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Checklist for Subfamily Aphodiinae Leach, 1815 (Aphodiine Dung Beetles)
and region Far East (Asia)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
1257
36320
255777
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
1251
36273
255683
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
1149
35660
253017
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
671
26495
204903
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
186
10830
97491
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
152
9729
84162
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
18
1072
13577
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
18
1072
13577
Families
Genera
Species
 
Taxom
 
FamilyScarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles
797
10488
Genera
Species
 
Taxom
 
SubfamilyAphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles
153
965
Genera
Species
 
Taxom
 


Genera for Far East
153 Genera with 965 Species
#ABCDEGHKLMNOPQRSTV
Acanthobodilus Dellacasa, 19222SpeciesTaxom
Acrossidius Schmidt, 18924SpeciesTaxom
Acrossus Mulsant, 196140SpeciesTaxom
Adebrattia Bordat & Howden, 19951SpeciesTaxom
Adeloparius Schmidt, 19421SpeciesTaxom
Aegialia Latreille, 19869SpeciesTaxom
Aganocrossus Reitter, 18625SpeciesTaxom
Agoliinus Schmidt, 198113SpeciesTaxom
Agolius Mulsant & Rey, 20143SpeciesTaxom
Agrilinellus Dellacasa, ..., 19111SpeciesTaxom
Agrilinus Mulsant & Rey, 196226SpeciesTaxom
Airapus Stebnicka & ..., 201920SpeciesTaxom
Alocoderus Schmidt, 19356SpeciesTaxom
Amidorus Mulsant & Rey, 19134SpeciesTaxom
Ammoecius Mulsant, 19602SpeciesTaxom
Aparammoecius Petrovitz, 201710SpeciesTaxom
Aphodaulacus Kozhantschikov, 18758SpeciesTaxom
Aphodiellus Schmidt, 18751SpeciesTaxom
Aphodiites Heer, 18651SpeciesTaxom
Aphodius Hellwig, 1951103SpeciesTaxom
Aphodobius Péringuey, 19451SpeciesTaxom
Ataenius Harold, 199824SpeciesTaxom
Australaphodius Balthasar, 18921SpeciesTaxom
Blackburneus Schmidt, 19081SpeciesTaxom
Bodiloides1SpeciesTaxom
Bodilopsis Adám, 07SpeciesTaxom
Bodilus Mulsant & Rey, 197510SpeciesTaxom
Boucomontiellus Balthasar, 19132SpeciesTaxom
Brachiaphodius Koshantschikov, 200011SpeciesTaxom
Bruneixenus Howden & Storey, 19922SpeciesTaxom
Caelius Lewis, 18872SpeciesTaxom
Calamosternus Motschulsky, 19947SpeciesTaxom
Carinaulus Tesar, 200018SpeciesTaxom
Cephalocyclus Dellacasa, ..., 18871SpeciesTaxom
Cheleion Vårdal & For..., 20193SpeciesTaxom
Chilothorax Motschulsky, 200135SpeciesTaxom
Chiron MacLeay, 18311SpeciesTaxom
Cnematoplatys A.Schmidt, 19673SpeciesTaxom
Cnemisus Motschulsky, 19033SpeciesTaxom
Colobopterus Mulsant, 18474SpeciesTaxom
Coprimorphus Mulsant, 17891SpeciesTaxom
Coptochiroides Balthasar, 04SpeciesTaxom
Craterocyphus Schmidt, 18472SpeciesTaxom
Cryptoscatomaseter Gordon and S..., 20011SpeciesTaxom
