 1971)⇒ Liothorax alternatus Horn, 1870
Aphodius muchei Petrovitz, 1962
Aphodius multiplex Reitter, 1897
Aphodius nanus (Horn, 1887)⇒ Aphodius asellus Schmidt, 1907
Aphodius nasutus (Dellacasa, 1983)⇒ Chilothorax melanostictus Schmidt, 1840
Aphodius neodistinctus (Brown, 1928)⇒ Pardalosus neodistinctus Brown, 1928
Aphodius nevadensis Horn, 1870
Aphodius nigerrimus (Dallatorre, 1879)⇒ Oromus alpinus Scopoli, 1763
Aphodius nigricans (Paulian, 1934)⇒ Neocolobopterus maculicollis Reiche, 1847
Aphodius nigripennis (Miksic, 1962)⇒ Loraphodius suarius Faldermann, 1835
Aphodius nigromarginatus (Cockerell, 1888)⇒ Aphodius vittatus Mulsant, 1842
Aphodius nigrotessellatus (Nakane, 1972)⇒ Chilothorax punctatus Waterhouse, 1875
Aphodius nitidus (Ballion, 1870)⇒ Plagiogonus arenarius Olivier, 1789
Aphodius obsoleteguttatus (Waterhouse, 1875)⇒ Aphodaulacus nigrotessellatus Motschulsky, 1866
Aphodius obtusus (LeConte, 1878)⇒ Ballucus obtusus LeConte, 1878
Aphodius occidentalis (Horn, 1870)⇒ Aphodius hyperboreus LeConte, 1850
Aphodius ochreipennis (Horn, 1871)⇒ Cryptoscatomaseter ochreipennis Horn, 1871
Aphodius ochripennis (Reitter, 1892)⇒ Esymus pusillus roubali Balthasar, 1932
Aphodius odocoilis (Robinson, 1939)⇒ Oscarinus odocoilis Robinson, 1939
Aphodius odocollis
Aphodius omissus LeConte, 1850
Aphodius opisthinus (Bates, 1887)⇒ Gonaphodiellus opisthius Bates, 1887
Aphodius oviformis Robinson, 1946
Aphodius palmetincolus (Karsch, 1881)⇒ Mendidius palmetincola Karsch, 1881
Aphodius panamensis (Harold, 1859)⇒ Aidophus panamensis Harold, 1859
Aphodius parcus (Horn, 1887)⇒ Aidophus parcus Horn, 1887
Aphodius partitus (Endrödi, 1957)⇒ Adeloparius luridipes Harold, 1861
Aphodius parvoniger (Lea, 1923)⇒ Ozodius neglectus Schmidt, 1913
Aphodius parvulus (Hochhuth, 1873)⇒ Esymus pusillus Herbst, 1789
Aphodius pennsylvanicus (Robinson, 1940)⇒ Pardalosus serval Say, 1835
Aphodius peregrinus (Boheman, 1857)⇒ Nobius hepaticus Roth, 1851
Aphodius perpolitus (Gordon, 1973)⇒ Aphodius subaeneus LeConte, 1857
Aphodius phaeopterus (Stephens, 1830)⇒ Esymus pusillus Herbst, 1789
Aphodius phaleroides (Horn, 1870)⇒ Flaviellus phaleroides Horn, 1870
Aphodius pholetus (Skelley & Wo..., 1991)⇒ Cryptoscatomaseter pholetus Skelley & Wo..., 1991
Aphodius piceatus (Robinson, 1946)⇒ Agoliinus piceatus Robinson, 1946
Aphodius pictipennis (Fairmaire, 1897)⇒ Didactylia pictipennis Fairmaire, 1897
Aphodius pingellus (Brown, 1929)⇒ Diapterna pinguella Brown, 1929
Aphodius pinguis Haldeman, 1848
Aphodius plasoni Kaufel, 1914
Aphodius platypleurus (Skelley & Wo..., 1991)⇒ Cryptoscatomaseter platypleurus Skelley & Wo..., 1991
Aphodius politus (Horn, 1870)⇒ Aphodius haldemani Horn, 1887
Aphodius posticus (Boheman, 1857)⇒ Pharaphodius russatus Erichson, 1842
Aphodius postpilosus Reitter, 1895
Aphodius praeemptor H. F. Wickham, 1913
Aphodius precursor G. H. Horn, 1876
Aphodius principalis (Harold, 1861)⇒ Calocolobopterus principalis Harold, 1861
Aphodius probes (Péringuey, 1901)⇒ Mesontoplatys dorsalis Klug, 1855
Aphodius productus (Boheman, 1857)⇒ Bodilus laetus Wiedemann, 1823
Aphodius protervus (Balthasar, 1973)⇒ Pseudopharaphodius rufomaculatus Petrovitz, 1958
Aphodius przewalskyi Reitter, 1887
Aphodius pseudabusus (Cartwright, 1957)⇒ Oscarinus pseudabusus Cartwright, 1957
Aphodius pseudocalcaratus (Paulian, 1954)⇒ Trichaphodius copulatus Schmidt, 1916
Aphodius pseudotasmaniae (Given, 1950)⇒ Acrossidius pseudotasmaniae Given, 1950
Aphodius pullmani Saylor, 1940
Aphodius pumilus (Horn, 1887)⇒ Pardalosus pumilio Schmidt, 1907
Aphodius punctatissimus (Dejean, 1833)⇒ Amidorus cribrarius Brullé, 1832
Aphodius puncticeps (Harold, 1862)⇒ Cephalocyclus puncticeps Harold, 1862
Aphodius punctissimus (Brown, 1928)⇒ Cryptoscatomaseter punctissimus Brown, 1928
Aphodius pygmaeus Boheman, 1857
Aphodius pyriformis Brown, 1929
Aphodius rainieri Hatch, 1972
Aphodius reginae Kral, 1997
Aphodius rehni (Robinson, 1938)⇒ Flaviellus subtruncatus LeConte, 1878
Aphodius rendalli Wollaston, 1867
Aphodius restrictus (Mulsant, 1842)⇒ Melinopterus prodromus Brahm, 1790
Aphodius rhinocerontis A. M. Lomnicki, 1894
Aphodius rhododactylus (Marsham, 1802)⇒ Aphodius arenarius Olivier, 1789
Aphodius richardsonae (Hatch, 1971)⇒ Agoliinus congregatus Mannerheim, 1853
Aphodius rogersi (Hatch, 1971)⇒ Dialytodius rogersi Hatch, 1971
Aphodius rubidus (Sturm, 1843)⇒ Alloblackburneus rubeolus Palisot de B..., 1809
Aphodius rubricatus (Paulian, 1936)⇒ Plagiogonus oberthuri Paulian, 1936
Aphodius rudii Endrodi, 1968
Aphodius ruficrus (Marsham, 1802)⇒ Sigorus porcus ruficrus Marsham, 1802
Aphodius rufipennis (Motschulsky, 1863)⇒ Parammoecius corvinus Erichson, 1848
Aphodius rugoclypeus Hinton, 1934
Aphodius rugostriatus (Reitter, 1892)⇒ Pharaphodius rugosostriatus Waterhouse, 1875
Aphodius ruricola (Melsheimer, 1845)⇒ Oscarinus rusicola Melsheimer, 1845
Aphodius russelli (Hatch, 1971)⇒ Agoliinus canadensis Garnett, 1920
Aphodius russeus (Brown, 1928)⇒ Geomyphilus russeus Brown, 1928
Aphodius ruthenus A. M. Lomnicki, 1894
Aphodius sallaei (Bates, 1887)⇒ Haroldiellus sallei Harold, 1863
Aphodius sareptanus (Balthasar, 1935)⇒ Mendidaphodius linearis Reiche & Saulcy, 1856
Aphodius saylorea (Robinson, 1940)⇒ Blackburneus saylorea Robinson, 1940
Aphodius scabriceps LeConte, 1878
Aphodius scalaris (Dellacasa, 1983)⇒ Chilothorax conspurcatus Linnaeus, 1758
Aphodius scapularis (Mulsant, 1842)⇒ Euorodalus coenosus Panzer, 1798
Aphodius schlickumi G. Statz, 1952
Aphodius schuhi (Gordon, 1975)⇒ Caligodorus tuberosus Barret, 1932
Aphodius sedulus (Harold, 1871)⇒ Planolinoides borealis Gyllenhal, 1827
Aphodius senex H. F. Wickham, 1914
Aphodius septemmaculatus (Mulsant, 1842)⇒ Chilothorax melanostictus Schmidt, 1840
Aphodius serenus (Péringuey, 1901)⇒ Haroldaphodius binodulus Harold, 1866
Aphodius setiger (Mulsant, 1842)⇒ Trichonotulus scrofa Fabricius, 1787
Aphodius sharpi Harold, 1874
Aphodius shoshonis H. F. Wickham, 1912
Aphodius silvanicus (Cartwright, 1972)⇒ Oscarinus silvanicus Cartwright, 1972
Aphodius silvaticus (Ahrens, 1812)⇒ Teuchestes fossor Linnaeus, 1758
Aphodius sinuaticeps (Boucomont, 1932)⇒ Koshantschikovius haematicus Boheman, 1857
Aphodius smithi (Brown, 1930)⇒ Cryptoscatomaseter browni Hinton, 1934
Aphodius snowi (Brown, 1929)⇒ Agoliinus cruentatus LeConte, 1878
Aphodius socialis (Brown, 1928)⇒ Cryptoscatomaseter punctissimus Brown, 1928
Aphodius spadix Schmidt, 1916
Aphodius spectabilis Péringuey, 1901
Aphodius spermophili (Gordon, 1973)⇒ Cinacanthus crenicollis Fall, 1932
Aphodius spermophilis
Aphodius spretus (Haldeman, 1848)⇒ Calamosternus granarius Linnaeus, 1767
Aphodius subaeneus LeConte, 1857
Aphodius subater A. M. Lomnicki, 1894
Aphodius subcarinatus (Robinson, 1946)⇒ Dellacasiellus ruficlarus Fall, 1932
Aphodius subcostatus H. J. Kolbe, 1886
Aphodius sublividus Balthasar, 1941
Aphodius subluteus (Mulsant, 1842)⇒ Aphodius fimetarius Linnaeus, 1758
Aphodius subsericeus Ballion, 1878
Aphodius subtruncatus (LeConte, 1878)⇒ Flaviellus subtruncatus LeConte, 1878
Aphodius sus Kugelann, 1792
Aphodius suspectus (Sharp, 1876)⇒ Setodius diogenes Gordon & Ske..., 2007
Aphodius swaneticus Reitter, 1892
Aphodius syriacus (Mulsant & Rey, 1870)⇒ Melinopterus prodromus Brahm, 1790
Aphodius talpoidesi (Brown, 1928)⇒ Cryptoscatomaseter talpoidesi Brown, 1928
Aphodius tanytarsus Skelley & Wo..., 1991
Aphodius tarsalis (Schmidt, 1907)⇒ Australaphodius frenchi Blackburn, 1892
Aphodius tenax (Balthasar, 1932)⇒ Agrilinus tenax Balthasar, 1932
Aphodius tenella (Say, 1823)⇒ Planolinus tenellus Say, 1823
Aphodius tenuitarsis (Fairmaire, 1897)⇒ Mesontoplatys dorsalis Klug, 1855
Aphodius terminatus (Marseul, 1878)⇒ Ammoecius terminatus Harold, 1869
Aphodius testaceicolor (Balthasar, 1937)⇒ Pseudopharaphodius anthrax Gerstaecker, 1871
Aphodius testudinarius Fabricius, 1775
Aphodius texana
Aphodius theobaldi Krell, 2000
Aphodius theryi (Clouët, 1896)⇒ Anomius permixtus Clément & Pe..., 1971
Aphodius thomomysi (Brown, 1928)⇒ Geomyphilus thomomysi Brown, 1928
Aphodius thoracicus (Roth, 1851)⇒ Calamosternus granarius Linnaeus, 1767
Aphodius tiligranicus (Nakane, 1966)⇒ Calamosternus granarius Linnaeus, 1767
Aphodius timidus (Boheman, 1857)⇒ Nobius hepaticus Roth, 1851
Aphodius tragicus (Schmidt, 1922)⇒ Dialytellus humeralis LeConte, 1868
Aphodius transversus (Robinson, 1940)⇒ Cephalocyclus hogei Bates, 1887
Aphodius tristis Zenker, 1801
Aphodius tuberosus Barrett, 1932
Aphodius turbatus Baudi di Selve, 1870
Aphodius ungulatus (Fall, 1901)⇒ Geomyphilus ungulatus Fall, 1901
Aphodius unifasciatus Nomura & Nakane, 1951
Aphodius utahensis (Robinson, 1938)⇒ Ballucus obtusus LeConte, 1878
Aphodius validus (Horn, 1870)⇒ Aphodius pinguis Haldeman, 1848
Aphodius vectis Krell, 2019
Aphodius verrucosus Bordat, 1996
Aphodius vestiarius (Horn, 1870)⇒ Nialaphodius nigrita Fabricius, 1801
Aphodius villarreali Baraud, 1975
Aphodius villosus (Gyllenhal, 1806)⇒ Euheptaulacus villosus Gyllenhal, 1806
Aphodius washtucha
Aphodius washtucna Robinson, 1938
Aphodius wickhami (Brown, 1928)⇒ Agoliinus wickhami Brown, 1928
Aphodius windsori (Cartwright, 1939)⇒ Oscarinus windsori Cartwright, 1939
Aphodius wolcotti (Blatchley, 1910)⇒ Aidophus parcus Horn, 1887
Aphodius yavapai (Robinson, 1946)⇒ Cryptoscatomaseter yavapai Robinson, 1946
Aphodius yukonensis (Robinson, 1948)⇒ Agoliinus corruptor Brown, 1929
Aphodius zangi A. SCHMIDT, 1906
Aphodius zebrinus (Petrovitz, 1958)⇒ Aganocrossus fugitivus Péringuey, 1901
Aphodobius amplipennis Balthasar, 1945
Aphodobius bernoni Endrödi, 1980
Aphodobius bicoloratus Schmidt, 1908
Aphodobius erugatus Harold, 1871
Aphodobius fentoni Endrödi, 1982
Aphodobius heynei Schmidt, 1911
Aphodobius innocens Schmidt, 1911
Aphodobius inornatus Schmidt, 1908
Aphodobius mulciber Balthasar, 1933
Aphodobius partitus Endrödi, 1956
Aphodobius peringueyi Schmidt, 1909
Aphodobius rubrothorax Bordat, 2005
Aphodobius tessmanni Petrovitz, 1969
Aphodobius transvaalicus Petrovitz, 1964
Aphodobius ugandanus Balthasar, 1963
Aphodobius villosulus Péringuey, 1901
Aphodobius zumpti Balthasar, 1937
Aphodoharmogaster consocius Endrödi, 1964
Aphodoharmogaster costata Endrodi, 1983
Aphodoharmogaster costipennis Boheman, 1857
Aphodopsammobius australicus Blackburn, 1904
Aphodopsammobius demarzi Petrovitz, 1964
Aphodopsammobius matthewsi (Rakovic, 1981)⇒ Aphodopsammobius rugicollis MacLeay, 1888
Aphodopsammobius mirabilis Endrodi, 1964
Aphodopsammobius rugicollis MacLeay, 1888
Aphodopsammobius zietzi Blackburn, 1895
Aphotaenius cambeforti Chalumeau, 1983
Aphotaenius carolinus Van Dyke, 1928
Aphotaenius colombiensis (Cartwright, 1952)⇒ Aphotaenius convexus Harold, 1880
Aphotaenius convexus Harold, 1880
Aphotaenius elegans Skelley & Va..., 2020
Aphotaenius gaucho Skelley & Va..., 2020
Aphotaenius hamelae Skelley & Va..., 2020
Aphotaenius howdeni Cartwright, 1963
Aphotaenius plaumanni Cartwright, 1963
Apsteiniella aralica Nikolajev, 1979
Apsteiniella aralicus Nikolajev, 1979
Apsteiniella georgii W.Koshantsch..., 1912
Apsteiniella naviauxi Baraud, 1977
Argeremazus neuquen Stebnicka & ..., 2003
Arrowiellus mulciber Balthasar, 1933
Arupaia friedenreichi Harold, 1870
Aschnarhyparus peregrinus Hinton, 1934
Ataeniopsis armasi Chalumeau, 1982
Ataeniopsis carupanoi Stebnicka, 2003
Ataeniopsis duncani Cartwright, 1974
Ataeniopsis edistoi Cartwright, 1974
Ataeniopsis figurator Harold, 1874
Ataeniopsis haroldi Steinheil, 1872
Ataeniopsis jaltipani Stebnicka, 2003
Ataeniopsis notabilis Petrovitz, 1973
Ataeniopsis parallelus Petrovitz, 1961
Ataeniopsis parkeri Cartwright, 1974
Ataeniopsis pusillus Burmeister, 1877
Ataeniopsis regulus Balthasar, 1947
Ataeniopsis rugopygus Cartwright, 1974
Ataeniopsis saxatilis Cartwright, 1944Monkfish
Ataeniopsis vinacoensis Stebnicka, 2003
Ataenius abancay Stebnicka, 2005
Ataenius abditoides Chapin, 1940
Ataenius abditus Haldeman, 1848
Ataenius abditus exiguus (Brown, 1932)⇒ Ataenius exiguus Brown, 1932
Ataenius aciculus Hinton, 1937
Ataenius acutulus Schmidt, 1913
Ataenius aequalis Harold, 1880
Ataenius aequatorialis Petrovitz, 1961
Ataenius africanus Endrödi, 1967
Ataenius alternatus Melsheimer, 1845
Ataenius ambaritae Stebnicka, 1988
Ataenius angulatus Petrovitz, 1964
Ataenius annanus Stebnicka & ..., 1997
Ataenius anticus (Fall, 1930)⇒ Ataenius platensis Blanchard, 1846
Ataenius aphodiiformis Bordat & ThÃ..., 2012
Ataenius apicalis Hinton, 1937Red pricklypoppy
Ataenius arenosus Harold, 1868
Ataenius argillaceus (Schmidt, 1916)⇒ Ataenius imbricatus Melsheimer, 1845
Ataenius armasi Chalumeau, 1982
Ataenius arrowi Hinton, 1936
Ataenius asper Petrovitz, 1975
Ataenius atramentarius Erichson, 1847
Ataenius attenuator (Harold, 1874)⇒ Ataenius abditus Haldeman, 1848
Ataenius auberti Paulian, 1937
Ataenius australasiae Boheman, 1858
Ataenius australis Harold, 1875
Ataenius balthasari Petrovitz, 1973
Ataenius barberi Cartwright, 1974
Ataenius basiceps Lea, 1923
Ataenius beattyi Chapin, 1940
Ataenius benarabyensis Stebnicka & ..., 1997
Ataenius bicolor Petrovitz, 1963
Ataenius biroi Endrödi, 1951
Ataenius bispinulosus Schmidt, 1911
Ataenius blapoides Balthasar, 1947
Ataenius bolivarensis Stebnicka, 2007
Ataenius bolivari Stebnicka, 2001
Ataenius bolivianus (Hinton, 1936)⇒ Ataenius nugator Harold, 1880
Ataenius borgmeieri Hinton, 1936
Ataenius borgmeiri
Ataenius borjae Stebnicka, 2005
Ataenius borneensis Minkina & Be..., 2023
Ataenius brevicollis Wollaston, 1854
Ataenius brevinotus (Chapin, 1940)⇒ Odontolytes denominatus Chevrolat, 1864
Ataenius brevis Fall, 1930
Ataenius brevitarsis Petrovitz, 1964
Ataenius brouni Sharp, 1876Broun's Scarab
Ataenius brunneus (Schmidt, 1922)⇒ Parataenius brunneus Schmidt, 1922
Ataenius buenavistae Stebnicka, 2001
Ataenius caesoides Fairmaire, 1899
Ataenius caesoides anjouanensis Bordat, 1990
Ataenius caesoides caesoides
Ataenius caicarae Stebnicka, 2005
Ataenius californicus Horn, 1887
Ataenius canoasus Stebnicka, 2007
Ataenius capitosus (Harold, 1867)⇒ Auperia capitosa Harold, 1867
Ataenius carinatipennis Petrovitz, 1973
Ataenius carinator Harold, 1874
Ataenius cartago Stebnicka, 2007
Ataenius cartwrighti Chalumeau & ..., 1974
Ataenius castanicolor Motschulsky, 1858
Ataenius castaniellus Bates, 1887
Ataenius catarinaensis Stebnicka, 2007
Ataenius catenulatus Erichson, 1847
Ataenius cavagnaroi Cartwright, 1970
Ataenius ceylonensis Schmidt, 1912
Ataenius chapini Hinton, 1937
Ataenius chilensis Solier, 1851
Ataenius chinacotae Stebnicka, 2007
Ataenius circulusbrumalis Petrovitz, 1964
Ataenius clavatus Schmidt, 1916
Ataenius clitellarius Petrovitz, 1973
Ataenius cochabambae Stebnicka, 2005
Ataenius cognata LeConte, 1858
Ataenius cognatus (LeConte, 1858)⇒ Ataenius liogaster Bates, 1887
Ataenius columbicus Harold, 1880
Ataenius communis Hinton, 1936
Ataenius complicatus Harold, 1869
Ataenius confertus Fall, 1909
Ataenius consors (Blackburn, 1904)⇒ Australammoecius goyderensis Blackburn, 1896
Ataenius convexus (Robinson, 1940)⇒ Haroldiataenius convexus Robinson, 1940
Ataenius coriarius Schmidt, 1912
Ataenius corporaali Boucomont, 1924
Ataenius corrosus Chapin, 1940
Ataenius costulifer Balthasar, 1951
Ataenius crenaticollis Petrovitz, 1973
Ataenius crenatipennis MacLeay, 1871
Ataenius crenator Harold, 1876
Ataenius crenatostriatus Blanchard, 1843
Ataenius crenulatus (Schmidt, 1910)⇒ Parataenius derbesis Solier, 1851
Ataenius cribrithorax Bates, 1887
Ataenius cristobalensis Cook & Peck, 2000
Ataenius cucutae Stebnicka, 2007
Ataenius cylindrus (Horn, 1871)⇒ Ataenius ovatulus Horn, 1871
Ataenius damzeni Bukejs & Ale..., 2018
Ataenius dentatus Koshantschikov, 1916
Ataenius depilis Petrovitz, 1976
Ataenius deserti Blackburn, 1894
Ataenius desertus Horn, 1871Scratchbush
Ataenius duncani (Cartwright, 1974)⇒ Ataeniopsis duncani Cartwright, 1974
Ataenius duplex (Hinton, 1936)⇒ Ataenius nugator Harold, 1880
Ataenius ecruensis Stebnicka & ..., 2005Black cod
Ataenius edistoi (Cartwright, 1974)⇒ Ataeniopsis edistoi Cartwright, 1974
Ataenius edungalbae Stebnicka & ..., 1997
Ataenius edwardsi (Chapin, 1940)⇒ Ataenius liogaster Bates, 1887
Ataenius elegans Harold, 1868
Ataenius elisaensis Stebnicka, 2002
Ataenius elongatulus MacLeay, 1888
Ataenius elongatus (Palisot de B..., 1805)⇒ Ataenius picinus Harold, 1861
Ataenius endroedyyoungai Endrödi, 1973
Ataenius erinaceus Petrovitz, 1961
Ataenius eringundae Stebnicka & ..., 1997
Ataenius erratus Fall, 1930
Ataenius europaeus G. Quiel, 1911
Ataenius eurynotus Lea, 1923
Ataenius excisicollis Petrovitz, 1961
Ataenius exiguus Brown, 1932
Ataenius fattigi Cartwright, 1948
Ataenius figurator (Harold, 1874)⇒ Ataeniopsis figurator Harold, 1874
Ataenius floreanae Cook & Peck, 2000
Ataenius forgiei Bordat & ThÃ..., 2012
Ataenius forsteri Balthasar, 1960
Ataenius frater Arrow, 1903
Ataenius freyi Petrovitz, 1961
Ataenius fukiensis (Balthasar, 1942)⇒ Ataenius australasiae Boheman, 1858
Ataenius gammonensis Stebnicka & ..., 1997
Ataenius garamas Peyerimhoff, 1929
Ataenius gascoyneensis Stebnicka & ..., 1997
Ataenius gibbus Blackburn, 1904
Ataenius gilesi Stebnicka & ..., 1997
Ataenius glabriventris Schmidt, 1911
Ataenius glaseri Cartwright, 1974
Ataenius gracilis Melsheimer, 1845
Ataenius granchacoensis (Balthasar, 1939)⇒ Ataenius platensis Blanchard, 1846
Ataenius granocostatus Schmidt, 1912
Ataenius griffini (Cartwright, 1974)⇒ Haroldiataenius griffini Cartwright, 1974
Ataenius gruneri Chalumeau, 1979
Ataenius guanacastae Stebnicka, 2005
Ataenius guayasi Stebnicka, 2001
Ataenius gungareei Stebnicka & ..., 1997
Ataenius guriensis Stebnicka, 2005
Ataenius haitianus Hinton, 1937
Ataenius haroldi Steinheil, 1872
Ataenius havanensis (Balthasar, 1938)⇒ Ataenius scabrellus Schmidt, 1909
Ataenius heinekeni Wollaston, 1854
Ataenius hesperius Cartwright, 1974
Ataenius hirsutus Horn, 1871Scratchbush
Ataenius hispaniolae Chalumeau, 1982
Ataenius hispidus Harold, 1867
Ataenius holopubescens Hinton, 1938
Ataenius horticola Harold, 1869
Ataenius howdeni Chalumeau, 1978
Ataenius hrubanti Balthasar, 1967
Ataenius huanus Stebnicka, 2007
Ataenius humptydooensis Stebnicka & ..., 1997
Ataenius hygrophilus Paulian, 1947
Ataenius icanus Balthasar, 1941
Ataenius illaetabilis Lea, 1923
Ataenius imbricatoides Schmidt, 1909
Ataenius imbricatus Melsheimer, 1845
Ataenius imparilis Blackburn, 1904
Ataenius impiger Schmidt, 1916
Ataenius impressus Petrovitz, 1963
Ataenius indutus Petrovitz, 1961
Ataenius inops (Horn, 1887)⇒ Parapsammodius puncticollis LeConte, 1858
Ataenius inquisitus Horn, 1887
Ataenius insculptus Horn, 1887
Ataenius insolitus Schmidt, 1909
Ataenius insulae Chalumeau & ..., 1974
Ataenius insularis Lea, 1923
Ataenius insulicola (Chapin, 1940)⇒ Ataenius aequalis Harold, 1880
Ataenius integer (Harold, 1868)⇒ Ataenius platensis Blanchard, 1846
Ataenius integricollis Lea, 1923
Ataenius intermedius (Bates, 1887)⇒ Ataenius polyglyptus Bates, 1887
Ataenius iquitosae Stebnicka, 2007
Ataenius isabelae Franz, 1985
Ataenius jalapensis (Bates, 1887)⇒ Ataenius peregrinator Harold, 1877
Ataenius jamaicensis Chapin, 1940
Ataenius jardinensis Stebnicka, 2002
Ataenius jelineki Chalumeau, 1982
Ataenius kapalgaensis Stebnicka & ..., 1997
Ataenius klapperichi Howden, 1978
Ataenius kochi Balthasar, 1941
Ataenius koebelei Blackburn, 1904
Ataenius koelleri Balthasar, 1963
Ataenius koghianus Paulian, 1991
Ataenius koghiensis
Ataenius kratochvili (Balthasar, 1942)⇒ Ataenius garamas Peyerimhoff, 1929
Ataenius laevifrons (Lea, 1923)⇒ Ataenius semicoecus MacLeay, 1888
Ataenius laeviventris (Horn, 1887)⇒ Ataenius lobatus Horn, 1887
Ataenius lamarensis Stebnicka, 2007
Ataenius lamotteiroyi Bordat, 1992
Ataenius lanei Petrovitz, 1973
Ataenius languidus Schmidt, 1911Antarctic dragonfishes
Ataenius latus Petrovitz, 1963
Ataenius lenkoi Petrovitz, 1973
Ataenius limbatus (Bates, 1887)⇒ Haroldiataenius limbatus Bates, 1887
Ataenius linelli (Cartwright, 1944)⇒ Ataenius languidus Schmidt, 1911
Ataenius liogaster Bates, 1887
Ataenius lobatus Horn, 1887Scratchbush
Ataenius londrinae Stebnicka, 2007
Ataenius longiclavus Petrovitz, 1970
Ataenius lorettii (Martínez, 1952)⇒ Ataenius complicatus Harold, 1869
Ataenius lucanus (Horn, 1871)⇒ Haroldiataenius lucanus Horn, 1871
Ataenius luctuosus Burmeister, 1877
Ataenius ludovicianus (Fall, 1930)⇒ Ataenius wenzelii Horn, 1887
Ataenius lueomargo
Ataenius luteomargo Chapin, 1940
Ataenius macilentus (Blackburn, 1904)⇒ Ataenius brouni Sharp, 1876
Ataenius madagassicus Bordat, 1990
Ataenius maghribinicus Baraud, 1985
Ataenius malayianus (Petrovitz, 1961)⇒ Ataenius australasiae Boheman, 1858
Ataenius martinezi Petrovitz, 1973
Ataenius miamii (Cartwright, 1934)⇒ Ataenius scabrellus Schmidt, 1909
Ataenius michelii Chalumeau, 1978
Ataenius microtrichopterus Lea, 1923
Ataenius millstreamae Stebnicka & ..., 1997
Ataenius montanus Schmidt, 1911
Ataenius monteithi Paulian, 1991
Ataenius monticola Paulian, 1940
Ataenius montreuili Bordat & ThÃ..., 2012
Ataenius morator Harold, 1869
Ataenius murchisoni Stebnicka & ..., 1997
Ataenius nakpandurii Endrödi, 1973
Ataenius nanus DeGeer, 1774
Ataenius napoensis Stebnicka, 2007
Ataenius nigricans Paulian, 1933
Ataenius nigrolineatus (Hinton, 1938)⇒ Lomanoxoides nigrolineatus Hinton, 1938
Ataenius nocturnus Nomura, 1943
Ataenius noques Stebnicka, 2007
Ataenius noronhai Stebnicka, 2007
Ataenius nudus Blackburn, 1904
Ataenius nugator Harold, 1880
Ataenius nunenmacheri (Cartwright, 1974)⇒ Ataenius peregrinator Harold, 1877
Ataenius oaxacaensis Stebnicka & ..., 2005
Ataenius oblongus (Horn, 1871)⇒ Ataenius sculptor Harold, 1868
Ataenius ocumarensis Stebnicka, 2001
Ataenius okinawensis (Nakane, 1960)⇒ Ataenius australasiae Boheman, 1858
Ataenius oklahomensis Brown, 1930
Ataenius onkonensis Stebnicka, 2005
Ataenius opacipennis Schmidt, 1910
Ataenius opatrinus Harold, 1867Antarctic blennies
Ataenius opatroides Blanchard, 1846
Ataenius orbicularis (Schmidt, 1914)⇒ Ataenius liogaster Bates, 1887
Ataenius ovatulus Horn, 1871Scratchbush
Ataenius pacificus Sharp, 1879
Ataenius palmaritoensis Stebnicka, 2007
Ataenius palmerstoni Blackburn, 1891
Ataenius palustris Montrouzier, 1860
Ataenius panamensis Hinton, 1936
Ataenius parallelipennis Petrovitz, 1973
Ataenius parkeri (Cartwright, 1974)⇒ Ataeniopsis parkeri Cartwright, 1974
Ataenius parvus Lea, 1923
Ataenius patescens S. H. Scudder, 1900
Ataenius pearlensis Stebnicka, 2006
Ataenius peregianensis Stebnicka & ..., 1997
Ataenius peregrinator Harold, 1877Birthworts
Ataenius pereirai Petrovitz, 1970
Ataenius perforatus Harold, 1867
Ataenius pertuga Balthasar, 1961
Ataenius peruvianus Minkina, 2020
Ataenius petrovitzi Balthasar, 1960
Ataenius picinus Harold, 1861Antarctic blennies
Ataenius picipes (Fleutiaux & ..., 1889)⇒ Ataenius morator Harold, 1869
Ataenius platensis Blanchard, 1846
Ataenius plaumanni Petrovitz, 1973
Ataenius polyglyptus Bates, 1887
Ataenius pseudimparilis Stebnicka & ..., 1997
Ataenius pseudocarinator Balthasar, 1947
Ataenius pseudoclavatus Stebnicka, 2005
Ataenius pseudocommunis Stebnicka, 2001
Ataenius pseudohirsutus (Cartwright, 1974)⇒ Ataenius setiger Bates, 1887
Ataenius pseudostercorator Stebnicka, 2003
Ataenius pseudousingeri Galante, Ste..., 2003
Ataenius punctatohirsutus Schmidt, 1909
Ataenius puncticollis (LeConte, 1868)⇒ Parapsammodius puncticollis LeConte, 1858
Ataenius punctifrons (Cartwright, 1974)⇒ Ataenius texanus Harold, 1874
Ataenius punctipennis Harold, 1868
Ataenius purator Harold, 1868
Ataenius quintanaroo Stebnicka, 2006
Ataenius raccurti Chalumeau, 1978
