Checklist for Subfamily Aphodiinae Leach, 1815 (Aphodiine Dung Beetles)
and region Europe (Eurasia)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 1408
35417
222091 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 1401
35364
221754 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 1211
34497
219210 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 647
26961
190296 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 169
9191
59961 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 143
8084
50035 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 14
778
3568 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 14
778
3568 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 616
3000 | Genera
Species |
Taxom
| | Subfamily | Aphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles | 136
719 | Genera
Species |
Taxom
|
Genera for Europe
136 Genera with 719 Species #ABCDEGHLMNOPRSTVX
| Acanthobodilus Dellacasa, 1799 | | 2 | Species | Taxom | | Acrossidius Schmidt, 1847 | | 1 | Species | Taxom | | Acrossus Mulsant, 0 | | 12 | Species | Taxom | | Adeloparius Schmidt, 1847 | | 2 | Species | Taxom | | Aegialia Latreille, 1878 | | 10 | Species | Taxom | | Aganocrossus Reitter, 1862 | | 1 | Species | Taxom | | Agoliinus Schmidt, 1892 | | 11 | Species | Taxom | | Agolius Mulsant & Rey, 0 | | 10 | Species | Taxom | | Agrilinus Mulsant & Rey, 0 | | 19 | Species | Taxom | | Ahermodontus Baguena, 1930 | | 3 | Species | Taxom | | Aidophus Balthasar, 1859 | | 1 | Species | Taxom | | Airapus Stebnicka & ..., 2017 | | 1 | Species | Taxom | | Alloblackburneus Bordat, 1887 | | 5 | Species | Taxom | | Alocoderus Schmidt, 1870 | | 4 | Species | Taxom | | Amerisaprus | | 1 | Species | Taxom | | Amidorus Mulsant & Rey, 1800 | | 9 | Species | Taxom | | Ammoecius Mulsant, 1850 | | 10 | Species | Taxom | | Anomius Mulsant & Rey, 1983 | | 4 | Species | Taxom | | Aphodiites Heer, 1865 | | 1 | Species | Taxom | | Aphodius Hellwig, 1975 | | 226 | Species | Taxom | | Aphodoharmogaster Endrodi, 1857 | | 1 | Species | Taxom | | Aphotaenius Cartwright, 1928 | | 1 | Species | Taxom | | Aschnarhyparus Makhan, 1934 | | 1 | Species | Taxom | | Ataeniopsis Petrovitz, 1974 | | 4 | Species | Taxom | | Ataenius Harold, 1864 | | 36 | Species | Taxom | | Auperia Chevrolat, 1864 | | 1 | Species | Taxom | | Australaphodius Balthasar, 1892 | | 1 | Species | Taxom | | Biralus Mulsant & Rey, 1789 | | 3 | Species | Taxom | | Blackburneus Schmidt, 1845 | | 2 | Species | Taxom | | Bodiloides | | 3 | Species | Taxom | | Bodilopsis Adám, 0 | | 8 | Species | Taxom | | Bodilus Mulsant & Rey, 1862 | | 15 | Species | Taxom | | Brindalus Landin, 1980 | | 4 | Species | Taxom | | Caelius Lewis, 1887 | | 1 | Species | Taxom | | Calamosternus Motschulsky, 1789 | | 7 | Species | Taxom | | Chilothorax Motschulsky, 1798 | | 25 | Species | Taxom | | Chiron MacLeay, 1798 | | 1 | Species | Taxom | | Cinacanthus Schmidt, 1901 | | 1 | Species | Taxom | | Cnemisus Motschulsky, 1845 | | 1 | Species | Taxom | | Colobopterus Mulsant, 0 | | 3 | Species | Taxom | | Coprimorphus Mulsant, 1789 | | 1 | Species | Taxom | | Coptochiroides Balthasar, 1886 | | 1 | Species | Taxom | | Coptochirus Harold, 1868 | | 2 | Species | Taxom | | Dellacasiellus Gordon and S..., 1922 | | 1 | Species | Taxom | | Dialytes Harold, 1875 | | 3 | Species | Taxom | | Diapterna Horn, 1850 | | 1 | Species | Taxom | | Diastictus Mulsant, 1805 | | 2 | Species | Taxom | | Didactylia Orbigny, 1859 | | 1 | Species | Taxom | | Drepanocanthoides Schmidt, 1870 | | 1 | Species | Taxom | | Drepanocanthus Péringuey, 1823 | | 2 | Species | Taxom | | Eremazus Mulsant, 1851 | | 1 | Species | Taxom | | Erytus Mulsant & Rey, 1892 | | 4 | Species | Taxom | | Esymus Mulsant & Rey, 1916 | | 7 | Species | Taxom | | Eudolus Mulsant & Rey, 1892 | | 2 | Species | Taxom | | Euheptaulacus Dellacasa, 1806 | | 7 | Species | Taxom | | Euorodalus Dellacasa, 1848 | | 5 | Species | Taxom | | Euparixoides Hinton, 1936 | | 1 | Species | Taxom | | Eupleurus Mulsant, 1758 | | 2 | Species | Taxom | | Geopsammodius Gordon & Pit..., 1991 | | 1 | Species | Taxom | | Gonaphodiellus Schmidt, 1945 | | 1 | Species | Taxom | | Harmogaster Harold, 1901 | | 4 | Species | Taxom | | Haroldiataenius Chalumeau, 1939 | | 1 | Species | Taxom | | Haroldiellus Gordon and S..., 1863 | | 1 | Species | Taxom | | Heptaulacus Mulsant, 1806 | | 9 | Species | Taxom | | Labarrus Mulsant and Rey, 1941 | | 3 | Species | Taxom | | Lechorodius Gordon and S..., 1823 | | 1 | Species | Taxom | | Leiopsammodius Rakovic, 1983 | | 3 | Species | Taxom | | Limarus Mulsant & Rey, 1813 | | 2 | Species | Taxom | | Liothorax Motschulsky, 0 | | 9 | Species | Taxom | | Loraphodius Reitter, 1835 | | 2 | Species | Taxom | | Loraspis Mulsant & Rey, 1870 | | 1 | Species | Taxom | | Mecynodes Mulsant & Rey, 1835 | | 2 | Species | Taxom | | Megatelus Reitter, 1845 | | 1 | Species | Taxom | | Melinopterus Mulsant, 1975 | | 17 | Species | Taxom | | Mendidaphodius Reitter, 1963 | | 3 | Species | Taxom | | Mendidius Harold, 1881 | | 5 | Species | Taxom | | Mesontoplatys Motschulsky, 1855 | | 2 | Species | Taxom | | Mothon Semenov & Me..., 1927 | | 1 | Species | Taxom | | Neagolius Kozhantschikov, 1938 | | 16 | Species | Taxom | | Nialaphodius Kolbe, 1801 | | 2 | Species | Taxom | | Nialus Mulsant & Rey, 1969 | | 3 | Species | Taxom | | Nimbus Mulsant & Rey, 1984 | | 13 | Species | Taxom | | Nobiellus | | 2 | Species | Taxom | | Nobius Mulsant & Rey, 1799 | | 5 | Species | Taxom | Odontolochus Schmidt, 1916 | | 0 | Species | Taxom | | Odontolytes Koshantschikov, 1963 | | 2 | Species | Taxom | | Orodaliscus Reitter, 1906 | | 3 | Species | Taxom | | Oromus Mulsant & Rey, 1968 | | 3 | Species | Taxom | | Oscarinus Gordon and S..., 2007 | | 2 | Species | Taxom | | Osmanius Branco & Baraud, 1963 | | 1 | Species | Taxom | | Otophorus Mulsant, 1758 | | 1 | Species | Taxom | | Oxyomoides Dellacasa, D..., 1944 | | 1 | Species | Taxom | | Oxyomus Dejean, 1763 | | 2 | Species | Taxom | | Parabodilus | | 1 | Species | Taxom | | Paracoptochirus Balthasar, 1990 | | 3 | Species | Taxom | | Paracrossidius Balthasar, 2017 | | 1 | Species | Taxom | | Parammoecius Seidlitz, 1863 | | 5 | Species | Taxom | | Parapsammodius Verdú, Steb..., 1858 | | 1 | Species | Taxom | | Parataenius Balthasar, 1868 | | 