Danielssonia Bordat & Howden, 19951SpeciesTaxom
Dialytes Harold, 18452SpeciesTaxom
Diastictus Mulsant, 18051SpeciesTaxom
Diasticus Mulsant, 1842
0SpeciesTaxom
Didactylia Orbigny, 19611SpeciesTaxom
Emadiellus Schmidt, 03SpeciesTaxom
Eocorythoderus Maruyama, 20121SpeciesTaxom
Erytus Mulsant & Rey, 18923SpeciesTaxom
Esymus Mulsant & Rey, 19324SpeciesTaxom
Eudolus Mulsant & Rey, 17831SpeciesTaxom
Euheptaulacus Dellacasa, 18064SpeciesTaxom
Euorodalus Dellacasa, 19452SpeciesTaxom
Eupleurus Mulsant, 17583SpeciesTaxom
Gilletianus Balthasar, 201539SpeciesTaxom
Gonaphodius Reitter, 18771SpeciesTaxom
Granulopsammodius Rakovic, 19844SpeciesTaxom
Guanyinaphodius1SpeciesTaxom
Heptaulacus Mulsant, 17752SpeciesTaxom
Koreoxyomus1SpeciesTaxom
Koshantschikovius Schmidt, 19831SpeciesTaxom
Labarrus Mulsant and Rey, 18606SpeciesTaxom
Leiopsammodius Rakovic, 20128SpeciesTaxom
Limarus Mulsant & Rey, 18132SpeciesTaxom
Liothorax Motschulsky, 17673SpeciesTaxom
Loraphodius Reitter, 18781SpeciesTaxom
Loraspis Mulsant & Rey, 18701SpeciesTaxom
Megateloides Landin, 18601SpeciesTaxom
Megatelus Reitter, 18791SpeciesTaxom
Melinopterus Mulsant, 17988SpeciesTaxom
Mendidius Harold, 19418SpeciesTaxom
Mesontoplatys Motschulsky, 19812SpeciesTaxom
Messyrhus Rakovič & K..., 19971SpeciesTaxom
Microtermitodius Pittino, 20061SpeciesTaxom
Mozartius Nomura & Nakane, 011SpeciesTaxom
Myrhessus Balthasar, 20065SpeciesTaxom
Neagolius Kozhantschikov, 199710SpeciesTaxom
Neocalaphodius Bordat, 18012SpeciesTaxom
Neotrichiorhyssemus Rakovič & K..., 19689SpeciesTaxom
Nettelislasia Martínez, 19451SpeciesTaxom
Nialus Mulsant & Rey, 18052SpeciesTaxom
Nimbus Mulsant & Rey, 18233SpeciesTaxom
Nipponaphodius Nakane, 19511SpeciesTaxom
Nipponoagoliinus1SpeciesTaxom
Nobius Mulsant & Rey, 17992SpeciesTaxom
Odochilus Harold, 199724SpeciesTaxom
Orodaliscus Reitter, 19062SpeciesTaxom
Oromus Mulsant & Rey, 17631SpeciesTaxom
Otophorus Mulsant, 17581SpeciesTaxom
Oxyomus Dejean, 199116SpeciesTaxom
Palnia Stebnicka, 20202SpeciesTaxom
Paracrossidius Balthasar, 196310SpeciesTaxom
Paradidactylia Balthasar, 194110SpeciesTaxom
Parammoecius Seidlitz, 19604SpeciesTaxom
Paulianellus Balthasar, 20156SpeciesTaxom
Petrovitzius Rakovič, 1979
0SpeciesTaxom
Phaeaphodius Reitter, 18665SpeciesTaxom
Phalacronothus Motschulsky, 19525SpeciesTaxom
Pharaphodius Reitter, 194611SpeciesTaxom
Plagiogonus Mulsant, 195113SpeciesTaxom
Planolinellus Dellacasa an..., 18252SpeciesTaxom
Planolinoides Dellacasa an..., 18272SpeciesTaxom
Planolinus Mulsant and Rey, 19513SpeciesTaxom
Platyderides Schmidt, 19957SpeciesTaxom
Platytomus Mulsant, 20063SpeciesTaxom
Pleuraphodius Schmidt, 19937SpeciesTaxom
Pleurophorus Mulsant, 193313SpeciesTaxom
Proctophanes Harold, 19101SpeciesTaxom
Psammodius Fallen, 198016SpeciesTaxom
Psammoporus Thomson, 198611SpeciesTaxom
Pseudacrossus Reitter, 192213SpeciesTaxom