Ataenius rakovici Bordat & ThÃ..., 2012
Ataenius raslani Bordat & ThÃ..., 2012
Ataenius raucus Schmidt, 1908
Ataenius restructus H. F. Wickham, 1912
Ataenius rhyticephalus Chevrolat, 1864
Ataenius rickardsi Hinton, 1938
Ataenius robustus Horn, 1871Scratchbush
Ataenius rosinae Balthasar, 1967
Ataenius rubrotessellatus Blanchard, 1846
Ataenius rudellus (Fall, 1930)⇒ Ataenius wenzeli Horn, 1887
Ataenius rugopygus (Cartwright, 1974)⇒ Ataeniopsis rugopygus Cartwright, 1974
Ataenius sabinoi (Cartwright, 1974)⇒ Haroldiataenius lucanus Horn, 1871
Ataenius saltae Stebnicka, 2007
Ataenius salutator (Fall, 1930)⇒ Ataenius picinus Harold, 1861
Ataenius santarosae Stebnicka, 2007
Ataenius saramari (Cartwright, 1939)⇒ Haroldiataenius saramari Cartwright, 1939
Ataenius saulensis Stebnicka, 2006
Ataenius saxatilis (Cartwright, 1944)⇒ Ataeniopsis saxatilis Cartwright, 1944
Ataenius scabrelloides Petrovitz, 1962Blennies
Ataenius scabrellus Schmidt, 1909Scarab beetle
Ataenius scalptifrons Bates, 1887
Ataenius schmidti Stebnicka, 2003
Ataenius sciurus (Cartwright, 1974)⇒ Odontolytes denominatus Chevrolat, 1864
Ataenius sculptilis (Harold, 1868)⇒ Ataenius insculptus Horn, 1887
Ataenius sculptor Harold, 1868
Ataenius scutellaris (Harold, 1867)⇒ Ataenius atramentarius Erichson, 1847
Ataenius seaforthensis Stebnicka & ..., 1997
Ataenius semicoecus MacLeay, 1888
Ataenius semicornutus MacLeay, 1871
Ataenius semipilosus (Van Dyke, 1928)⇒ Haroldiataenius semipilosus Van Dyke, 1928
Ataenius setiger Bates, 1887
Ataenius setosus Schmidt, 1909
Ataenius seydeli Endrödi, 1964
Ataenius similis (Petrovitz, 1961)⇒ Ataenius freyi Petrovitz, 1961
Ataenius siminasus Petrovitz, 1973
Ataenius simplicipes Mulsant & Rey, 1870
Ataenius simulator (Harold, 1868)⇒ Parataenius simulator Harold, 1868
Ataenius skelleyi Stebnicka, 2007
Ataenius socialis (Harold, 1874)⇒ Pseudataenius socialis G. Horn, 1871
Ataenius sparsicollis Blackburn, 1904
Ataenius speculator Blackburn, 1891
Ataenius spinipennis Lea, 1923
Ataenius spretulus Haldeman, 1848Black turfgrass ataenius
Ataenius stebnickae Bordat & ThÃ..., 2012
Ataenius steinheili Harold, 1874
Ataenius stephani Cartwright, 1974
Ataenius stercorator (Fabricius, 1775)⇒ Ataenius strigatus Say, 1823
Ataenius striatocrenatus Fairmaire, 1889
Ataenius strigatus Say, 1823Black turfgrass ataenius
Ataenius strigicauda Bates, 1887
Ataenius strigicaudus Bates, 1887
Ataenius strigifrons Schmidt, 1920
Ataenius stroheckeri (Cartwright, 1974)⇒ Ataenius peregrinator Harold, 1877
Ataenius subopacus (Chapin, 1940)⇒ Ataenius punctipennis Harold, 1868
Ataenius suineata Deloya, 2012
Ataenius superficialis (Cartwright, 1974)⇒ Ataenius scabrelloides Petrovitz, 1962
Ataenius synnotensis Stebnicka & ..., 1997
Ataenius talpoides Stebnicka, 2001
Ataenius tambopatae Stebnicka, 2001
Ataenius tarumensis Stebnicka, 2007
Ataenius temperei Chalumeau & ..., 1974
Ataenius tenebrosus (Arrow, 1903)⇒ Ataenius morator Harold, 1869
Ataenius terminalis (Chevrolat, 1864)⇒ Ataenius luteomargo Chapin, 1940
Ataenius texanus Harold, 1874
Ataenius thomasi Schnepp & As..., 2020
Ataenius tindalensis Stebnicka & ..., 1997
Ataenius tomentosus Petrovitz, 1969
Ataenius torridus Blackburn, 1892
Ataenius tovarensis Stebnicka, 2001
Ataenius tuberculatus Schmidt, 1911
Ataenius tweedensis (Blackburn, 1904)⇒ Ataenius castanicolor Motschulsky, 1858
Ataenius uriarrae Stebnicka & ..., 1997
Ataenius usingeri Hinton, 1937
Ataenius utahensis Cartwright, 1974
Ataenius vandykei (Cartwright, 1974)⇒ Martineziana vandykei Hinton, 1936
Ataenius variopunctatus Schmidt, 1922
Ataenius versicolor (Schmidt, 1916)⇒ Ataenius luteomargo Chapin, 1940
Ataenius vethianus Schmidt, 1909
Ataenius vinacoensis Stebnicka, 2006
Ataenius vincentiae (Arrow, 1903)⇒ Ataenius carinator Harold, 1874
Ataenius walkeri Blackburn, 1904
Ataenius walterhorni Balthasar, 1938
Ataenius waltherhorni Balthasar, 1938Thornfish
Ataenius warisensis Stebnicka, 1998
Ataenius wenzeli Horn, 1887
Ataenius wenzelii Horn, 1887
Ataenius windjanae Stebnicka & ..., 1997
Ataenius woodruffi (Cartwright, 1974)⇒ Ataenius opatrinus Harold, 1867
Ataenius yasamatsui
Ataenius yasumatsui (Nomura, 1943)⇒ Airapus interstitialis Fairmaire, 1883
Ataenius yungasus Stebnicka, 2001
Aulonocnemis alluaudi Cambefort, 1987
Aulonocnemis andohahelica Paulian, 1975
Aulonocnemis angusta Cambefort, 1987
Aulonocnemis anobioides Harold, 1869
Aulonocnemis basalis Schaufuss, 1890
Aulonocnemis basalis basalis
Aulonocnemis basalis perineta Cambefort, 1987
Aulonocnemis camilloi Cambefort, 1987
Aulonocnemis comorica Paulian, 1975
Aulonocnemis comorica comorica
Aulonocnemis comorica mayottensis Cambefort, 1987
Aulonocnemis crassecostata Fairmaire, 1886
Aulonocnemis crassecostata africana Boucomont, 1937
Aulonocnemis crassecostata crassecostata
Aulonocnemis crassecostata fairmairei Boucomont, 1931
Aulonocnemis crassecostata moheliensis Cambefort, 1987
Aulonocnemis cylindrica Schaufuss, 1902
Aulonocnemis diego Cambefort, 1987
Aulonocnemis electrina Cambefort, 1987
Aulonocnemis elongata (Schaufuss, 1902)⇒ Aulonocnemis anobioides Harold, 1869
Aulonocnemis exarata Klug, 1838
Aulonocnemis exaratoides Cambefort, 1987
Aulonocnemis griveaudi Cambefort, 1987
Aulonocnemis grossa Schaufuss, 1902
Aulonocnemis grossepunctata Paulian, 1975
Aulonocnemis irregularis Harold, 1869
Aulonocnemis laevis Schaufuss, 1890
Aulonocnemis meridionalis Boucomont, 1937
Aulonocnemis micheli Cambefort, 1987
Aulonocnemis milloti Cambefort, 1987
Aulonocnemis nana Paulian, 1978
Aulonocnemis nana ambrensis Cambefort, 1987
Aulonocnemis nana nana
Aulonocnemis nitida Schaufuss, 1890
Aulonocnemis nitidula Cambefort, 1987
Aulonocnemis oberthueri Schaufuss, 1902
Aulonocnemis olsoufieffi Boucomont, 1937
Aulonocnemis opatrina Klug, 1838
Aulonocnemis pauliani Cambefort, 1987
Aulonocnemis peyrierasi Cambefort, 1987
Aulonocnemis pilosa Cambefort, 1987
Aulonocnemis pygidialis Boucomont, 1931
Aulonocnemis robinsoni Cambefort, 1987
Aulonocnemis schaufussi Boucomont, 1931
Aulonocnemis sellata Schaufuss, 1890
Aulonocnemis sicardi Cambefort, 1987
Aulonocnemis tenebrionoides Paulian, 1977
Aulonocnemis tenuis Schaufuss, 1890
Aulonocnemis thoracica Schaufuss, 1890
Aulonocnemis torquata Péringuey, 1901
Aulonocnemis vadoni Cambefort, 1987
Aulonocnemis valida Schaufuss, 1890
Aulonocnemis vulgaris Schaufuss, 1891
Auperia andamanensis Koshantschikov, 1916
Auperia capitosa Harold, 1867
Auperia denominata Chevrolat, 1864
Auperia iquitosae Stebnicka, 2002
Auperia landai Balthasar, 1963
Auperia panamensis Stebnicka, 2002
Auperia squamosa (Petrovitz, 1976)⇒ Auperia landai Balthasar, 1963
Australagolius sauerovae Mencl, 2008
Australammoecius aphodioides Lea, 1923
Australammoecius coloratus Blackburn, 1904
Australammoecius goyderensis Blackburn, 1896
Australammoecius occidentalis MacLeay, 1888
Australammoecius peckorum Stebnicka & ..., 1996
Australammoecius persimilis Lea, 1923
Australaphodius accola Kolbe, 1908
Australaphodius catulus Balthasar, 1946
Australaphodius frenchi Blackburn, 1892Little pasture cockchafer
Australaphodius muellerae Bordat, 1990
Australoxenella bathurstensis Howden & Storey, 1992
Australoxenella concinna Storey & Howden, 1996
Australoxenella humptydooensis Howden & Storey, 1992
Australoxenella kalpara Storey & Howden, 1996
Australoxenella midgee Storey & Howden, 1996
Australoxenella mirreen Storey & Howden, 1996
Australoxenella moogoon Storey & Howden, 1996
Australoxenella peckorum Storey & Howden, 1996
Australoxenella teeta Storey & Howden, 1996
Australoxenella wurrook Storey & Howden, 1996
Australoxenella zborowskii Storey & Howden, 1996
Ballucus barri Gordon, 2006Greater weever
Ballucus bruneaui Gordon & Ske..., 2007Lesser weever
Ballucus obtusus LeConte, 1878
Ballucus oregonus Gordon & Ske..., 2007Lesser weever
Basilewskyanus elisabethae Endrödi, 1956
Basilewskyanus insignis Schmidt, 1910
Basilewskyanus iridescens Péringuey, 1901
Batesiana chamorroi Skelley & Va..., 2022
Batesiana tuberculata Bates, 1887
Biralus equinus Faldermann, 1835
Biralus jhelumensis Rakovic, 1991
Biralus mahunkaorum Adám, 1983
Biralus menetriesi Ménétriés, 1849
Biralus satellitius Herbst, 1789
Blackburneus aegrotus Horn, 1870
Blackburneus amazonicus Dellacasa, D..., 2011
Blackburneus argentinensis Schmidt, 1909
Blackburneus banari Minkina & Bo..., 2019
Blackburneus biroi Endrödi, 1956
Blackburneus bongorensis Petrovitz, 1964
Blackburneus calvus Schmidt, 1909
Blackburneus caracaensis Petrovitz, 1970
Blackburneus cavidomus Brown, 1927
Blackburneus charmionus Bates, 1887
Blackburneus clipealis Endrödi, 1964
Blackburneus cobi Endrödi, 1979
Blackburneus collarti Paulian, 1942
Blackburneus consonus Endrödi, 1967
Blackburneus cynomysi Brown, 1927
Blackburneus dellacasarum Bordat, 2005
Blackburneus detruncatus Schmidt, 1908
Blackburneus diminutus Bates, 1887
Blackburneus dubitabilis Endrödi, 1967
Blackburneus erythrinus Bates, 1887
Blackburneus funestus Endrödi, 1955
Blackburneus furcatus Schmidt, 1909
Blackburneus geomysi Cartwright, 1939
Blackburneus ghanaensis Endrödi, 1973
Blackburneus gnu Endrödi, 1964
Blackburneus guatemalensis Bates, 1887
Blackburneus improvisus Bordat, 2017
Blackburneus indio Petrovitz, 1973
Blackburneus ineptus Petrovitz, 1964
Blackburneus iringanus Balthasar, 1946
Blackburneus jugalis Péringuey, 1901
Blackburneus koenigsbaueri Petrovitz, 1973
Blackburneus laxepunctatus Schmidt, 1910
Blackburneus levis Schmidt, 1908
Blackburneus luciphilus Endrödi, 1973
Blackburneus microreticulatus Landin, 1967
Blackburneus minutissimus Schmidt, 1908
Blackburneus mollis Endrödi, 1964
Blackburneus morogoroensis Petrovitz, 1969
Blackburneus niunzuensis Paulian, 1942
Blackburneus nyassicus Petrovitz, 1969
Blackburneus ohopohoensis Endrödi, 1991
Blackburneus optatoides Landin, 1967
Blackburneus optatus Endrödi, 1964
Blackburneus paragnu Balthasar, 1971
Blackburneus pseudocalvus Bordat, 2005
Blackburneus puncticollis Petrovitz, 1969
Blackburneus radamus Petrovitz, 1958
Blackburneus richteri (Schmidt, 1911)⇒ Nobius hepaticus Roth, 1851
Blackburneus rubeolus Palisot de B..., 1805
Blackburneus sanfilippoi Dellacasa, D..., 2011
Blackburneus saylorea Robinson, 1940
Blackburneus saylori Hinton, 1934
Blackburneus stebnickae Pittino, 1988
Blackburneus stercorosus Melsheimer, 1845
Blackburneus strandi Balthasar, 1935
Blackburneus surinamensis Dellacasa, D..., 2011
Blackburneus tachyoryctis Endrödi, 1960
Blackburneus telekii Bordat, 1992
Blackburneus teposcolulaensis Dellacasa, D..., 2011
Blackburneus thomasi Dellacasa, D..., 2011
Blackburneus truncaticollis Petrovitz, 1967
Blackburneus vacca Endrödi, 1964
Blackburneus vadoni Bordat, 1986
Blackburneus villiersianus Clement, 1969
Blackburneus vixpunctatus Endrödi, 1980
Blackburneus vojnitsi Bordat, 1992
Blackburneus walteri Endrödi, 1976
Blackburneus xanthus Bates, 1887
Blackburneus zairensis Endrödi, 1979
Blackburneus zernyi Petrovitz, 1964
Bobiricola endroedyyoungai Endrodi, 1973
Bodiloides asiricus Pittino, 1984
Bodiloides ictericus Laicharting, 1781
Bodiloides ictericus ghardimaouensis Balthasar, 1929
Bodiloides ictericus ictericus Laicharting, 1781
Bodilopsis aquila Schmidt, 1916
Bodilopsis aquilus Schmidt, 1916
Bodilopsis ogloblini Semenov & Me..., 1928
Bodilopsis rufa Moll, 1782
Bodilopsis rufus Moll, 1782
Bodilopsis sordida Fabricius, 1775
Bodilopsis sordida sordida Fabricius, 1775
Bodilopsis sordidus Fabricius, 1775
Bodilopsis sordidus changajicus Endrödi, 1965
Bodilopsis sordidus sordidus
Bodilus acutifrons Petrovitz, 1958
Bodilus amabilis Boheman, 1857
Bodilus angolensis Petrovitz, 1958
Bodilus apertus Schmidt, 1909
Bodilus ardens Harold, 1866
Bodilus ardescens Schmidt, 1916
Bodilus arsissaensis Petrovitz, 1968
Bodilus barbarus Fairmaire, 1860
Bodilus beduinus Reitter, 1892
Bodilus buxeipennis Harold, 1871
Bodilus caspius Menestries, 1832
Bodilus cathedralensis Endrödi, 1991
Bodilus circumcinctus Schmidt, 1840
Bodilus comptifer Petrovitz, 1958
Bodilus crassus Schmidt, 1916
Bodilus crimensis Koshantschikov, 1913
Bodilus damarinus Kolbe, 1908
Bodilus flagrans Erichson, 1843
Bodilus furvus Schmidt, 1916
Bodilus gregarius Harold, 1871
Bodilus hammeri Petrovitz, 1958
Bodilus hastatus Reitter, 1892
Bodilus holgati Endrödi, 1983
Bodilus ictericus (Laicharting, 1781)⇒ Bodiloides ictericus Laicharting, 1781
Bodilus ictericus ictericus (Laicharting, 1781)⇒ Bodiloides ictericus ictericus Laicharting, 1781
Bodilus immundus
Bodilus insperatus Petrovitz, 1967
Bodilus inylchekensis Frolov, 2001
Bodilus karroensis Endrödi, 1983
Bodilus karrooensis Endrödi, 1983
Bodilus kaznakovi Frolov, 2001
Bodilus kermanschahensis Petrovitz, 1959
Bodilus kububanus Kolbe, 1908
Bodilus laetus Wiedemann, 1823
Bodilus laterosetosus Petrovitz, 1958
Bodilus liesenfeldti Petrovitz, 1958
Bodilus loeffleri Petrovitz, 1963
Bodilus longeciliatus Reitter, 1887
Bodilus longispina Küster, 1854
Bodilus lucidulus Boheman, 1857
Bodilus lugens Creutzer, 1799
Bodilus marani Balthasar, 1929
Bodilus marshalli Paulian, 1942
Bodilus nigriventris (Reitter, 1892)⇒ Bodilus lugens Creutzer, 1799
Bodilus noxius Schmidt, 1913
Bodilus nylsvleyicus Endrödi, 1980
Bodilus oblatus Bordat, 1996
Bodilus pantherinus Petrovitz, 1963
Bodilus punctipennis Erichson, 1848
Bodilus rudii Endrödi, 1968
Bodilus rufus
Bodilus sabaeus Peyerimhoff, 1907
Bodilus saoudi Paulian, 1980
Bodilus sordescens Harold, 1869
Bodilus sordidus
Bodilus strigilatus Roth, 1851
Bodilus terminassianae Kolov & Niko..., 2011
Bodilus transvaalensis Endrödi, 1979
Bodilus veselyi Tesar, 1969
Bodilus wollastoni Harold, 1862
Bodilus zarudnyi Frolov, 2001
Bodilus zunbajanicus Stebnicka, 1975
Bordatius australis Pittino & Ma..., 1986
Bordatius capensis Pittino & Ma..., 1986
Bordatius mthatha Maté, 2015
Bordatius muellerae Pittino, 1996
Bordatius tingitanus Pittino & Ma..., 1986
Boucomontiellus pelliculosus Balthasar, 1932
Boucomontiellus umbrinus Koshantschikov, 1913
Brachiaphodius ahrensi Kral, 2000
Brachiaphodius aureopilosus Boucomont, 1930
Brachiaphodius babori Balthasar, 1938
Brachiaphodius bellmanni Bordat, 2005
Brachiaphodius eccoptus Bates, 1889
Brachiaphodius erinaceus Balthasar, 1935
Brachiaphodius jeanneli Paulian, 1938
Brachiaphodius jendeki Kral, 2000
Brachiaphodius langeri Bellmann, 2009
Brachiaphodius nainiensis Petrovitz, 1963
Brachiaphodius pilifer Paulian, 1934
Brachiaphodius pilosus Harold, 1874
Brachiaphodius ruupbai Masumoto, 1991
Brachiaphodius taiwanicus Petrovitz, 1976
Brachiaphodius weibao Kral, 2000
Brachiaphodius yaaunitnoi Masumoto, 1991
Brachiaphodius yuan Kral, 2000
Brindalus granulicollis Pittino, 1980
Brindalus maderae Pittino, 1983
Brindalus porcicollis Illiger, 1803
Brindalus rotundipennis Reitter, 1892
Brindalus schatzmayri Pittino, 1980
Bruchaphodius bruchi A.Schmidt, 1922
Bruchaphodius ovalipennis Harold, 1871
Bruchaphodius shannoni Bruch, 1938
Bruneixenus reidi Howden & Storey, 2000
Bruneixenus squamosus Howden & Storey, 1992
Caelius browni Saylor, 1934
Caelius denticollis (Lewis, 1895)⇒ Silluvia elongata Landin, 1949
Caelius humeralis Brown, 1931
Caelius montana Brown, 1931
Caelius montanus Brown, 1931
Caelius rufescens Horn, 1887
Calamosternus acriculus Balthasar, 1938
Calamosternus algiricus Mariani & Pi..., 1983
Calamosternus boreosinicus Cervenka, 1994
Calamosternus ciliaticollis Petrovitz, 1973
Calamosternus clermonti Reitter, 1907
Calamosternus colimaensis Hinton, 1934
Calamosternus descarpentriesi Petrovitz, 1966
Calamosternus granarius Linnaeus, 1767Granular Small Dung Beetle
Calamosternus hyxos Petrovitz, 1962
Calamosternus hyxos algiricus (Mariani & Pi..., 1983)⇒ Calamosternus algiricus Mariani & Pi..., 1983
Calamosternus machulkai Balthasar, 1935
Calamosternus magnicollis Petrovitz, 1969
Calamosternus mayeri Pilleri, 1953
Calamosternus paludani Petrovitz, 1955
Calamosternus prusai Tesar, 1945
Calamosternus pseudolucidus Rakovic, 1977
Calamosternus sauteri Petrovitz, 1961
Calamosternus srinagarensis Pittino, 2001
Calamosternus stercorarius Mulsant & Rey, 1870
Calamosternus subdolus Petrovitz, 1962
Calamosternus tricornifrons Reitter, 1909
Calamosternus trucidatus Harold, 1863
Calamosternus unicolor Olivier, 1789
Calamosternus uniplagiatus Waterhouse, 1875
Calamosternus vexator Balthasar, 1933
Calamosternus zmoskev Cervenka, 1994
Caligodorus opacus LeConte, 1872Sand divers
Caligodorus tuberosus Barret, 1932
Caligodorus vandykei Barret, 1931Filamented sanddiver
Calocolobopterus caprai Dellacasa, 1984
Calocolobopterus diabolus Endrödi, 1957
Calocolobopterus egregius Petrovitz, 1958
Calocolobopterus magnicornis Bordat, 1992
Calocolobopterus magnus Endrödi, 1957
Calocolobopterus principalis Harold, 1861
Calocolobopterus senegalensis Klug, 1835
Calocolobopterus szekessyi Endrödi, 1960
Candezeollus bicrenulatus Lea, 1923
Candezeollus candezei Harold, 1868
Candezeollus pseudocandezei Stebnicka & ..., 1995
Carinaulus armandidavidi Cervenka, 2000
Carinaulus coreensis Kim, 1986
Carinaulus dierli Balthasar, 1967
Carinaulus dundarum Cervenka, 2000
Carinaulus gansuicus Cervenka, 2000
Carinaulus imitabilis Cervenka, 2000
Carinaulus inexpectatus Balthasar, 1935
Carinaulus jakli Cervenka, 2000
Carinaulus krali Cervenka, 2000
Carinaulus kucerai Král, Mencl ..., 2021
Carinaulus meghalayensis Cervenka, 2000
Carinaulus milaninikodymi Cervenka, 2000
Carinaulus nanxiensis Masumoto, Ki..., 2018
Carinaulus pseudovseteckai Cervenka, 2003
Carinaulus pseudozhanglaensis Cervenka, 2003
Carinaulus pucholti Balthasar, 1961
Carinaulus sikkimensis Balthasar, 1965
Carinaulus tryznai Cervenka, 2000
Carinaulus vseteckai Tesar, 1945
Carinaulus wrzecionkoi Cervenka, 2000
Carinaulus zhanglaensis Cervenka, 2000
Cartwrightia cartwrighti Cartwright, 1967
Cartwrightia intertribalis Islas, 1958
Cartwrightia islasi O.L.Cartwright, 1967
Cephalocyclus bordati Dellacasa, D..., 2007Yellowface pikeblenny
Cephalocyclus carmenae Dellacasa, D..., 2007Yellowface pikeblenny
Cephalocyclus cartagoensis Dellacasa, D..., 2007
Cephalocyclus costaricensis Dellacasa, D..., 2007
Cephalocyclus durangoensis Bates, 1887
Cephalocyclus fuliginosus Harold, 1863
Cephalocyclus gravidus Harold, 1863
Cephalocyclus halffteri Dellacasa, D..., 2007
Cephalocyclus hogei Bates, 1887
Cephalocyclus howdenorum Dellacasa, D..., 2007Yellowface pikeblenny
Cephalocyclus iohannisgabrielis Dellacasa & ..., 2020
Cephalocyclus lagoi Dellacasa, D..., 2007Yellowface pikeblenny
Cephalocyclus luridiventris Harold, 1862
Cephalocyclus luteolus Horn, 1887
Cephalocyclus majomaensis Dellacasa, D..., 2013
Cephalocyclus mexicanus Harold, 1862
Cephalocyclus moroni Minor, Della..., 2015
Cephalocyclus ordonezi Dellacasa, D..., 2007
Cephalocyclus potosinus Dellacasa, D..., 2007Yellowface pikeblenny
Cephalocyclus pseudofuliginosus Dellacasa, D..., 2011
Cephalocyclus pullatus Schmidt, 1913
Cephalocyclus puncticeps Harold, 1862
Cephalocyclus rockefelleri Dellacasa, D..., 2007Yellowface pikeblenny
Cephalocyclus stebnickae Deloya & Iba..., 2000
Cephalocyclus villosipes Harold, 1862
Cesamexico constricticollis Bates, 1890
Chaetopisthes (Chaetopisthes) fulvus Westwood, 1847
Chaetopisthes (Chaetopisthides) sulciger Wasmann, 1899
Chaetopisthes (Miochaetopisthes) cincinnatus Tangelder & ..., 1982
Chaetopisthes (Neochaetopisthes) assmuthi asmann, 1911
Chaetopisthes angustipes Tangelder & ..., 1982
Chaetopisthes brevipes Champion & W..., 1923
Chaetopisthes brunneus Wasmann, 1918
Chaetopisthes fulvoides Tangelder & ..., 1982
Chaetopisthes heimi Wasmann, 1903
Chaetopisthes latipes Tangelder & ..., 1982
Chaetopisthes longulus Champion & W..., 1923
Chaetopisthes meridionalis Tangelder & ..., 1982
Chaetopisthes oberthueri Tangelder & ..., 1982
Chaetopisthes popei Tangelder & ..., 1982
Chaetopisthes saetiger Tangelder & ..., 1982
Chaetopisthes semisulcatus Tangelder & ..., 1982
Chaetopisthes septentrionalis Tangelder & ..., 1982
Chaetopisthes simplicipes Gestro, 1891
Chaetopisthes singalensis Champion & W..., 1923
Chaetopisthes ventriosus Tangelder & ..., 1982
Cheleion jendeki Král & Hájek, 2015
Cheleion malayanum Vårdal & For..., 2010
Cheleion watanabei Kakizoe, Mar..., 2019
Chilothorax albosetosus Pittino, 1995
Chilothorax alexis Frolov, 2001
Chilothorax aljibei Blanco, 1986
Chilothorax altaicus Nikolajev, 1984
Chilothorax auliensis Balthasar, 1938
Chilothorax badenkoi Nikolajev, 1987
Chilothorax baghlanicus Frolov, 1997
Chilothorax bistriga Reitter, 1900
Chilothorax brancoi Baraud, 1981
Chilothorax cervorum Fairmaire, 1871
Chilothorax chandmanicus Frolov, 2001
Chilothorax clathratus Reitter, 1892
Chilothorax clausula Koshantschikov, 1910
Chilothorax comma Reitter, 1892
Chilothorax conspurcatus Linnaeus, 1758
Chilothorax discedens Schmidt, 1907
Chilothorax distinctus (Müller, 1776)⇒ Aphodius distinctus O.F. Müller, 1776
Chilothorax distinctus distinctus O. F. Müller, 1776
Chilothorax distinctus sevanicus Rakovic, 1991
Chilothorax dobrovljanskyi Koshantschikov, 1913
Chilothorax dzongensis Petrovitz, 1976
Chilothorax equitis Koshantschikov, 1912
Chilothorax exclamationis Motschulsky, 1849
Chilothorax exilimanus Kabakov, 1996
Chilothorax figuratus Schmidt, 1906
Chilothorax flammulatus Harold, 1876
Chilothorax flavimargo Reitter, 1901
Chilothorax fritschi Balthasar, 1933
Chilothorax glebi Frolov, 2006
Chilothorax grafi Reitter, 1901
Chilothorax hahni Reitter, 1907
Chilothorax hieroglyphicus Klug, 1845
Chilothorax hozuensis Ochi, Tsukam..., 2015
Chilothorax huangyuanensis Cervenka, 1994
Chilothorax ivanovi Lebedev, 1911
Chilothorax jacobsoni Koshantschikov, 1911
Chilothorax kandaharicus Balthasar, 1961
Chilothorax kardonensis Huang & Du, 1999
Chilothorax kerzhneri Nikolajev, 1984
Chilothorax kukunorensis Semenov, 1898
Chilothorax lieni Masumoto, Ki..., 2014
Chilothorax lineolatus Illiger, 1803
Chilothorax logunovi Zinchenko, 2003
Chilothorax longetarsalis Pittino, 1988
Chilothorax melanostictus Schmidt, 1840
Chilothorax melanostictus melanostictus W. Schmidt, 1840
Chilothorax melanostictus simulans Koshantschikov, 1913
Chilothorax melanosticus
Chilothorax mongolaltaicus Nikolajev, 1984
Chilothorax mossulensis Mulsant & Go..., 1879
Chilothorax naevuliger Reitter, 1894
Chilothorax nigrivittis Solsky, 1876
Chilothorax nigrivittis impubis Koshantschikov, 1913
Chilothorax nigrivittis kongurensis Ochi, Tsukam..., 2015
Chilothorax nigrivittis nigrivittis
Chilothorax ohishii Masumoto, 1975
Chilothorax okadai Nakane, 1951
Chilothorax okadai kiritachiensis Ochi, Kawaha..., 2012
Chilothorax okadai okadai
Chilothorax ornatus Schmidt, 1911
Chilothorax pamirensis Medvedev, 1928
Chilothorax paykulli Bedel, 1907
Chilothorax pictus Sturm, 1805
Chilothorax planus Koshantschikov, 1894
Chilothorax plutschewskyi Koshantschikov, 1894
Chilothorax potanini Frolov, 2001
Chilothorax praenubilus Balthasar, 1933
Chilothorax propola Balthasar, 1946
Chilothorax punctatus Waterhouse, 1875
Chilothorax pustulifer Reitter, 1892
Chilothorax scuticollis Semenov, 1898
Chilothorax sesquivittatus Fairmaire, 1883
Chilothorax sexmaculosus Schmidt, 1911
Chilothorax shansianus Nakane & Shi..., 1957
Chilothorax signifer Mulsant & Rey, 1870
Chilothorax sinicus Cervenka, 1994
Chilothorax sticticus (Panzer, 1798)⇒ Volinus sticticus Panzer, 1798
Chilothorax subpolitus Motschulsky, 1860
Chilothorax tanhensis Frolov, 2001
Chilothorax tenuimanus Sharp, 1878
Chilothorax variicolor Koshantschikov, 1894
Chilothorax variipennis Schmidt, 1922
Chilothorax xanthellus Frolov, 2002
Chilothorax xanthopterus Balthasar, 1938
Chilothorax zaissanicus Nikolajev, 1987
Chiron aberlenci Huchet, 2019
