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Checklist for Subfamily Aphodiinae Leach, 1815 (Aphodiine Dung Beetles)
and region Caucasus (Eurasia)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
523
6140
18845
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
521
6137
18842
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
497
6044
18637
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
331
4886
16042
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
98
1952
8628
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
93
1701
7123
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
11
207
655
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
11
207
655
Families
Genera
Species
 
Taxom
 
FamilyScarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles
177
565
Genera
Species
 
Taxom
 
SubfamilyAphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles
69
176
Genera
Species
 
Taxom
 


Genera for Caucasus
69 Genera with 176 Species
#ABCEHLMNOPRSTV
Acanthobodilus Dellacasa, 17991SpeciesTaxom
Acrossus Mulsant, 17586SpeciesTaxom
Agoliinus Schmidt, 18923SpeciesTaxom
Agrilinus Mulsant & Rey, 18484SpeciesTaxom
Alloblackburneus Bordat, 18875SpeciesTaxom
Alocoderus Schmidt, 18702SpeciesTaxom
Amidorus Mulsant & Rey, 18005SpeciesTaxom
Ammoecius Mulsant, 19752SpeciesTaxom
Aparammoecius Petrovitz, 19681SpeciesTaxom
Aphodius Hellwig, 18925SpeciesTaxom
Ataenius Harold, 184610SpeciesTaxom
Biralus Mulsant & Rey, 17893SpeciesTaxom
Bodiloides1SpeciesTaxom
Bodilopsis Adám, 04SpeciesTaxom
Bodilus Mulsant & Rey, 18484SpeciesTaxom
Calamosternus Motschulsky, 18632SpeciesTaxom
Chilothorax Motschulsky, 18949SpeciesTaxom
Colobopterus Mulsant, 17582SpeciesTaxom
Coprimorphus Mulsant, 17891SpeciesTaxom
Eremazus Mulsant, 18512SpeciesTaxom
Erytus Mulsant & Rey, 18602SpeciesTaxom
Esymus Mulsant & Rey, 19165SpeciesTaxom
Eudolus Mulsant & Rey, 18922SpeciesTaxom
Euheptaulacus Dellacasa, 18064SpeciesTaxom
Euorodalus Dellacasa, 19602SpeciesTaxom
Eupleurus Mulsant, 17581SpeciesTaxom
Heptaulacus Mulsant, 17751SpeciesTaxom
Labarrus Mulsant and Rey, 19413SpeciesTaxom
Limarus Mulsant & Rey, 18001SpeciesTaxom
Liothorax Motschulsky, 17673SpeciesTaxom
Loraphodius Reitter, 18352SpeciesTaxom
Loraspis Mulsant & Rey, 18701SpeciesTaxom
Mecynodes Mulsant & Rey, 18761SpeciesTaxom
Melinopterus Mulsant, 17988SpeciesTaxom
Mendidaphodius Reitter, 18991SpeciesTaxom
Mendidius Harold, 18976SpeciesTaxom
Neagolius Kozhantschikov, 18483SpeciesTaxom
Neopsammodius Rakovic, 18351SpeciesTaxom
Nialaphodius Kolbe, 18011SpeciesTaxom
Nialus Mulsant & Rey, 18051SpeciesTaxom
Nimbus Mulsant & Rey, 18233SpeciesTaxom
Nobiellus2SpeciesTaxom
Nobius Mulsant & Rey, 17992SpeciesTaxom
Oromus Mulsant & Rey, 19681SpeciesTaxom
Otophorus Mulsant, 17581SpeciesTaxom
Oxyomus Dejean, 17631SpeciesTaxom
Parammoecius Seidlitz, 19723SpeciesTaxom
Parataenius Balthasar, 18681SpeciesTaxom
Phaeaphodius Reitter, 18631SpeciesTaxom
Phalacronothus Motschulsky, 19175SpeciesTaxom
Plagiogonus Mulsant, 18634SpeciesTaxom
Planolinellus Dellacasa an..., 18251SpeciesTaxom
Planolinoides Dellacasa an..., 18272SpeciesTaxom
Planolinus Mulsant and Rey, 17891SpeciesTaxom
Platytomus Mulsant, 18462SpeciesTaxom
Pleurophorus Mulsant, 19614SpeciesTaxom
Psammodius Fallen, 18026SpeciesTaxom
Pseudacrossus Reitter, 18921SpeciesTaxom
Rhodaphodius1SpeciesTaxom
Rhyssemodes Reitter, 18741SpeciesTaxom
Rhyssemus Mulsant, 20187SpeciesTaxom
Scabrostomus Gordon and S..., 19911SpeciesTaxom
Sigorus Mulsant & Rey, 17921SpeciesTaxom
Stenotothorax Schmidt, 20182SpeciesTaxom
Subrinus Mulsant & Rey, 18702SpeciesTaxom
Teuchestes Mulsant, 17581SpeciesTaxom
Trichonotulus Bedel, 17872SpeciesTaxom
Turanella Semenov, 18871SpeciesTaxom
Volinus Mulsant & Rey, 17981SpeciesTaxom