1 | Species | Taxom | | Pardalosus Gordon and S..., 1857 | | 1 | Species | Taxom | | Passaliolla Balthasar, 1887 | | 1 | Species | Taxom | | Phaeaphodius Reitter, 1911 | | 7 | Species | Taxom | | Phalacronothus Motschulsky, 1917 | | 10 | Species | Taxom | | Pharaphodius Reitter, 1781 | | 2 | Species | Taxom | | Plagiogonus Mulsant, 1863 | | 4 | Species | Taxom | | Planolinellus Dellacasa an..., 1825 | | 1 | Species | Taxom | | Planolinoides Dellacasa an..., 1827 | | 2 | Species | Taxom | | Planolinus Mulsant and Rey, 1825 | | 5 | Species | Taxom | | Platytomus Mulsant, 1846 | | 7 | Species | Taxom | | Pleuraphodius Schmidt, 1851 | | 1 | Species | Taxom | | Pleurophorus Mulsant, 1961 | | 9 | Species | Taxom | | Podotenus Schmidt, 1904 | | 1 | Species | Taxom | | Psammodius Fallen, 1802 | | 15 | Species | Taxom | | Psammoporus Thomson, 1931 | | 7 | Species | Taxom | | Pseudacrossus Reitter, 1906 | | 14 | Species | Taxom | | Pseudesymus Orbigny, 1845 | | 1 | Species | Taxom | | Pubinus Mulsant & Rey, 1776 | | 1 | Species | Taxom | | Rhodaphodius | | 1 | Species | Taxom | | Rhysothorax Bedel, 1792 | | 2 | Species | Taxom | | Rhyssemodes Reitter, 1874 | | 1 | Species | Taxom | | Rhyssemus Mulsant, 1963 | | 30 | Species | Taxom | | Rugaphodius Gordon and S..., 1907 | | 1 | Species | Taxom | | Saprosites Redtenbacher, 1905 | | 5 | Species | Taxom | | Scabrostomus Gordon and S..., 1991 | | 1 | Species | Taxom | | Sigorus Mulsant & Rey, 1802 | | 2 | Species | Taxom | | Stenotothorax Schmidt, 2018 | | 3 | Species | Taxom | | Subrinus Mulsant & Rey, 1845 | | 3 | Species | Taxom | | Tesarius RakoviÄ, 1955 | | 2 | Species | Taxom | | Tetraclipeoides Schmidt, 1861 | | 1 | Species | Taxom | | Teuchestes Mulsant, 0 | | 3 | Species | Taxom | | Trichaphodius Schmidt, 1916 | | 1 | Species | Taxom | | Trichiorhyssemus Clouët, 1892 | | 2 | Species | Taxom | | Trichonotulus Bedel, 1787 | | 2 | Species | Taxom | | Trigonoscelus Petrovitz, 2017 | | 1 | Species | Taxom | | Volinus Mulsant & Rey, 1798 | | 1 | Species | Taxom | | Xeropsamobeus Saylor, 1907 | | 1 | Species | Taxom |
Species list for Europe
719 Species and 63 Subspecies | Acanthobodilus crimensis W.Koshantsch..., 1913 | | | Acanthobodilus immundus Creutzer, 1799 | | | Acrossidius tasmaniae Hope, 1847 | Black-headed pasture cockchafer | | Acrossus atratus Waterhouse, 1875 | | | Acrossus bimaculatus Laxmann, 1770 | | | Acrossus carpetanus Graells, 1846 | | | Acrossus depressus Kugelann, 1792 | | | Acrossus gagatinus Ménétriés, 1832 | | | Acrossus laticollis Baudi, 1870 | | | Acrossus luridus Fabricius, 1775 | | | Acrossus planicollis Reitter, 1890 | | | Acrossus rufipes Linnaeus, 1758 | Night-Flying Dung Beetle | | Acrossus siculus Harold, 1862 | | | Acrossus siculus buturensis Dellacasa, 1983 | | | Acrossus siculus siculus | | | Adeloparius overlaeti Boucomont, 1936 | | | Adeloparius pulcherrimus Reiche, 1847 | | | Aegialia (Aegialia) arenaria Fabricius, 1787 | Dune sand scarab beetle | | Aegialia (Aegialia) crassa LeConte, 1857 | | | Aegialia (Psammoporus) insularis (Pittino, 2006) | ⇒ | | Aegialia (Rhysothorax) rufa (Fabricius, 1792) | ⇒ Caelius rufescens Horn, 1887 | | Aegialia latipunctus Gredler, 1863 | | | Aegialia mimica Pittino, 2006 | | | Aegialia mimicus Pittino, 2006 | | | Aegialia rufina (Silfverberg, 1977) | ⇒ Rhysothorax rufa Fabricius, 1792 | | Aegialia sabuleti Panzer, 1797 | | | Aegialia spissipes (LeConte, 1878) | ⇒ Rhysothorax rufa Fabricius, 1792 | | Aganocrossus urostigma Harold, 1862 | | | Agoliinus aleutus Eschscholtz, 1822 | | | Agoliinus canadensis Garnett, 1920 | Streaked weeverfish | | Agoliinus isajevi Kabakov, 1996 | | | Agoliinus lapponum Gyllenhal, 1808 | Northern Dung Beetle | | Agoliinus leopardus Horn, 1870 | | | Agoliinus nemoralis Erichson, 1848 | | | Agoliinus piceus Gyllenhal, 1808 | | | Agoliinus pittinoi Carpaneto, 1986 | | | Agoliinus ragusae Reitter, 1892 | | | Agoliinus ragusai Reitter, 1892 | | | Agoliinus satyrus Reitter, 1892 | | | Agolius abdominalis Bonelli, 1812 | | | Agolius abdominalis abdominalis Bonelli, 1812 | | | Agolius abdominalis balazuci Nicolas, 1971 | | | Agolius abdominalis emilianus Dellacasa, 1983 | | | Agolius abdominalis pecoudi Hoffmann, 1954 | | | Agolius abdominalis tatrensis Dellacasa & ..., 2005 | | | Agolius bonvouloiri Harold, 1860 | | | Agolius bonvouloiri bonvouloiri | | | Agolius bonvouloiri cantabricus Pittino, 1981 | | | Agolius limbolarius | | | Agrilinus aquilus Schmidt, 1916 | | | Agrilinus ater De Geer, 1774 | | | Agrilinus ater ascendens Reiche, 1863 | | | Agrilinus ater lucasi Harold, 1859 | | | Agrilinus bellus (Schmidt, 1916) | ⇒ Agrilinus aquilus Schmidt, 1916 | | Agrilinus borealis | | | Agrilinus constans Duftschmid, 1805 | | | Agrilinus convexus Erichson, 1848 | | | Agrilinus ibericus Harold, 1874 | | | Agrilinus ibericus basilicatus Fiori, 1907 | | | Agrilinus ibericus ibericus | | | Agrilinus nemoralis Erichson, 1848 | | | Agrilinus nikolajevi Berlov & Kal..., 1989 | | | Agrilinus piceus | | | Agrilinus pirinensis Balthasar, 1946 | | | Agrilinus rufus (Moll, 1782) | ⇒ Bodilopsis rufa Moll, 1782 | | Agrilinus sordidus (Fabricius, 1775) | ⇒ Bodilopsis sordida Fabricius, 1775 | | Agrilinus sordidus sordidus (Fabricius, 1775) | ⇒ Bodilopsis sordida sordida Fabricius, 1775 | | Agrilinus uliginosus | | | Ahermodontus ambrosi Pardo Alcaide, 1936 | | | Ahermodontus bischoffi Vsetecka, 1939 | | | Ahermodontus marini Baguena, 1930 | | | Aidophus notatus Harold, 1859 | | | Airapus lithuanicus Tamutis, Ale..., 2017 | | | Alloblackburneus aegrotus Horn, 1870 | | | Alloblackburneus geomysi Cartwright, 1939 | | | Alloblackburneus lentus Horn, 1870 | | | Alloblackburneus rubeolus Palisot de B..., 1809 | Surf blenny | | Alloblackburneus tenuistriatus Horn, 1887 | | | Alocoderus carinifrons Reitter, 1892 | | | Alocoderus holdereri Reitter, 1900 | | | Alocoderus hydrochaeris Fabricius, 1798 | | | Alocoderus turbatus Baudi, 1870 | | | Amerisaprus valdivia Stebnicka & ..., 2003 | | | Amidorus cribrarius Brullé, 1832 | | | Amidorus cribricollis Lucas, 1846 | | | Amidorus immaturus Mulsant, 1842 | | | Amidorus koshantschikovi Jacobson, 1911 | | | Amidorus obscurus Fabricius, 1792 | | | Amidorus obscurus latinus Dellacasa, 1983 | | | Amidorus obscurus obscurus Fabricius, 1792 | | | Amidorus porcus | | | Amidorus