Pseudoahermodontus1SpeciesTaxom
Pseudoheptaulacus Dellacasa & ..., 20144SpeciesTaxom
Pseudostereomera Bordat & Howden, 19951SpeciesTaxom
Pterobius Maruyama, 20101SpeciesTaxom
Pubinus Mulsant & Rey, 17761SpeciesTaxom
Qingaphodius2SpeciesTaxom
Rakovicius Pittino, 19658SpeciesTaxom
Renaudius Balthasar, 19912SpeciesTaxom
Rhinocerotopsis Maruyama, 20091SpeciesTaxom
Rhodaphodius1SpeciesTaxom
Rhyparus Westwood, 201969SpeciesTaxom
Rhyssemodes Reitter, 20176SpeciesTaxom
Rhyssemus Mulsant, 198426SpeciesTaxom
Saprosites Redtenbacher, 201939SpeciesTaxom
Saprovisca2SpeciesTaxom
Sariangus Rakovič & M..., 20111SpeciesTaxom
Scabrostomus Gordon and S..., 19911SpeciesTaxom
Setylaides6SpeciesTaxom
Siamaphodius1SpeciesTaxom
Sigorus Mulsant & Rey, 17921SpeciesTaxom
Silluvia Landin, 20088SpeciesTaxom
Sinaphodius6SpeciesTaxom
Sinodiapterna Dellacasa, 19916SpeciesTaxom
Stenotothorax Schmidt, 02SpeciesTaxom
Stereomera Arrow, 19052SpeciesTaxom
Subrinus Mulsant & Rey, 18701SpeciesTaxom
Sugrames Reitter, 1894
0SpeciesTaxom
Sybacodes Fairmaire, 08SpeciesTaxom
Sybax Boheman, 18571SpeciesTaxom
Termitodiellus Nakane, 19877SpeciesTaxom
Termitodius Wasmann, 1894
0SpeciesTaxom
Termitopisthes Wasmann, 19112SpeciesTaxom
Termitotrox Reichensperger, 20153SpeciesTaxom
Teuchestes Mulsant, 20067SpeciesTaxom
Trichiorhyssemus Clouët, 200714SpeciesTaxom
Trichonotulus Bedel, 19416SpeciesTaxom
Tristaphodius Balthasar, 19322SpeciesTaxom
Turanella Semenov, 18871SpeciesTaxom
Volinus Mulsant & Rey, 17981SpeciesTaxom


Species list for Far East
965 Species and 46 Subspecies
Acanthobodilus immundus Creutzer, 1799
Acanthobodilus languidulus Schmidt, 1922
Acrossidius delavayi Koshantschikov, 1916
Acrossidius kozlovi Koshantschikov, 1916
Acrossidius tasmaniae Hope, 1847Black-headed pasture cockchafer
Acrossidius yorkensis Blackburn, 1892
Acrossus atratus Waterhouse, 1875
Acrossus belousovi Frolov, 2001
Acrossus binaevulus Heyden, 1887
Acrossus byki Minkina, 2018
Acrossus delavayi Koshantschikov, 1916
Acrossus depressus Kugelann, 1792
Acrossus formosanus Nomura & Nakane, 1951
Acrossus haliki Balthasar, 1933
Acrossus histrio Balthasar, 1932
Acrossus hrubanti Balthasar, 1961
Acrossus humerospinosus Petrovitz, 1958
Acrossus igai Nakane, 1956
Acrossus igai pseudoigai Nakane & Mas..., 1967
Acrossus igai sakuraii Nakane & Mas..., 1967
Acrossus igai tsurugianus Nakane & Mas..., 1967
Acrossus impressiusculus Fairmaire, 1888
Acrossus japonicus Nomura & Nakane, 1951
Acrossus jedlickai Balthasar, 1932
Acrossus jeloneki Minkina, 2018
Acrossus klickai Balthasar, 1932
Acrossus koreanensis Kim, 1986
Acrossus kwanhsiensis Balthasar, 1945
Acrossus laticapitus Zhang, 1992
Acrossus luridus Fabricius, 1775
Acrossus melanodiscus Zhang, 1992
Acrossus milloti Paulian, 1945
Acrossus obenbergeri Balthasar, 1932
Acrossus opacipennis Schmidt, 1910
Acrossus pseudoobenbergeri Cervenka, 1995