Chiron assamensis Hope, 1845
Chiron bartolozzii Huchet, 2019
Chiron cylindrus Fabricius, 1798
Chiron demirei Huchet, 2019
Chiron elegans Huchet, 2020
Chiron grandis (Gory, 1838)⇒ Theotimius grandis Gory, 1838
Chiron hovanus Fairmaire, 1901
Chiron kelleri Fairmaire, 1893
Chiron lucifer Huchet, 2019
Chiron mariannae Huchet, 2019
Chiron massaicus Huchet, 2019
Chiron oddurensis Huchet, 2019
Chiron senegalensis Hope, 1845
Chiron stuempkei Huchet, 2019
Chiron sulcithorax Perty, 1831
Chiron volvulus Klug, 1855
Chittius anatolicus Petrovitz, 1963
Cinacanthus anomalipus Péringuey, 1901
Cinacanthus coquilletti Linell, 1896
Cinacanthus crenicollis Fall, 1932
Cinacanthus crescentus Gordon & Ske..., 2007Lesser weever
Cinacanthus cunninghami Warner, 2021
Cinacanthus freudei Endrödi, 1957
Cinacanthus freudei freudei
Cinacanthus freudei longidoensis Endrödi, 1960
Cinacanthus hirsutus Brown, 1928
Cinacanthus hottentotus Petrovitz, 1964
Cinacanthus militaris LeConte, 1858Berg-salmon gill worm
Cinacanthus penetrans Schmidt, 1911
Cinacanthus solus Bordat, 1990
Cinacanthus tenerus Petrovitz, 1959
Cinacanthus tibialis Schmidt, 1908
Cinacanthus ulkei Gordon, 2006Greater weever
Cnemargulus alkhubraicus Pittino, 1984
Cnemargulus krulikovskii Semenov, 1903
Cnemargulus krulikovskyi Semenov, 1903
Cnemargulus pusio Semenov, 1903
Cnematoplatys benardi A.Schmidt, 1914
Cnematoplatys cylindrica Endrödi, 1951
Cnematoplatys numensis Stebnicka, 1986
Cnematoplatys sumatrana Balthasar, 1967
Cnematoplatys torricellensis Stebnicka, 1998
Cnematoplatys tozerensis Stebnicka & ..., 1996
Cnematoplatys yorkensis Stebnicka, 2006
Cnemisus ahngeri Semenov, 1903
Cnemisus kaszabi (Endrödi, 1964)⇒ Cnemisus kaznakovi Semenov, 1903
Cnemisus kaznakovi Semenov, 1903
Cnemisus kozlovi Semenov, 1903
Cnemisus rufescens Motschulsky, 1845
Coelotrachelus burgosi Deloya & Mcc..., 1992
Coelotrachelus ixtapalapaensis Dellacasa M...., 2009
Coelotrachelus kuntzeni Schmidt, 1913
Coelotrachelus macgregori Islas, 1955
Coelotrachelus rudis LeConte, 1878
Coelotrachelus symbius Gordon & Howden, 1972Southern sandfishes
Coelotrachelus venustus Schmidt, 1911
Colobopteridius renaudi Clement, 1958
Colobopteroides latus Schmidt, 1911
Colobopteroides latus coryndoni Paulian, 1939
Colobopteroides latus latus
Colobopterus brignolii Carpaneto, 1973
Colobopterus erraticus Linnaeus, 1758Erratic small dung beetle
Colobopterus indagator Mannerheim, 1849
Colobopterus notabilipennis Petrovitz, 1972
Colobopterus propraetor Balthasar, 1932
Colobopterus quadratus Reiche, 1847
Colobopterus scrutator
Colobopterus subterraneus Linnaeus, 0
Coprimorphus scrutator Herbst, 1789
Coptochiroides laotianus Cervenka, 2005
Coptochiroides subcostatus Kolbe, 1886
Coptochiroides subcostatus perroti Paulian, 1945
Coptochiroides subcostatus subcostatus
Coptochirus alternatus Bordat, 1989
Coptochirus brachypterus Harold, 1868
Coptochirus cognatus Péringuey, 1901
Coptochirus emarginatus Germar, 1824
Coptochirus endroedyyoungai Bordat, 1989
Coptochirus excisus Harold, 1868
Coptochirus pallidipennis Harold, 1868
Coptochirus perlucidus Bordat, 1989
Coptochirus pteropus Harold, 1859
Coptochirus reticulatus Bordat, 1989
Coptochirus viduus
Coptochirus vulgatus Harold, 1868
Corythoderus (Corythoderus) loripes Klug, 1845
Corythoderus (Eurycorythoderus) gibbiger Wasmann, 1899
Corythoderus (Stenocorythoderus) braminus Wasmann, 1918
Corythoderus marshalli (Brauns, 1900)⇒ Paracorythoderus marshalli Brauns, 1900
Craterocyphus abscissus Petrovitz, 1958
Craterocyphus bituberculatus Endrödi, 1960
Craterocyphus forticornis Schmidt, 1910
Craterocyphus ikuthanus Balthasar, 1935
Craterocyphus illustris Schmidt, 1920
Craterocyphus magnificus Schmidt, 1909
Craterocyphus rhinoceros Reiche, 1847
Craterocyphus rhinoceros corniculatus Schmidt, 1920
Craterocyphus rhinoceros kifaru Endrödi, 1956
Craterocyphus rhinoceros mirus Schmidt, 1917
Craterocyphus rhinoceros rhinoceros
Craterocyphus rhinocerus Reiche, 1847
Craterocyphus rhinocerus corniculatus Schmidt, 1920
Craterocyphus rhinocerus kifaru Endrödi, 1956
Craterocyphus rhinocerus mirus Schmidt, 1916
Craterocyphus serratodentatus Petrovitz, 1964
Craterocyphus simplicicornis Endrödi, 1960
Cryptaphodius entellus Endrödi, 1967
Cryptaphodius excavatus Schmidt, 1916
Cryptoscatomaseter acuminatus Cartwright, 1944Monkfish
Cryptoscatomaseter brevicollis LeConte, 1878
Cryptoscatomaseter browni Hinton, 1934Sand stargazers
Cryptoscatomaseter caccabatus Warner & Ske..., 2006
Cryptoscatomaseter captivus Cartwright, 1944Monkfish
Cryptoscatomaseter criddlei Brown, 1928
Cryptoscatomaseter depressiusculus Schmidt, 1907Spotted weever
Cryptoscatomaseter durangoi Gordon & Howden, 1972Arrow stargazer
Cryptoscatomaseter ejectus Gordon & Ske..., 2007Lesser weever
Cryptoscatomaseter epulus Gordon & Howden, 1972Arrow stargazer
Cryptoscatomaseter excavatus Robinson, 1940
Cryptoscatomaseter gambrinus Skelley & Go..., 2001Warteye stargazer
Cryptoscatomaseter haldemani Horn, 1887
Cryptoscatomaseter hubbeli Skelley & Wo..., 1991
Cryptoscatomaseter hubbelli Skelley and ..., 1991Masked stargazer
Cryptoscatomaseter incomptus Gordon & Ske..., 2007Lesser weever
Cryptoscatomaseter iowensis Wickham, 1913
Cryptoscatomaseter magnificens Robinson, 1940
Cryptoscatomaseter ochreipennis Horn, 1871
Cryptoscatomaseter oklahomensis Brown, 1928
Cryptoscatomaseter paulseni Warner & Ske..., 2006Bigeye stargazer
Cryptoscatomaseter pholetus Skelley & Wo..., 1991Masked stargazer
Cryptoscatomaseter platypleurus Skelley & Wo..., 1991Masked stargazer
Cryptoscatomaseter punctissimus Brown, 1928
Cryptoscatomaseter rossi Cartwright, 1944Monkfish
Cryptoscatomaseter salsburyi Gordon, 2006Greater weever
Cryptoscatomaseter talpoidesi Brown, 1928
Cryptoscatomaseter umbricollis Fall, 1907
Cryptoscatomaseter utopensis Warner & Ske..., 2006
Cryptoscatomaseter yavapai Robinson, 1946Goby duckbill
Daintreeola grovei Howden & Storey, 2000
Danielssonia minuta Bordat & Howden, 1995
Dellacasiellus carpinterius Gordon & Ske..., 2007Lesser weever
Dellacasiellus claudus Fall, 1932
Dellacasiellus concavus Say, 1823
Dellacasiellus fucosus Schmidt, 1922Common sand stargazer
Dellacasiellus geomynidomus Godwin, 2016
Dellacasiellus glamisensis Gordon & Ske..., 2007Lesser weever
Dellacasiellus griffini Gordon & Ske..., 2007Lesser weever
Dellacasiellus humboldti Gordon & Ske..., 2007Lesser weever
Dellacasiellus kirni Cartwright, 1944Monkfish
Dellacasiellus laevigatus Haldeman, 1848
Dellacasiellus milleri Gordon & Ske..., 2007Lesser weever
Dellacasiellus pseudofucosus Gordon & Ske..., 2007Lesser weever
Dellacasiellus ruficlarus Fall, 1932
Dialytellus dialytoides Fall & Cocke..., 1907
Dialytellus humeralis LeConte, 1868
Dialytellus tragicus Schmidt, 1916
Dialytes criddlei Brown, 1929
Dialytes foveatus (Schmidt, 1909)⇒ Setylaides foveatus Schmidt, 1909
Dialytes striatulus Say, 1825
Dialytes truncatus Melsheimer, 1845
Dialytes ulkei Horn, 1875
Dialytes umbratus Balthasar, 1941
Dialytodius carri Brown, 1929
Dialytodius decipiens Horn, 1887
Dialytodius olympicus Gordon & Ske..., 2007Lesser weever
Dialytodius rogersi Hatch, 1971
Diapterna dugesi Bates, 1887
Diapterna hamata Say, 1824
Diapterna hyperborea LeConte, 1850
Diapterna omissa LeConte, 1850
Diapterna pinguella Brown, 1929
Diapterna pinguis Haldeman, 1848
Diastictus californicus (Petrovitz, 1963)⇒ Platytomus micros Bates, 1887
Diastictus fracturalis⇒ Diastictis fracturalis Zeller, 1872
Diastictus hawaiiensis (Rakovic, 1981)⇒ Platytomus hawaiiensis Rakovic, 1981
Diastictus tibialis Fabricius, 1798
Diastictus vulneratus Sturm, 1805
Didactylia arabica Pittino, 1984
Didactylia asbenica Villiers, 1950
Didactylia binotata Müller, 1942
Didactylia buxtoni Paulian, 1939
Didactylia capitulata Clouët, 1898
Didactylia castanea Petrovitz, 1973
Didactylia cicatricosa Schmidt, 1908
Didactylia clementi Peyerimhoff, 1929
Didactylia exsecta Schmidt, 1911
Didactylia gilva Petrovitz, 1973
Didactylia gilvoides Endrödi, 1979
Didactylia macarthuri Paulian, 1942
Didactylia mara Krikken & Kaas, 1984
Didactylia murati Paulian, 1939
Didactylia notata (Harold, 1859)⇒ Aidophus notatus Harold, 1859
Didactylia pallicolor Fairmaire, 1886
Didactylia picta Müller, 1942
Didactylia pictipennis Fairmaire, 1897
Didactylia pittinoi Bordat, 1986
Didactylia puncticeps Paulian, 1942
Didactylia pura Schmidt, 1913
Didactylia rosickyi Balthasar, 1963
Didactylia somalicus Balthasar, 1941
Didactylia szunyoghyi Endrödi, 1961
Didactylia tenuipunctata Bordat, 1988
Didactylia turbida Erichson, 1843
Didactylia varia Schmidt, 1908
Didactylia volkonskyi Paulian, 1942
Dilortomaeus angolensis Bordat, 1994
Dilortomaeus deformis Bordat, 2009
Dilortomaeus excultus Péringuey, 1908
Dilortomaeus senegalensis Bordat, 2013
Dilortomaeus setulosus Schmidt, 1908
Doaphius batekensis Walter & End..., 1981
Doaphius clarissimus Petrovitz, 1973
Doaphius comptus Schmidt, 1911
Doaphius gracilioides Paulian, 1942
Doaphius gracilis Boheman, 1857
Doaphius gracilis gracilis
Doaphius gracilis lefiniensis Endrödi, 1967
Doaphius penicilarius Petrovitz, 1963
Doaphius rufopolitus Balthasar, 1941
Drepanacanthoides larreae Horn, 1887
Drepanacanthoides walshii Horn, 1870
Drepanocanthoides canberrae (Stebnicka & ..., 1995)⇒ Ozodius canberrae Stebnicka & ..., 1995
Drepanocanthoides larreae Horn, 1887
Drepanocanthoides neglectus (Schmidt, 1913)⇒ Ozodius neglectus Schmidt, 1913
Drepanocanthoides walpolensis (Stebnicka & ..., 1995)⇒ Ozodius walpolensis Stebnicka & ..., 1995
Drepanocanthoides walshii Horn, 1870
Drepanocanthoides windyensis (Stebnicka & ..., 1995)⇒ Ozodius windyensis Stebnicka & ..., 1995
Drepanocanthus cochranae
Drepanocanthus davisi Bordat, 1996
Drepanocanthus eximius (Péringuey, 1901)⇒ Pseudoxyomus eximius Péringuey, 1901
Drepanocanthus intrusus (Péringuey, 1901)⇒ Pseudoxyomus intrusus Péringuey, 1901
Drepanocanthus lineatus Wiedemann, 1823
Drepanocanthus mineti Bordat, 1993
Drepanocanthus obsurus
Dudleyellus angusticeps Bellmann, 2012
Dudleyellus desaegeri Bordat, 1996
Dudleyellus goniocephalus Roth, 1851
Dudleyellus lassallei Bordat, 2014
Dudleyellus limbourgi Bordat, 2014
Dudleyellus ndoki Bordat, 2014
Dudleyellus nigripennis Petrovitz, 1958
Dudleyellus pensus Petrovitz, 1964
Dudleyellus pulawskii Balthasar, 1970
Dudleyellus rakovicianus Bordat, 2014
Dudleyellus strangulatus Bordat, 2009
Emadiellus cruentus Klug, 1855
Emadiellus equestris Endrödi, 1964
Emadiellus rufopustulatus Wiedemann, 1823
Emadiellus rufopustulatus pistor Balthasar, 1945
Emadiellus rufopustulatus rufopustulatus
Emadiellus zonatus Endrödi, 1964
Eocorythoderus incredibilis Maruyama, 2012
Eremazus cribratus Semenov, 1893
Eremazus giganteus Král, 2017
Eremazus marmottani Fairmaire, 1870
Eremazus punctatus Harold, 1869
Eremazus sefrensis Clouët, 1897
Eremazus unistriatus Mulsant, 1851
Erytodes confirmatus Schmidt, 1916
Erytus aequalis Schmidt, 1907
Erytus bucharicus Petrovitz, 1961
Erytus cognatus Fairmaire, 1860
Erytus hormonzensis Petrovitz, 1980
Erytus lindemannae Balthasar, 1960
Erytus opacior Koshantschikov, 1894
Erytus persicus Petrovitz, 1961
Erytus pruinosus Reitter, 1892
Erytus psammophilus Balthasar, 1941
Erytus tekkensis Petrovitz, 1961
Erytus transcaspicus Petrovitz, 1961
Esymus alaiensis Reitter, 1892
Esymus alkani Petrovitz, 1963
Esymus evanescens Boheman, 1857
Esymus filitarsis Reitter, 1898
Esymus finitimus Schmidt, 1922
Esymus fumigatulus Reitter, 1892
Esymus helenaeliviae Dellacasa & ..., 1985
Esymus hladili Chromy, 1993
Esymus hyalinus Petrovitz, 1961
Esymus indulgens Balthasar, 1960
Esymus kudukensis Obenberger, 1914
Esymus merdarius Fabricius, 1775
Esymus ornatulus Harold, 1866
Esymus pokornyi Chromy, 1993
Esymus pusillus Herbst, 1789
Esymus pusillus pusillus Herbst, 1789
Esymus pusillus roubali Balthasar, 1932
Esymus pyreti Penecke, 1911
Esymus sculpturatus Reitter, 1892
Esymus sesquivittatus Fairmaire, 1883
Esymus sicardi Reitter, 1892
Esymus suturinigra Schmidt, 1916
Eudolus quadriguttatus Herbst, 1783
Eudolus quadrinaevulus Reitter, 1892
Euhemicyclium demeyeri Bordat, 2003
Euhemicyclium planatum Schmidt, 1909
Euheptaulacus atlantis Peyerimhoff, 1925
Euheptaulacus carinatus Germar, 1824
Euheptaulacus carinatus brutius Luigioni, 1920
Euheptaulacus carinatus carinatus Germar, 1824
Euheptaulacus carinatus esuriens Dellacasa, 1983
Euheptaulacus nemethi Théry, 1925
Euheptaulacus porcellus Frivaldszky, 1879
Euheptaulacus sus Herbst, 1783
Euheptaulacus villosus Gyllenhal, 1806
Euorodalus baraudi (Villarreal, 1975)⇒ Euorodalus boiteli Théry, 1918
Euorodalus boiteli Théry, 1918
Euorodalus bredoi Endrödi, 1964
Euorodalus coenosus Panzer, 1798
Euorodalus elephanthinus Petrovitz, 1967
Euorodalus himalayanus Koshantschikov, 1916
Euorodalus longevittatus Schmidt, 1922
Euorodalus lucifer Koshantschikov, 1894
Euorodalus marlieri Endrödi, 1960
Euorodalus paracoenosus Balthasar & ..., 1960
Euorodalus signifer Mulsant & Rey, 1870
Euorodalus stribrnyi Tesar, 1945
Euorodalus subtilitarsis Petrovitz, 1963
Euorodalus tersus Erichson, 1848
Euparia africana Schmidt, 1909
Euparia ambrymensis Paulian, 1941
Euparia baraudi Chalumeau & ..., 1974
Euparia bitubericollis (Schmidt, 1909)⇒ Lomanoxoides bitubericollis A.Schmidt, 1909
Euparia bruchi (Schmidt, 1922)⇒ Bruchaphodius bruchi A.Schmidt, 1922
Euparia castanea Lepeletier &..., 1828Butterfly blenny
Euparia consimilis Balthasar, 1945
Euparia cribricollis (Burmeister, 1877)⇒ Parataenius derbesis Solier, 1851
Euparia luctuosa (Burmeister, 1877)⇒ Ataenius luctuosus Burmeister, 1877
Euparia mirabilis Balthasar, 1945
Euparia pusilla (Burmeister, 1877)⇒ Ataeniopsis pusillus Burmeister, 1877
Euparia shannoni (Bruch, 1938)⇒ Bruchaphodius shannoni Bruch, 1938
Euparixia boliviana Gordon & Mcc..., 2003
Euparixia bruneri Chapin, 1940
Euparixia campbelli Gordon & Mcc..., 2003
Euparixia costaricaensis
Euparixia costaricensis Hinton, 1936
Euparixia duncani Brown, 1927
Euparixia formica Hinton, 1936
Euparixia isthmia Gordon & Mcc..., 2003
Euparixia mexicana Gordon & Mcc..., 2003
Euparixia moseri Woodruff & C..., 1967
Euparixia panamaensis Gordon & Mcc..., 2003
Euparixia panamensis
Euparixoides araguaensis Stebnicka & ..., 2005
Euparixoides cribratus Hinton, 1936
Euparixoides johnsoni Stebnicka, 1998
Euparixoides papilio Stebnicka & ..., 2005
Euparixoides tachirensis Stebnicka & ..., 2005
Euparotrix squamosa Lea, 1923
Eupleurus antiquus Faldermann, 1835
Eupleurus subterraneus Linnaeus, 1758Subterranean small dung beetle
Eupleurus subterraneus krasnojarskicus Dellacasa, 1986
Eupleurus subterraneus subterraneus Linnaeus, 1758
Exaphodius excellens Endrodi, 1960
Exoxyomus curvus Schmidt, 1909
Ferrerianus biimpressus Schmidt, 1909
Flaviellus aggregatus Gordon, 1977Dartfish
Flaviellus consentaneus LeConte, 1850
Flaviellus gordoni Ratcliffe, 1988
Flaviellus kluanensis Smith & Skelley, 2020
Flaviellus perfimbriatus Gordon, 1977Dartfish
Flaviellus phalerioides Horn, 1870
Flaviellus phalerius Gordon & Ske..., 2007Lesser weever
Flaviellus phaleroides Horn, 1870
Flaviellus subtruncatus LeConte, 1878
Flechtmanniella laticollis Petrovitz, 1973
Geomyphilus alabama Skelley & Go..., 1995Saddle stargazer
Geomyphilus barrerai Islas, 1955
Geomyphilus coronadoi Islas, 1955
Geomyphilus essigi Saylor, 1935
Geomyphilus geronimo Gordon & Ske..., 2007Lesser weever
Geomyphilus insolitus Brown, 1928
Geomyphilus kiowensis Gordon & Sal..., 1999Stargazers
Geomyphilus leptotarsis Brown, 1928
Geomyphilus macgregori Islas, 1955
Geomyphilus pierai Lobo & Deloya, 1995
Geomyphilus pinolus Gordon & Ske..., 2007Lesser weever
Geomyphilus rubiginosus Horn, 1870
Geomyphilus russeus Brown, 1928
Geomyphilus thomomysi Brown, 1928
Geomyphilus tuzincola Dellacasa, D..., 2017
Geomyphilus ungulatus Fall, 1901
Geomyphilus viceversus Paulsen, 2006Electric stargazers
Geopsammodius atlantida Skelley, 2006
Geopsammodius fuscus Skelley, 2006Striped blenny
Geopsammodius hydropicus Horn, 1887
Geopsammodius morrisi Skelley, 2006Striped blenny
Geopsammodius ohoopee Skelley, 2006Striped blenny
Geopsammodius relictillus Deyrup & Woo..., 1991Redspeckled blenny
Geopsammodius rileyi Skelley, 2006Striped blenny
Geopsammodius sabinae Lavalette, 1999
Geopsammodius subpedalis Skelley, 2006Striped blenny
Geopsammodius unsidensis Skelley, 2006Striped blenny
Geopsammodius withlacooche Skelley, 2006
Geopsammodius withlacoochee Skelley, 2006Striped blenny
Gilletianus aganocrossoides Kral, Rakovi..., 2014
Gilletianus asinensis Ochi, Kon & ..., 2016
Gilletianus assamensis Petrovitz, 1976
Gilletianus atsushii Ochi, 1986
Gilletianus banteng Král & Masum..., 2020
Gilletianus bolm Kral, Rakovi..., 2014
Gilletianus cavitensis Ochi, Kon & ..., 2016
Gilletianus comatus Schmidt, 1920
Gilletianus commatoides Balthasar, 1961
Gilletianus costatulus Schmidt, 1908
Gilletianus fukiensis Balthasar, 1952
Gilletianus imasakai Ochi, Kon & ..., 2016
Gilletianus kanchil Král & Masum..., 2020
Gilletianus kazirangensis Stebnicka, 1981
Gilletianus kinabuhayan kinabuhayan
Gilletianus kinabuhayan lucenaensis Ochi, Kon & ..., 2016
Gilletianus kratochvili Balthasar, 1941
Gilletianus lomsakensis Stebnicka, 1992
Gilletianus luzonensis Ochi, Kon & ..., 2016
Gilletianus matotomensis Ochi, Kon & ..., 2010
Gilletianus miksici Balthasar, 1960
Gilletianus mindorensis Ochi, Kon & ..., 2010
Gilletianus miurai Minkina, 2022
Gilletianus negrosensis Ochi, Kon & ..., 2016
Gilletianus nigrovirgatus Schmidt, 1911
Gilletianus poggii Ochi, Kon & ..., 2014
Gilletianus proclivis Balthasar, 1933
Gilletianus pseudoreichei Stebnicka, 1998
Gilletianus rajawatorum Kral & Sipek, 2013
Gilletianus rangoonensis Petrovitz, 1970
Gilletianus reichei Harold, 1859
Gilletianus schoolmeestersi Ochi, Kon & ..., 2016
Gilletianus segmentarius Harold, 1879
Gilletianus segmentaroides Schmidt, 1909
Gilletianus seguyi Paulian, 1945
Gilletianus sundainsulanus Ochi, Kon & ..., 2014
Gilletianus superreichei Ochi, Kon & ..., 2010
Gilletianus takeshii Ochi, Kon & ..., 2010
Gilletianus therondi Balthasar, 1963
Gilletianus timorensis Ochi, Kon & ..., 2016
Gilletianus tonkineus Paulian, 1933
Gilletianus yunnanensis Král, Rakovi..., 2015
Goiginus howdeni Endrödi, 1982
Goiginus zumpti Petrovitz, 1967
Gonaphodiellus arcanus Dellacasa, D..., 2012
Gonaphodiellus ataeniodes Hinton, 1938
Gonaphodiellus bimaculosus Schmidt, 1909
Gonaphodiellus cerropuntanus Dellacasa, D..., 2012
Gonaphodiellus hoffmanni Islas, 1945
Gonaphodiellus manantlanicus Dellacasa, D..., 2012
Gonaphodiellus martinpierai Dellacasa, D..., 2012
Gonaphodiellus nigrinus (Schmidt, 1916)⇒ Gonaphodiellus pacatus Harold, 1880
Gonaphodiellus omiltemicus Dellacasa, D..., 2012
Gonaphodiellus opisthius Bates, 1887
Gonaphodiellus pacatus Harold, 1880
Gonaphodiellus sacatepequezis Dellacasa, D..., 2012
Gonaphodiellus sexguttatus Schmidt, 1916
Gonaphodioides acutecernans Balthasar, 1960
Gonaphodioides cartwrighti Dellacasa, D..., 2013
Gonaphodioides chapini Hinton, 1934
Gonaphodioides columbicus Harold, 1880
Gonaphodioides newtoni Dellacasa, D..., 2012
Gonaphodioides ratcliffei Dellacasa, D..., 2012
Gonaphodioides sincerus Petrovitz, 1973
Gonaphodioides skelleyi Dellacasa, D..., 2012
Gonaphodioides zempoaltepetlensis Dellacasa, D..., 2018
Gonaphodiopsis deloyai Dellacasa, D..., 2012
Gonaphodiopsis hypogea Dellacasa, D..., 2012
Gonaphodiopsis montesdeocai Dellacasa, D..., 2012
Gonaphodiopsis pachecoi Dellacasa, D..., 2012
Gonaphodiopsis teopiscaensis Dellacasa, D..., 2012
Gonaphodius scoparius Harold, 1877
Gongrolophus storeyi Stebnicka & ..., 1996
Gordonius rhinocerillus Skelley, Del..., 2009
Grandinaphodius inferorum Ziani, 2002
Grandinaphodius smoliki Kaufel, 1914
Granulopsammodius centralasiae Rakovic, 1978
Granulopsammodius loebli Pittino, 1980
Granulopsammodius mesopotamicus Petrovitz, 1971
Granulopsammodius mongol Endrödi, 1969
Granulopsammodius persicus Rakovic, 1998
Granulopsammodius petrovitzi Rakovic, 1978
Granulopsammodius plicatuloides Pittino, 1984
Granulopsammodius plicatulus Fairmaire, 1892
Granulopsammodius sinicus Rakovic, 1986
Granulopsammodius tienshanicus Pittino, 1984
Granulopsammodius transcaspicus Petrovitz, 1961
Guanyinaphodius kishimotoi Masumoto & K..., 2001
Hadrorhyparus pecki Howden, 1995
Harmodactylus oscitans Péringuey, 1901
Harmodactylus solus Bordat, 1990
Harmogaster cochranae
Harmogaster exarata Harold, 1861
Harmogaster geminata (Schmidt, 1911)⇒ Pseudoxyomus geminata Schmidt, 1911
Harmogaster intrusus
Harmogaster oblongopunctata (Balthasar, 1941)⇒ Pseudoxyomus oblongopunctata Balthasar, 1941
Harmogaster opacula (Harold, 1869)⇒ Pseudoxyomus opaculus Harold, 1869
Harmogaster opatroides (Balthasar, 1941)⇒ Pseudoxyomus opatroides Balthasar, 1941
Harmogaster quadridentella Schmidt, 1911
Harmogaster scobina (Petrovitz, 1956)⇒ Pseudoxyomus scobina Petrovitz, 1956
Harmogaster spectabilis
Harmogaster sulcatula Schmidt, 1911
Harmogaster transitoria Péringuey, 1901
Harmogaster tuberculata Endrödi, 1991
Haroldaphodius binodulus Harold, 1866
Haroldaphodius panticosus Bordat, 2003
Haroldaphodius procerus Harold, 1862
Haroldiataenius (Haroldiataenius) mariarum Bates, 1887
Haroldiataenius (Sayloria) hintoni Saylor, 1933
Haroldiataenius buvexus Stebnicka & ..., 2009Caneva's blenny
Haroldiataenius convexus Robinson, 1940
Haroldiataenius griffini Cartwright, 1974
Haroldiataenius limbatus Bates, 1887
Haroldiataenius lucanus Horn, 1871
Haroldiataenius saramari Cartwright, 1939
Haroldiataenius semipilosus Van Dyke, 1928
Haroldiellus lansbergei Harold, 1874
Haroldiellus sallei Harold, 1863
Haroldiellus woodruffi Skelley & Ke..., 2022
Hemicorythoderus vaneyeni Paulian, 1947
Hemicyclium lateritium Roth, 1851
Hemicyclium lateritius Roth, 1851
Hemicyclium unicornutum Schmidt, 1909
Hemicyclium unicornutus Schmidt, 1909
Hemycorythoderus vaneyeni Paulian, 1947
Heptaulacus algarbiensis Branco & Baraud, 1984
Heptaulacus alpinus Drapiez, 1819
Heptaulacus brancoi Baraud, 1976
Heptaulacus carinatus (Gebler, 1833)⇒ Euheptaulacus carinatus Germar, 1824
Heptaulacus gadetinus Baraud, 1973
Heptaulacus koshantschikoffi Schmidt, 1911
Heptaulacus pirazzolii Fairmaire, 1881
Heptaulacus puberulus (Boheman, 1857)⇒ Neoheptaulacus puberulus Boheman, 1857
Heptaulacus rasettii Carpaneto, 1978
Heptaulacus sus (Herbst, 1783)⇒ Euheptaulacus sus Herbst, 1783
Heptaulacus syrticola Fairmaire, 1882
Heptaulacus testudinarius Fabricius, 1775
Heptaulacus villosus (Gyll., 1806)⇒ Euheptaulacus villosus Gyllenhal, 1806
Hornietus ventralis Horn, 1887
Iarupea goias Stebnicka, 2007
Iarupea guimaraesi (Martínez, 1955)⇒ Arupaia friedenreichi Harold, 1870
Iarupea lopeteguii Martínez, 1953
Iarupea luisae Stebnicka, 2007
Iarupea nigricans Westwood, 1843
Iarupea serratipennis Petrovitz, 1973
Iarupea yungana
Iguazua blackwelderi Chapin, 1940
Iguazua costaricensis Stebnicka & ..., 2005
Iguazua lilloana Stebnicka, 1997
Ingogius leleupi Endrodi, 1976
Irrasinus dumasi Gordon & Ske..., 2007Lesser weever
Irrasinus stupidus Horn, 1870
Jalisco plumipes Dellacasa, G..., 2003
Koreoxyomus koreanus Kim, 1996
Koshantschikovius andrewesi Schmidt, 1908
Koshantschikovius anthonyi Bordat, 1989
Koshantschikovius astacus Boucomont, 1932
Koshantschikovius bifidus Endrödi, 1967
Koshantschikovius buchamaensis Bordat, 2005
Koshantschikovius chovdgolensis Endrödi, 1983
Koshantschikovius crassigena Bordat, 2009
Koshantschikovius doguetianus Bordat, 2018
Koshantschikovius erytoides Endrödi, 1983
Koshantschikovius genieri Bordat, 2009
Koshantschikovius haematicus Boheman, 1857
Koshantschikovius haematiticus Boheman, 1857
Koshantschikovius hartwigi Petrovitz, 1967