thermicola Sturm, 1800 | | | Ammoecius brevis Erichson, 1848 | | | Ammoecius dentatus Schmidt, 1908 | | | Ammoecius elevatus Olivier, 1789 | | | Ammoecius frigidus Brisout, 1866 | | | Ammoecius lugubris Boheman, 1857 | | | Ammoecius lusitanicus Erichson, 1848 | | | Ammoecius muchei Petrovitz, 1963 | | | Ammoecius numidicus Mulsant, 1851 | | | Ammoecius pannonicus (Petrovitz, 1961) | ⇒ Loraspis frater Mulsant & Rey, 1870 | | Ammoecius rugifrons Aubé, 1850 | | | Anomius annamariae Baraud, 1982 | | | Anomius baeticus Mulsant & Rey, 1869 | | | Anomius castaneus Illiger, 1803 | | | Anomius crovettii Dellacasa, 1983 | | | Aphodiites protogaeus ♰ Heer, 1865 | | | Aphodius (Acanthobodilus) immundus (Creutzer, 1799) | ⇒ Pubinus tomentosus O.F.Muller, 1776 | | Aphodius (Acrossus) bimaculatus (Laxman, 1778) | ⇒ Teuchestes brachysomus Solsky, 1874 | | Aphodius (Acrossus) carpetanus Graells, 1847 | | | Aphodius (Acrossus) depressus (Kugelann, 1792) | ⇒ Acrossus depressus Kugelann, 1792 | | Aphodius (Acrossus) laticollis (Baudi di Selve, 1870) | ⇒ Acrossus laticollis Baudi, 1870 | | Aphodius (Acrossus) luridus (Fabricius, 1775) | ⇒ Acrossus luridus Fabricius, 1775 | | Aphodius (Acrossus) rufipes (Linnaeus, 1758) | ⇒ Acrossus rufipes Linnaeus, 1758 | | Aphodius (Acrossus) siculus Harold, 1862 | | | Aphodius siculus buturensis Dellacasa, 1983 | | | Aphodius siculus siculus Harold, 1862 | | | Aphodius (Agoliinus) lapponum (Gyllenhal, 1806) | ⇒ Agoliinus lapponum Gyllenhal, 1808 | | Aphodius (Agoliinus) piceus (Gyllenhal, 1808) | ⇒ Agoliinus piceus Gyllenhal, 1808 | | Aphodius (Agoliinus) ragusae Reitter, 1892 | | | Aphodius (Agoliinus) satyrus (Reitter, 1892) | ⇒ Agoliinus satyrus Reitter, 1892 | | Aphodius (Agolius) abdominalis (Bonelli, 1812) | ⇒ Agolius abdominalis Bonelli, 1812 | | Aphodius abdominalis abdominalis Bonelli, 1812 | | | Aphodius abdominalis emilianus Dellacasa, 1983 | | | Aphodius (Agolius) mixtus (Motschulsky, 1858) | ⇒ Agolius abdominalis Bonelli, 1812 | | Aphodius (Agrilinus) ater (DeGeer, 1774) | ⇒ Agrilinus ater De Geer, 1774 | | Aphodius (Agrilinus) constans (Duftschmid, 1805) | ⇒ Agrilinus constans Duftschmid, 1805 | | Aphodius (Agrilinus) convexus (Erichson, 1848) | ⇒ Agrilinus convexus Erichson, 1848 | | Aphodius (Agrilinus) ibericus Harold, 1874 | | | Aphodius (Agrilinus) nemoralis (Erichson, 1848) | ⇒ Agoliinus nemoralis Erichson, 1848 | | Aphodius (Agrilinus) rufus Moll, 1782 | | | Aphodius (Agrilinus) sordidus (Fabricius, 1775) | ⇒ Bodilopsis sordida Fabricius, 1775 | | Aphodius (Ahermodontus) marini Baguena, 1930 | | | Aphodius (Alocoderus) hydrochaeris | | | Aphodius (Amidorus) cribrarius Brullé, 1832 | | | Aphodius (Amidorus) immaturus (Mulsant, 1842) | ⇒ Amidorus immaturus Mulsant, 1842 | | Aphodius (Amidorus) obscurus (Fabricius, 1792) | ⇒ Amidorus obscurus Fabricius, 1792 | | Aphodius obscurus latinus Dellacasa, 1983 | | | Aphodius obscurus obscurus | | | Aphodius (Ammoecius) brevis (Erichson, 1848) | ⇒ Ammoecius brevis Erichson, 1848 | | Aphodius (Ammoecius) elevatus (Olivier, 1789) | ⇒ Ammoecius elevatus Olivier, 1789 | | Aphodius (Ammoecius) rugifrons (Horn, 1871) | ⇒ Aphodius fucosus Schmidt, 1922 | | Aphodius (Anomius) castaneus (Illiger, 1803) | ⇒ Anomius castaneus Illiger, 1803 | | Aphodius (Anomius) crovettii G. Dellacasa, 1983 | | | Aphodius (Aphodius) aestivalis (Stephens, 1839) | ⇒ Rhodaphodius foetens Fabricius, 1787 | | Aphodius (Aphodius) coniugatus Panzer, 1795 | | | Aphodius (Aphodius) conjugatus (Mulsant, 1842) | ⇒ Aphodius coniugatus Panzer, 1795 | | Aphodius (Aphodius) fimetarius Linnaeus, 1758 | Common dung beetle | | Aphodius (Aphodius) foetens (Fabricius, 1787) | ⇒ Rhodaphodius foetens Fabricius, 1787 | | Aphodius (Aphodius) foetidus Herbst, 1783 | | | Aphodius foetidus foetidus | | | Aphodius (Aphodius) pedellus (DeGeer, 1774) | ⇒ Aphodius fimetarius Linnaeus, 1758 | | Aphodius (Aphodius) pedrosi Wollaston, 1854 | | | Aphodius (Aphodius) scybalarius (Fabricius, 1781) | ⇒ Bodilopsis rufa Moll, 1782 | | Aphodius (Biralus) satellitius (Herbst, 1789) | ⇒ Biralus satellitius Herbst, 1789 | | Aphodius (Bodilus) arvernicus (Hoffmann, 1928) | ⇒ Bodilopsis rufus Moll, 1782 | | Aphodius (Bodilus) beduinus Reitter, 1892 | | | Aphodius (Bodilus) ghardimaouensis (Balthasar, 1929) | ⇒ Bodiloides ictericus ghardimaouensis Balthasar, 1929 | | Aphodius (Bodilus) gregarius Harold, 1871 | | | Aphodius (Bodilus) hydrochaeris (Fabricius, 1798) | ⇒ Alocoderus hydrochaeris Fabricius, 1798 | | Aphodius (Bodilus) ictericus (Laicharting, 1781) | ⇒ Bodiloides ictericus Laicharting, 1781 | | Aphodius (Bodilus) longispina Küster, 1854 | | | Aphodius (Bodilus) lugens (Creutzer, 1799) | ⇒ Bodilus lugens Creutzer, 1799 | | Aphodius (Bodilus) nitidulus (C. O. Waterh..., 1875) | ⇒ Bodiloides ictericus ictericus Laicharting, 1781 | | Aphodius (Bodilus) punctipennis Erichson, 1848 | | | Aphodius (Bodilus) rufus (Moll, 1782) | ⇒ Bodilopsis rufa Moll, 1782 | | Aphodius (Bodilus) sordidus (Fabricius, 1775) | ⇒ Bodilopsis rufus Moll, 1782 | | Aphodius (Bodilus) wollastoni Harold, 1862 | | | Aphodius (Calamosternus) granarius (Linnaeus, 1767) | ⇒ Calamosternus granarius Linnaeus, 1767 | | Aphodius (Calamosternus) hyxos Petrovitz, 1962 | | | Aphodius hyxos algiricus Mariani & Pi..., 1983 | | | Aphodius (Calamosternus) mayeri (Pilleri, 1953) | ⇒ Calamosternus mayeri Pilleri, 1953 | | Aphodius (Calamosternus) trucidatus (Chevrolat, 1863) | ⇒ Calamosternus trucidatus Harold, 1863 | | Aphodius (Calamosternus) unicolor (Lucas, 1846) | ⇒ Anomius castaneus Illiger, 1803 | | Aphodius (Calomosternus) granarius Linnaeus, 1767 | | | Aphodius (Chilothorax) cervorum (Fairmaire, 1871) | ⇒ Chilothorax cervorum Fairmaire, 1871 | | Aphodius (Chilothorax) conspurcatus (Linnaeus, 1758) | ⇒ Chilothorax conspurcatus Linnaeus, 1758 | | Aphodius (Chilothorax) distinctus O.F. Müller, 1776 | Maculated Dung Beetle | | Aphodius (Chilothorax) hieroglyphicus (Klug, 1845) | ⇒ Chilothorax hieroglyphicus Klug, 1845 | | Aphodius (Chilothorax) lineolatus (Illiger, 1803) | ⇒ Chilothorax lineolatus Illiger, 1803 | | Aphodius (Chilothorax) melanostictus (W.L.E. Schmidt, 1840) | ⇒ Chilothorax melanostictus Schmidt, 1840 | | Aphodius (Chilothorax) paykulli Bedel, 1907 | | | Aphodius (Chilothorax) pictus (D. Koshantsc..., 1894) | ⇒ Chilothorax pictus Sturm, 