Acrossus ritsemai Schmidt, 1909
Acrossus rufipes Linnaeus, 1758Night-Flying Dung Beetle
Acrossus semiopacus Reitter, 1887
Acrossus sichuanensis Zhang, 1992
Acrossus superatratus Nomura & Nakane, 1951
Acrossus tiabi Masumoto, 1988
Acrossus transmontanus Balthasar, 1932
Acrossus unifasciatus Nomura & Nakane, 1951
Acrossus unifasciatus kiiensis Nakane, 1960
Acrossus viturati Reitter, 1907
Acrossus yunlingensis Petrovitz, 1961
Adebrattia depressa Bordat & Howden, 1995
Adeloparius formosus Paulian, 1942
Aegialia abdita Nikritin, 1975
Aegialia comis Lewis, 1895
Aegialia friebi (Balthasar, 1935)⇒ Psammoporus friebi Balthasar, 1935
Aegialia kamtschaticus Motschulsky, 1860
Aegialia koreana Kim, 1979
Aegialia nakanei Masumoto, 1986
Aegialia nakanei horii Ochi & Kawahara, 2012
Aegialia nitida Waterhouse, 1875
Aegialia tsukamotoi Masumoto, 1986
Aganocrossus amoenus Boheman, 1857
Aganocrossus bordatianus Masumoto & K..., 2003
Aganocrossus borneensis Petrovitz, 1962
Aganocrossus postpilosus Reitter, 1895
Aganocrossus urostigma Harold, 1862
Agoliinus anthracus Gordon & Ske..., 2007Lesser weever
Agoliinus cruentatus LeConte, 1878
Agoliinus kiuchii Masumoto, 1984
Agoliinus lapponum Gyllenhal, 1808Northern Dung Beetle
Agoliinus morii Nakane, 1983
Agoliinus parastorkani Cervenka, 1995
Agoliinus piceus Gyllenhal, 1808
Agoliinus pseudostorkani Stebnicka, 1982
Agoliinus satyrus Reitter, 1892
Agoliinus setchan Masumoto, 1984
Agoliinus shibatai Nakane, 1983
Agoliinus storkani Balthasar, 1932
Agoliinus tanakai Masumoto, 1981
Agolius abdominalis Bonelli, 1812
Agolius moronensis Frolov, 2009
Agolius petri Frolov, 2014
Agrilinellus ornatus Schmidt, 1911Banded pipefish
Agrilinus ater De Geer, 1774
Agrilinus bardus Balthasar, 1946
Agrilinus bollowi Balthasar, 1941
Agrilinus breviusculus Motschulsky, 1866
Agrilinus convexus Erichson, 1848
Agrilinus hasegawai Nomura & Nakane, 1951
Agrilinus hasegawai aka Nakane, 1960
Agrilinus hasegawai akahane Nakane, 1960
Agrilinus hayachinensis Nomura & Nakane, 1951
Agrilinus ishidai Masumoto & K..., 1987
Agrilinus lungaiensis Petrovitz, 1962
Agrilinus madara Nakane, 1960
Agrilinus matusiaki Minkina, 2022
Agrilinus monikae Kral, 2013
Agrilinus obliviosus Reitter, 1892
Agrilinus pseudolungaiensis Mencl & Rakovic, 2012
Agrilinus punctator Reitter, 1892
Agrilinus ritsukoae Kawai, 2004
Agrilinus sordidus (Fabricius, 1775)⇒ Bodilopsis sordida Fabricius, 1775
Agrilinus spinulosus Schmidt, 1910
Agrilinus striatus Schmidt, 1910
Agrilinus surdus Boucomont, 1929
Agrilinus tenax Balthasar, 1932
Agrilinus uniformis Waterhouse, 1875
Agrilinus virginalis Reitter, 1900
Agrilinus wassuensis Petrovitz, 1962
Airapus aegialiformis Minkina, 2020
Airapus burrundieae Stebnicka & ..., 1996
Airapus cechovskyi Král, Mencl ..., 2019
Airapus granulator Harold, 1877
Airapus halmaheraensis Minkina & Be..., 2023