Koshantschikovius holmi Endrödi, 1983
Koshantschikovius intractatus Balthasar, 1973
Koshantschikovius latecinctus Fairmaire, 1903
Koshantschikovius motoi Paulian, 1939
Koshantschikovius nagporensis Petrovitz, 1968
Koshantschikovius neohaematiticus Landin, 1967
Koshantschikovius oscari Bordat, 2015
Koshantschikovius peculator Balthasar, 1971
Koshantschikovius praedabundus Balthasar, 1939
Koshantschikovius schmitzi Bordat, 1990
Koshantschikovius schusteri Balthasar, 1935
Koshantschikovius snizeki Bordat, 2005
Koshantschikovius splendens Balthasar, 1935
Koshantschikovius strandi Balthasar, 1935
Koshantschikovius substriatoides Bordat, 1990
Koshantschikovius substriatus Schmidt, 1911
Koshantschikovius sulcifrons Bordat, 1990
Koshantschikovius szunyoghyi Endrödi, 1971
Koshantschikovius ventriosus Bordat, 2014
Koshantschikovius youngai Endrödi, 1967
Koshantschikovius zambiensis Bordat, 2016
Labarrus afrolividus Rakovic, 1991
Labarrus cincticulus Hope, 1847
Labarrus digitatus Harold, 1871
Labarrus evachromae Chromy, 1993
Labarrus hoabinhensis Balthasar, 1946
Labarrus innumerabilis Schmidt, 1910
Labarrus kasyi Petrovitz, 1965
Labarrus lividus Olivier, 1789Scarab beetle
Labarrus lividus lividus
Labarrus lividus matusitai Nakane, 1961
Labarrus madagassicus Petrovitz, 1961
Labarrus meruensis Petrovitz, 1961
Labarrus nabeleki Balthasar, 1936
Labarrus paludani Petrovitz, 1955
Labarrus paralividus Balthasar, 1941
Labarrus parnaguaensis Petrovitz, 1961
Labarrus pseudolividus Balthasar, 1941
Labarrus rigidus Balthasar, 1936
Labarrus rugosicapita Mittal, 1993
Labarrus scintillius Nikritin & K..., 1979
Labarrus splendidus Petrovitz, 1955
Labarrus sublimbatus Motschulsky, 1860
Labarrus sublividus (Balthasar, 1941)⇒ Aphodius sublividus Balthasar, 1941
Labarrus translucidus Petrovitz, 1961
Labarrus umbratus Petrovitz, 1961
Labarrus witboii Petrovitz, 1961
Lechorodius lutulentus Haldeman, 1843
Lechorodius terminalis Say, 1823
Leiopsammodius abyssinicus Müller, 1942
Leiopsammodius acei Harpootlian ..., 2000
Leiopsammodius aegialius Adám, 1986
Leiopsammodius balthasari Rakovic, 1995
Leiopsammodius belloi Pierotti, 1980
Leiopsammodius bolivianus Cartwright, 1955
Leiopsammodius chifengleei Kiuchi & Tsai, 2018
Leiopsammodius degallieri Rakovi�, Me..., 2017
Leiopsammodius desertorum Fairmaire, 1868
Leiopsammodius deyrupi Harpootlian ..., 2000
Leiopsammodius endroedii Rakovic, 1981
Leiopsammodius evanidus Péringuey, 1901
Leiopsammodius freyi Petrovitz, 1961
Leiopsammodius gestroi Clouët, 1900
Leiopsammodius globatus Petrovitz, 1972
Leiopsammodius haruspex Adám, 1980
Leiopsammodius horaki Rakovic & Kral, 1996
Leiopsammodius implicatus Schmidt, 1925
Leiopsammodius indefensus Schmidt, 1909
Leiopsammodius indicus Harold, 1877
Leiopsammodius inflatus Cartwright, 1955
Leiopsammodius japonicus Harold, 1878
Leiopsammodius jelineki Rakovic, 1977
Leiopsammodius kenyensis Rakovic, 1978
Leiopsammodius laevicollis Klug, 1845
Leiopsammodius laevis Paulian, 1942
Leiopsammodius litoralis Lea, 1923
Leiopsammodius liviae Pittino, 1978
Leiopsammodius malindii Rakovi�, Me..., 2017
Leiopsammodius malkini Cartwright, 1946Hen-like blenny
Leiopsammodius manaosi Cartwright, 1955
Leiopsammodius martinezi Cartwright, 1955
Leiopsammodius modestus Péringuey, 1901
Leiopsammodius newcastleensis Stebnicka & ..., 1996
Leiopsammodius nomurai Masumoto, 2012
Leiopsammodius ocmulgeei Harpootlian ..., 2000
Leiopsammodius pellucens Petrovitz, 1961
Leiopsammodius placidus Schmidt, 1911
Leiopsammodius rakovici Král, 2017
Leiopsammodius rufus Rakovic, 1981
Leiopsammodius santaremi Cartwright, 1955
Leiopsammodius scabrifrons Walker, 1871
Leiopsammodius seychellensis Rakovic, 1979
Leiopsammodius soledadei Petrovitz, 1961
Leiopsammodius somalicus Petrovitz, 1961
Leiopsammodius strumae Chromy, 1983
Leiopsammodius subciliatus Harold, 1869
Leiopsammodius subcililatus Harold, 1869
Leiopsammodius substriatus Balthasar, 1941
Leiopsammodius viti Chalumeau, 1983
Leptorhyparus brasiliensis Minkina, 2020
Leptorhyparus gilli Howden, 2003
Leptorhyparus peruanus Minkina, 2020
Leptorhyparus quadricornis Skelley, 2021
Limarus hirtipennis Lucas, 1846
Limarus maculatus Sturm, 1800
Limarus zenkeri Germar, 1813
Lindbergianus solitarius Petrovitz, 1961
Lioglyptoxenus mirabilis Pittino, 2006
Liothorax alternatus Horn, 1870
Liothorax consociatus Horn, 1887
Liothorax innexus Say, 1835
Liothorax isikdagensis Balthasar, 1952
Liothorax kraatzi Harold, 1868
Liothorax levatus Schmidt, 1907
Liothorax lividus
Liothorax longiartus (Girault, 1932)⇒ Copidosoma longiartus Girault, 1932
Liothorax melanocera (Ashmead, 1900)⇒ Copidosoma melanocerum Ashmead, 1900
Liothorax muscorum (Adám, 1994)⇒ Liothorax niger Illiger, 1798
Liothorax niger Illiger, 1798Beaulieu dung beetle
Liothorax plagiatus Linnaeus, 1767
Liothorax rusakovi Gusakov, 2004
Liothorax sturmi
Liothorax subaeneus LeConte, 1857
Liothorax varians
Lomanoxia alternata Krikken, 1972
Lomanoxia canthonopsis Skelley, 2003
Lomanoxia chacocola Krikken, 1972
Lomanoxia costulata Harold, 1867
Lomanoxia ituensis Stebnicka, 1999
Lomanoxia melloi Stebnicka, 1999
Lomanoxia ovalis Schmidt, 1911
Lomanoxoides bitubericollis A.Schmidt, 1909
Lomanoxoides herediae Stebnicka & ..., 2005
Lomanoxoides mapitunari Stebnicka & ..., 2005
Lomanoxoides nigrolineatus Hinton, 1938
Lomanoxoides selviriaensis Stebnicka, 1999
Lomanoxoides setosus Balthasar, 1941
Lomanoxoides tesari Balthasar, 1963
Longaphodius namaquarum Endrödi, 1980
Loraphodius corrugatus Petrovitz, 1954
Loraphodius kashmirensis Sharp, 1878
Loraphodius latisulcus Reitter, 1892
Loraphodius lebruni Paulian, 1939
Loraphodius suarius Faldermann, 1835
Loraspis frater Mulsant & Rey, 1870
Lorditomaeus aequus Schmidt, 1908
Lorditomaeus basilewskyi Balthasar, 1965
Lorditomaeus bifidus Schmidt, 1908
Lorditomaeus bifidus bifidus
Lorditomaeus bifidus occidentalis Bordat, 1996
Lorditomaeus blattoides Petrovitz, 1969
Lorditomaeus cambeforti Bordat, 1983
Lorditomaeus ellenbergeri Paulian, 1942
Lorditomaeus excultus (Péringuey, 1908)⇒ Dilorditomaeus excultus Péringuey, 1908
Lorditomaeus ferreri Bordat, 1997
Lorditomaeus fornicatus Schmidt, 1908
Lorditomaeus fratris Bordat, 1996
Lorditomaeus gridellii (Endrödi, 1964)⇒ Dilorditomaeus setulosus Schmidt, 1908
Lorditomaeus harpalinus Gerstaecker, 1883
Lorditomaeus horni Balthasar, 1937
Lorditomaeus infuscatus Schmidt, 1908
Lorditomaeus invenustus Schmidt, 1908
Lorditomaeus magnus Balthasar, 1965
Lorditomaeus mirandus Balthasar, 1965
Lorditomaeus morettoi Bordat, 2000
Lorditomaeus opatroides Klug, 1851
Lorditomaeus orthochaetus Balthasar, 1965
Lorditomaeus proditor Gestro, 1895
Lorditomaeus setulosus (Schmidt, 1908)⇒ Dilorditomaeus setulosus Schmidt, 1908
Lorditomaeus similis Bordat, 1996
Lorditomaeus tanganyicanus Balthasar, 1965
Lorditomaeus tenuis Schmidt, 1908
Lorditomaeus uhligi Bordat, 1995
Lorditomaeus youngai Endrödi, 1967
Lunaphodius edgardi Solsky, 1876
Lunaphodius longipennis Rakovic, 1984
Lunaphodius lunifer Solsky, 1876
Luxolinus luxatus Horn, 1887
Macroretroides lumareti Bordat, Dell..., 2000
Macroretrus confusus Harold, 1862
Macroretrus singularis Péringuey, 1908
Macroretrus thilliezi Bordat & Del..., 2000
Macroretrus zavattarii Müller, 1940
Macrotetrus singularis Péringuey, 1908
Maculaphodius conspersus Horn, 1887
Manjarivolo andohahelicus Paulian, 1974
Manjarivolo crypticus Paulian, 1974
Manjarivolo mirabilis Paulian, 1976
Manjarivolo randrianirinai Ballerio & M..., 2019
Martineziana argentina Harold, 1867
Martineziana dutertrei Chalumeau, 1983
Martineziana excavaticollis Blanchard, 1846Beetle
Martineziana separata Schmidt, 1909
Martineziana simplex Balthasar, 1963
Martineziana vandykei Hinton, 1936
Mecynodes anemurensis Petrovitz, 1968
Mecynodes angulosus Harold, 1869
Mecynodes asmaricus Balthasar, 1955
Mecynodes becvari Cervenka, 1995
Mecynodes demoflysi Baraud, 1977
Mecynodes gilvipennis Balthasar, 1946
Mecynodes kisilkumi Solsky, 1876
Mecynodes leucopterus Klug, 1845
Mecynodes stereotypus Koshantschikov, 1894
Mecynodes striatulus Waltl, 1835
Mecynodes trochilus Reitter, 1892
Mecynodes zagrosensis Minkina, Mie..., 2021
Megateloides bostrichoides Harold, 1860
Megateloides dimidiatus Roth, 1851
Megateloides dimidiatus diadimitus Petrovitz, 1962
Megateloides dimidiatus dimidiatus
Megatelus brahminus Harold, 1879
Megatelus contractus Klug, 1845
Megatelus doriae Harold, 1871
Melinopterus abeillei Sietti, 1903
Melinopterus circumcintus
Melinopterus consentaneus LeConte, 1850
Melinopterus consputus Creutzer, 1799
Melinopterus dellacasai Avila, 1986
Melinopterus edithae Reitter, 1906
Melinopterus femoralis Say, 1823
Melinopterus feryi Rössner, 2018
Melinopterus gissaricus Akhmetova & ..., 2012
Melinopterus guillebeaui Reitter, 1892
Melinopterus imamae Stebnicka, 1988
Melinopterus longipes Landin, 1949
Melinopterus makowskyi Koshantschikov, 1891
Melinopterus meuseli Reitter, 1906
Melinopterus prodromus Brahm, 1790Spring small dung beetle
Melinopterus pubescens Sturm, 1800
Melinopterus punctatosulcatus Sturm, 1805
Melinopterus punctatosulcatus hirtipes Fischer von ..., 1844
Melinopterus punctatosulcatus ponticus Petrovitz, 1967
Melinopterus punctatosulcatus punctatosulcatus Sturm, 1805
Melinopterus rapax Faldermann, 1836
Melinopterus reyi Reitter, 1892
Melinopterus scuticollis Semenov, 1898
Melinopterus sertavulensis Pittino, 1988
Melinopterus sphacelatus Panzer, 1798
Melinopterus sphacelatus ponticus Petrovitz, 1967
Melinopterus stolzi Reitter, 1906
Melinopterus tingens Reitter, 1892
Melinopterus villarreali Baraud, 1975
Melinopterus wittmeri Petrovitz, 1975
Mendidaphodius adustus Klug, 1855
Mendidaphodius angustatus Klug, 1845
Mendidaphodius armatulus Fairmaire, 1893
Mendidaphodius armiger Harold, 1871
Mendidaphodius biangulatus Fairmaire, 1893
Mendidaphodius brancsiki Reitter, 1899
Mendidaphodius calamitosus Balthasar, 1935
Mendidaphodius ellenbergeri Paulian, 1942
Mendidaphodius fischeri Petrovitz, 1962
Mendidaphodius ganabi Endrödi, 1977
Mendidaphodius leleupi Endrödi, 1960
Mendidaphodius lepidulus Harold, 1866
Mendidaphodius linearis Reiche & Saulcy, 1856
Mendidaphodius minusculus Fairmaire, 1893
Mendidaphodius nodulifer Koshantschikov, 1894
Mendidaphodius paganettii Petrovitz, 1963
Mendidaphodius porrectus (Schmidt, 1913)⇒ Mendidaphodius adustus Klug, 1855
Mendidaphodius pseudoadustus Petrovitz, 1958
Mendidaphodius ruandanus Endrödi, 1955
Mendidaphodius rufovittatus Müller, 1941
Mendidaphodius rugulicollis Fairmaire, 1893
Mendidaphodius rutilior (Endrödi, 1960)⇒ Mendidaphodius rutilus Klug, 1845
Mendidaphodius rutilus Klug, 1845
Mendidaphodius steinbergi Nikritin, 1973
Mendidaphodius timurkirgizi Dellacasa & ..., 2009
Mendidaphodius uaniombae Paulian, 1942
Mendidaphodius weidholzi Petrovitz, 1964
Mendidius abdurakhmanovi Shokhin, 2018
Mendidius aculeatus Robinson, 1940
Mendidius assectator Balthasar, 1961
Mendidius atricolor Reitter, 1892
Mendidius baigakumi Koshantschikov, 1911
Mendidius beluchistanicus Petrovitz, 1962
Mendidius berbericus Balthasar, 1941
Mendidius bidens Solsky, 1876
Mendidius bispinifrons Reitter, 1889
Mendidius bivittatus Schmidt, 1922
Mendidius calliger Sahlberg, 1908
Mendidius crassulus Horn, 1870
Mendidius curtulus Harold, 1866
Mendidius diffidens Reitter, 1892
Mendidius endroedii Balthasar, 1967
Mendidius fimbriolatus Mannerheim, 1849
Mendidius gobiensis Endrödi, 1964
Mendidius granulifrons (Fairmaire, 1883)⇒ Mendidius multiplex Reitter, 1897
Mendidius heikertingeri Balthasar, 1938
Mendidius ivanovi (Lebedev, 1911)⇒ Chilothorax ivanovi Lebedev, 1911
Mendidius kobakovi Nikritin, 1979
Mendidius kulikulanus (Balthasar, 1970)⇒ Nobiellus inclusus Reitter, 1892
Mendidius laevicollis Harold, 1866
Mendidius mamajevi Medvedev, 1968
Mendidius margianus Semenov & Me..., 1928
Mendidius multiplex Reitter, 1897
Mendidius nelsinae Medvedev, 1968
Mendidius ogloblini Semenov & Me..., 1928
Mendidius olexai Kral, 1996
Mendidius osiris Koshantschikov, 1916
Mendidius palmetincola Karsch, 1881
Mendidius palmetincolus Karsch, 1881
Mendidius pseudatricolor Nikritin & K..., 1979
Mendidius pseudobidens Balthasar, 1941
Mendidius repetekensis Nikritin, 1971
Mendidius rutilinus Reitter, 1892
Mendidius samarcandicus Nikritin, 1979
Mendidius sijazovi Lebedev, 1932
Mendidius willbergi Reitter, 1891
Mendidius xerxes Petrovitz, 1963
Mendidius zhantievi Gusakov, 1997
Merogyrus rotundiceps Fall, 1907
Mesontoplatys arabicus Harold, 1875
Mesontoplatys bolzi Petrovitz, 1967
Mesontoplatys dorsalis Klug, 1855
Mesontoplatys durandi Petrovitz, 1973
Mesontoplatys effetus Kolbe, 1908
Mesontoplatys indorsalis Endrödi, 1960
Mesontoplatys malzyi Paulian, 1939
Mesontoplatys maoensis (Endrödi, 1964)⇒ Mesontoplatys dorsalis Klug, 1855
Mesontoplatys marlieri Endrödi, 1960
Mesontoplatys mbaoensis Petrovitz, 1969
Mesontoplatys mungo Balthasar, 1946
Mesontoplatys offensus Petrovitz, 1967
Mesontoplatys ovamboensis Petrovitz, 1962
Mesontoplatys parvulus Harold, 1871
Mesontoplatys politipennis Müller, 1941
Mesontoplatys probes Péringuey, 1901
Mesontoplatys remedellii Müller, 1941
Mesontoplatys rougoni Petrovitz, 1974
Mesontoplatys rubrus Bordat, 1992
Mesontoplatys rufolaterus Motschulsky, 1863
Mesontoplatys scutatus Petrovitz, 1961
Mesontoplatys simplicipennis Petrovitz, 1972
Mesontoplatys simplicius Petrovitz, 1974
Mesontoplatys sunantae Stebnicka, 1981
Mesontoplatys triangularis Schmidt, 1907
Mesontoplatys zavadili Balthasar, 1941
Mesontoplatys zulu Petrovitz, 1962
Messyrhus brachypterus Rakovic & Kral, 1997
Micraegialia pusilla G. Horn, 1887
Microtermitodius atomus Pittino, 2006
Microteuchestes brusewitzi Landin, 1974
Monteitheolus fijiensis Howden & Storey, 2000
Mothon sarmaticus Semenov & Me..., 1927
Mozartius jugosus Lewis, 1895
Mozartius jugosus horii Ochi, Kawaha..., 2006
Mozartius jugosus jugosus
Mozartius jugosus shingoi Ochi, Kawaha..., 2006
Mozartius kyushuensis Ochi & Kawah..., 2002
Mozartius testaceus Nomura & Nakane, 1951
Mozartius testaceus shikokuensis Masumoto, 1984
Mozartius testaceus testaceus
Mozartius uenoi Masumoto, 1984
Mozartius uenoi hadai Kawai, 2003
Mozartius uenoi uenoi
Myrhessus afghanus Balthasar, 1939
Myrhessus kubotai Ochi, Kawaha..., 2012
Myrhessus mirabilis Balthasar, 1955
Myrhessus nishinoi Ochi, Kawaha..., 2012
Myrhessus samurai Balthasar, 1941
Myrhessus sunadai Ochi, Kawaha..., 2019
Myrhessus yorikoae Ochi, Kawaha..., 2006
Myrmecaphodius proseni Martiznez, 1952
Mysarus peruanus Petrovitz, 1962
Nanotermitodius andersoni Skelley, Smi..., 2022
Nanotermitodius peckorum Howden, 2003
Napoa peckorum Stebnicka, 1999
Neagolius abchasicus Reitter, 1892
Neagolius amblyodon Daniel, 1900
Neagolius aragatsi Shokhin & Ka..., 2015
Neagolius bilimecki Seidlitz, 1891
Neagolius bilimeckii Seidlitz, 1891
Neagolius dzhungaricus Nikolajev & ..., 1996
Neagolius falcispinis Koshantschikov, 1912
Neagolius haroldi Koshantschikov, 1894
Neagolius hengduanensis Kral, 1997
Neagolius heydeni Harold, 1871
Neagolius interstitialis Koshantschikov, 1894
Neagolius liguricus Daniel, 1902
Neagolius limbolarius (Semenov, 1902)⇒ Neagolius bilimecki Seidlitz, 1891
Neagolius montanus Erichson, 1848
Neagolius montisjuldi Pittino, 1988
Neagolius montivagus Erichson, 1848
Neagolius moronensis Frolov, 2009
Neagolius orinus Koshantschikov, 1912
Neagolius ovitensis Pittino & Ba..., 1994
Neagolius ovitensis laticeps Pittino & Ba..., 1996
Neagolius ovitensis ovitensis
Neagolius penninus Daniel, 1902
Neagolius petri Frolov, 2014
Neagolius pollicatus Erichson, 1848
Neagolius praecox Erichson, 1848
Neagolius savitskiyi Gusakov, 2004
Neagolius schlumbergeri Seidlitz, 1891
Neagolius schlumbergeri consobrinus Daniel, 1900
Neagolius schlumbergeri samniticus Daniel, 1902
Neagolius schlumbergeri schlumbergeri Seidlitz, 1888
Neagolius schlumbergeri temperei Hoffmann, 1938
Neagolius suschkini Koshantschikov, 1913
Neagolius takin Kral, 1995
Neagolius zangbo Kral, 1997
Neocalaphodius karolinae Minkina, 2015
Neocalaphodius moestus Fabricius, 1801
Neocolobopterus abyssinicus Harold, 1861
Neocolobopterus ballioni Schmidt, 1906
Neocolobopterus kivuanus Endrödi, 1956
Neocolobopterus maculicollis Reiche, 1847
Neocolobopterus maculicollis kenyensis Müller, 1939
Neocolobopterus maculicollis maculicollis
Neocolobopterus marginicollis Harold, 1859
Neocolobopterus spangleri Dellacasa, 1990
Neocolobopterus stefenelli Dellacasa, 1984
Neodiapterna erichsoni Harold, 1861
Neodiapterna tognoni Dellacasa & ..., 1997
Neoemadiellus andringitraensis Maté, 2007
Neoemadiellus didactylioides Bordat, 1990
Neoemadiellus didactyloides Bordat, 1990
Neoemadiellus humerosanguineum Maté, 2007
Neoemadiellus perinetensis Bordat, 1990
Neoemadiellus peyrierasi Bordat, 1990
Neoemadiellus ranomafanaensis Maté, 2007
Neoemadiellus ranomandryensis Bordat, 1990
Neoemadiellus unctus Bordat, 1990
Neoemadiellus uxoris Clement, 1986
Neoemadiellus viljanenae Maté, 2007
Neoheptaulacus hepaticolor Quedenfeldt, 1884
Neoheptaulacus puberulus Boheman, 1857
Neoheptaulacus puberulus puberulus
Neoheptaulacus puberulus scotti Paulian, 1939
Neoheptaulacus pygmaeus Boheman, 1857
Neopsammodius blandus Fall, 1932
Neopsammodius canoensis Cartwright, 1955
Neopsammodius culminatus Bates, 1887
Neopsammodius interruptus Say, 1835
Neopsammodius mimeticus Fall, 1932
Neopsammodius quinqueplicatus G. Horn, 1871
Neopsammodius saltilloensis Cartwright, 1955
Neopsammodius veraecrucis Bates, 1887
Neopsammodius werneri O.L.Cartwright, 1955
Neorhyssemus beccarii Balthasar, 1939
Neorhyssemus intrinsecus Gordon & Car..., 1980
Neorhyssemus lineatus Gordon & Car..., 1980
Neorhyssemus quinquecostatus Schmidt, 1911
Neotrichaphodioides caracanus Balthasar, 1970
Neotrichaphodioides ecuadoriensis Petrovitz, 1961
Neotrichaphodioides forsterianus Balthasar, 1960
Neotrichaphodioides volxemi Harold, 1876
Neotrichaphodioides woytkowskii Dellacasa, D..., 2010
Neotrichiorhyssemus babaulti Benard, 1917
Neotrichiorhyssemus balthasari Rakovic, 1987
Neotrichiorhyssemus boninensis Nakane, 1960
Neotrichiorhyssemus esakii Nomura, 1943
Neotrichiorhyssemus expansicollis Benard, 1930
Neotrichiorhyssemus ghati Rakovi�, Me..., 2023
Neotrichiorhyssemus hauseri Balthasar, 1933
Neotrichiorhyssemus hegeri Petrovitz, 1968
Neotrichiorhyssemus hirsutus Clouët, 1901
Neotrichiorhyssemus kentingensis Masumoto & al., 2013
Neotrichiorhyssemus klapaleki Balthasar, 1963
Neotrichiorhyssemus malabaricus Balthasar, 1963
Neotrichiorhyssemus malkini Rakovic, 1987
Neotrichiorhyssemus matthewsi Rakovic, 1983
Neotrichiorhyssemus setiventris Petrovitz, 1968
Neotrichiorhyssemus umbilicatus Petrovitz, 1968
Neotrichonotulus inurbanus Gordon & Howden, 1972Arrow stargazer
Neotrichonotulus perotensis Deloya & Lobo, 1995
Neotrichonotulus urangai Islas, 1955
Nettelislasia euparinoides Islas, 1945
Nialaphodius calidus Harold, 1871
Nialaphodius nigrita Fabricius, 1801
Nialaphodius nigritus Fabricius, 1801
Nialosternus rendalli Wollaston, 1867
Nialosternus sitiphoides d´Orbigny, 1896
Nialus alienus Schmidt, 1922
Nialus consimilis Boheman, 1857
Nialus discelis Endrödi, 1964
Nialus feculentus Fairmaire, 1892
Nialus fingo Petrovitz, 1971
Nialus insignis Schmidt, 1911
Nialus jhavanicus Balthasar, 1971
Nialus motschulskyi Nikritin, 1969
Nialus nefandus Reitter, 1907
Nialus politus Mulsant & Rey, 1870
Nialus tanganyicensis Balthasar, 1967
Nialus tetricus Harold, 1871
Nialus turbidus Bordat, 1992
Nialus varians Duftschmid, 1805
Nialus venyigei Endrödi, 1969
Nimbus affinis Panzer, 1823
Nimbus affinis affinis
Nimbus affinis dorbignyi Clouët, 1896
Nimbus anyerae Ruiz, 1998
Nimbus cartalinius Olsoufieff, 1918
Nimbus contaminatus Herbst, 1783
Nimbus franzinii Pittino, 1978
Nimbus harpagonis Reitter, 1890
Nimbus hoberlandti Tesar, 1945
Nimbus johnsoni Baraud, 1976
Nimbus lederi Harold, 1876
Nimbus libanonensis Petrovitz, 1958
Nimbus makowskyi Koshantschikov, 1891
Nimbus marianii Pittino, 1978
Nimbus obliteratus Panzer, 1823
Nimbus orbignyi Clouët, 1896
Nimbus proximus Adám, 1994
Nimbus richardi Veiga, 1984
Nipponaphodius gotoi Nomura & Nakane, 1951
Nipponoagoliinus yasutakai Ochi & Kawahara, 2001
Nobiellus bonnairei Reitter, 1892
Nobiellus circumductus Solsky, 1876
Nobiellus inclusus Reitter, 1892
Nobiellus pustulifer Reitter, 1892
Nobius bonnairei (Reitter, 1892)⇒ Nobiellus bonnairei Reitter, 1892
Nobius bosaki Tesar, 1945
Nobius chaldaeus Petrovitz, 1971
Nobius costatellus Schmidt, 1922
Nobius diens Balthasar, 1946
Nobius dosangi Akhmetova & ..., 2008
Nobius gresseri Semenov, 1899
Nobius hepaticus Roth, 1851
Nobius rhodiensis Baraud, 1976
Nobius serotinus Panzer, 1799
Nobius zumpti Balthasar, 1937
Nolicus areroensis Bordat & Min..., 2019
Nolicus veteranus Petrovitz, 1962
Notocaulus abessinicus Machatschke, 1955
Notocaulus basilewskyi Endrödi, 1957
Notocaulus felschei Schmidt, 1911
Notocaulus grilloti Walter, 1979
Notocaulus hurneri Machatschke, 1955
Notocaulus laticollis Arrow, 1906
Notocaulus machatschkei Endrödi, 1957
Notocaulus muansicus Machatschke, 1955
Notocaulus muansicus kilimandjarensis Machatschke, 1955
Notocaulus muansicus muansicus
Notocaulus muansicus uhaensis Machatschke, 1955
Notocaulus neumanni Felsche, 1909
Notocaulus nigropiceus Quedenfeldt, 1884
Notocaulus sachtlebeni Balthasar, 1937
Notocaulus schoutedeni Boucomont, 1928
Notocaulus sculpturatus Boheman, 1848
Obaphodius silvestris Bordat, 1988
Odochilus bacchusi Rakovic, 1987
Odochilus balthasari Rakovic, 1987
Odochilus borneensis Rakovi� & A..., 2021
Odochilus buruensis Boucomont, 1926
Odochilus convexus Nomura, 1971
Odochilus haroldi Rakovic, 1987
Odochilus jani Bezdek & Krivan, 2001
Odochilus johnsoni Rakovic, 1987
Odochilus kashizakii Ochi, Kon & ..., 2017
Odochilus laotianus Minkina, 2020
Odochilus montanus Masumoto & K..., 2019
Odochilus parajohnsoni Rakovic, 1997
Odochilus parasetosus Rakovic, 1997
Odochilus parasyntheticus Rakovic, 1997
Odochilus philippinensis Rakovic, 1987
Odochilus pittinoi Rakovic, 1987
Odochilus sarawakensis Rakovic, 1997
Odochilus setosus Boucomont, 1914
Odochilus shavrini Rakovi� & A..., 2021
Odochilus sulawesiensis Minkina, Bez..., 2023
Odochilus syntheticus Harold, 1877
Odochilus syntheticus philippinensis (Rakovic, 1987)⇒ Odochilus philippinensis Rakovic, 1987
Odochilus taiwanus Masumoto, La..., 2016
Odochilus thailandicus Rakovic, 1997
Odontolochus basilewskyi Endrödi, 1957
Odontolochus burgeoni Boucomont, 1930
Odontolochus costatus Endrödi, 1964
Odontolochus ferenczi Endrödi, 1976
Odontolochus gestroi Clouët, 1900
Odontolochus lackneri Rakovic, Men..., 2014
Odontolochus monteithi Stebnicka & ..., 1996
Odontolochus oberthueri Clouët des P..., 1900
Odontolochus oberthuri Clouët, 1900
Odontolochus paucus Schmidt, 1916
Odontolochus raffrayi Paulian, 1942
Odontolochus setulosus Endrödi, 1964
Odontolochus spinicollis Harold, 1871
Odontolochus villiersi Petrovitz, 1966
Odontolochus weiri Stebnicka & ..., 1996
Odontolytes andamanensis (W. Koshantsc..., 1916)⇒ Auperia andamanensis Koshantschikov, 1916
Odontolytes angusticollis Schmidt, 1909
Odontolytes capitosus Harold, 1867
Odontolytes denominatus Chevrolat, 1864
Odontolytes domingo Stebnicka, 2002
Odontolytes guayara Stebnicka, 2002
Odontolytes huebneri Petrovitz, 1970
Odontolytes iquitosae Stebnicka, 2002
Odontolytes landai Balthasar, 1963
Odontolytes loretoensis Stebnicka, 2002
Odontolytes minutus Petrovitz, 1973
Odontolytes panamensis Stebnicka, 2002
Odontolytes puyoensis Stebnicka, 2002
Odontolytes rondoniae Stebnicka, 2002
Odontolytes tectipennis Stebnicka & ..., 2005