1805 | | Aphodius (Chilothorax) sticticus (Panzer, 1798) | ⇒ Volinus sticticus Panzer, 1798 | | Aphodius (Chilothorax) tessulatus (Paykull, 1798) | ⇒ Chilothorax paykulli Bedel, 1907 | | Aphodius (Colobopterus) erraticus (Linnaeus, 1758) | ⇒ Colobopterus erraticus Linnaeus, 1758 | | Aphodius (Coprimorphus) scrutator (Herbst, 1789) | ⇒ Coprimorphus scrutator Herbst, 1789 | | Aphodius (Dellacasiellus) fucosus Schmidt, 1922 | | | Aphodius (Erytus) aequalis (A. SCHMIDT, 1907) | ⇒ Erytus aequalis Schmidt, 1907 | | Aphodius (Erytus) cognatus (Fairmaire, 1860) | ⇒ Alocoderus hydrochaeris Fabricius, 1798 | | Aphodius (Esymus) merdarius (Fabricius, 1775) | ⇒ Esymus merdarius Fabricius, 1775 | | Aphodius (Esymus) pusillus Herbst, 1789 | | | Aphodius pusillus pusillus | | | Aphodius (Eudolus) quadriguttatus (Herbst, 1783) | ⇒ Eudolus quadriguttatus Herbst, 1783 | | Aphodius (Euorodalus) coenosus (Panzer, 1798) | ⇒ Euorodalus coenosus Panzer, 1798 | | Aphodius (Euorodalus) paracoenosus (Balthasar & ..., 1960) | ⇒ Euorodalus paracoenosus Balthasar & ..., 1960 | | Aphodius (Euorodalus) pusillus (Herbst, 1789) | ⇒ Esymus pusillus Herbst, 1789 | | Aphodius (Euorodalus) tersus (Rosenhauer, 1856) | ⇒ Euorodalus tersus Erichson, 1848 | | Aphodius (Eupleurus) subterraneus (Linnaeus, 1758) | ⇒ Eupleurus subterraneus Linnaeus, 1758 | | Aphodius subterraneus subterraneus | | | Aphodius (Labarrus) lividus (Olivier, 1789) | ⇒ Labarrus lividus Olivier, 1789 | | Aphodius (Labarrus) pseudolividus (Balthasar, 1941) | ⇒ Labarrus pseudolividus Balthasar, 1941 | | Aphodius (Limarus) maculatus (Sturm, 1800) | ⇒ Limarus maculatus Sturm, 1800 | | Aphodius (Limarus) zenkeri (Germar, 1813) | ⇒ Limarus zenkeri Germar, 1813 | | Aphodius (Liothorax) niger (Panzer, 1797) | ⇒ Planolinellus vittatus Say, 1825 | | Aphodius (Liothorax) plagiatus (Linnaeus, 1767) | ⇒ Liothorax plagiatus Linnaeus, 1767 | | Aphodius (Lithorax) linearis (Reiche & Saulcy, 1856) | ⇒ Mendidaphodius linearis Reiche & Saulcy, 1856 | | Aphodius (Loraphodius) suarius (Faldermann, 1835) | ⇒ Loraphodius suarius Faldermann, 1835 | | Aphodius (Lorapsis) frater (Mulsant & Rey, 1869) | ⇒ Loraspis frater Mulsant & Rey, 1870 | | Aphodius (Loraspis) frater Mulsant & Rey, 1870 | | | Aphodius (Mecynodes) striatulus (Waltl, 1835) | ⇒ Mecynodes striatulus Waltl, 1835 | | Aphodius (Megatelus) contractus Klug, 1845 | | | Aphodius contractus scolytoides Lucas, 1846 | | | Aphodius (Melinopterus) consputus (Creutzer, 1799) | ⇒ Melinopterus consputus Creutzer, 1799 | | Aphodius (Melinopterus) prodromus (Braham, 1790) | ⇒ Melinopterus prodromus Brahm, 1790 | | Aphodius (Melinopterus) pubescens (Sturm, 1800) | ⇒ Melinopterus pubescens Sturm, 1800 | | Aphodius (Melinopterus) punctatosulcatus (Sturm, 1805) | ⇒ Melinopterus punctatosulcatus punctatosulcatus Sturm, 1805 | | Aphodius (Melinopterus) reyi (Reitter, 1892) | ⇒ Melinopterus reyi Reitter, 1892 | | Aphodius (Melinopterus) sabulicola (Thomson, 1868) | ⇒ Melinopterus punctatosulcatus hirtipes Fischer von ..., 1844 | | Aphodius (Melinopterus) sphacelatus (Panzer, 1798) | ⇒ Melinopterus sphacelatus Panzer, 1798 | | Aphodius (Melinopterus) stolzi (Reitter, 1906) | ⇒ Melinopterus stolzi Reitter, 1906 | | Aphodius (Melinopterus) tingens (Reitter, 1892) | ⇒ Melinopterus tingens Reitter, 1892 | | Aphodius (Neagolius) amblyodon (K. Daniel, 1900) | ⇒ Neagolius amblyodon Daniel, 1900 | | Aphodius (Neagolius) heydeni Harold, 1871 | | | Aphodius (Neagolius) liguricus (J. Daniel, 1902) | ⇒ Neagolius liguricus Daniel, 1902 | | Aphodius (Neagolius) limbolarius Reitter, 1892 | | | Aphodius limbolarius danielorum A. Semenov, 1902 | | | Aphodius limbolarius limbolarius Reitter, 1892 | | | Aphodius (Neagolius) montanus (Schmidt, 1840) | ⇒ Neagolius montanus Erichson, 1848 | | Aphodius (Neagolius) montivagus (Erichson, 1848) | ⇒ Neagolius montivagus Erichson, 1848 | | Aphodius (Neagolius) penninus J. Daniel, 1902 | | | Aphodius (Neagolius) pollicatus Erichson, 1848 | | | Aphodius (Neagolius) praecox Erichson, 1848 | | | Aphodius (Neagolius) schlumbergeri (Seidlitz, 1891) | ⇒ Neagolius schlumbergeri Seidlitz, 1891 | | Aphodius schlumbergeri consobrinus | | | Aphodius schlumbergeri samniticus J. Daniel, 1902 | | | Aphodius (Nialus) lividus (Olivier, 1789) | ⇒ Acrossus luridus Fabricius, 1775 | | Aphodius (Nialus) plagiatus (Linnaeus, 1758) | ⇒ Liothorax plagiatus Linnaeus, 1767 | | Aphodius (Nialus) sturmi (Harold, 1870) | ⇒ Subrinus sturmi Harold, 1870 | | Aphodius (Nialus) varians (Duftschmid, 1805) | ⇒ Nialus varians Duftschmid, 1805 | | Aphodius (Nimbus) affinis (Lucas, 1846) | ⇒ Agrilinus ater De Geer, 1774 | | Aphodius (Nimbus) contaminatus (Herbst, 1783) | ⇒ Nimbus contaminatus Herbst, 1783 | | Aphodius (Nimbus) franzinii (Pittino, 1978) | ⇒ Nimbus franzinii Pittino, 1978 | | Aphodius (Nimbus) johnsoni Baraud, 1976 | | | Aphodius (Nimbus) obliteratus (Panzer, 1823) | ⇒ Nimbus obliteratus Panzer, 1823 | | Aphodius (Nobius) bonnairei (Reitter, 1892) | ⇒ Nobiellus bonnairei Reitter, 1892 | | Aphodius (Nobius) serotinus Creutzer, 1799 | | | Aphodius (Oromus) alpinus (Scopoli, 1763) | ⇒ Euheptaulacus carinatus carinatus Germar, 1824 | | Aphodius (Otophorus) haemorrhoidalis (Linnaeus, 1758) | ⇒ Otophorus haemorrhoidalis Linnaeus, 1758 | | Aphodius (Parammoecius) corvinus (Haldeman, 1848) | ⇒ Parammoecius corvinus Erichson, 1848 | | Aphodius (Parammoecius) gibbus (Germar, 1817) | ⇒ Parammoecius gibbus Germar, 1817 | | Aphodius (Parammoecius) pyrenaeus (Jacquelin du..., 1860) | ⇒ Parammoecius pyrenaeus Jacquelin du..., 1863 | | Aphodius (Phalacronothus) biguttatus Dallatorre, 1879 | | | Aphodius (Phalacronothus) quadrimaculatus Linné, 1761 | | | Aphodius quadrimaculatus quadrimaculatus | | | Aphodius (Phalocronothus) biguttatus (Dallatorre, 1879) | ⇒ Phalacronothus biguttatus Germar, 1824 | | Aphodius (Phalocronothus) quadrimaculatus (Linnaeus, 1761) | ⇒ Teuchestes brachysomus Solsky, 1874 | | Aphodius (Pharaphodius) priscus Motschulsky, 1858 | | | Aphodius (Plagiogonus) arenarius Olivier, 1789 | | | Aphodius (Plagiogonus) putridus (Herbst, 1789) | ⇒ Planolinoides borealis Gyllenhal, 1827 | | Aphodius (Planolinellus) vittatus Mulsant, 1842 | | | Aphodius (Planolinoides) borealis (Gyllenhal, 1827) | ⇒ Planolinoides borealis Gyllenhal, 1827 | | Aphodius (Planolinus) borealis (Gyllenhal, 1827) | ⇒ Planolinoides borealis Gyllenhal, 1827 | | Aphodius (Planolinus) fasciatus (Olivier, 1789) | ⇒ Limarus maculatus Sturm, 1800 | | Aphodius (Planolinus) uliginosus (Hardy, 1847) | ⇒ Planolinus fasciatus Olivier, 1789 | | Aphodius (Pseudacrossus) cribricollis Lucas, 1849 | | | Aphodius (Pseudacrossus) suffertus A. SCHMIDT, 1916 | | | Aphodius suffertus ampliatus Reitter, 1892 | | | Aphodius suffertus defiorei Fiori, 1907 | | | Aphodius (Pseudacrossus) thermicola (Sturm, 1800) | ⇒ Amidorus thermicola Sturm, 1800 | | Aphodius (Pseudocalamosternus) rigidus Balthasar, 1936 | | | Aphodius (Pubinus) tomentosus (O.F.Muller, 1776) | ⇒ Pubinus tomentosus O.F.Muller, 1776 | | Aphodius (Sigorius) porcus Fabricius, 1..., 1869 | | | Aphodius (Sigorus) porcus (Fabricius, 1792) | ⇒ Sigorus porcus Fabricius, 1792 | | Aphodius (Subrinus) sturmi (Harold, 1870) | ⇒ Subrinus sturmi Harold, 1870 | | Aphodius (Subrinus) vitellinus (Klug, 1845) | ⇒ Subrinus vitellinus Klug, 1845 | | Aphodius (Teuchestes) fossor (Linnaeus, 1758) | ⇒ Teuchestes fossor Linnaeus, 1758 | | Aphodius (Trichonotulus) scrofa (Fabricius, 1787) | ⇒ Trichonotulus scrofa Fabricius, 1787 | | Aphodius (Volinus) sticticus Panzer, 1798 | | | Aphodius abeillei Sietti, 1903 | | | Aphodius annamariae Baraud, 1982 | | | Aphodius anteactus ♰ Krell, 2000 | | | Aphodius ardoei (Landin, 1954) | ⇒ Agoliinus nemoralis Erichson, 1848 | | Aphodius armaticeps Péringuey, 1901 | | | Aphodius asellus Schmidt, 1907 | | | Aphodius baeticus Mulsant & Rey, 1870 | | | Aphodius baraudi (Villarreal, 1975) | ⇒ Euorodalus boiteli Théry, 1918 | | Aphodius bonvouloiri (Harold, 1860) | ⇒ Agolius bonvouloiri Harold, 1860 | | Aphodius boryslavicus ♰ A. M. Lomnicki, 1894 | | | Aphodius bosniaskii ♰ A. Handlirsch, 1907 | | | Aphodius brevipennis ♰ (O. Heer, 1862) | ⇒ Aphodius foetidus Herbst, 1783 | | Aphodius caminarius (Faldermann, 1835) | ⇒ Pseudacrossus tenebricosus Schmidt, 1922 | | Aphodius cardinalis (Reitter, 1892) | ⇒ Aphodius fimetarius Linnaeus, 1758 | | Aphodius charauxi ♰ L. Piton, 1940 | | | Aphodius circumcinctus W. L. E. Sch..., 1840 | | | Aphodius citellorum Semenov & S...., 1929 | | | Aphodius conigatus Panzer, 1795 | | | Aphodius connexus (Klug, 1855) | ⇒ Trichaphodius copulatus Schmidt, 1916 | | Aphodius cylindricus (Dejean, 1833) | ⇒ Mendidaphodius linearis Reiche & Saulcy, 1856 | | Aphodius dubiosus Péringuey, 1901 | | | Aphodius equestris (Panzer, 1798) | ⇒ Volinus sticticus Panzer, 1798 | | Aphodius fimitarius C. Linnaeus, 1758 | | | Aphodius gracilis Boheman, 1857 | | | Aphodius guillebeaui (Reitter, 1891) | ⇒ Melinopterus guillebeaui Reitter, 1892 | | Aphodius helvolus ♰ G. Statz, 1952 | | | Aphodius holdereri Reitter, 1900 | | | Aphodius inquinatus (Creutzer, 1799) | ⇒ Aphodius distinctus O.F. Müller, 1776 | | Aphodius intercalaris Péringuey, 1901 | | | Aphodius krantzi ♰ von Heyden &..., 1866 | | | Aphodius limbatus (Germar, 1824) | ⇒ Rhodaphodius foetens Fabricius, 1787 | | Aphodius marginalis (Stephens, 1830) | ⇒ Melinopterus sphacelatus Panzer, 1798 | | Aphodius meyeri ♰ Heer, 1847 | | | Aphodius nanus (Horn, 1887) | ⇒ Aphodius asellus Schmidt, 1907 | | Aphodius nitidus (Ballion, 1870) | ⇒ Plagiogonus arenarius Olivier, 1789 | | Aphodius palmetincolus (Karsch, 1881) | ⇒ Mendidius palmetincola Karsch, 1881 | | Aphodius rhinocerontis ♰ A. M. Lomnicki, 1894 | | | Aphodius rhododactylus (Marsham, 1802) | ⇒ Aphodius arenarius Olivier, 1789 | | Aphodius ruthenus ♰ A. M. Lomnicki, 1894 | | | Aphodius scalaris (Dellacasa, 1983) | ⇒ Chilothorax conspurcatus Linnaeus, 1758 | | Aphodius scapularis (Mulsant, 1842) | ⇒ Euorodalus coenosus Panzer, 1798 | | Aphodius schlickumi ♰ G. Statz, 1952 | | | Aphodius sharpi Harold, 1874 | | | Aphodius subater ♰ A. M. Lomnicki, 1894 | | | Aphodius sus Kugelann, 1792 | | | Aphodius tenella (Say, 1823) | ⇒ Planolinus tenellus Say, 1823 | | Aphodius testudinarius Fabricius, 1775 | | | Aphodius theobaldi ♰ Krell, 2000 | | | Aphodius tristis Zenker, 1801 | | | Aphodius vectis Krell, 2019 | | | Aphodius villarreali Baraud, 1975 | | | Aphodoharmogaster costipennis Boheman, 1857 | | | Aphotaenius carolinus Van Dyke, 1928 | | | Aschnarhyparus peregrinus Hinton, 1934 | | | Ataeniopsis duncani Cartwright, 1974 | | | Ataeniopsis figurator Harold, 1874 | | | Ataeniopsis parkeri Cartwright, 1974 | | | Ataeniopsis rugopygus Cartwright, 1974 | | | Ataenius alternatus Melsheimer, 1845 | | | Ataenius apicalis Hinton, 1937 | Red pricklypoppy | | Ataenius basiceps Lea, 1923 | | | Ataenius brevicollis Wollaston, 1854 | | | Ataenius californicus Horn, 1887 | | | Ataenius carinator Harold, 1874 | | | Ataenius cognatus (LeConte, 1858) | ⇒ Ataenius liogaster Bates, 1887 | | Ataenius communis Hinton, 1936 | | | Ataenius complicatus Harold, 1869 | | | Ataenius damzeni Bukejs & Ale..., 2018 | | | Ataenius exiguus Brown, 1932 | | | Ataenius glabriventris Schmidt, 1911 | | | Ataenius gracilis Melsheimer, 1845 | | | Ataenius heinekeni Wollaston, 1854 | | | Ataenius horticola Harold, 1869 | | | Ataenius imbricatus Melsheimer, 1845 | | | Ataenius insculptus Horn, 1887 | | | Ataenius languidus Schmidt, 1911 | Antarctic dragonfishes | | Ataenius liogaster Bates, 1887 | | | Ataenius luteomargo Chapin, 1940 | | | Ataenius nugator Harold, 1880 | | | Ataenius opatroides Blanchard, 1846 | | | Ataenius ovatulus Horn, 1871 | Scratchbush | | Ataenius parvus Lea, 1923 | | | Ataenius picinus Harold, 1861 | Antarctic blennies | | Ataenius platensis Blanchard, 1846 | | | Ataenius pseudousingeri Galante, Ste..., 2003 | | | Ataenius puncticollis (LeConte, 1868) | ⇒ Parapsammodius puncticollis LeConte, 1858 | | Ataenius robustus Horn, 1871 | Scratchbush | | Ataenius scabrellus Schmidt, 1909 | Scarab beetle | | Ataenius scalptifrons Bates, 1887 | | | Ataenius simplicipes Mulsant & Rey, 1870 | | | Ataenius spretulus Haldeman, 1848 | Black turfgrass ataenius | | Ataenius stercorator (Fabricius, 1775) | ⇒ Ataenius strigatus Say, 1823 | | Ataenius strigatus Say, 1823 | Black turfgrass ataenius | | Ataenius terminalis (Chevrolat, 1864) | ⇒ Ataenius luteomargo Chapin, 1940 | | Auperia denominata Chevrolat, 1864 | | | Australaphodius frenchi Blackburn, 1892 | Little pasture cockchafer | | Biralus mahunkaorum Adám, 1983 | | | Biralus menetriesi Ménétriés, 1849 | | | Biralus satellitius Herbst, 