Airapus hollandiae Stebnicka, 1998
Airapus idenburgii Paulian, 1937
Airapus jenisi Král, Mencl ..., 2019
Airapus nondugli Stebnicka, 1998
Airapus parvispinus Schmidt, 1920
Airapus popondettae Stebnicka, 1998
Airapus rakovici Král & Lu, 2021
Airapus rugulopunctatus Krikken, 1970
Airapus sicardi Paulian, 1945
Airapus spinator Harold, 1877
Airapus sulawesianus Rakovi�, Me..., 2021
Airapus sumatrae Fairmaire, 1896
Airapus tyri Král, Mencl ..., 2019
Airapus vangviengensis Minkina & Be..., 2023
Airapus wallaceanus Rakovi�, Me..., 2019
Alocoderus elongatulus Fabricius, 1801
Alocoderus holdereri Reitter, 1900
Alocoderus imitator Koshanschikov, 1916
Alocoderus luangensis Emberson & S..., 2001
Alocoderus semenowi Reitter, 1887
Alocoderus teyrovskyi Balthasar, 1935
Amidorus koshantschikovi Jacobson, 1911
Amidorus obscurus Fabricius, 1792
Amidorus serrimargo Koshantschikov, 1913
Amidorus umbrinus Koshantschikov, 1913
Ammoecius brevis Erichson, 1848
Ammoecius yamato Nakane, 1960
Aparammoecius balangensis Petrovitz, 1958
Aparammoecius chuangkuohungi Masumoto, Ki..., 2017
Aparammoecius gansuensis Stebnicka, 1997
Aparammoecius isaburoi Nakane, 1956
Aparammoecius mizo Nakane, 1967
Aparammoecius nomurai Stebnicka, 1986
Aparammoecius pallidiligonis Waterhouse, 1875
Aparammoecius tarokensis Stebnicka, 1994
Aparammoecius yenpingensis Stebnicka, 1986
Aparammoecius yushanus Masumoto, Ki..., 2017
Aphodaulacus grumi Frolov, 2002
Aphodaulacus ignobilis Reitter, 1887
Aphodaulacus kabaki Frolov, 2001
Aphodaulacus koltzei Reitter, 1892
Aphodaulacus nigrotessellatus Motschulsky, 1866
Aphodaulacus perblandus Balthasar, 1933
Aphodaulacus turkestanicus Heyden, 1881
Aphodaulacus variabilis Waterhouse, 1875
Aphodiellus impunctatus Waterhouse, 1875
Aphodiites protogaeus Heer, 1865
Aphodius (Acanthobodilus) immundus (Creutzer, 1799)⇒ Pubinus tomentosus O.F.Muller, 1776
Aphodius (Acrossoides) yorkensis (Blackburn, 1892)⇒ Acrossidius yorkensis Blackburn, 1892
Aphodius (Acrossus) formosanus Nomura & Nakane, 1951
Aphodius (Acrossus) rufipes (Linnaeus, 1758)⇒ Acrossus rufipes Linnaeus, 1758
Aphodius (Aganocrossus) urostigma Harold, 1862
Aphodius (Ahermodontus) marini Baguena, 1930
Aphodius (Aparammoecius) balangensis Petrovitz, 1958
Aphodius (Aparammoecius) nishikawai (Masumoto, 1996)⇒ Aparammoecius tarokensis Stebnicka, 1994
Aphodius (Aparammoecius) tarokensis Stebnicka, 1994
Aphodius (Aphodiellus) impunctatus (Waterhouse, 1875)⇒ Melinopterus consputus Creutzer, 1799
Aphodius (Aphodius) elegans Allibert, 1847
Aphodius (Aphodius) fimetarius Linnaeus, 1758Common dung beetle
Aphodius (Aphodius) foetidus Herbst, 1783
Aphodius (Aphodius) pedellus (DeGeer, 1774)⇒ Aphodius fimetarius Linnaeus, 1758
Aphodius (Aphodius) variabilis (C. O. Waterh..., 1875)⇒ Aphodaulacus nigrotessellatus Motschulsky, 1866