Odontolytes teutoniae Stebnicka, 2002
Odontolytes transversaria Schmidt, 1909
Odontolytes transversarius Schmidt, 1909
Odontolytes viejoae Stebnicka & ..., 2005
Odontolytes waoraniae Stebnicka & ..., 2005
Odontopsammodius aenictus Cartwright, 1955
Odontopsammodius armaticeps Fall, 1932
Odontopsammodius atopus Cartwright, 1955
Odontopsammodius bidens (Horn, 1871)⇒ Parapsammodius bidens Horn, 1871
Odontopsammodius brunneus Balthasar, 1961
Odontopsammodius cameneni Chalumeau, 1976
Odontopsammodius chipiririi Cartwright, 1955
Odontopsammodius cruentus Harold, 1867Scarab beetle
Odontopsammodius decuiella Chalumeau, 1979
Odontopsammodius fimbriatus Cartwright, 1955
Odontopsammodius formosus Cartwright, 1955
Odontopsammodius insulcatus Schmidt, 1916
Odontopsammodius mapirii Cartwright, 1955
Orocanthus helenae Endrodi, 1973
Orodaliscoides fimbripes Brown, 1928
Orodaliscoides giulianii Gordon, 1977
Orodaliscoides reflexus Schmidt, 1910
Orodaliscoides rugosiceps Harold, 1859
Orodaliscus angulatulus Schmidt, 1916
Orodaliscus rotundangulus Reitter, 1900
Orodaliscus spalacophilus Novikov, 1996
Orodaliscus transaralicus Nikolajev, 1983
Orodaliscus zangi Schmidt, 1906
Oromus alpinus Scopoli, 1763
Oromus corvinus
Oromus leisteri Medvedev, 1968
Oromus tragicus Schmidt, 1920
Oscarinus abusus Fall, 1907
Oscarinus bottimeri Cartwright, 1957Spotted stargazer
Oscarinus brimleyi Cartwright, 1939
Oscarinus cabreroi Dellacasa, D..., 2013
Oscarinus crassuloides Fall, 1907
Oscarinus crassulus Horn, 1870
Oscarinus floridanus Robinson, 1947
Oscarinus indutilis Harold, 1874
Oscarinus lodingi Cartwright, 1957Spotted stargazer
Oscarinus matiganae Paulsen, 2006Electric stargazers
Oscarinus odocoilis Robinson, 1939Mottled stargazer
Oscarinus pseudabusus Cartwright, 1957Spotted stargazer
Oscarinus rusicola Melsheimer, 1845
Oscarinus silvanicus Cartwright, 1972
Oscarinus spiniclypeus Hinton, 1934
Oscarinus stuessyi Gordon & Ske..., 2007Lesser weever
Oscarinus texensis Cartwright, 1972
Oscarinus welderi Gordon & Ske..., 2007Lesser weever
Oscarinus windsori Cartwright, 1939
Osmanius balthasari Petrovitz, 1963
Osmanius dellacasai Petrovitz, 1970
Osmanius persicus Branco & Baraud, 1988
Otophorus haemorrhoidalis Linnaeus, 1758Bloody small dung beetle
Oxyataenius morosus Harold, 1869
Oxycorythus morawitzi Solsky, 1876
Oxyomoides baileyi Skelley & Go..., 2001
Oxyomoides dyspistus Skelley & Wo..., 1991
Oxyomoides sepultus Cartwright, 1944
Oxyomus aciculatus Schmidt, 1909
Oxyomus alligator Ho, Hu & Fik..., 2022
Oxyomus arunae Stebnicka, 1985
Oxyomus bremeri Stebnicka, 1981
Oxyomus cameratus Schmidt, 1908
Oxyomus costipennis Boheman, 1857
Oxyomus costulatus Fairmaire, 1849
Oxyomus curvus Schmidt, 1909
Oxyomus debilis Harold, 1877
Oxyomus dschagga Petrovitz, 1969
Oxyomus inaequalis Lansberge, 1886
Oxyomus ishidai Nakane, 1977
Oxyomus jucundulus Péringuey, 1901
Oxyomus jugalis (Péringuey, 1901)⇒ Blackburneus jugalis Péringuey, 1901
Oxyomus kiuchii Masumoto, 1991
Oxyomus kocoti Minkina, 2018
Oxyomus lineatosulcatus Harold, 1859
Oxyomus malgorzatae Minkina, 2016
Oxyomus mariateresae Dellacasa, D..., 2014
Oxyomus masumotoi Nomura, 1973
Oxyomus medioximus (Péringuey, 1901)⇒ Pleuraphodius medioximus Péringuey, 1901
Oxyomus mencli Minkina, 2016
Oxyomus miliaris Schmidt, 1908
Oxyomus nanxiensis Masumoto, Ki..., 2018
Oxyomus nearcticus H. F. Wickham, 1914
Oxyomus nowaki Minkina, 2016
Oxyomus nubigenus Petrovitz, 1968
Oxyomus opacifrons (Horn, 1871)⇒ Oxyomus silvestris Scopoli, 1763
Oxyomus setosopunctatus Schmidt, 1911
Oxyomus silvestris Scopoli, 1763
Oxyomus simillimus Schmidt, 1908
Oxyomus sulcipennis (Dejean, 1833)⇒ Xenoheptaulacus tricostatus Harold, 1896
Oxyomus sylvestris Scopoli, 1763Forest small dung beetle
Oxyomus taipingensis Masumoto, Ki..., 2014
Oxyomus thailandicus Masumoto, 1991
Ozodius canberrae Stebnicka & ..., 1995
Ozodius neglectus Schmidt, 1913
Ozodius walpolensis Stebnicka & ..., 1995
Ozodius windyensis Stebnicka & ..., 1995
Palnia cambeforti Kral, 1999
Palnia loebli Stebnicka, 1985
Palnia stebnickae Cambefort, 1989
Palnia thailandica Bellmann, 2020
Parabodilus wollastoni Harold, 1862
Parabodilus wollastoni iranicus Balthasar, 1946
Parabodilus wollastoni wollastoni
Paracoptochirus kozanensis Pittino, 2001
Paracoptochirus petrovitzi Branco & Baraud, 1988
Paracoptochirus singularis Harold, 1868
Paracoptochirus vignai Carpaneto & ..., 1990
Paracorythoderus casperi Kolbe, 1909
Paracorythoderus endroedyi Endrödi, 1991
Paracorythoderus marshalli Brauns, 1900
Paracorythoderus ridens Tangelder & ..., 1982
Paracrossidius impressiusculus Fairmaire, 1888
Paracrossidius instigator Balthasar, 1932
Paracrossidius kozlovi Koshantschikov, 1916
Paracrossidius laticapitus Zhang, 1992
Paracrossidius melanodiscus Zhang, 1992
Paracrossidius planipennis Petrovitz, 1961
Paracrossidius radekcervenkai Cervenka, 2017
Paracrossidius sichuanensis Zhang, 1992
Paracrossidius suvorovi Kabakov, 1996
Paracrossidius transmontanus Balthasar, 1932
Paracrossidius truculentus Balthasar, 1963
Paradeloparius brincki Landin, 1959
Paradeloparius cavicollis Endrödi, 1964
Paradeloparius circumdatus Klug, 1855
Paradeloparius emeritus Péringuey, 1901
Paradidactylia avuncula Balthasar, 1946
Paradidactylia avunculus Balthasar, 1946
Paradidactylia avus Petrovitz, 1966
Paradidactylia biseriata Schmidt, 1908
Paradidactylia biseriatus Schmidt, 1908
Paradidactylia brivioi Petrovitz, 1961
Paradidactylia carinulata Motschulsky, 1863
Paradidactylia carinulatus Motschulsky, 1863
Paradidactylia continua Petrovitz, 1972
Paradidactylia continuus Petrovitz, 1972
Paradidactylia dionysii Clement, 1958
Paradidactylia edithae Endrödi, 1964
Paradidactylia fastigata Schmidt, 1908
Paradidactylia flavescens Balthasar, 1961
Paradidactylia haafi Petrovitz, 1961
Paradidactylia hiekei Endrödi, 1983
Paradidactylia incomparabilis Bordat, 2005
Paradidactylia jattaensis Petrovitz, 1969
Paradidactylia kathmandui Stebnicka, 1985
Paradidactylia kindianus Petrovitz, 1958
Paradidactylia klasinskii Minkina, 2016
Paradidactylia madibirana Petrovitz, 1964
Paradidactylia madibiranus Petrovitz, 1964
Paradidactylia montana (Clement, 1958)⇒ Paradidactylia pierreclementi Dellacasa, 1988
Paradidactylia ovatula Harold, 1861
Paradidactylia ovatulus Harold, 1861
Paradidactylia pierreclementi Dellacasa, 1988
Paradidactylia semiangulus Balthasar, 1952
Paradidactylia shantungensis Balthasar, 1941
Paradidactylia subflavus Petrovitz, 1969
Paradidactylia umfolozica Endrödi, 1976
Paradidactylia umfolozicus Endrödi, 1976
Paradidactylia variolosa Clément, 1958
Paradidactylia variolosus Clement, 1958
Paradidactylia venalis Schmidt, 1911
Paradidactylia venaloides Endrödi, 1960
Paradidactylia wichei Petrovitz, 1961
Parammoecius amanicus Stebnicka, 1978
Parammoecius asphaltinus Kolenati, 1846
Parammoecius brevithorax Sumakov, 1903
Parammoecius corvinus Erichson, 1848
Parammoecius gibbus Germar, 1817
Parammoecius osseticus Iablokov-Khn..., 1972
Parammoecius pyrenaeus Jacquelin du..., 1863
Parammoecius yamato (Nakane, 1960)⇒ Ammoecius yamato Nakane, 1960
Paranimbus longitarsis Harold, 1860
Paranimbus penai Petrovitz, 1970
Paranimbus peruanus Erichson, 1834
Paranimbus zoiai Dellacasa, 1990
Paraplesiataenius catarinaensis Stebnicka, 2003
Paraplesiataenius genieri Stebnicka, 2003
Paraplesiataenius tremolerasi Schmidt, 1911
Parapsammodius bidens Horn, 1871
Parapsammodius integer Bates, 1887
Parapsammodius pseudointeger Verdú, Steb..., 2006
Parapsammodius puncticollis LeConte, 1858Berg-salmon gill worm
Pararhyssemus coluber Mayet, 1887
Parataenius brunneus Schmidt, 1922
Parataenius derbesis Solier, 1851
Parataenius estero Stebnicka & ..., 2009
Parataenius martinezi Stebnicka & ..., 2009
Parataenius selvae Stebnicka & ..., 2009
Parataenius simulator Harold, 1868
Paratrichaphodius badius Boheman, 1857
Paratrichaphodius foveiventris Raffray, 1877
Paratrichaphodius sericeus Schmidt, 1909
Pardalosus neodistinctus Brown, 1928
Pardalosus pardalis LeConte, 1857
Pardalosus pseudopardalis Gordon & Ske..., 2007Lesser weever
Pardalosus pumilio Schmidt, 1907Spotted weever
Pardalosus sayi Gordon, 2006Greater weever
Pardalosus serval Say, 1835
Pardalosus slevini Van Dyke, 1928
Passalaphodius cartwrighti (Martínez, 1953)⇒ Saprosites sulcatus Harold, 1869
Passaliolla aspericeps Harold, 1876
Passaliolla cancellata Bates, 1887
Passaliolla corticalis Bates, 1887
Passaliolla cossonoides Bates, 1887
Passaliolla eugastrica Harold, 1869
Paulianellus andrei Minkina & Král, 2018
Paulianellus asahinai Nakane, 1951
Paulianellus decoctor Balthasar, 1965
Paulianellus jacksoni Petrovitz, 1975
Paulianellus khandui Stebnicka, 1981
Paulianellus laoticus Mencl, Král ..., 2015
Paulianellus maderi Balthasar, 1938
Paulianellus mureensis Stebnicka, 1981
Paulianellus neomaderi Mencl, Král ..., 2015
Paulianellus nepalensis Balthasar, 1965
Paulianellus taiwanensis Mencl, Král ..., 2015
Paulianellus trisuliensis Stebnicka, 1986
Petrovitzius tesari Rakovic, 1986
Phaeaphodius costalis Gebler, 1848
Phaeaphodius dauricus Harold, 1863
Phaeaphodius fusculus Reitter, 1892
Phaeaphodius himalorectus Ahrens & Ste..., 1997
Phaeaphodius jouravliowi Reitter, 1907
Phaeaphodius kiulungensis Balthasar, 1932
Phaeaphodius motoi Paulian, 1939
Phaeaphodius novikovi Kabakov, 1998
Phaeaphodius plutenkoi Kral, 2002
Phaeaphodius pupillus Balthasar, 1961
Phaeaphodius rectus Motschulsky, 1866
Phaeaphodius roddi Koshantschikov, 1911
Phaeaphodius tauricola Hrubant, 1961
Phaeaphodius youngai Endrödi, 1967
Phalacronothus ambulans Petrovitz, 1971
Phalacronothus atlanticus Balthasar, 1960
Phalacronothus biguttatus Germar, 1824
Phalacronothus botulus Balthasar, 1945
Phalacronothus carinulatus Motschulsky, 1863
Phalacronothus ceylonensis Schmidt, 1912
Phalacronothus citellorum Semenov & Me..., 1929
Phalacronothus coenosus
Phalacronothus flavolimbatus Petrovitz, 1963
Phalacronothus fumigatulus Reitter, 1892
Phalacronothus hilaris Harold, 1869
Phalacronothus paracoenosus
Phalacronothus pseudoavunculus Masumoto, 1977
Phalacronothus pusillus
Phalacronothus putoni Reitter, 1894
Phalacronothus quadriguttatus
Phalacronothus quadrimaculatus Linnaeus, 1761
Phalacronothus semiangulus Balthasar, 1952
Phalacronothus suturinigra Schmidt, 1917
Pharaphodius acutidens Petrovitz, 1964
Pharaphodius acutus Schmidt, 1908
Pharaphodius aegyptiacus Petrovitz, 1964
Pharaphodius aprilis Bordat, 1988
Pharaphodius archettii Müller, 1942
Pharaphodius argutus Schmidt, 1911
Pharaphodius atroscutellatus Schmidt, 1909
Pharaphodius attritus (Balthasar, 1933)⇒ Pharaphodius marginellus Fabricius, 1781
Pharaphodius bechuanus Petrovitz, 1958
Pharaphodius bigibber Bordat, 1992
Pharaphodius birmanicus Petrovitz, 1965
Pharaphodius bodiliformis Balthasar, 1970
Pharaphodius calo Balthasar, 1971
Pharaphodius campestris Mittal, 1993
Pharaphodius chausibensis Endrödi, 1980
Pharaphodius coiffaiti Clement, 1985
Pharaphodius congolanus Endrödi, 1964
Pharaphodius corax Balthasar, 1937
Pharaphodius cornix Balthasar, 1933
Pharaphodius cornutus Wiedemann, 1823
Pharaphodius costatus Endrödi, 1964
Pharaphodius crenatus Harold, 1862
Pharaphodius curvodilatatus Schmidt, 1909
Pharaphodius desertoides Endrödi, 1964
Pharaphodius desertus Klug, 1845
Pharaphodius devabhumi Mittal, 1993
Pharaphodius diadema Wiedemann, 1823
Pharaphodius difficilis Endrödi, 1955
Pharaphodius discoidalis Boheman, 1857
Pharaphodius discoidalis discoidalis
Pharaphodius discoidalis neghellinus Balthasar, 1939
Pharaphodius discolor Erichson, 1859
Pharaphodius dubiosus Péringuey, 1901
Pharaphodius dubitosus Endrödi, 1964
Pharaphodius dubitosus dubitosus
Pharaphodius dubitosus hautkatangae Endrödi, 1964
Pharaphodius evae Endrödi, 1973
Pharaphodius falzonii Petrovitz, 1973
Pharaphodius ferenczi Bordat, 1990
Pharaphodius ferreirae Petrovitz, 1971
Pharaphodius fiechteri Balthasar, 1941
Pharaphodius guineensis Klug, 1835
Pharaphodius hastulifer Petrovitz, 1958
Pharaphodius hauserianus Balthasar, 1935
Pharaphodius hetaerus Petrovitz, 1958
Pharaphodius impressipennis Schmidt, 1908
Pharaphodius impressithorax Bordat, 1992
Pharaphodius impurus Roth, 1851
Pharaphodius incommodus Bordat, 1989
Pharaphodius indicus Petrovitz, 1958
Pharaphodius infinitus Schmidt, 1920
Pharaphodius intercalaris Péringuey, 1901
Pharaphodius intonsus Petrovitz, 1964
Pharaphodius javanus Schmidt, 1909
Pharaphodius jodhpurensis Petrovitz, 1958
Pharaphodius katanai Paulian, 1939
Pharaphodius lacunosus Schmidt, 1911
Pharaphodius levithorax Bordat, 1992
Pharaphodius lewisi
Pharaphodius mabiranus Balthasar, 1933
Pharaphodius mangoroensis Bordat, 1990
Pharaphodius marginellus Fabricius, 1781
Pharaphodius massaicus Gerstaecker, 1884
Pharaphodius medius Schmidt, 1920
Pharaphodius moderatus Petrovitz, 1967
Pharaphodius musculus Petrovitz, 1969
Pharaphodius mussooriensis Mittal, 1993
Pharaphodius negus Petrovitz, 1964
Pharaphodius okatumbanus Balthasar, 1942
Pharaphodius oleosus Harold, 1871
Pharaphodius orientalis Harold, 1862
Pharaphodius pallescens Walker, 1871
Pharaphodius panchrestus Balthasar, 1971
Pharaphodius paratschakai Clement, 1985
Pharaphodius parvatia Mittal, 1993
Pharaphodius pereirai Balthasar, 1955
Pharaphodius perfidus Schmidt, 1908
Pharaphodius perplicatus Bordat, 1996
Pharaphodius pongensis Endrödi, 1983
Pharaphodius posticus Boheman, 1857
Pharaphodius priscus Motschulsky, 1858
Pharaphodius profundus Schmidt, 1911
Pharaphodius pseudignotus Bordat, 1986
Pharaphodius pseudimpurus Balthasar, 1941
Pharaphodius pseudodesertus Bordat, 1996
Pharaphodius pseudotibialis Bordat, 1992
Pharaphodius putearius Reitter, 1895
Pharaphodius ratnapurensis
Pharaphodius redargutus Balthasar, 1971
Pharaphodius resplendens Balthasar, 1941
Pharaphodius robustus Walker, 1858
Pharaphodius rougeoti Paulian, 1984
Pharaphodius rubricosus Boheman, 1857
Pharaphodius rugosicapita Mittal, 1993
Pharaphodius rugosostriatus Waterhouse, 1875
Pharaphodius russatus Erichson, 1842
Pharaphodius schatzmayri Balthasar, 1939
Pharaphodius seicherti Rakovic, 1991
Pharaphodius spinatus Schmidt, 1920
Pharaphodius sudanicus Petrovitz, 1964
Pharaphodius sundaicus Balthasar, 1946
Pharaphodius tamalensis Endrödi, 1973
Pharaphodius taufikijanus Balthasar, 1971
Pharaphodius tavetanus Dellacasa, 1983
Pharaphodius tenerus Petrovitz, 1959
Pharaphodius tschakai Petrovitz, 1964
Pharaphodius ugongoanus Petrovitz, 1964
Pharaphodius unicornis Petrovitz, 1973
Pharaphodius vastus Petrovitz, 1964
Pharaphodius vernilis Schmidt, 1911
Pharaphodius vividus Petrovitz, 1964
Pharaphodius zavattarii Müller, 1941
Pholeoaphodius duplicatus Schmidt, 1907
Pholeoaphodius kalungwensis Paulian, 1954
Pholeoaphodius peckorum Endrödi, 1982
Pholeoaphodius sphinx Paulian, 1942
Pholeoaphodius ukamiensis Petrovitz, 1962
Phycochus graniceps Broun, 1886
Phycocus graniceps Broun, 1883
Pittinius farkaci Rakovic, Kra..., 2014
Pittinius omnisetosus Rakovic & Kral, 1997
Plagiogonus arenarius Olivier, 1789
Plagiogonus bortkevitschi Koshantschikov, 1916
Plagiogonus burgaltaicus Csiki, 1901
Plagiogonus chan Endrödi, 1967
Plagiogonus clypeodentatus Bordat, 1990
Plagiogonus critchlowi Bordat, 2005
Plagiogonus culminarius Reitter, 1900
Plagiogonus duporti Paulian, 1945
Plagiogonus esimoides Reitter, 1892
Plagiogonus farai Balthasar, 1955
Plagiogonus hirticeps Péringuey, 1901
Plagiogonus inflatus Balthasar, 1938
Plagiogonus krataay Masumoto, 1992
Plagiogonus martensi Stebnicka, 1986
Plagiogonus nanoides Balthasar, 1961
Plagiogonus nanus Fairmaire, 1860
Plagiogonus oberthueri Paulian, 1936
Plagiogonus oberthuri Paulian, 1936
Plagiogonus ohkurai Masumoto, 1992
Plagiogonus palea Balthasar, 1967
Plagiogonus praeustus Ballion, 1870
Plagiogonus putridus (Geoffroy, 1785)⇒ Plagiogonus arenarius Olivier, 1789
Plagiogonus quadriclypealis Cervenka, 1995
Plagiogonus ramamiensis Stebnicka, 1981
Plagiogonus reitteri Koshantschikov, 1894
Plagiogonus separatus Petrovitz, 1962
Plagiogonus sextus Bordat, 2019
Plagiogonus shirahatai Nakane, 1951
Plagiogonus syriacus Harold, 1863
Plagiogonus tertius Bordat, 1992
Plagiogonus zobeidae Petrovitz, 1980
Planolinellus grzegorzi Minkina, 2017
Planolinellus rufoanalis Petrovitz, 1961
Planolinellus vittatus Say, 1825
Planolinoides aenictus Cooper & Gordon, 1987Benttooths
Planolinoides borealis Gyllenhal, 1827
Planolinoides duplex LeConte, 1878
Planolinoides neotomae Fall & Cocke..., 1907
Planolinoides pectoralis LeConte, 1857
Planolinus borealis (Gyllenhal, 1827)⇒ Planolinoides borealis Gyllenhal, 1827
Planolinus fasciatus Olivier, 1789
Planolinus khaoensis Stebnicka, 1988
Planolinus pratensis Nomura & Nakane, 1951
Planolinus tenellus Say, 1823
Planolinus uliginosus (Hardy, 1847)⇒ Planolinus fasciatus Olivier, 1789
Planolinus vittatus Say, 1825
Platyderides arrowi Schmidt, 1910
Platyderides doiangkhangensis Masumoto, 1991
Platyderides jengi Masumoto, Ki..., 2014
Platyderides klapperichi Balthasar, 1942
Platyderides planipennis Petrovitz, 1961
Platyderides suvorovi Kabakov, 1996
Platyderides truculentus Balthasar, 1963
Platyderides vietnamicus Cervenka, 1995
Platytomus antipodum Fauvel, 1903
Platytomus ashantii Endrödi, 1973
Platytomus atlanticus Cartwright, 1948Horned blenny
Platytomus bordati Rakovi�, Me..., 2020
Platytomus brasiliensis Rakovi�, Me..., 2020
Platytomus caelicollis Cartwright, 1948Horned blenny
Platytomus darwini Cartwright, 1970
Platytomus freudei Balthasar, 1960
Platytomus gomyi Pittino & Ma..., 1986
Platytomus gregalis Cartwright, 1948
Platytomus grisoli Paulian, 1942
Platytomus grundeli Van Dyke, 1918
Platytomus hawaiiensis Rakovic, 1981
Platytomus indicus Balthasar, 1941
Platytomus jailensis Apostolov & ..., 1986
Platytomus laevistriatus Perris, 1869
Platytomus longulus Cartwright, 1948Horned blenny
Platytomus micros Bates, 1887Beetle
Platytomus mongolicus Medvedev, 1974
Platytomus nagporensis Pittino & Ma..., 1986
Platytomus nathani Pittino & Ma..., 1986
Platytomus notialis Cartwright, 1948Horned blenny
Platytomus obscurior Blackburn, 1904
Platytomus pachypus Lea, 1923
Platytomus parvulus Chevrolat, 1864Dung beetle
Platytomus pumilio Balthasar, 1966
Platytomus sabulosus (Mulsant, 1842)⇒ Platytomus tibialis Fabricius, 1798
Platytomus tibialis Fabricius, 1798
Platytomus variolosus Kolenati, 1846
Platytomus yadai Ochi, Kawaha..., 2006
Pleuraphodius abax Endrödi, 1960
Pleuraphodius aequatorialis Petrovitz, 1966
Pleuraphodius agnoscibilis Bordat, 1997
Pleuraphodius anceps Schmidt, 1911
Pleuraphodius arabiaefelicis Král, Rakovi..., 2016
Pleuraphodius assimilis Petrovitz, 1962
Pleuraphodius berlandi Paulian, 1945
Pleuraphodius bibatillatus Petrovitz, 1958
Pleuraphodius bovis Endrödi, 1960
Pleuraphodius breuningi Endrödi, 1964
Pleuraphodius brunneus Thunberg, 1818
Pleuraphodius burgeoni Paulian, 1939
Pleuraphodius burorum Endrödi, 1980
Pleuraphodius buschirii Petrovitz, 1964
Pleuraphodius capillatus Bordat, 2016
Pleuraphodius carinatipennis Bordat, 1994
Pleuraphodius chaboti Paulian, 1942
Pleuraphodius clementi Bordat, 1986
Pleuraphodius confinis Schmidt, 1911
Pleuraphodius conformis Petrovitz, 1967
Pleuraphodius consocius Endrödi, 1964
Pleuraphodius darfurensis Balthasar, 1963
Pleuraphodius decellei Balthasar, 1966
Pleuraphodius dingaani Petrovitz, 1964
Pleuraphodius dispar Bordat, 2005
Pleuraphodius ducorpsi Paulian, 1942
Pleuraphodius ebeninus Endrödi, 1960
Pleuraphodius elgonensis Balthasar, 1967
Pleuraphodius fosseyae Bordat, Camb..., 1991
Pleuraphodius funestus Endrödi, 1955
Pleuraphodius gnomus Kozhantschikov, 1916
Pleuraphodius hamoriae Endrödi, 1973
Pleuraphodius hanstroemi Landin, 1956
Pleuraphodius hospes Endrödi, 1964
Pleuraphodius humator Endrödi, 1964
Pleuraphodius hybridus Endrödi, 1964
Pleuraphodius ilamensis Stebnicka, 1990
Pleuraphodius impudicus Fabricius, 1801
Pleuraphodius javanus Balthasar, 1945
Pleuraphodius jombaensis Bordat, Camb..., 1991
Pleuraphodius kaffa Petrovitz, 1964
Pleuraphodius kaokoveldensis Endrödi, 1991
Pleuraphodius kavani Balthasar, 1935
Pleuraphodius kavani aethiopicus Müller, 1942
Pleuraphodius kilimandjaroensis Petrovitz, 1965
Pleuraphodius kuijteni Petrovitz, 1969
Pleuraphodius landini Endrödi, 1956
Pleuraphodius laterociliatus Petrovitz, 1958
Pleuraphodius latistriatus Bordat, 1990
Pleuraphodius leo Paulian, 1942
Pleuraphodius letabus Landin, 1956
Pleuraphodius lewisi Waterhouse, 1875
Pleuraphodius lineatosulcatus Harold, 1859
Pleuraphodius luzonensis Petrovitz, 1972
Pleuraphodius malangensis Balthasar, 1960
Pleuraphodius malawiensis Bordat, 1997
Pleuraphodius matamataensis Endrödi, 1991
Pleuraphodius maynei Paulian, 1939
Pleuraphodius medanii Endrödi, 1979
Pleuraphodius medioximus Péringuey, 1901
Pleuraphodius mime Paulian, 1942
Pleuraphodius montuosus Schmidt, 1911
Pleuraphodius multicarinatus Endrödi, 1964
Pleuraphodius mutilus Schmidt, 1911
Pleuraphodius mwingi Bordat, 2005
Pleuraphodius natalensis Petrovitz, 1964
Pleuraphodius olkokolae Endrödi, 1971
Pleuraphodius paranceps Endrödi, 1979
Pleuraphodius pumilus Quedenfeldt, 1884
Pleuraphodius purkynei Balthasar, 1933
Pleuraphodius pusio Kolbe, 1908
Pleuraphodius pygmaeus Boheman, 1857
Pleuraphodius ratnapurensis Stebnicka, 1988
Pleuraphodius reticulatus Endrödi, 1980
Pleuraphodius reverendus Bordat, 2014
Pleuraphodius rothschildi Schmidt, 1911
Pleuraphodius rudis Endrödi, 1971
Pleuraphodius sebastianellus Endrödi, 1973
Pleuraphodius simillimus Petrovitz, 1964
Pleuraphodius singularis Petrovitz, 1969
Pleuraphodius socius Endrödi, 1964
Pleuraphodius soudanensis Petrovitz, 1958
Pleuraphodius sparsepunctatus Endrödi, 1964
Pleuraphodius stehliki Endrödi, 1977
Pleuraphodius stipulus Petrovitz, 1962
Pleuraphodius subteter Petrovitz, 1962
Pleuraphodius sulcipennis Boheman, 1857
Pleuraphodius tectipennis Endrödi, 1964
Pleuraphodius tectoformibus Schmidt, 1909
Pleuraphodius teter Roth, 1851
Pleuraphodius thomasi Endrödi, 1973
Pleuraphodius umbilicatus Bordat, 1996
Pleuraphodius utae Endrödi, 1973
Pleuraphodius vespuccii Petrovitz, 1972
Pleuraphodius viscaensis Stebnicka, 1993
Pleuraphodius wardi Bordat, 1994
Pleurophorus africanus Pittino, 1984
Pleurophorus akamasicus Pittino & Mi..., 2007
Pleurophorus anatolicus Petrovitz, 1961
Pleurophorus apicipennis Reitter, 1892
Pleurophorus arabicus Pittino & Ma..., 1986
Pleurophorus atlanticus (Cartwright, 1948)⇒ Platytomus atlanticus Cartwright, 1948
Pleurophorus banksi Pittino & Ma..., 1986
Pleurophorus batesi (Arrow, 1903)⇒ Platytomus micros Bates, 1887
Pleurophorus beccarii Pittino & Ma..., 1986
Pleurophorus bengalensis Pittino & Ma..., 1986
Pleurophorus besucheti Pittino & Ma..., 1986
Pleurophorus caelicollis (Cartwright, 1948)⇒ Platytomus caelicollis Cartwright, 1948
Pleurophorus caesus Creutzer, 1796Cut small dung beetle
Pleurophorus cambeforti Pittino & Ma..., 1986
Pleurophorus capicola Péringuey, 1901
Pleurophorus cracens Motschulsky, 1863
Pleurophorus dinajpurensis Pittino & Ma..., 1986
Pleurophorus dubius
Pleurophorus edithae Endrödi, 1973
Pleurophorus formosanus Pittino & Ma..., 1986
Pleurophorus gregalis (Cartwright, 1948)⇒ Platytomus gregalis Cartwright, 1948
Pleurophorus impressicollis Boheman, 1858
Pleurophorus laticollis Pittino & Ma..., 1986
Pleurophorus ledouxi Pittino & Ma..., 1986
Pleurophorus longulus (Cartwright, 1948)⇒ Platytomus longulus Cartwright, 1948
Pleurophorus madagassus Pittino & Ma..., 1986
Pleurophorus maghrebinicus Pittino & Ma..., 1986
Pleurophorus mediterranicus Pittino & Ma..., 1986
Pleurophorus micros Bates, 1887
Pleurophorus natalensis (Péringuey, 1901)⇒ Saprosites natalensis Péringuey, 1901