Species list for Caucasus
176 Species and 12 Subspecies
Acanthobodilus immundus Creutzer, 1799
Acrossus bolognai Carpaneto & ..., 1989
Acrossus depressus Kugelann, 1792
Acrossus gagatinus Ménétriés, 1832
Acrossus luridus Fabricius, 1775
Acrossus planicollis Reitter, 1890
Acrossus rufipes Linnaeus, 1758Night-Flying Dung Beetle
Agoliinus piceus Gyllenhal, 1808
Agoliinus satunini Olsoufieff, 1918
Agoliinus satyrus Reitter, 1892
Agrilinus ater De Geer, 1774
Agrilinus ater falsarius Reitter, 1892
Agrilinus constans Duftschmid, 1805
Agrilinus convexus Erichson, 1848
Alloblackburneus aegrotus Horn, 1870
Alloblackburneus geomysi Cartwright, 1939
Alloblackburneus lentus Horn, 1870
Alloblackburneus rubeolus Palisot de B..., 1809Surf blenny
Alloblackburneus tenuistriatus Horn, 1887
Alocoderus hydrochaeris Fabricius, 1798
Alocoderus turbatus Baudi, 1870
Amidorus alagoezi Olsoufieff, 1918
Amidorus cribrarius Brullé, 1832
Amidorus obscurus Fabricius, 1792
Amidorus obscurus obscurus Fabricius, 1792
Amidorus thermicola Sturm, 1800
Ammoecius brevis Erichson, 1848
Ammoecius meurguesae Clement, 1975
Aparammoecius leisteri Medvedev, 1968
Aphodius (Aphodius) coniugatus Panzer, 1795
Aphodius (Aphodius) fimetarius Linnaeus, 1758Common dung beetle
Aphodius (Lorapsis) frater (Mulsant & Rey, 1869)⇒ Loraspis frater Mulsant & Rey, 1870
Aphodius (Nialus) lividus (Olivier, 1789)⇒ Acrossus luridus Fabricius, 1775
Aphodius swaneticus Reitter, 1892
Ataenius alternatus Melsheimer, 1845
Ataenius brevis Fall, 1930
Ataenius cylindrus (Horn, 1871)⇒ Ataenius ovatulus Horn, 1871
Ataenius fattigi Cartwright, 1948
Ataenius gracilis Melsheimer, 1845
Ataenius horticola Harold, 1869
Ataenius imbricatus Melsheimer, 1845
Ataenius opatroides Blanchard, 1846
Ataenius picinus Harold, 1861Antarctic blennies
Ataenius platensis Blanchard, 1846
Biralus equinus Faldermann, 1835
Biralus menetriesi Ménétriés, 1849
Biralus satellitius Herbst, 1789
Bodiloides ictericus ictericus Laicharting, 1781
Bodilopsis ogloblini Semenov & Me..., 1928
Bodilopsis rufus Moll, 1782
Bodilopsis sordidus Fabricius, 1775
Bodilopsis sordidus sordidus
Bodilus caspius Menestries, 1832
Bodilus gregarius Harold, 1871
Bodilus lugens Creutzer, 1799
Bodilus punctipennis Erichson, 1848
Calamosternus clermonti Reitter, 1907
Calamosternus trucidatus Harold, 1863
Chilothorax clathratus Reitter, 1892
Chilothorax distinctus (Müller, 1776)⇒ Aphodius distinctus O.F. Müller, 1776
Chilothorax distinctus sevanicus Rakovic, 1991
Chilothorax dobrovljanskyi Koshantschikov, 1913
Chilothorax flammulatus Harold, 1876
Chilothorax lineolatus Illiger, 1803