Aphodius (Balthasarianus) babori Balthasar, 1938
Aphodius (Balthasarianus) pilifer Paulian, 1934
Aphodius (Bodilus) gregarius Harold, 1871
Aphodius (Bodilus) punctipennis Erichson, 1848
Aphodius (Bodilus) sordidus (Fabricius, 1775)⇒ Bodilopsis rufus Moll, 1782
Aphodius (Calamosternus) granarius (Linnaeus, 1767)⇒ Calamosternus granarius Linnaeus, 1767
Aphodius (Calamosternus) sublimbatus Motschulsky, 1860
Aphodius (Calamosternus) trucidatus (Chevrolat, 1863)⇒ Calamosternus trucidatus Harold, 1863
Aphodius (Calaphodius) moestus (Fabricius, 1801)⇒ Calamosternus granarius Linnaeus, 1767
Aphodius (Carinaulus) pucholti Balthasar, 1961
Aphodius (Colobopteroides) latus A.Schmidt, 1911
Aphodius (Colobopterus) erraticus (Linnaeus, 1758)⇒ Colobopterus erraticus Linnaeus, 1758
Aphodius (Diapterna) hamatus Say, 1824
Aphodius (Euorodalus) pusillus (Herbst, 1789)⇒ Esymus pusillus Herbst, 1789
Aphodius (Eupleurus) subterraneus (Linnaeus, 1758)⇒ Eupleurus subterraneus Linnaeus, 1758
Aphodius (Gonaphodius) scoparius Harold, 1877
Aphodius (Loboparius) kotoensis Miwa, 1930
Aphodius (Melinopterus) prodromus (Braham, 1790)⇒ Melinopterus prodromus Brahm, 1790
Aphodius (Nialus) insularis (Petrovitz, 1961)⇒ Labarrus hoabinhensis Balthasar, 1946
Aphodius (Nialus) plagiatus (Linnaeus, 1758)⇒ Liothorax plagiatus Linnaeus, 1767
Aphodius (Nialus) sturmi (Harold, 1870)⇒ Subrinus sturmi Harold, 1870
Aphodius (Oromus) alpinus (Scopoli, 1763)⇒ Euheptaulacus carinatus carinatus Germar, 1824
Aphodius (Otophorus) haemorrhoidalis (Linnaeus, 1758)⇒ Otophorus haemorrhoidalis Linnaeus, 1758
Aphodius (Paremadus) isaburoi Nakane, 1967
Aphodius (Paulianellus) maderi Balthasar, 1938
Aphodius (Phaeaphodius) rectus Motschulsky, 1866
Aphodius (Phalacronothus) pseudoavunculus Masumoto, 1977
Aphodius (Pharaphodius) marginellus (Fabricius, 1781)⇒ Ataenius luteomargo Chapin, 1940
Aphodius (Pharaphodius) priscus Motschulsky, 1858
Aphodius (Plagiogonus) arenarius Olivier, 1789
Aphodius (Plagiogonus) putridus (Herbst, 1789)⇒ Planolinoides borealis Gyllenhal, 1827
Aphodius (Planolinellus) vittatus Mulsant, 1842
Aphodius (Planolinoides) borealis (Gyllenhal, 1827)⇒ Planolinoides borealis Gyllenhal, 1827
Aphodius (Pleuraphodius) lewisi C. O. Waterh..., 1875
Aphodius (Sinodiapterna) troitzkyi Jacobson, 1897
Aphodius (Sinodiapterna) zeni Ochi, 1991
Aphodius (Teuchestes) analis (Gistel, 1857)⇒ Rhodaphodius foetens Fabricius, 1787
Aphodius (Trichaphodius) reichei Harold, 1859
Aphodius (Trichonotulus) scrofa (Fabricius, 1787)⇒ Trichonotulus scrofa Fabricius, 1787
Aphodius anteactus Krell, 2000
Aphodius apicalis (Schilsky, 1888)⇒ Calamosternus granarius Linnaeus, 1767
Aphodius armatus (Roth, 1851)⇒ Craterocyphus rhinocerus Reiche, 1847
Aphodius australasiae (Boheman, 1858)⇒ Ataenius castanicolor Motschulsky, 1858
Aphodius bifasciatus (Dalla Torre, 1879)⇒ Chilothorax melanostictus Schmidt, 1840