1789 | | | Blackburneus morogoroensis Petrovitz, 1969 | | | Blackburneus stercorosus Melsheimer, 1845 | | | Bodiloides ictericus Laicharting, 1781 | | | Bodiloides ictericus ghardimaouensis Balthasar, 1929 | | | Bodiloides ictericus ictericus Laicharting, 1781 | | | Bodilopsis aquila Schmidt, 1916 | | | Bodilopsis aquilus Schmidt, 1916 | | | Bodilopsis rufa Moll, 1782 | | | Bodilopsis rufus Moll, 1782 | | | Bodilopsis sordida Fabricius, 1775 | | | Bodilopsis sordida sordida Fabricius, 1775 | | | Bodilopsis sordidus Fabricius, 1775 | | | Bodilopsis sordidus sordidus | | | Bodilus barbarus Fairmaire, 1860 | | | Bodilus beduinus Reitter, 1892 | | | Bodilus caspius Menestries, 1832 | | | Bodilus circumcinctus Schmidt, 1840 | | | Bodilus crimensis Koshantschikov, 1913 | | | Bodilus gregarius Harold, 1871 | | | Bodilus ictericus (Laicharting, 1781) | ⇒ Bodiloides ictericus Laicharting, 1781 | | Bodilus ictericus ictericus (Laicharting, 1781) | ⇒ Bodiloides ictericus ictericus Laicharting, 1781 | | Bodilus immundus | | | Bodilus longispina Küster, 1854 | | | Bodilus lugens Creutzer, 1799 | | | Bodilus punctipennis Erichson, 1848 | | | Bodilus rufus | | | Bodilus sordidus | | | Bodilus wollastoni Harold, 1862 | | | Brindalus maderae Pittino, 1983 | | | Brindalus porcicollis Illiger, 1803 | | | Brindalus rotundipennis Reitter, 1892 | | | Brindalus schatzmayri Pittino, 1980 | | | Caelius rufescens Horn, 1887 | | | Calamosternus algiricus Mariani & Pi..., 1983 | | | Calamosternus granarius Linnaeus, 1767 | Granular Small Dung Beetle | | Calamosternus hyxos Petrovitz, 1962 | | | Calamosternus hyxos algiricus (Mariani & Pi..., 1983) | ⇒ Calamosternus algiricus Mariani & Pi..., 1983 | | Calamosternus mayeri Pilleri, 1953 | | | Calamosternus trucidatus Harold, 1863 | | | Calamosternus unicolor Olivier, 1789 | | | Chilothorax aljibei Blanco, 1986 | | | Chilothorax brancoi Baraud, 1981 | | | Chilothorax cervorum Fairmaire, 1871 | | | Chilothorax clathratus Reitter, 1892 | | | Chilothorax conspurcatus Linnaeus, 1758 | | | Chilothorax distinctus (Müller, 1776) | ⇒ Aphodius distinctus O.F. Müller, 1776 | | Chilothorax distinctus distinctus O. F. Müller, 1776 | | | Chilothorax exclamationis Motschulsky, 1849 | | | Chilothorax exilimanus Kabakov, 1996 | | | Chilothorax hahni Reitter, 1907 | | | Chilothorax hieroglyphicus Klug, 1845 | | | Chilothorax ivanovi Lebedev, 1911 | | | Chilothorax jacobsoni Koshantschikov, 1911 | | | Chilothorax lineolatus Illiger, 1803 | | | Chilothorax melanostictus Schmidt, 1840 | | | Chilothorax melanostictus melanostictus W. Schmidt, 1840 | | | Chilothorax melanostictus simulans Koshantschikov, 1913 | | | Chilothorax melanosticus | | | Chilothorax paykulli Bedel, 1907 | | | Chilothorax pictus Sturm, 1805 | | | Chilothorax planus Koshantschikov, 1894 | | | Chilothorax propola Balthasar, 1946 | | | Chilothorax pustulifer Reitter, 1892 | | | Chilothorax sesquivittatus Fairmaire, 1883 | | | Chilothorax sticticus (Panzer, 1798) | ⇒ Volinus sticticus Panzer, 1798 | | Chiron cylindrus Fabricius, 1798 | | | Cinacanthus anomalipus Péringuey, 1901 | | | Cnemisus rufescens Motschulsky, 1845 | | | Colobopterus erraticus Linnaeus, 1758 | Erratic small dung beetle | | Colobopterus scrutator | | | Colobopterus subterraneus Linnaeus, 0 | | | Coprimorphus scrutator Herbst, 1789 | | | Coptochiroides subcostatus Kolbe, 1886 | | | Coptochirus brachypterus Harold, 1868 | | | Coptochirus excisus Harold, 1868 | | | Dellacasiellus fucosus Schmidt, 1922 | Common sand stargazer | | Dialytes striatulus Say, 1825 | | | Dialytes truncatus Melsheimer, 1845 | | | Dialytes ulkei Horn, 1875 | | | Diapterna omissa LeConte, 1850 | | | Diastictus tibialis Fabricius, 1798 | | | Diastictus vulneratus Sturm, 1805 | | | Didactylia notata (Harold, 1859) | ⇒ Aidophus notatus Harold, 1859 | | Drepanocanthoides walshii Horn, 1870 | | | Drepanocanthus eximius (Péringuey, 1901) | ⇒ Pseudoxyomus eximius Péringuey, 1901 | | Drepanocanthus lineatus Wiedemann, 1823 | | | Eremazus unistriatus Mulsant, 1851 | | | Erytus aequalis Schmidt, 1907 | | | Erytus cognatus Fairmaire, 1860 | | | Erytus opacior Koshantschikov, 1894 | | | Erytus pruinosus Reitter, 1892 | | | Esymus merdarius Fabricius, 1775 | | | Esymus pusillus Herbst, 1789 | | | Esymus pusillus pusillus Herbst, 1789 | | | Esymus pyreti Penecke, 1911 | | | Esymus sculpturatus Reitter, 1892 | | | Esymus sesquivittatus Fairmaire, 1883 | | | Esymus suturinigra Schmidt, 1916 | | | Eudolus quadriguttatus Herbst, 1783 | | | Eudolus quadrinaevulus Reitter, 1892 | | | Euheptaulacus carinatus Germar, 1824 | | | Euheptaulacus carinatus brutius Luigioni, 1920 | | | Euheptaulacus carinatus carinatus Germar, 1824 | | | Euheptaulacus carinatus esuriens Dellacasa, 1983 | | | Euheptaulacus porcellus Frivaldszky, 1879 | | | Euheptaulacus sus Herbst, 1783 | | | Euheptaulacus villosus Gyllenhal, 1806 | | | Euorodalus baraudi (Villarreal, 1975) | ⇒ Euorodalus boiteli Théry, 1918 | | Euorodalus boiteli Théry, 1918 | | | Euorodalus coenosus Panzer, 1798 | | | Euorodalus paracoenosus Balthasar & ..., 1960 | | | Euorodalus tersus Erichson, 1848 | | | Euparixoides cribratus Hinton, 1936 | | | Eupleurus subterraneus Linnaeus, 1758 | Subterranean small dung beetle | | Eupleurus subterraneus subterraneus Linnaeus, 1758 | | | Geopsammodius relictillus Deyrup & Woo..., 1991 | Redspeckled blenny | | Gonaphodiellus hoffmanni Islas, 1945 | | | Harmogaster exarata Harold, 1861 | | | Harmogaster intrusus | | | Harmogaster spectabilis | | | Harmogaster transitoria Péringuey, 1901 | | | Haroldiataenius saramari Cartwright, 1939 | | | Haroldiellus sallei Harold, 1863 | | | Heptaulacus algarbiensis Branco & Baraud, 1984 | | | Heptaulacus alpinus Drapiez, 1819 | | | Heptaulacus brancoi Baraud, 1976 | | | Heptaulacus carinatus (Gebler, 1833) | ⇒ Euheptaulacus carinatus Germar, 1824 | | Heptaulacus gadetinus Baraud, 1973 | | | Heptaulacus rasettii Carpaneto, 1978 | | | Heptaulacus sus (Herbst, 1783) | ⇒ Euheptaulacus sus Herbst, 1783 | | Heptaulacus testudinarius Fabricius, 1775 | | | Heptaulacus villosus (Gyll., 1806) | ⇒ Euheptaulacus villosus Gyllenhal, 1806 | | Labarrus lividus Olivier, 1789 | Scarab beetle | | Labarrus lividus lividus | | | Labarrus pseudolividus Balthasar, 1941 | | | Lechorodius terminalis Say, 1823 | | | Leiopsammodius