Pleurophorus nepalensis Pittino & Ma..., 1986
Pleurophorus notialis (Cartwright, 1948)⇒ Platytomus notialis Cartwright, 1948
Pleurophorus opacus Reitter, 1892
Pleurophorus pannonicus Petrovitz, 1961
Pleurophorus parvulus Chevrolat, 1864
Pleurophorus raffrayi Pittino & Ma..., 1986
Pleurophorus schereri Pittino & Ma..., 1986
Pleurophorus setosus Pittino & Ma..., 1986
Pleurophorus thailandicus Pittino & Ma..., 1986
Pleurophorus tonkinensis Balthasar, 1933
Pleurophorus torretassoi (Schatzmayer, 1930)⇒ Pleurophorus opacus Reitter, 1892
Pleurophorus ventralis (Horn, 1887)⇒ Hornietus ventralis Horn, 1887
Pleurophorus villiersi Bordat, 1984
Podotenoides jeannelianus Balthasar, 1945
Podotenus (Podotenus) gracilipes Harold, 1867
Podotenus (Pseudopodotenus) fulviventris Fairmaire & ..., 1860
Podotenus allynensis Stebnicka & ..., 1994
Podotenus arowsmithensis Stebnicka & ..., 1994
Podotenus arrowsmithensis (Stebnicka & ..., 1994)⇒ Podotenus arowsmithensis Stebnicka & ..., 1994
Podotenus athertonensis Stebnicka & ..., 1994
Podotenus badgingarrae Stebnicka & ..., 1994
Podotenus baldiensis Blackburn, 1904
Podotenus barringtonensis Stebnicka & ..., 1994
Podotenus bourkensis Stebnicka & ..., 1994
Podotenus callabonnensis Blackburn, 1895
Podotenus channonensis Stebnicka & ..., 1994
Podotenus coffensis Stebnicka & ..., 1994
Podotenus dilgryensis Stebnicka & ..., 1994
Podotenus erosus Erichson, 1842
Podotenus forrestensis Stebnicka & ..., 1994
Podotenus gallagheri Stebnicka & ..., 1994
Podotenus insignior Blackburn, 1904
Podotenus integrifrons Lea, 1923
Podotenus kulkynensis Stebnicka & ..., 1995
Podotenus lindensis Blackburn, 1892
Podotenus mattewsi (Stebnicka & ..., 1994)⇒ Podotenus matthewsi Stebnicka & ..., 1994
Podotenus matthewsi Stebnicka & ..., 1994
Podotenus nigrosetosus Stebnicka & ..., 1994
Podotenus oodlawirraensis Stebnicka & ..., 1994
Podotenus otwayensis Stebnicka & ..., 1994
Podotenus pseudovictoriae Stebnicka & ..., 1994
Podotenus schmidti Stebnicka & ..., 1994
Podotenus storeyi Stebnicka & ..., 1994
Podotenus suberosus Blackburn, 1904
Podotenus subfuscus Petrovitz, 1975
Podotenus tennantensis Stebnicka & ..., 1995
Podotenus toowoombaensis Stebnicka & ..., 1994
Podotenus tricolor Lea, 1923
Podotenus victoriae Blackburn, 1897
Podotenus wauchopensis Stebnicka & ..., 1994
Podotenus williamsi Stebnicka & ..., 1994
Proctophanes anneae Stebnicka & ..., 1995
Proctophanes caldwellensis Stebnicka & ..., 1995
Proctophanes distinguendus Schmidt, 1910
Proctophanes heterodoxus Lea, 1923
Proctophanes metasternalis Lea, 1923
Proctophanes minor Blackburn, 1897
Proctophanes scrubensis Stebnicka & ..., 1995
Proctophanes sculptus Hope, 1846
Psammodaphodius kochi Endrodi, 1976
Psammodius aegialioides (Haldeman, 1848)⇒ Psammodius laevipennis Costa, 1844
Psammodius aenictus (Cartwright, 1955)⇒ Odontopsammodius aenictus Cartwright, 1955
Psammodius ambiguus (Fall, 1907)⇒ Xeropsamobeus ambiguus Fall & Cocke..., 1907
Psammodius asper (Fabricius, 1775)⇒ Psammodius sulcicollis Illiger, 1802
Psammodius asperoides Pittino, 2007
Psammodius atopus (Cartwright, 1955)⇒ Odontopsammodius atopus Cartwright, 1955
Psammodius basalis Mulsant & Rey, 1870Crested blenny
Psammodius besucheti Petrovitz, 1975
Psammodius bidens (Horn, 1871)⇒ Parapsammodius bidens Horn, 1871
Psammodius bilyi Rakovic, 1994
Psammodius boreoitalicus Mencl, Král ..., 2022
Psammodius bulgaricus Mencl, 1982
Psammodius cameneni (Chalumeau, 1976)⇒ Odontopsammodius cameneni Chalumeau, 1976
Psammodius caucasicus Pittino & Sh..., 2006
Psammodius chipiririi (Cartwright, 1955)⇒ Odontopsammodius chipiririi Cartwright, 1955
Psammodius clypeatus Say, 1823
Psammodius convexus Waterhouse, 1875
Psammodius coreanus (Kim, 1980)⇒ Rakovicius coreanus Kim, 1980
Psammodius danubialis Adám, 1989
Psammodius evanidus (Péringuey, 1901)⇒ Leiopsammodius evanidus Péringuey, 1901
Psammodius fimbriatus (Cartwright, 1955)⇒ Odontopsammodius fimbriatus Cartwright, 1955
Psammodius flavolittoralis Kim, 1980
Psammodius formosus (Cartwright, 1955)⇒ Odontopsammodius formosus Cartwright, 1955
Psammodius generosus (Reitter, 1892)⇒ Psammodius nocturnus Reitter, 1892
Psammodius hangangnensis Kim, 1980
Psammodius hydropicus (Horn, 1887)⇒ Geopsammodius hydropicus Horn, 1887
Psammodius inscitus (Walker, 1858)⇒ Rhyssemus inscitus Walker, 1858
Psammodius japonicus (Harold, 1878)⇒ Leiopsammodius japonicus Harold, 1878
Psammodius jordanus Rakovi�, Me..., 2022
Psammodius kobayashii Nomura, 1973
Psammodius kondoi (Masumoto, 1984)⇒ Psammodius kobayashii Nomura, 1973
Psammodius laevicollis (Klug, 1845)⇒ Leiopsammodius laevicollis Klug, 1845
Psammodius laevipennis Costa, 1844Awkward-winged small dung beetle
Psammodius maceki Rakovic, 1994
Psammodius macnamarae Pittino, 1984
Psammodius malkini (Cartwright, 1946)⇒ Leiopsammodius malkini Cartwright, 1946
Psammodius mapirii (Cartwright, 1955)⇒ Odontopsammodius mapirii Cartwright, 1955
Psammodius maruyamai Ochi, Kawaha..., 2011
Psammodius mcclayi (Cartwright, 1955)⇒ Tesarius mcclayi Cartwright, 1955
Psammodius modestus (Péringuey, 1901)⇒ Leiopsammodius modestus Péringuey, 1901
Psammodius nepalensis Balthasar, 1971
Psammodius nocturnus Reitter, 1892
Psammodius oregonensis (Cartwright, 1955)⇒ Tesarius oregonensis Cartwright, 1955
Psammodius ovatus Pittino, 1983
Psammodius pierottii Pittino, 1979Mud blenny
Psammodius plicicollis Erichson, 1848
Psammodius porcicollis (Fairmaire, 1870)⇒ Platytomus laevistriatus Perris, 1869
Psammodius quinqueplicatus (Horn, 1871)⇒ Neopsammodius quinqueplicatus G. Horn, 1871
Psammodius rotundipennis (Reitter, 1892)⇒ Brindalus rotundipennis Reitter, 1892
Psammodius schmidti Gistel, 1857
Psammodius schmidtii Gistel, 1857
Psammodius schwarzi (Linell, 1896)⇒ Parataenius simulator Harold, 1868
Psammodius sculpticollis (Fairmaire, 1897)⇒ Leiopsammodius indicus Harold, 1877
Psammodius sefrensis Petrovitz, 1961
Psammodius seiferti Rakovic, 1994
Psammodius shermani (Cartwright, 1946)⇒ Odontopsammodius cruentus Harold, 1867
Psammodius sulcicollis Illiger, 1802
Psammodius sungshinarum Kim, 1980
Psammodius tesari Rakovic, 1977
Psammodius viti Chalumeau, 1983
Psammoporus abdita Nikritin, 1975
Psammoporus comis Lewis, 1895
Psammoporus criddlei Brown, 1931
Psammoporus cylindricus Eschscholtz, 1822
Psammoporus exaratus Mannerheim, 1853
Psammoporus friebi Balthasar, 1935
Psammoporus hybridus Reitter, 1892
Psammoporus insularis Pittino, 2006
Psammoporus kamtschaticus Motschulsky, 1860
Psammoporus lacustris LeConte, 1850
Psammoporus latipunctus Gredler, 1866
Psammoporus matalini Gusakov, 2003
Psammoporus mimica Pittino, 2006
Psammoporus mimicus Pittino, 2006
Psammoporus nakanei (Masumoto, 1986)⇒ Aegialia kamtschaticus Motschulsky, 1860
Psammoporus nakanei horii Ochi & Kawahara, 2012
Psammoporus nakanei nakanei
Psammoporus nanus Brown, 1931
Psammoporus opacus Brown, 1931
Psammoporus sabuleti (Panzer, 1797)⇒ Psammoporus mimicus Pittino, 2006
Psammoporus sibirica Stebnicka, 1977
Psammoporus terminalis Brown, 1931
Psammoporus tsukamotoi Masumoto, 1986
Psammorpha lawrencei Stebnicka, 1994
Pseudacrossus absconditus Balthasar, 1932
Pseudacrossus beringi Berlov, 1989
Pseudacrossus caspius Menetries, 1832
Pseudacrossus circassicus Reitter, 1892
Pseudacrossus cribricollis (Lucas, 1846)⇒ Amidorus cribricollis Lucas, 1846
Pseudacrossus grebenschikovi Balthasar, 1961
Pseudacrossus grebenscikovi Balthasar, 1961
Pseudacrossus grombczewskyi Koshantschikov, 1891
Pseudacrossus juxtus Petrovitz, 1972
Pseudacrossus kalabi Kral, 1997
Pseudacrossus koshantschikovi Jacobson, 1911
Pseudacrossus kuskai Stebnicka, 1981
Pseudacrossus nasutus Reitter, 1887
Pseudacrossus przewalskyi Reitter, 1887
Pseudacrossus qinghaiensis Kral, 1997
Pseudacrossus serrimargo Koshantschikov, 1913
Pseudacrossus sharpi Harold, 1874
Pseudacrossus smetanai Kral, 2011
Pseudacrossus spalacophilus Novikov, 1996
Pseudacrossus subsericeus Ballion, 1878
Pseudacrossus suffertus Schmidt, 1922
Pseudacrossus suffertus ampliatus Reitter, 1892
Pseudacrossus suffertus defiorei Fiori, 1907
Pseudacrossus suffertus suffertus
Pseudacrossus tenebricosus Schmidt, 1922
Pseudacrossus wewalkai Petrovitz, 1971
Pseudacrossus zangi A.Schmidt, 1906
Pseudacrossus zuercheri Reitter, 1908
Pseudagolius bicolor Say, 1823
Pseudagolius caballeroi Islas, 1945
Pseudagolius coloradensis Horn, 1870
Pseudagolius warneri Skelley & Go..., 2007Lesser weever
Pseudataenius contortus Cartwright, 1974
Pseudataenius gracilitarsis Petrovitz, 1973
Pseudataenius socialis G. Horn, 1871
Pseudataenius waltherhorni Balthasar, 1938
Pseuderytus chobauti Clouët, 1896
Pseudesymus lucidus Klug, 1845
Pseudoahermodontus benesi Dellacasa & ..., 1997
Pseudocoelotrachelus peckorum Dellacasa, D..., 2013
Pseudocoelotrachelus tristaobrancoi Dellacasa, D..., 2013
Pseudodrepanocanthus nasutus Harold, 1869
Pseudodrepanocanthus obscurus Bordat, 1993
Pseudodrepanocanthus spinitarsis Péringuey, 1901
Pseudogonaphodiellus zdzislawae Dellacasa, G..., 2007
Pseudoheptaulacus azumai Ochi, Kon & ..., 2014
Pseudoheptaulacus iniquus Schmidt, 1909
Pseudoheptaulacus nepalensis Pittino, 2004
Pseudoheptaulacus reikoae Ochi, Kon & ..., 2014
Pseudoheptaulacus riekoae Ochi, Kon & ..., 2014
Pseudomothon arabicus Pittino, 1984
Pseudomothon pittinoi Kral, 2017
Pseudopharaphodius abstrusus Bordat, 2018
Pseudopharaphodius admirandus Bordat, 2018
Pseudopharaphodius annoyeri Bordat, 2018
Pseudopharaphodius anthrax Gerstaecker, 1871
Pseudopharaphodius apicesetosus Clement, 1969
Pseudopharaphodius bucolicus Bordat, 2011
Pseudopharaphodius burundiensis Bordat, 2011
Pseudopharaphodius dalabaensis Petrovitz, 1973
Pseudopharaphodius janssensi Paulian, 1939
Pseudopharaphodius kaszabi Endrödi, 1957
Pseudopharaphodius novus Schmidt, 1911
Pseudopharaphodius phalacronotoides Balthasar, 1960
Pseudopharaphodius propinquus Bordat, 2011
Pseudopharaphodius rufomaculatus Petrovitz, 1958
Pseudopharaphodius tai Bordat, 2011
Pseudopharaphodius tumulosus Schmidt, 1911
Pseudopharaphodius urostigmoides Schmidt, 1913
Pseudopharaphodius verecundus Bordat, 2011
Pseudopharaphodius zinqueae Bordat, 2015
Pseudopodotenus fulviventris Fairmaire, 1860
Pseudosaprosites deplanatus Balthasar, 1967
Pseudostereomera mirabilis Bordat & Howden, 1995
Pseudoteuchestes schirparakensis Petrovitz, 1955
Pseudoxyomus armaticeps Péringuey, 1901
Pseudoxyomus cochraneae Bordat, 1996
Pseudoxyomus davisi Bordat, 1996
Pseudoxyomus eximius Péringuey, 1901
Pseudoxyomus geminata Schmidt, 1911
Pseudoxyomus intrusus Péringuey, 1901
Pseudoxyomus maniculus Petrovitz, 1961
Pseudoxyomus oblongopunctata Balthasar, 1941
Pseudoxyomus opaculus Harold, 1869
Pseudoxyomus opatroides Balthasar, 1941
Pseudoxyomus rubescens Petrovitz, 1962
Pseudoxyomus scobina Petrovitz, 1956
Pseudoxyomus setosus Petrovitz, 1964
Pseudoxyomus strydomi Endrödi, 1976
Pterobius itiokai Maruyama, 2010
Pubinus tomentosus O.F.Muller, 1776
Qingaphodius nikodymi Kral, 1997
Qingaphodius ochotonarum Kral, 1997
Rakovicius ainu Lewis, 1895
Rakovicius coreanus Kim, 1980
Rakovicius kawaii (Pittino, 2006)⇒ Rakovicius subopacus Nomura, 1973
Rakovicius kyotoensis Ochi, Kawaha..., 2019
Rakovicius lanae Masumoto, 2012
Rakovicius subopacus Nomura, 1973
Rakovicius takashii Ochi, Kawaha..., 2019
Rakovicius thailandicus Balthasar, 1965
Renaudius tesari Paulian, 1942
Renaudius thailandicus (Masumoto, 1991)⇒ Oxyomus thailandicus Masumoto, 1991
Rhinocerotopsis nakasei Maruyama, 2009
Rhodaphodius foetens Fabricius, 1787
Rhyparus accessoricostatus Mencl, Rakov..., 2013
Rhyparus adebratti Bordat, 1996
Rhyparus amamianus (Nakane, 1956)⇒ Rhyparus helophoroides Fairmaire, 1893
Rhyparus annamariae Minkina, 2017
Rhyparus anneae Stebnicka, 1998
Rhyparus apoensis Ochi & Kakizoe, 2021
Rhyparus apomontanus Ochi & Kakizoe, 2021
Rhyparus approximans Fairmaire, 1893
Rhyparus australiae (Lea, 1923)⇒ Rhyparus helophoroides Fairmaire, 1893
Rhyparus azumai Nakane, 1956
Rhyparus azumai loebli Paulian, 1983
Rhyparus bacanensis Minkina, Ani..., 2022
Rhyparus baracatanensis Ochi & Kakizoe, 2021
Rhyparus barclayi Masumoto, Oc..., 2019
Rhyparus basilanensis Ochi & Kakizoe, 2021
Rhyparus birmanicus Fairmaire, 1897
Rhyparus blantoni Cartwright &..., 1969
Rhyparus breviceps Paulian, 1984
Rhyparus burckhardti Paulian, 1989
Rhyparus buruensis Minkina, Ani..., 2022
Rhyparus chimalapensis Mora-Aguilar..., 2019
Rhyparus chinensis Balthasar, 1952
Rhyparus clavipes Arrow, 1935
Rhyparus comorianus Fairmaire, 1896
Rhyparus costaricensis Cartwright &..., 1969
Rhyparus costatus
Rhyparus danielssoni Bordat, 1996
Rhyparus denieri Martínez, 1950
Rhyparus dentatus Fairmaire, 1896
Rhyparus denticollis Fairmaire, 1893
Rhyparus desjardinsi Westwood, 1845
Rhyparus edieae Stebnicka, 1998
Rhyparus fijiensis Minkina, Ani..., 2022
Rhyparus flavohirtus Anichtchenko..., 2021
Rhyparus gracilis Arrow, 1905
Rhyparus helephoroides (Fairmaire, 1893)⇒ Rhyparus helophoroides Fairmaire, 1893
Rhyparus helophoroides Fairmaire, 1893
Rhyparus henryi Stebnicka, 1998
Rhyparus huaphanensis Ochi, Kon & ..., 2019
Rhyparus ifugaoensis Ochi & Kakizoe, 2021
Rhyparus ironensis Stebnicka & ..., 1996
Rhyparus isidroi Cartwright &..., 1969
Rhyparus jakli Minkina & An..., 2022
Rhyparus kinabalu Paulian, 1989
Rhyparus kitanoi Miyake, 1982
Rhyparus kitanoi kitanoi
Rhyparus kitanoi taiwanus Ochi, 2001
Rhyparus klapperichorum Paulian, 1983
Rhyparus kudrnai Mencl & Rakovic, 2013
Rhyparus laosensis Ochi, Kon & ..., 2018
Rhyparus loebli (Paulian, 1983)⇒ Rhyparus azumai loebli Paulian, 1983
Rhyparus luzonensis Ochi & Kakizoe, 2021
Rhyparus magnus Schmidt, 1911
Rhyparus merkli Minkina & An..., 2022
Rhyparus mexicanus Cartwright &..., 1969
Rhyparus micros Bordat, 1996
Rhyparus mindanaoensis Anichtchenko..., 2021
Rhyparus minor Paulian, 1989
Rhyparus mokaiensis Stebnicka, 1998
Rhyparus multipunctatus Paulian, 1984
Rhyparus myslenickorum Minkina, 2019
Rhyparus nahangensis Minkina & Král, 2022
Rhyparus negrosensis Ochi & Kakizoe, 2021
Rhyparus nepalensis Balthasar, 1971
Rhyparus nilgirensis Arrow, 1909
Rhyparus obsoletus Fairmaire, 1893
Rhyparus occidentalis Paulian, 1990
Rhyparus octovirgatus Schmidt, 1916
Rhyparus opacus Cartwright &..., 1969
Rhyparus orousseti (Paulian, 1981)⇒ Rhyparus helophoroides Fairmaire, 1893
Rhyparus paraloebli Ochi, Kon & ..., 2018
Rhyparus paraphilippinensis Ochi & Kakizoe, 2021
Rhyparus peninsularis Arrow, 1905
Rhyparus philippinensis Arrow, 1905
Rhyparus pseudominor Bordat, 1996
Rhyparus punctithoracicus Ochi, Kon & Bai, 2019
Rhyparus risbeci (Paulian, 1934)⇒ Rhyparus helophoroides Fairmaire, 1893
Rhyparus rugatus Arrow, 1935
Rhyparus saundersi Pascoe, 1866
Rhyparus schachti Balthasar, 1971
Rhyparus schoolmeestersi Ochi, Kon & ..., 2018
Rhyparus sculpturatus Cartwright &..., 1969
Rhyparus semikitanoi Ochi, Kon & ..., 2018
Rhyparus sepikensis Stebnicka, 1998
Rhyparus sinewitensis Stebnicka, 1998
Rhyparus snizeki Mencl & Rakovic, 2013
Rhyparus sogai Paulian, 1959
Rhyparus spangleri Cartwright &..., 1969
Rhyparus spilmani Cartwright &..., 1977
Rhyparus stebnickae Minkina, Ani..., 2022
Rhyparus striatus Arrow, 1935
Rhyparus sumatrensis Fairmaire, 1893
Rhyparus suspiciosis
Rhyparus suspiciosus Cartwright &..., 1969
Rhyparus suturalis Schmidt, 1911
Rhyparus thailandicus Masumoto & Ochi, 2018
Rhyparus verrucosus Schmidt, 1916
Rhyparus vietnamicus Ochi, Kusui ..., 2018
Rhyparus vitikaboureki Mencl & Rakovic, 2013
Rhyparus wushensis Ochi & Masumoto, 2018
Rhyparus xanthi Frivaldszky, 1883
Rhyparus yangi Ochi, Kon & Bai, 2019
Rhyparus zayasi Cartwright &..., 1969
Rhysothorax rufa Fabricius, 1792
Rhysothorax rufus Fabricius, 1792Red sand scarab beetle
Rhyssemodes alutaceus Reitter, 1892
Rhyssemodes bouvieri Clouët, 1901
Rhyssemodes malyi Rakovic, 1982
Rhyssemodes ningxia Rakovi�, Kr..., 2017
Rhyssemodes nox Iablokov-khn..., 1971
Rhyssemodes orientalis Mulsant & Go..., 1874
Rhyssemodes sindicus Pittino, 1984
Rhyssemodes taklamakan Rakovi�, Kr..., 2017
Rhyssemodes tenuisculptus Reitter, 1892
Rhyssemodes transcaspicus (Rakovic, 1982)⇒ Rhyssemodes tenuisculptus Reitter, 1892
Rhyssemodes turcicus Pittino & Ra..., 1984
Rhyssemodes wittmeri Pittino, 1984
Rhyssemorphus clementi Pittino, 1990
Rhyssemorphus mocquerysi Clouët, 1900
Rhyssemorphus nitidus Pittino, 1990
Rhyssemorphus rakovici Pittino, 1990
Rhyssemorphus vadoni Pittino, 1990
Rhyssemus africanus Petrovitz, 1963
Rhyssemus ahrensi Rakovic & Kral, 2001
Rhyssemus algiricus Lucas, 1846
Rhyssemus algiricus marqueti (Reiche, 1863)⇒ Rhyssemus marqueti Reiche, 1863
Rhyssemus algiricus meridionalis Reitter, 1890
Rhyssemus alluaudi Clouët, 1901
Rhyssemus ambovombensis Pittino, 1990
Rhyssemus amplicollis Pittino, 1984
Rhyssemus andreinii Pittino, 1984
Rhyssemus annaedicatus Pierotti, 1980
Rhyssemus archambaulti Benard, 1929
Rhyssemus arenarius (A. Costa, 1844)⇒ Rhyssemus plicatus Germar, 1817
Rhyssemus asper Fabricius, 1775
Rhyssemus aspericollis Petrovitz, 1965
Rhyssemus asperocostatus Fairmaire, 1892
Rhyssemus atramentarius Péringuey, 1901
Rhyssemus aurivillii Clouët, 1901
Rhyssemus aurivilliusi (Clouët, 1901)⇒ Rhyssemus aurivillii Clouët, 1901
Rhyssemus australis (Petrovitz, 1963)⇒ Rhyssemus inscitus Walker, 1858
Rhyssemus bacchusi Pittino, 1984
Rhyssemus balteatus Petrovitz, 1966
Rhyssemus bechuanus Petrovitz, 1956
Rhyssemus bedeli Clouët, 1901
Rhyssemus berytensis Marseul, 1878
Rhyssemus bicolor (Clouët, 1901)⇒ Trichiorhyssemus bicolor Clouët, 1901
Rhyssemus bilyi Rakovic, 2001
Rhyssemus biovatus Clouët, 1901
Rhyssemus birmensis Clouët, 1901
Rhyssemus blackburnei Clouët, 1901
Rhyssemus bordati Pittino, 1984
Rhyssemus brevis Pittino, 1990
Rhyssemus brevitarsis Pittino, 1984
Rhyssemus brownwoodi Gordon & Car..., 1980Fringe blenny
Rhyssemus bucciarellii Pittino, 1990
Rhyssemus buettikeri Pittino, 1984
Rhyssemus bufonis Boucomont, 1935
Rhyssemus caelatus (LeConte, 1881)⇒ Trichiorhyssemus riparius G. Horn, 1871
Rhyssemus californicus Horn, 1871Scratchbush
Rhyssemus canaliculatus Clement, 1969
Rhyssemus cantabricus (Balthasar, 1961)⇒ Rhyssemus germanus Linnaeus, 1767
Rhyssemus capensis Clouët, 1901
Rhyssemus carinatipennis Péringuey, 1901
Rhyssemus carinatus Pittino, 1984
Rhyssemus chinensis Pittino, 1996
Rhyssemus congolanus (Clouët, 1901)⇒ Trichiorhyssemus congolanus Clouët, 1901
Rhyssemus convexus Benard, 1911
Rhyssemus crispus Schmidt, 1922
Rhyssemus decellei Pittino, 1984
Rhyssemus descarpentriesi Pittino, 1990
Rhyssemus dispar Pittino, 1990
Rhyssemus endroedyyoungai Petrovitz, 1975
Rhyssemus evae Endrödi, 1964
Rhyssemus exaratus Marseul, 1878
Rhyssemus fairmairei Clouët, 1901
Rhyssemus falcatus Petrovitz, 1961
Rhyssemus feae Clouët, 1901
Rhyssemus ferenczi Endrödi, 1976
Rhyssemus frankenbergeri Balthasar, 1942
Rhyssemus franzi Petrovitz, 1963
Rhyssemus freudei Balthasar, 1960
Rhyssemus freyi Petrovitz, 1963
Rhyssemus germanus Linnaeus, 1767Germany small dung beetle
Rhyssemus gestroi Clouët, 1901
Rhyssemus ghanaensis Petrovitz, 1975
Rhyssemus goudoti Harold, 1868
Rhyssemus granosus (Klug & Erichson, 1842)⇒ Rhyssemus granulosus Ballion, 1871
Rhyssemus granulosocostatus Clouët, 1901
Rhyssemus granulosus Ballion, 1871
Rhyssemus grossepunctatus Balthasar, 1961
Rhyssemus guineensis Petrovitz, 1969
Rhyssemus haafi Petrovitz, 1964
Rhyssemus haafi occidentalis Endrödi, 1976
Rhyssemus hamatus Petrovitz, 1963
Rhyssemus haroldi Clouët, 1901
Rhyssemus hauseri Balthasar, 1933
Rhyssemus helenae Pittino, 1983
Rhyssemus histrio Balthasar, 1961
Rhyssemus histrioides Petrovitz, 1963
Rhyssemus hottentottus Petrovitz, 1963
Rhyssemus imitator Pittino, 1990
Rhyssemus indicus Clouët, 1901
Rhyssemus inermis Clouët, 1901
Rhyssemus infidus Pittino, 1984
Rhyssemus inscitus Walker, 1858
Rhyssemus insignicollis (Lea, 1923)⇒ Rhyssemus inscitus Walker, 1858
Rhyssemus insularis Pittino, 1983
Rhyssemus interruptus Reitter, 1892
Rhyssemus karnatakaensis Pittino, 1984
Rhyssemus keisseri Benard, 1910
Rhyssemus koreanus Stebnicka, 1980
Rhyssemus laesifrons (Fairmaire, 1892)⇒ Pararhyssemus coluber Mayet, 1887
Rhyssemus laevinasus Petrovitz, 1964
Rhyssemus lamyensis Petrovitz, 1963
Rhyssemus laoticus Pittino, Rak..., 2013
Rhyssemus limbolarius Petrovitz, 1963
Rhyssemus limborarius (Petrovitz, 1963)⇒ Rhyssemus limbolarius Petrovitz, 1963
Rhyssemus linnavuorii Balthasar, 1972
Rhyssemus loebli Petrovitz, 1975
Rhyssemus lunatus (Petrovitz, 1963)⇒ Rhyssemus feae Clouët, 1901
Rhyssemus macedonicus Benard, 1923
Rhyssemus madagassus Harold, 1879
Rhyssemus malasiacus (Lansberg, 1886)⇒ Rhyssemus inscitus Walker, 1858
Rhyssemus marqueti Reiche, 1863
Rhyssemus maximus Clouët, 1901
Rhyssemus mayeti Clouët, 1901
Rhyssemus meridionalis (Reitter, 1890)⇒ Rhyssemus algiricus meridionalis Reitter, 1890
Rhyssemus meruensis Pittino, 1983
Rhyssemus mesopotamicus Petrovitz, 1963
Rhyssemus mexicanus Hinton, 1934
Rhyssemus mimus Balthasar, 1961
Rhyssemus mirus Petrovitz, 1967
Rhyssemus mongol (Endrödi, 1969)⇒ Granulopsammodius mongol Endrödi, 1969
Rhyssemus morgani (Benard, 1911)⇒ Rhyssemus berytensis Marseul, 1878
Rhyssemus murghabensis Balthasar, 1967
Rhyssemus mussardi (Petrovitz, 1975)⇒ Rhyssemus inscitus Walker, 1858
Rhyssemus namorokae Pittino, 1990
Rhyssemus nanshanchicus Masumoto, 1977
Rhyssemus neglectus Brown, 1929
Rhyssemus nitidus Petrovitz, 1972
Rhyssemus obliviosus Petrovitz, 1963
Rhyssemus olympiae Pittino, 1990
Rhyssemus osmanlis Koshantschikov, 1916
Rhyssemus papuanus (Petrovitz, 1965)⇒ Rhyssemus inscitus Walker, 1858
Rhyssemus parallelicollis Clouët, 1901
Rhyssemus parallelus Reitter, 1892
Rhyssemus pauliani Pittino, 1990
Rhyssemus pectoralis Clouët, 1901
Rhyssemus perissinottoi Pittino, 1983
Rhyssemus perlatus Petrovitz, 1975
Rhyssemus pertinax (Balthasar, 1961)⇒ Rhyssemus germanus Linnaeus, 1767
Rhyssemus peyrierasi Pittino, 1990
Rhyssemus pfefferi Balthasar, 1972
Rhyssemus philippineus (Masumoto, 1980)⇒ Rhyssemus inscitus Walker, 1858
Rhyssemus plicatus Germar, 1817
Rhyssemus polycolpus Fairmaire, 1886
Rhyssemus pondoensis Petrovitz, 1967
Rhyssemus ponticus Petrovitz, 1962
Rhyssemus procerus Petrovitz, 1973
Rhyssemus promontorii Péringuey, 1901
Rhyssemus propinquus Petrovitz, 1964
Rhyssemus psammobiiformis Petrovitz, 1963
Rhyssemus punctatissimus Pittino, 1984
Rhyssemus puncticollis (Brown, 1929)⇒ Rhyssemus germanus Linnaeus, 1767
Rhyssemus punctiventris Balthasar, 1961
Rhyssemus purkynei Balthasar, 1961
Rhyssemus rajasthani Rakovi�, Me..., 2017
Rhyssemus reitteri (Clouët, 1901)⇒ Rhyssemus sculptilipennis Schmidt, 1922
Rhyssemus relegatus Balthasar, 1965
Rhyssemus ressli Petrovitz, 1965
Rhyssemus riparius (Horn, 1871)⇒ Trichiorhyssemus riparius G. Horn, 1871
Rhyssemus ritsemae Clouët, 1901
Rhyssemus rohani Benard, 1920
Rhyssemus rotschildi Benard, 1909
Rhyssemus rubeolus Harold, 1871
Rhyssemus saldaitisi Rakovi�, Kr..., 2016
Rhyssemus samurai (Balthasar, 1941)⇒ Myrhessus samurai Balthasar, 1941
Rhyssemus saoudi Pittino, 1984
Rhyssemus sardous Pierotti, 1980
Rhyssemus scaber Haldeman, 1848
Rhyssemus scabrosus Pittino, 1990
Rhyssemus schaeuffelei Petrovitz, 1964
Rhyssemus sculptilipennis Schmidt, 1922
Rhyssemus sculptilipennis falsus Balthasar, 1965
Rhyssemus sculptilipennis sculptilipennis
Rhyssemus sculptipennis (Schmidt, 1916)⇒ Rhyssemus sculptilipennis Schmidt, 1922
Rhyssemus seineri Petrovitz, 1969
Rhyssemus senegalensis Petrovitz, 1972
Rhyssemus severini (Clouët, 1901)⇒ Rhyssemus carinatipennis Péringuey, 1901
Rhyssemus sexcostatus Schmidt, 1909
Rhyssemus similis Petrovitz, 1963
Rhyssemus sinuaticollis Pittino, 1984
Rhyssemus sonatus LeConte & Horn, 1881
Rhyssemus spangleri Gordon & Car..., 1980