Aphodius bonariensis Klug, 1829
Aphodius calichromus Balthasar, 1932
Aphodius caminarius (Faldermann, 1835)⇒ Pseudacrossus tenebricosus Schmidt, 1922
Aphodius cancelliventris (Motschulsky, 1858)⇒ Ataenius australasiae Boheman, 1858
Aphodius castanicolor (Motschulsky, 1858)⇒ Ataenius castanicolor Motschulsky, 1858
Aphodius catulus (Balthasar, 1946)⇒ Australaphodius frenchi Blackburn, 1892
Aphodius chokaiensis (Nomura & Nakane, 1951)⇒ Gilletianus comatus Schmidt, 1920
Aphodius corallifer Koshantschikov, 1913
Aphodius crenatus (Dejean, 1833)⇒ Pharaphodius crenatus Harold, 1862
Aphodius cruentatus (Dejean, 1833)⇒ Planolinellus vittatus Say, 1825
Aphodius ealaensis (Endrödi, 1956)⇒ Adeloparius formosus Paulian, 1942
Aphodius edai (Balthasar, 1956)⇒ Phaeaphodius rectus Motschulsky, 1866
Aphodius epotus (Dellacasa, 1983)⇒ Chilothorax conspurcatus Linnaeus, 1758
Aphodius fasciger Harold, 1881
Aphodius gestroi (Harold, 1877)⇒ Ataenius australasiae Boheman, 1858
Aphodius guangdongensis Maté, 2008
Aphodius haroldianus (Balthasar, 1932)⇒ Colobopterus quadratus Reiche, 1847
Aphodius hibernalis Nakane & Tsu..., 1956
Aphodius igai Nakane, 1956
Aphodius illigeri (Dallatorre, 1879)⇒ Bodilopsis rufus Moll, 1782
Aphodius informis (Mulsant, 1842)⇒ Acrossus luridus Fabricius, 1775
Aphodius irakensis (Baraud, 1977)⇒ Erytus opacior Koshantschikov, 1894
Aphodius irregularis (Westwood, 1839)⇒ Chilothorax paykulli Bedel, 1907
Aphodius jengi Masumoto, Ki..., 2014
Aphodius lapponicus (Villa, 1833)⇒ Oromus alpinus Scopoli, 1763
Aphodius madari (Tesar, 1945)⇒ Nimbus makowskyi Koshantschikov, 1891
Aphodius mara (Krikken & Kaas, 1984)⇒ Didactylia szunyoghyi Endrödi, 1961
Aphodius marginatus (LeConte, 1878)⇒ Cryptoscatomaseter depressiusculus Schmidt, 1907
Aphodius miniatulus (Balthasar, 1931)⇒ Agoliinus lapponum Gyllenhal, 1808
Aphodius mizo Nakane, 1967
Aphodius multiplex Reitter, 1897
Aphodius nasutus (Dellacasa, 1983)⇒ Chilothorax melanostictus Schmidt, 1840
Aphodius nigerrimus (Dallatorre, 1879)⇒ Oromus alpinus Scopoli, 1763
Aphodius nigrotessellatus (Nakane, 1972)⇒ Chilothorax punctatus Waterhouse, 1875
Aphodius obsoleteguttatus (Waterhouse, 1875)⇒ Aphodaulacus nigrotessellatus Motschulsky, 1866
Aphodius plasoni Kaufel, 1914
Aphodius postpilosus Reitter, 1895
Aphodius przewalskyi Reitter, 1887
Aphodius reginae Kral, 1997
Aphodius rudii Endrodi, 1968
Aphodius rugostriatus (Reitter, 1892)⇒ Pharaphodius rugosostriatus Waterhouse, 1875
Aphodius snowi (Brown, 1929)⇒ Agoliinus cruentatus LeConte, 1878
Aphodius subcostatus H. J. Kolbe, 1886
Aphodius unifasciatus Nomura & Nakane, 1951
Aphodobius amplipennis Balthasar, 1945
Ataenius acutulus Schmidt, 1913
Ataenius ambaritae Stebnicka, 1988
Ataenius atramentarius Erichson, 1847
Ataenius australasiae Boheman, 1858
Ataenius borneensis Minkina & Be..., 2023