Chilothorax melanostictus melanostictus W. Schmidt, 1840
Chilothorax paykulli Bedel, 1907
Chilothorax variicolor Koshantschikov, 1894
Colobopterus brignolii Carpaneto, 1973
Colobopterus erraticus Linnaeus, 1758Erratic small dung beetle
Coprimorphus scrutator Herbst, 1789
Eremazus cribratus Semenov, 1893
Eremazus unistriatus Mulsant, 1851
Erytus aequalis Schmidt, 1907
Erytus cognatus Fairmaire, 1860
Esymus fumigatulus Reitter, 1892
Esymus merdarius Fabricius, 1775
Esymus pusillus pusillus Herbst, 1789
Esymus sculpturatus Reitter, 1892
Esymus suturinigra Schmidt, 1916
Eudolus quadriguttatus Herbst, 1783
Eudolus quadrinaevulus Reitter, 1892
Euheptaulacus carinatus Germar, 1824
Euheptaulacus carinatus carinatus Germar, 1824
Euheptaulacus sus Herbst, 1783
Euheptaulacus villosus Gyllenhal, 1806
Euorodalus coenosus Panzer, 1798
Euorodalus paracoenosus Balthasar & ..., 1960
Eupleurus subterraneus subterraneus Linnaeus, 1758
Heptaulacus testudinarius Fabricius, 1775
Labarrus lividus Olivier, 1789Scarab beetle
Labarrus lividus lividus
Labarrus pseudolividus Balthasar, 1941
Limarus maculatus Sturm, 1800
Liothorax kraatzi Harold, 1868
Liothorax niger Illiger, 1798Beaulieu dung beetle
Liothorax plagiatus Linnaeus, 1767
Loraphodius latisulcus Reitter, 1892
Loraphodius suarius Faldermann, 1835
Loraspis frater Mulsant & Rey, 1870
Mecynodes kisilkumi Solsky, 1876
Melinopterus consputus Creutzer, 1799
Melinopterus edithae Reitter, 1906
Melinopterus prodromus Brahm, 1790Spring small dung beetle
Melinopterus pubescens Sturm, 1800
Melinopterus punctatosulcatus hirtipes Fischer von ..., 1844
Melinopterus rapax Faldermann, 1836
Melinopterus reyi Reitter, 1892
Melinopterus sphacelatus Panzer, 1798
Mendidaphodius brancsiki Reitter, 1899
Mendidius abdurakhmanovi Shokhin, 2018
Mendidius bidens Solsky, 1876
Mendidius bispinifrons Reitter, 1889
Mendidius diffidens Reitter, 1892
Mendidius heikertingeri Balthasar, 1938
Mendidius multiplex Reitter, 1897
Neagolius abchasicus Reitter, 1892
Neagolius aragatsi Shokhin & Ka..., 2015
Neagolius montanus Erichson, 1848
Neopsammodius interruptus Say, 1835
Nialaphodius nigrita Fabricius, 1801
Nialus varians Duftschmid, 1805
Nimbus cartalinius Olsoufieff, 1918
Nimbus lederi Harold, 1876
Nimbus obliteratus Panzer, 1823
Nobiellus circumductus Solsky, 1876
Nobiellus inclusus Reitter, 1892
Nobius gresseri Semenov, 1899
Nobius serotinus Panzer, 1799
Oromus leisteri Medvedev, 1968
Otophorus haemorrhoidalis Linnaeus, 1758Bloody small dung beetle
Oxyomus sylvestris Scopoli, 1763Forest small dung beetle
Parammoecius asphaltinus Kolenati, 1846
Parammoecius brevithorax Sumakov, 1903