Rhyssemus spiniger Pittino, 1983
Rhyssemus subdolus Balthasar, 1961
Rhyssemus sulcatus Olivier, 1789
Rhyssemus syriacus Petrovitz, 1971
Rhyssemus tarsalis (Waterhouse, 1876)⇒ Rhyssemus inscitus Walker, 1858
Rhyssemus testudo Pittino, 1990
Rhyssemus thailandicus Pittino, 1996
Rhyssemus thomasi Endrödi, 1964
Rhyssemus tonkineus (Balthasar, 1945)⇒ Rhyssemus feae Clouët, 1901
Rhyssemus transcaucasicus Minkina & Král, 2018
Rhyssemus tristis Petrovitz, 1969
Rhyssemus trisulcatus Petrovitz, 1956
Rhyssemus tschadensis Petrovitz, 1963
Rhyssemus tsihombensis Pittino, 1990
Rhyssemus tuberculatus (Nomura & Nakane, 1951)⇒ Myrhessus samurai Balthasar, 1941
Rhyssemus tuberculicollis Petrovitz, 1965
Rhyssemus uncispinis Pittino, 1984
Rhyssemus vaulogeri Clouët, 1901
Rhyssemus verrucosus Mulsant, 1842
Rhyssemus villosus Pittino, 1990
Rhyssemus vinodolensis Petrovitz, 1963
Rhyssemus waboniensis Petrovitz, 1964
Rhyssemus xerxes Pittino, 1983
Rhyssemus zumpti Petrovitz, 1956
Rugaphodius rugatus Schmidt, 1907Spotted weever
Sahlbergianus longissimus Sahlberg, 1908
Saprolochus bolivarensis Stebnicka & ..., 2007
Saprolochus lobatus Skelley, 2007
Saprolochus tambopatae Stebnicka & ..., 2007
Saprolochus tridentatus Skelley, 2007
Saprositellus ariquemes Stebnicka, 2003
Saprositellus denticulatus Balthasar, 1967
Saprositellus kenodontus Skelley, 2007
Saprositellus peruanus Stebnicka, 2003
Saprositellus santaritae Stebnicka, 2003
Saprosites abyssinicus Bordat, 1990
Saprosites africanus Endrödi, 1967
Saprosites arabicus Pittino, 1984
Saprosites armatus Pittino, 2008
Saprosites balthasari Endrödi, 1951
Saprosites bangueyensis Paulian, 1944
Saprosites barombii Petrovitz, 1961
Saprosites basilewskyi (Endrödi, 1955)⇒ Ataenius garamas Peyerimhoff, 1929
Saprosites blackwelderi (Chapin, 1940)⇒ Iguazua blackwelderi Chapin, 1940
Saprosites bolivianus Petrovitz, 1961
Saprosites brevitarsis Endrödi, 1964
Saprosites breviusculus Harold, 1867
Saprosites bunyaensis Stebnicka & ..., 1996
Saprosites burgeoni Paulian, 1939
Saprosites calvus Schmidt, 1911
Saprosites camerounensis Paulian, 1940
Saprosites capitalis Fairmaire, 1883
Saprosites carinatus Bordat, 1992
Saprosites cariniceps Pittino, 2008
Saprosites cascus E. B. Britton, 1960
Saprosites castaneus Lea, 1923
Saprosites catenatus Fauvel, 1903
Saprosites cavus Schmidt, 1908
Saprosites celebicus Paulian, 1944
Saprosites cheesmani Paulian, 1939
Saprosites chyuluensis Paulian, 1939
Saprosites clydensis Stebnicka & ..., 1996
Saprosites communis Broun, 1880
Saprosites comorianus Bordat, 1990
Saprosites congoana (Petrovitz, 1956)⇒ Saprosites schoutedeni Paulian, 1939
Saprosites consonus Schmidt, 1911
Saprosites crockerensis Stebnicka, 1991
Saprosites cycloporum Paulian, 1944
Saprosites declivis Schmidt, 1911
Saprosites degallieri Bordat & ThÃ..., 2012
Saprosites deharvengi Bordat & ThÃ..., 2012
Saprosites dentipes Harold, 1867
Saprosites difficilis Harold, 1877
Saprosites dilutus Fairmaire, 1849
Saprosites distans Sharp, 1876
Saprosites dubius Bordat, 1990
Saprosites dudichi Endrödi, 1951
Saprosites dufaui Paulian, 1947
Saprosites dynastoides Walker, 1858
Saprosites enarotadii Stebnicka, 1998
Saprosites exaratus Fleutiaux & ..., 1889
Saprosites excisus Petrovitz, 1976
Saprosites explanatus Bordat, 1990
Saprosites exsculptus White, 1846
Saprosites falcatus Schmidt, 1908
Saprosites fastus Petrovitz, 1965
Saprosites fodori Endrödi, 1951
Saprosites formosensis (Nomura, 1939)⇒ Saprosites japonicus Waterhouse, 1875
Saprosites fortipes Broun, 1886
Saprosites freyi Petrovitz, 1956
Saprosites gestroi Schmidt, 1911
Saprosites girardi Bordat, 2003
Saprosites gnomus Balthasar, 1941
Saprosites grenadensis Arrow, 1903
Saprosites guineensis Petrovitz, 1956
Saprosites haafi Petrovitz, 1956
Saprosites holzschuhi Pittino, 2008
Saprosites implicatus Schmidt, 1922
Saprosites japonicus Waterhouse, 1875
Saprosites javanus Balthasar, 1941
Saprosites kaimai Stebnicka, 2005
Saprosites kamai Stebnicka, 2005
Saprosites kapitensis Petrovitz, 1973
Saprosites kinabalu Pittino, 2008
Saprosites kingsensis Stebnicka, 2001
Saprosites komumi Stebnicka, 1998
Saprosites laeviceps Harold, 1877
Saprosites laticeps Fairmaire, 1871
Saprosites laticeps laticeps
Saprosites laticeps seychellensis Bordat, 1990
Saprosites laticollis Pittino, 2008
Saprosites lepersonnei Paulian, 1942
Saprosites lodoicea Scott, 1912
Saprosites loebli Balthasar, 1972
Saprosites longethorax Paulian, 1944
Saprosites madagascariensis Schmidt, 1911
Saprosites mahavus Stebnicka, 2012
Saprosites malkini Paulian, 1958
Saprosites mansuetus Blackburn, 1904
Saprosites marchionalis (Harold, 1877)⇒ Saprosites cycloporum Paulian, 1944
Saprosites matangi Stebnicka, 2012
Saprosites meditans Harold, 1867
Saprosites mendax Blackburn, 1892
Saprosites merkli Pittino, 2008
Saprosites mesosternalis Lea, 1923
Saprosites mistakensis Stebnicka & ..., 1996
Saprosites mjoebergi Gillet, 1924
Saprosites monteverdeae Stebnicka, 2001
Saprosites narae Lewis, 1895
Saprosites natalensis Péringuey, 1901
Saprosites nepalensis Pittino, 2008
Saprosites nitidicollis (MacLeay, 1871)⇒ Saprosites sternalis Blackburn, 1904
Saprosites nocturnus (Nomura, 1943)⇒ Ataenius gracilis Melsheimer, 1845
Saprosites obscurus Endrödi, 1964
Saprosites pahangensis Stebnicka, 2012
Saprosites palmarum Scott, 1912
Saprosites papuanus Stebnicka, 1984
Saprosites parallelicollis Bordat, 1990
Saprosites parallelus (Harold, 1867)⇒ Saprosites mendax Blackburn, 1892
Saprosites pauliani Bordat, 1990
Saprosites penrisseni Stebnicka, 2012
Saprosites perbrevitarsis Schmidt, 1909
Saprosites peregrinus Redtenbacher, 1858
Saprosites perforatus Bordat, 1990
Saprosites perssoni Bordat, 1996
Saprosites pleurophoroides Balthasar, 1971
Saprosites porongurupae Stebnicka & ..., 1996
Saprosites puncticollis Harold, 1867
Saprosites punctiventris Petrovitz, 1961
Saprosites pygmaeus Harold, 1877Birthworts
Saprosites raffrayi Paulian, 1944
Saprosites raoulensis Broun, 1910
Saprosites rectus Bordat, 1990
Saprosites rougemonti Pittino, 2008
Saprosites rufopolitus Gillet, 1924
Saprosites rufus Paulian, 1944
Saprosites schoutedeni Paulian, 1939
Saprosites sicardi Bordat, 1990
Saprosites staudingeri Petrovitz, 1969
Saprosites sternalis Blackburn, 1904
Saprosites subterraneus Petrovitz, 1976
Saprosites succini R. Zang, 1905
Saprosites sulcatissimus Broun, 1911
Saprosites sulcatus Harold, 1869
Saprosites sulciceps Pittino, 2008
Saprosites sulcicollis Petrovitz, 1969
Saprosites tantulus Bordat, 1990
Saprosites tenuistriatus Bordat, 1990
Saprosites titschacki Balthasar, 1941
Saprosites topali Pittino, 2008
Saprosites verecundus Schmidt, 1909
Saprosites vosseleri Petrovitz, 1965
Saprosites watti Stebnicka, 2001
Saprosites wauensis Stebnicka, 1984
Saprosites weisei Petrovitz, 1969
Saprosites wirthi (Cartwright &..., 1977)⇒ Saprosites exaratus Fleutiaux & ..., 1889
Saprosites wittei Paulian, 1954
Saprosites yanoi Nomura, 1939
Saprosites yasutakai Ochi & Kawahara, 2019
Saprovisca leytensis Stebnicka, 1993
Saprovisca sarangay Skelley, 2015
Saprus griffithi Blackburn, 1904
Saprus lawrencei Stebnicka & ..., 1995
Saprus victoriae Stebnicka & ..., 1995
Saprus weiri Stebnicka & ..., 1995
Sariangus sariangi Rakovic & Mencl, 2011
Scabrostomus baileyi Skelley & Go..., 2001Warteye stargazer
Scabrostomus bakeri Skelley & Go..., 2001Warteye stargazer
Scabrostomus dyspistus Skelley & Wo..., 1991Masked stargazer
Scabrostomus kerni Gordon & Ske..., 2007Lesser weever
Scabrostomus parapeculiosus Gordon & Ske..., 2007Lesser weever
Scabrostomus peculiosus Schmidt, 1922Common sand stargazer
Scabrostomus sepultus Cartwright, 1944Monkfish
Scabrostomus siccatus Gordon & Ske..., 2007Lesser weever
Scabrostomus skillmani Warner & Ske..., 2006
Scabrostomus tanytarsus Skelley & Wo..., 1991Masked stargazer
Schaefferellus arizonensis Schaeffer, 1907
Selviria anneae Stebnicka, 2005
Selviria matogrossoensis Stebnicka, 1999
Setodius bryanti Brown, 1929
Setodius diogenes Gordon & Ske..., 2007Lesser weever
Setodius edmontonus Brown, 1929
Setylaides assamensis Maté, 2016
Setylaides canescens Balthasar, 1933
Setylaides chiangmai Stebnicka, 2008
Setylaides foveatus Schmidt, 1909
Setylaides monstrosus Harold, 1877
Setylaides nakanoi Ochi & Kawahara, 2019
Setylaides punctatus Schmidt, 1911
Setylaides tsujii Kakizoe, Kaw..., 2021
Siamaphodius naafon Masumoto, 1991
Sicardia boiteli Normand, 1938
Sicardia psammodiformis Reitter, 1896
Sigorus porcus Fabricius, 1792Mahogany Ridgeback scarab
Sigorus porcus ruficrus Marsham, 1802
Silluvia elongata Landin, 1949
Silluvia gansuensis Frolov, 2008
Silluvia gogona Stebnicka, 1977
Silluvia gosainkundae Stebnicka, 1986
Silluvia himalayanus Petrovitz, 1968
Silluvia igori Frolov, 2008
Silluvia kabaki Frolov, 2002
Silluvia petrovitzi Stebnicka, 1977
Silluvia shashi Stebnicka, 1977
Silluvia simbuae Stebnicka, 1990
Silluvia sinica Stebnicka, 1977
Silluvia wassuensis Petrovitz, 1963
Silluvia wittmeri Stebnicka, 1980
Silluvia yunnanica Frolov, 2008
Simogonius beccarii Harold, 1871
Simogonius janssensi Paulian, 1954
Simogonius minusculus Schmidt, 1920
Simogonius morettoi Bordat, 2017
Simogonius setosus Petrovitz, 1969
Sinaphodius chitwanensis Schoolmeeste..., 1999
Sinaphodius natalicus Mencl, Rakov..., 2013
Sinaphodius oharai Ochi & Kon, 2008
Sinaphodius philippinensis Cervenka, 1994
Sinaphodius sinuatus Harold, 1860
Sinaphodius sulawesiensis Ochi & Kon, 2004
Sinaphodius yunnanus Cervenka, 1994
Sinodiapterna gorodinskiyi Gusakov, 2006
Sinodiapterna hammondi Dellacasa, 1986
Sinodiapterna songrini Stebnicka & ..., 1992
Sinodiapterna troitzkyi Jacobson, 1897
Sinodiapterna walamiensis Masumoto, Wa..., 2017
Sinodiapterna yushana Li & Wang, 2017
Sinodiapterna zeni Ochi, 1991
Sitiphus ammodites Peyerimhoff, 1939
Sitiphus brevitarsis Reitter, 1894
Sitiphus sefrensis Petrovitz, 1958
Skelleyanus eremita Dellacasa, D..., 2011
Stebnickiella zosterixys Skelley, 2007
Stenotothorax badipes Melsheimer, 1845
Stenotothorax cadaverinus Mannerheim, 1843
Stenotothorax comosus Gordon & Ske..., 2007Lesser weever
Stenotothorax cribratulus Schmidt, 1916Common sand stargazer
Stenotothorax dilaticollis Saylor, 1935
Stenotothorax gardneri Gordon, 2006Greater weever
Stenotothorax gentilis Horn, 1887
Stenotothorax gravis Fall, 1927
Stenotothorax hibernalis Nakane & Tsu..., 1956
Stenotothorax hibernalis hibernalis
Stenotothorax hibernalis saghalinensis Nakane & Tsu..., 1956
Stenotothorax lahontanensis Skelley & Mc..., 2018
Stenotothorax lanei Saylor, 1940
Stenotothorax linsleyi Van Dyke, 1933
Stenotothorax martini Van Dyke, 1928
Stenotothorax mcpeaki Gordon & Ske..., 2007Lesser weever
Stenotothorax micellus Gordon & Ske..., 2007Lesser weever
Stenotothorax nevadensis Horn, 1870
Stenotothorax niviviator Skelley & Mc..., 2018
Stenotothorax odontomonteus Skelley & Mc..., 2018
Stenotothorax oriens Gordon & Ske..., 2007Lesser weever
Stenotothorax oviformis Robinson, 1946Goby duckbill
Stenotothorax ovipennis Horn, 1870
Stenotothorax parapyriformis Gordon & Ske..., 2007Lesser weever
Stenotothorax pyriformis Brown, 1929
Stenotothorax rugoclypeus Hinton, 1934Sand stargazers
Stenotothorax schneppi Skelley & Mc..., 2018
Stenotothorax smilodon Skelley & Mc..., 2018
Stenotothorax sparsus LeConte, 1878
Stenotothorax washtucna Robinson, 1938Black swallowers
Stenotothorax winnemucca Skelley & Mc..., 2018
Stenotothorax wintoni Skelley & Mc..., 2018
Stenotothorax woodleyi Gordon, 2006
Stereomera masumotoi Maruyama & N..., 2011
Stereomera pusilla Arrow, 1905
Strigodius robinsoni Cartwright, 1939
Subrinus clermonti Reitter, 1907
Subrinus scintillius Nikritin & K..., 1979
Subrinus sturmi Harold, 1870
Subrinus subtilis Koshantschikov, 1894
Subrinus vitellinus Klug, 1845
Subrinus yemeniticus Petrovitz, 1970
Sugrames auriculatus Reitter, 1892
Sugrames capensis Endrödi, 1983
Sugrames hauseri Reitter, 1894
Sugrames johni Kiseritzky, 1928
Sugrames reichardti Kiseritzky, 1928
Sussorca lusingae Paulian, 1954
Sybacodes borneensis Minkina, Men..., 2018
Sybacodes liliae Dellacasa, 1978
Sybacodes lutulentus Fairmaire, 1896
Sybacodes lutulentus alternatus Fairmaire, 1897
Sybacodes lutulentus lutulentus
Sybacodes simplicicollis Fairmaire, 1896
Sybacodes simplicicollis aureopilosus Schmidt, 1909
Sybacodes simplicicollis simplicicollis
Sybax distortus Schmidt, 1920
Sybax impressicollis Boheman, 1857
Sybax sulcicollis Boheman, 1857
Symphodon anomalum Harold, 1874
Symphodon anomalus Harold, 1874
Tanyana aniae Minkina, 2022
Tanyana guyanaensis Stebnicka, 2003
Tanyana guynaensis Stebnicka, 2003
Termitaxis holmgreni Krikken, 1970
Termitoderus genieri Bordat & Mor..., 2010
Termitoderus grassei Mateu, 1966
Termitoderus grassei grassei
Termitoderus grassei senegalensis Mateu, 1966
Termitoderus kistneri Howden & Gill, 1993
Termitodiellus besucheti Paulian, 1983
Termitodiellus esakii Nomura, 1943
Termitodiellus hainanensis Jiang & Wang, 2020
Termitodiellus hammondi Krikken & Hu..., 1987
Termitodiellus interruptus Krikken & Hu..., 1987
Termitodiellus luzonensis Howden, 1995
Termitodiellus mindanaoensis Minkina & Ka..., 2020
Termitodiellus minkinai Anichtchenko..., 2021
Termitodiellus monticola Krikken & Hu..., 1987
Termitodius araujoi Reyes-Castil..., 1979
Termitodius chaki Reyes-Castil..., 1979
Termitodius coronatus Wasmann, 1894
Termitodius denieri (Martínez, 1950)⇒ Rhyparus denieri Martínez, 1950
Termitodius peregrinus (Hinton, 1934)⇒ Aschnarhyparus peregrinus Hinton, 1934
Termitodius woodruffi Skelley, Cla..., 2022
Termitopisthes forticulus Tangelder & ..., 1982
Termitopisthes kistneri Tangelder & ..., 1982
Termitopisthes parallelus Champion & W..., 1923
Termitopisthes schmidti Champion & W..., 1923
Termitopisthes submissus Tangelder & ..., 1982
Termitopisthes termiticola Gestro, 1891
Termitopisthes wasmanni A.Schmidt, 1911
Termitotrox ancoroides Petrovitz, 1956
Termitotrox consobrinus Reichensperger, 1915
Termitotrox cupido Maruyama, 2012
Termitotrox icarus Kakizoe, Lia..., 2020
Termitotrox kenyensis Paulian, 1985
Termitotrox maynei Reichensperger, 1956
Termitotrox minutus Arrow, 1920
Termitotrox monodi Paulian, 1947
Termitotrox permirus Wasmann, 1918
Termitotrox turkanicus Krikken, 2008
Termitotrox turkestanicus Krikken, 2008
Termitotrox usambaricus Krikken, 2008
Termitotrox usumbaricus Krikken, 2008
Termitotrox vanbruggeni Krikken, 2008
Termitotrox venus Kakizoe & Ma..., 2015
Tesarius caelata LeConte, 1857
Tesarius caelatus LeConte, 1857
Tesarius doyeni Cartwright, 1977
Tesarius mcclayi Cartwright, 1955
Tesarius oregonensis Cartwright, 1955
Tesarius suclipennis Lea, 1904
Tesarius sulcipennis Lea, 1904
Tetraclipeoides acutissimus Gordon, 1976Black swallower
Tetraclipeoides aemulus Horn, 1887
Tetraclipeoides anomaliceps Brown, 1929
Tetraclipeoides calamitosus Balthasar, 1935
Tetraclipeoides denticulatus Haldeman, 1848
Tetraclipeoides dentiger LeConte, 1858Berg-salmon gill worm
Tetraclipeoides dentigerulus Brown, 1929
Tetraclipeoides fimbripes Brown, 1928
Tetraclipeoides formidatus Gordon, 1976Black swallower
Tetraclipeoides giulianii Gordon, 1977Dartfish
Tetraclipeoides henryi Gordon, 1976Black swallower
Tetraclipeoides moquinus Fall & Cocke..., 1907
Tetraclipeoides quadridentatus Harold, 1861
Tetraclipeoides quadridentulus Fairmaire, 1894
Tetraclipeoides testaceiventris Fall, 1932
Tetraclipeoides texanus Gordon & Ske..., 2007Lesser weever
Teuchestes analis Fabricius, 1787
Teuchestes brachysomus Solsky, 1874
Teuchestes caffer Wiedemann, 1823
Teuchestes caffer caffer
Teuchestes caffer scioensis Dellacasa & ..., 1983
Teuchestes dejeani Harold, 1862
Teuchestes fossor Linnaeus, 1758Digger small dung beetle
Teuchestes guangdong Rakovic & Mencl, 2012
Teuchestes haemorrhoidalis
Teuchestes hongson Rakovic & Mencl, 2012
Teuchestes sinofraternus Dellacasa & ..., 1983
Teuchestes uenoi Ochi, Kawaha..., 2006
Teuchestes wicheri Dellacasa & ..., 1983
Thinorycter balthasari Medvedev, 1969
Thinorycter chlamydatus Semenov & Re..., 1925
Thinorycter chorasmius Semenov & Me..., 1929
Thinorycter mamajevi Medvedev, 1969
Thinorycter medvedevi Nikolajev, 1975
Trichaphodiellus brasiliensis Castelnau, 1840
Trichaphodioides bellus Endrödi, 1961
Trichaphodioides calcaratus Boheman, 1857
Trichaphodioides cinerascens Klug, 1855
Trichaphodioides detruncatus Schmidt, 1908
Trichaphodioides dimorphus Petrovitz, 1966
Trichaphodioides incertepilosus Bordat, 1994
Trichaphodioides lackneri Bordat, 2015
Trichaphodioides manifestus Schmidt, 1911
Trichaphodioides marchi Bordat, 1994
Trichaphodioides multus Bordat, 1994
Trichaphodioides niloticus Petrovitz, 1967
Trichaphodioides pavo Balthasar, 1939
Trichaphodioides pulchellus Müller, 1941
Trichaphodioides schaumi Harold, 1859
Trichaphodioides securifer Petrovitz, 1958
Trichaphodioides seminitidus Quedenfeldt, 1884
Trichaphodioides simoni Petrovitz, 1959
Trichaphodioides simplicitarsis Müller, 1941
Trichaphodioides zborniki Balthasar, 1935
Trichaphodioides zicsii Endrödi, 1967
Trichaphodius amplitarsis Bordat, 1989
Trichaphodius bellonatus Balthasar, 1941
Trichaphodius calcaratoides Paulian, 1942
Trichaphodius cantaloubei Bordat, 1992
Trichaphodius cantoti Bordat, 1989
Trichaphodius cantoti cantoti
Trichaphodius cantoti mycetophilus Bordat, 1989
Trichaphodius copulatus Schmidt, 1916
Trichaphodius cubitus Petrovitz, 1966
Trichaphodius curvispina Bordat, 1989
Trichaphodius divisus Schmidt, 1908
Trichaphodius dudleyi Bordat, 1985
Trichaphodius flavus Endrödi, 1955
Trichaphodius fumosulus Endrödi, 1957
Trichaphodius globosus Bordat, 2013
Trichaphodius gorillae Bordat, 1989
Trichaphodius hindustanicus Balthasar, 1935
Trichaphodius hofferi Balthasar, 1943
Trichaphodius humilis Roth, 1851
Trichaphodius incisus Petrovitz, 1969
Trichaphodius jirianus (Balthasar, 1965)⇒ Trichaphodius hindustanicus Balthasar, 1935
Trichaphodius kachovskyi Koshantschikov, 1916
Trichaphodius kanemicus Endrödi, 1976
Trichaphodius lanuginosus Péringuey, 1901
Trichaphodius leoninus Schmidt, 1911
Trichaphodius longus Schmidt, 1912
Trichaphodius maldesi Bordat, 1989
Trichaphodius meruanus Balthasar, 1967
Trichaphodius misellus Boheman, 1857
Trichaphodius moorei Paulian, 1936
Trichaphodius mysticus Bordat, 1989
Trichaphodius niloticus Petrovitz, 1967
Trichaphodius obvius Endrödi, 1964
Trichaphodius okeani Bordat, 1992
Trichaphodius opisthius (Bates, 1887)⇒ Gonaphodiellus opisthius Bates, 1887
Trichaphodius paradivisus Balthasar, 1960
Trichaphodius pseudocopulatus Bordat, 1997
Trichaphodius pseudohumilis Bordat, 1989
Trichaphodius pseudoleoninus Bordat, 1997
Trichaphodius punctiger Endrödi, 1957
Trichaphodius quadripartitus Petrovitz, 1973
Trichaphodius schoutedeni Boucomont, 1928
Trichaphodius serrulatus Quedenfeldt, 1884
Trichaphodius seydeli Paulian, 1939
Trichaphodius sigwalti Bordat, 1989
Trichaphodius sipeki Rakovi�, Kr..., 2016
Trichaphodius viduus Petrovitz, 1966
Trichiopsammobius brasiliensis Petrovitz, 1963
Trichiopsammobius drumonti Rakovic & Mencl, 2013
Trichiorhyssemus adhabharicus Pittino, 1983
Trichiorhyssemus alternatus Hinton, 1938Maori chief
Trichiorhyssemus asperulus Waterhouse, 1875
Trichiorhyssemus balthasari (Rakovic, 1987)⇒ Neotrichiorhyssemus balthasari Rakovic, 1987
Trichiorhyssemus bicolor Clouët, 1901
Trichiorhyssemus bisigillatus Benard, 1924
Trichiorhyssemus boninensis (Nakane, 1960)⇒ Neotrichiorhyssemus esakii Nomura, 1943
Trichiorhyssemus cloueti Koshantschikov, 1916
Trichiorhyssemus congolanus Clouët, 1901
Trichiorhyssemus congolanus congolanus
Trichiorhyssemus congolanus mojwaeensis Pittino, 1984
Trichiorhyssemus congolanus mojwaensis Pittino, 1984
Trichiorhyssemus congolanus perfidus Pittino, 1984
Trichiorhyssemus cristatellus Bates, 1887
Trichiorhyssemus dalmatinus Petrovitz, 1967
Trichiorhyssemus decorsei Benard, 1914
Trichiorhyssemus elegans Petrovitz, 1963
Trichiorhyssemus expansicollis (Benard, 1930)⇒ Neotrichiorhyssemus expansicollis Benard, 1930
Trichiorhyssemus fokiensis Petrovitz, 1968
Trichiorhyssemus fruhstorferi Petrovitz, 1968
Trichiorhyssemus hatakei Ochi, Kawaha..., 2019
Trichiorhyssemus hauseri (Balthasar, 1933)⇒ Neotrichiorhyssemus hauseri Balthasar, 1933
Trichiorhyssemus hirsutus (Clouët, 1901)⇒ Neotrichiorhyssemus hirsutus Clouët, 1901
Trichiorhyssemus hispidus Pittino, 2007
Trichiorhyssemus immaturus Minkina, 2022
Trichiorhyssemus kitayamai Ochi & Kawah..., 2001
Trichiorhyssemus lasionotus Clouët, 1901
Trichiorhyssemus longetarsalis Benard, 1921
Trichiorhyssemus malabaricus (Balthasar, 1963)⇒ Neotrichiorhyssemus malabaricus Balthasar, 1963
Trichiorhyssemus malkini (Rakovic, 1987)⇒ Neotrichiorhyssemus malkini Rakovic, 1987
Trichiorhyssemus microtrichius Ochi, Kawaha..., 2011
Trichiorhyssemus nakoshii Ochi, Kawaha..., 2021
Trichiorhyssemus nepalensis Pittino, 1983
Trichiorhyssemus nishinoi Ochi, Kawaha..., 2020
Trichiorhyssemus occidentalis Endrödi, 1976
Trichiorhyssemus pseudoinscitus Pittino, 1984
Trichiorhyssemus riparius G. Horn, 1871
Trichiorhyssemus setulosus Reitter, 1892
Trichiorhyssemus taiwanus Ochi, Masumo..., 2015
Trichiorhyssemus yumikoae Pittino & Kawai, 2007
Trichonotuloides alfonsinae Dellacasa, D..., 2014
Trichonotuloides aphoderrans Skelley & Wa..., 2015
Trichonotuloides glyptus Bates, 1887
Trichonotuloides hansferyi Dellacasa, D..., 2014
Trichonotuloides latecrenatus Bates, 1887
Trichonotulus amdoensis Král, Rakovi..., 2015
Trichonotulus baloghi Endrödi, 1967
Trichonotulus brevisetosus Petrovitz, 1973
Trichonotulus castetsi Paulian, 1936
Trichonotulus cechovskyi Minkina & Mencl, 2018
Trichonotulus dakarensis Balthasar, 1960
Trichonotulus dzamosanicus Stebnicka, 1973
Trichonotulus erinaceus Balthasar, 1935
Trichonotulus frankkochi Bordat, 1995
Trichonotulus fulvescens Harold, 1874
Trichonotulus granulidorsis Bordat, 1990
Trichonotulus horaki Balthasar, 1971
Trichonotulus kachelekoensis Bordat, 2016
Trichonotulus khonensis Stebnicka, 1981
Trichonotulus mirei Bordat, 1984
Trichonotulus mubalensis Paulian, 1954
Trichonotulus osetinus Medvedev & D..., 1977
Trichonotulus pilosellus Schmidt, 1907
Trichonotulus scheuermanni Endrödi, 1973
Trichonotulus scrofa Fabricius, 1787Sow small dung beetle
Trichonotulus secquorum Bordat, 1990
Trichonotulus setosus Petrovitz, 1965
Trichonotulus shantungensis Balthasar, 1941
Trichonotulus stuckenbergi Petrovitz, 1967
Trichonotulus uganoensis Petrovitz, 1969
Trichonotulus upembanus Paulian, 1954
Trichonotulus verrucosus Bordat, 1996
Trichonotulus villiersi Petrovitz, 1968
Trichonotulus vultuosus Balthasar, 1971
Trigonoscelus afghanus Petrovitz, 1963
Trigonoscelus coelebs Petrovitz, 1963
Trigonoscelus elbursensis Stebnicka & ..., 1991
Trigonoscelus hypi Kral, 2017
Trigonoscelus narayaniensis Ilcikova & Kral, 2004
Tristaphodius asperulus Cervenka, 1994
Tristaphodius mortuorum Balthasar, 1932
Turanella latevittis Reitter, 1887
Vladimirellus socors Balthasar, 1967
Volinus sticticus Panzer, 1798
Xenoheptaulacus carinator (Hinton, 1934)⇒ Xenoheptaulacus tricostatus Harold, 1896
Xenoheptaulacus tricostatus Harold, 1896
Xeropsamobeus acerbus Horn, 1887
Xeropsamobeus ambiguus Fall & Cocke..., 1907
Xeropsamobeus arenicolus Gordon & Ske..., 2007Lesser weever
Xeropsamobeus asellus Schmidt, 1907Spotted weever
Xeropsamobeus brighti Gordon & Ske..., 2007Lesser weever
Xeropsamobeus desertus Van Dyke, 1918
Xeropsamobeus doyeni Gordon & Ske..., 2007Lesser weever
Xeropsamobeus mohavei Gordon & Ske..., 2007Lesser weever
Xeropsamobeus padrei Gordon & Ske..., 2007Lesser weever
Xeropsamobeus scabriceps LeConte, 1878
Youngaphodius agullasi Endrödi, 1983