belloi Pierotti, 1980 | | | Leiopsammodius haruspex Adám, 1980 | | | Leiopsammodius strumae Chromy, 1983 | | | Limarus maculatus Sturm, 1800 | | | Limarus zenkeri Germar, 1813 | | | Liothorax isikdagensis Balthasar, 1952 | | | Liothorax kraatzi Harold, 1868 | | | Liothorax lividus | | | Liothorax muscorum (Adám, 1994) | ⇒ Liothorax niger Illiger, 1798 | | Liothorax niger Illiger, 1798 | Beaulieu dung beetle | | Liothorax plagiatus Linnaeus, 1767 | | | Liothorax rusakovi Gusakov, 2004 | | | Liothorax sturmi | | | Liothorax varians | | | Loraphodius latisulcus Reitter, 1892 | | | Loraphodius suarius Faldermann, 1835 | | | Loraspis frater Mulsant & Rey, 1870 | | | Mecynodes leucopterus Klug, 1845 | | | Mecynodes striatulus Waltl, 1835 | | | Megatelus contractus Klug, 1845 | | | Melinopterus abeillei Sietti, 1903 | | | Melinopterus circumcintus | | | Melinopterus consputus Creutzer, 1799 | | | Melinopterus dellacasai Avila, 1986 | | | Melinopterus femoralis Say, 1823 | | | Melinopterus guillebeaui Reitter, 1892 | | | Melinopterus prodromus Brahm, 1790 | Spring small dung beetle | | Melinopterus pubescens Sturm, 1800 | | | Melinopterus punctatosulcatus Sturm, 1805 | | | Melinopterus punctatosulcatus hirtipes Fischer von ..., 1844 | | | Melinopterus punctatosulcatus ponticus Petrovitz, 1967 | | | Melinopterus punctatosulcatus punctatosulcatus Sturm, 1805 | | | Melinopterus reyi Reitter, 1892 | | | Melinopterus sphacelatus Panzer, 1798 | | | Melinopterus stolzi Reitter, 1906 | | | Melinopterus tingens Reitter, 1892 | | | Melinopterus villarreali Baraud, 1975 | | | Mendidaphodius armiger Harold, 1871 | | | Mendidaphodius linearis Reiche & Saulcy, 1856 | | | Mendidaphodius paganettii Petrovitz, 1963 | | | Mendidius curtulus Harold, 1866 | | | Mendidius ivanovi (Lebedev, 1911) | ⇒ Chilothorax ivanovi Lebedev, 1911 | | Mendidius multiplex Reitter, 1897 | | | Mendidius palmetincola Karsch, 1881 | | | Mendidius palmetincolus Karsch, 1881 | | | Mesontoplatys arabicus Harold, 1875 | | | Mesontoplatys dorsalis Klug, 1855 | | | Mothon sarmaticus Semenov & Me..., 1927 | | | Neagolius amblyodon Daniel, 1900 | | | Neagolius bilimecki Seidlitz, 1891 | | | Neagolius bilimeckii Seidlitz, 1891 | | | Neagolius heydeni Harold, 1871 | | | Neagolius liguricus Daniel, 1902 | | | Neagolius limbolarius (Semenov, 1902) | ⇒ Neagolius bilimecki Seidlitz, 1891 | | Neagolius montanus Erichson, 1848 | | | Neagolius montivagus Erichson, 1848 | | | Neagolius penninus Daniel, 1902 | | | Neagolius pollicatus Erichson, 1848 | | | Neagolius praecox Erichson, 1848 | | | Neagolius schlumbergeri Seidlitz, 1891 | | | Neagolius schlumbergeri consobrinus Daniel, 1900 | | | Neagolius schlumbergeri samniticus Daniel, 1902 | | | Neagolius schlumbergeri schlumbergeri Seidlitz, 1888 | | | Neagolius schlumbergeri temperei Hoffmann, 1938 | | | Nialaphodius nigrita Fabricius, 1801 | | | Nialaphodius nigritus Fabricius, 1801 | | | Nialus motschulskyi Nikritin, 1969 | | | Nialus varians Duftschmid, 1805 | | | Nialus venyigei Endrödi, 1969 | | | Nimbus affinis Panzer, 1823 | | | Nimbus affinis affinis | | | Nimbus cartalinius Olsoufieff, 1918 | | | Nimbus contaminatus Herbst, 1783 | | | Nimbus franzinii Pittino, 1978 | | | Nimbus hoberlandti Tesar, 1945 | | | Nimbus johnsoni Baraud, 1976 | | | Nimbus lederi Harold, 1876 | | | Nimbus marianii Pittino, 1978 | | | Nimbus obliteratus Panzer, 1823 | | | Nimbus orbignyi Clouët, 1896 | | | Nimbus proximus Adám, 1994 | | | Nimbus richardi Veiga, 1984 | | | Nobiellus bonnairei Reitter, 1892 | | | Nobiellus pustulifer Reitter, 1892 | | | Nobius bonnairei (Reitter, 1892) | ⇒ Nobiellus bonnairei Reitter, 1892 | | Nobius gresseri Semenov, 1899 | | | Nobius hepaticus Roth, 1851 | | | Nobius rhodiensis Baraud, 1976 | | | Nobius serotinus Panzer, 1799 | | | Odontolytes denominatus Chevrolat, 1864 | | | Odontolytes landai Balthasar, 1963 | | | Orodaliscus rotundangulus Reitter, 1900 | | | Orodaliscus spalacophilus Novikov, 1996 | | | Orodaliscus zangi Schmidt, 1906 | | | Oromus alpinus Scopoli, 1763 | | | Oromus corvinus | | | Oromus leisteri Medvedev, 1968 | | | Oscarinus rusicola Melsheimer, 1845 | | | Oscarinus stuessyi Gordon & Ske..., 2007 | Lesser weever | | Osmanius balthasari Petrovitz, 1963 | | | Otophorus haemorrhoidalis Linnaeus, 1758 | Bloody small dung beetle | | Oxyomoides sepultus Cartwright, 1944 | | | Oxyomus silvestris Scopoli, 1763 | | | Oxyomus sylvestris Scopoli, 1763 | Forest small dung beetle | | Parabodilus wollastoni Harold, 1862 | | | Paracoptochirus kozanensis Pittino, 2001 | | | Paracoptochirus singularis Harold, 1868 | | | Paracoptochirus vignai Carpaneto & ..., 1990 | | | Paracrossidius radekcervenkai Cervenka, 2017 | | | Parammoecius asphaltinus Kolenati, 1846 | | | Parammoecius corvinus Erichson, 1848 | | | Parammoecius gibbus Germar, 1817 | | | Parammoecius osseticus Iablokov-Khn..., 1972 | | | Parammoecius pyrenaeus Jacquelin du..., 1863 | | | Parapsammodius puncticollis LeConte, 1858 | Berg-salmon gill worm | | Parataenius simulator Harold, 1868 | | | Pardalosus pardalis LeConte, 1857 | | | Passaliolla cossonoides Bates, 1887 | | | Phaeaphodius costalis Gebler, 1848 | | | Phaeaphodius dauricus Harold, 1863 | | | Phaeaphodius jouravliowi Reitter, 1907 | | | Phaeaphodius novikovi Kabakov, 1998 | | | Phaeaphodius pupillus Balthasar, 1961 | | | Phaeaphodius rectus Motschulsky, 1866 | | | Phaeaphodius roddi Koshantschikov, 1911 | | | Phalacronothus biguttatus Germar, 1824 | | | Phalacronothus citellorum Semenov & Me..., 1929 | | | Phalacronothus coenosus | | | Phalacronothus fumigatulus Reitter, 1892 | | | Phalacronothus paracoenosus | | | Phalacronothus pusillus | | | Phalacronothus putoni Reitter, 1894 | | | Phalacronothus quadriguttatus | | | Phalacronothus quadrimaculatus Linnaeus, 1761 | | | Phalacronothus suturinigra Schmidt, 1917 | | | Pharaphodius impurus Roth, 1851 | | | Pharaphodius marginellus Fabricius, 1781 | | | Plagiogonus arenarius Olivier, 1789 | | | Plagiogonus nanus Fairmaire, 1860 | | | Plagiogonus putridus (Geoffroy, 1785) | ⇒ Plagiogonus arenarius Olivier, 1789 | | Plagiogonus syriacus Harold, 1863 | | | Planolinellus vittatus Say, 1825 | | | Planolinoides aenictus Cooper & Gordon, 1987 | Benttooths | | Planolinoides borealis Gyllenhal, 1827 | | | Planolinus borealis (Gyllenhal, 1827) | ⇒ Planolinoides borealis Gyllenhal, 1827 | | Planolinus fasciatus Olivier, 1789 | | | Planolinus tenellus Say, 1823 | | | Planolinus uliginosus (Hardy, 1847) | ⇒ Planolinus fasciatus Olivier, 1789 | | Planolinus vittatus Say, 1825 | | | Platytomus gregalis Cartwright, 1948 | | | Platytomus jailensis Apostolov & ..., 1986 | | | Platytomus laevistriatus Perris, 1869 | | | Platytomus parvulus Chevrolat, 1864 | Dung beetle | | Platytomus sabulosus (Mulsant, 1842) | ⇒ Platytomus tibialis Fabricius, 1798 | | Platytomus tibialis Fabricius, 1798 | | | Platytomus variolosus Kolenati, 1846 | | | Pleuraphodius teter Roth, 1851 | | | Pleurophorus anatolicus Petrovitz, 1961 | | | Pleurophorus apicipennis Reitter, 1892 | | | Pleurophorus arabicus Pittino & Ma..., 1986 | | | Pleurophorus caesus Creutzer, 1796 | Cut small dung beetle | | Pleurophorus capicola Péringuey, 1901 | | | Pleurophorus maghrebinicus Pittino & Ma..., 1986 | | | Pleurophorus mediterranicus Pittino & Ma..., 1986 | | | Pleurophorus opacus Reitter, 1892 | | | Pleurophorus pannonicus Petrovitz, 1961 | | | Podotenus suberosus Blackburn, 1904 | | | Psammodius asper (Fabricius, 1775) | ⇒ Psammodius sulcicollis Illiger, 1802 | | Psammodius asperoides Pittino, 2007 | | | Psammodius basalis Mulsant & Rey, 1870 | Crested blenny | | Psammodius boreoitalicus Mencl, Král ..., 2022 | | | Psammodius bulgaricus Mencl, 1982 | | | Psammodius danubialis Adám, 1989 | | | Psammodius laevipennis Costa, 1844 | Awkward-winged small dung beetle | | Psammodius nocturnus Reitter, 1892 | | | Psammodius pierottii Pittino, 1979 | Mud blenny | | Psammodius plicicollis Erichson, 1848 | | | Psammodius porcicollis (Fairmaire, 1870) | ⇒ Platytomus laevistriatus Perris, 1869 | | Psammodius rotundipennis (Reitter, 1892) | ⇒ Brindalus rotundipennis Reitter, 1892 | | Psammodius schmidti Gistel, 1857 | | | Psammodius schmidtii Gistel, 1857 | | | Psammodius sulcicollis Illiger, 1802 | | | Psammoporus insularis Pittino, 2006 | | | Psammoporus lacustris LeConte, 1850 | | | Psammoporus latipunctus Gredler, 1866 | | | Psammoporus mimica Pittino, 2006 | | | Psammoporus mimicus Pittino, 2006 | | | Psammoporus sabuleti (Panzer, 1797) | ⇒ Psammoporus mimicus Pittino, 2006 | | Psammoporus terminalis Brown, 1931 | | | Pseudacrossus caspius Menetries, 1832 | | | Pseudacrossus circassicus Reitter, 1892 | | | Pseudacrossus cribricollis (Lucas, 1846) | ⇒ Amidorus cribricollis Lucas, 1846 | | Pseudacrossus koshantschikovi Jacobson, 1911 | | | Pseudacrossus przewalskyi Reitter, 1887 | | | Pseudacrossus sharpi Harold, 1874 | | | Pseudacrossus spalacophilus Novikov, 1996 | | | Pseudacrossus subsericeus Ballion, 1878 | | | Pseudacrossus suffertus Schmidt, 1922 | | | Pseudacrossus suffertus ampliatus Reitter, 1892 | | | Pseudacrossus suffertus defiorei Fiori, 1907 | | | Pseudacrossus suffertus suffertus | | | Pseudacrossus tenebricosus Schmidt, 1922 | | | Pseudacrossus zangi A.Schmidt, 1906 | | | Pseudesymus lucidus Klug, 1845 | | | Pubinus tomentosus O.F.Muller, 1776 | | | Rhodaphodius foetens Fabricius, 1787 | | | Rhysothorax rufa Fabricius, 1792 | | | Rhysothorax rufus Fabricius, 1792 | Red sand scarab beetle | | Rhyssemodes orientalis Mulsant & Go..., 1874 | | | Rhyssemus algiricus Lucas, 1846 | | | Rhyssemus algiricus marqueti (Reiche, 1863) | ⇒ Rhyssemus marqueti Reiche, 1863 | | Rhyssemus algiricus meridionalis Reitter, 1890 | | | Rhyssemus annaedicatus Pierotti, 1980 | | | Rhyssemus arenarius (A. Costa, 1844) | ⇒ Rhyssemus plicatus Germar, 1817 | | Rhyssemus asper Fabricius, 1775 | | | Rhyssemus berytensis Marseul, 1878 | | | Rhyssemus cantabricus (Balthasar, 1961) | ⇒ Rhyssemus germanus Linnaeus, 1767 | | Rhyssemus convexus Benard, 1911 | | | Rhyssemus germanus Linnaeus, 1767 | Germany small dung beetle | | Rhyssemus insularis Pittino, 1983 | | | Rhyssemus interruptus Reitter, 1892 | | | Rhyssemus limbolarius Petrovitz, 1963 | | | Rhyssemus limborarius (Petrovitz, 1963) | ⇒ Rhyssemus limbolarius Petrovitz, 1963 | | Rhyssemus macedonicus Benard, 1923 | | | Rhyssemus marqueti Reiche, 1863 | | | Rhyssemus mayeti Clouët, 1901 | | | Rhyssemus meridionalis (Reitter, 1890) | ⇒ Rhyssemus algiricus meridionalis Reitter, 1890 | | Rhyssemus parallelus Reitter, 1892 | | | Rhyssemus pertinax (Balthasar, 1961) | ⇒ Rhyssemus germanus Linnaeus, 1767 | | Rhyssemus plicatus Germar, 1817 | | | Rhyssemus promontorii Péringuey, 1901 | | | Rhyssemus psammobiiformis Petrovitz, 1963 | | | Rhyssemus puncticollis (Brown, 1929) | ⇒ Rhyssemus germanus Linnaeus, 1767 | | Rhyssemus purkynei Balthasar, 1961 | | | Rhyssemus sardous Pierotti, 1980 | | | Rhyssemus sonatus LeConte & Horn, 1881 | | | Rhyssemus sulcatus Olivier, 1789 | | | Rhyssemus verrucosus Mulsant, 1842 | | | Rhyssemus vinodolensis Petrovitz, 1963 | | | Rugaphodius rugatus Schmidt, 1907 | Spotted weever | | Saprosites cascus ♰ E. B. Britton, 1960 | | | Saprosites excisus Petrovitz, 1976 | | | Saprosites mendax Blackburn, 1892 | | | Saprosites natalensis Péringuey, 1901 | | | Saprosites succini ♰ R. Zang, 1905 | | | Scabrostomus tanytarsus Skelley & Wo..., 1991 | Masked stargazer | | Sigorus porcus Fabricius, 1792 | Mahogany Ridgeback scarab | | Sigorus porcus ruficrus Marsham, 1802 | | | Stenotothorax badipes Melsheimer, 1845 | | | Stenotothorax gentilis Horn, 1887 | | | Stenotothorax schneppi Skelley & Mc..., 2018 | | | Subrinus sturmi Harold, 1870 | | | Subrinus subtilis Koshantschikov, 1894 | | | Subrinus vitellinus Klug, 1845 | | | Tesarius caelatus LeConte, 1857 | | | Tesarius mcclayi Cartwright, 1955 | | | Tetraclipeoides quadridentatus Harold, 1861 | | | Teuchestes brachysomus Solsky, 1874 | | | Teuchestes fossor Linnaeus, 1758 | Digger small dung beetle | | Teuchestes haemorrhoidalis | | | Trichaphodius copulatus Schmidt, 1916 | | | Trichiorhyssemus dalmatinus Petrovitz, 1967 | | | Trichiorhyssemus setulosus Reitter, 1892 | | | Trichonotulus pilosellus Schmidt, 1907 | | | Trichonotulus scrofa Fabricius, 1787 | Sow small dung beetle | | Trigonoscelus hypi Kral, 2017 | | | Volinus sticticus Panzer, 1798 | | | Xeropsamobeus asellus Schmidt, 1907 | Spotted weever |
Subnodes of Aphodiinae with occurrences in Europe
Links, sources and further information Aphodiinae at artfakta.se
Aphodiinae at dyntaxa.se
Aphodiinae at itis.gov
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