Parammoecius osseticus Iablokov-Khn..., 1972
Parataenius simulator Harold, 1868
Phaeaphodius dauricus Harold, 1863
Phalacronothus biguttatus Germar, 1824
Phalacronothus fumigatulus Reitter, 1892
Phalacronothus hilaris Harold, 1869
Phalacronothus quadrimaculatus Linnaeus, 1761
Phalacronothus suturinigra Schmidt, 1917
Plagiogonus arenarius Olivier, 1789
Plagiogonus praeustus Ballion, 1870
Plagiogonus putridus (Geoffroy, 1785)⇒ Plagiogonus arenarius Olivier, 1789
Plagiogonus syriacus Harold, 1863
Planolinellus vittatus Say, 1825
Planolinoides aenictus Cooper & Gordon, 1987Benttooths
Planolinoides borealis Gyllenhal, 1827
Planolinus fasciatus Olivier, 1789
Platytomus longulus Cartwright, 1948Horned blenny
Platytomus variolosus Kolenati, 1846
Pleurophorus anatolicus Petrovitz, 1961
Pleurophorus arabicus Pittino & Ma..., 1986
Pleurophorus caesus Creutzer, 1796Cut small dung beetle
Pleurophorus pannonicus Petrovitz, 1961
Psammodius asper (Fabricius, 1775)⇒ Psammodius sulcicollis Illiger, 1802
Psammodius caucasicus Pittino & Sh..., 2006
Psammodius generosus (Reitter, 1892)⇒ Psammodius nocturnus Reitter, 1892
Psammodius laevipennis Costa, 1844Awkward-winged small dung beetle
Psammodius nocturnus Reitter, 1892
Psammodius sulcicollis Illiger, 1802
Pseudacrossus circassicus Reitter, 1892
Rhodaphodius foetens Fabricius, 1787
Rhyssemodes orientalis Mulsant & Go..., 1874
Rhyssemus algiricus meridionalis Reitter, 1890
Rhyssemus annaedicatus Pierotti, 1980
Rhyssemus histrio Balthasar, 1961
Rhyssemus interruptus Reitter, 1892
Rhyssemus meridionalis (Reitter, 1890)⇒ Rhyssemus algiricus meridionalis Reitter, 1890
Rhyssemus morgani (Benard, 1911)⇒ Rhyssemus berytensis Marseul, 1878
Rhyssemus transcaucasicus Minkina & Král, 2018
Scabrostomus tanytarsus Skelley & Wo..., 1991Masked stargazer
Sigorus porcus Fabricius, 1792Mahogany Ridgeback scarab
Stenotothorax lanei Saylor, 1940
Stenotothorax smilodon Skelley & Mc..., 2018
Subrinus clermonti Reitter, 1907
Subrinus sturmi Harold, 1870
Teuchestes fossor Linnaeus, 1758Digger small dung beetle
Trichonotulus osetinus Medvedev & D..., 1977
Trichonotulus scrofa Fabricius, 1787Sow small dung beetle
Turanella latevittis Reitter, 1887
Volinus sticticus Panzer, 1798

Subnodes of Aphodiinae with occurrences in Caucasus
TribeAphodiini
Leach, 1815
26
70
Genera
Species

Taxom

TribeEremazini
Stebnicka, 1977
1
2
Genus
Species

Taxom

TribeEupariini
Schmidt, 1910
2
11
Genera
Species

Taxom

TribePsammodiini
Mulsant, 1842
6
21
Genera
Species

Taxom



Links, sources and further information
Aphodiinae at artfakta.se
Aphodiinae at dyntaxa.se
Aphodiinae at itis.gov