Subnodes of Aphodiinae with occurrences in Continents
TribeAegialiini
Laporte, 1840
10
107
Genera
Species

Taxom

TribeAphodiini
Leach, 1815
106
1814
Genera
Species

Taxom

TribeAulonocnemini
Janssens, 1946
4
56
Genera
Species

Taxom

TribeChironini
Harold, 1867
1
17
Genus
Species

Taxom

TribeCorythoderini
Schmidt, 1910
6
37
Genera
Species

Taxom

TribeEremazini
Stebnicka, 1977
1
6
Genus
Species

Taxom

TribeEupariini
Schmidt, 1910
46
749
Genera
Species

Taxom

TribeOdochilini
Rakovič, 1987
1
23
Genus
Species

Taxom

TribeOdontolochini
Stebnicka & Howden, 1996
5
22
Genera
Species

Taxom

TribePsammodiini
Mulsant, 1842
34
560
Genera
Species

Taxom

TribeRhyparini
Schmidt, 1910
11
130
Genera
Species

Taxom

TribeStereomerini
Howden & Storey, 1992
9
23
Genera
Species

Taxom

TribeTermitoderini
Tangelder & Krikken, 1982
2
4
Genera
Species

Taxom

TribeTermitotrogini
Wasmann, 1918
1
15
Genus
Species

Taxom

TribeThinorycterini
Semenov-Tian-Shanskij & Reichardt, 1925
1
5
Genus
Species

Taxom



Links, sources and further information
Aphodiinae at artfakta.se
Aphodiinae at dyntaxa.se
Aphodiinae at itis.gov