Ataenius castanicolor Motschulsky, 1858
Ataenius corporaali Boucomont, 1924
Ataenius fukiensis (Balthasar, 1942)⇒ Ataenius australasiae Boheman, 1858
Ataenius hrubanti Balthasar, 1967
Ataenius insularis Lea, 1923
Ataenius liogaster Bates, 1887
Ataenius luteomargo Chapin, 1940
Ataenius malayianus (Petrovitz, 1961)⇒ Ataenius australasiae Boheman, 1858
Ataenius nigricans Paulian, 1933
Ataenius okinawensis (Nakane, 1960)⇒ Ataenius australasiae Boheman, 1858
Ataenius orbicularis (Schmidt, 1914)⇒ Ataenius liogaster Bates, 1887
Ataenius peregrinator Harold, 1877Birthworts
Ataenius picinus Harold, 1861Antarctic blennies
Ataenius rosinae Balthasar, 1967
Ataenius scutellaris (Harold, 1867)⇒ Ataenius atramentarius Erichson, 1847
Ataenius striatocrenatus Fairmaire, 1889
Ataenius tweedensis (Blackburn, 1904)⇒ Ataenius castanicolor Motschulsky, 1858
Ataenius vethianus Schmidt, 1909
Ataenius warisensis Stebnicka, 1998
Australaphodius frenchi Blackburn, 1892Little pasture cockchafer
Blackburneus minutissimus Schmidt, 1908
Bodiloides ictericus Laicharting, 1781
Bodilopsis aquilus Schmidt, 1916
Bodilopsis rufa Moll, 1782
Bodilopsis rufus Moll, 1782
Bodilopsis sordida Fabricius, 1775
Bodilopsis sordidus Fabricius, 1775
Bodilopsis sordidus changajicus Endrödi, 1965
Bodilopsis sordidus sordidus
Bodilus caspius Menestries, 1832
Bodilus crassus Schmidt, 1916
Bodilus gregarius Harold, 1871
Bodilus kaznakovi Frolov, 2001
Bodilus longeciliatus Reitter, 1887
Bodilus nigriventris (Reitter, 1892)⇒ Bodilus lugens Creutzer, 1799
Bodilus punctipennis Erichson, 1848
Bodilus rudii Endrödi, 1968
Bodilus sordescens Harold, 1869
Bodilus zunbajanicus Stebnicka, 1975
Boucomontiellus pelliculosus Balthasar, 1932
Boucomontiellus umbrinus Koshantschikov, 1913
Brachiaphodius babori Balthasar, 1938
Brachiaphodius eccoptus Bates, 1889
Brachiaphodius jendeki Kral, 2000
Brachiaphodius langeri Bellmann, 2009
Brachiaphodius pilifer Paulian, 1934
Brachiaphodius pilosus Harold, 1874
Brachiaphodius ruupbai Masumoto, 1991
Brachiaphodius taiwanicus Petrovitz, 1976
Brachiaphodius weibao Kral, 2000
Brachiaphodius yaaunitnoi Masumoto, 1991
Brachiaphodius yuan Kral, 2000
Bruneixenus reidi Howden & Storey, 2000
Bruneixenus squamosus Howden & Storey, 1992
Caelius denticollis (Lewis, 1895)⇒ Silluvia elongata Landin, 1949
Caelius rufescens Horn, 1887
Calamosternus boreosinicus Cervenka, 1994
Calamosternus granarius Linnaeus, 1767Granular Small Dung Beetle
Calamosternus prusai Tesar, 1945
Calamosternus sauteri Petrovitz, 1961
Calamosternus trucidatus Harold, 1863
Calamosternus uniplagiatus Waterhouse, 1875
Calamosternus zmoskev Cervenka, 1994
Carinaulus armandidavidi Cervenka, 2000
Carinaulus coreensis Kim, 1986
Carinaulus dundarum Cervenka, 2000
Carinaulus gansuicus Cervenka, 2000
Carinaulus imitabilis Cervenka, 2000
Carinaulus inexpectatus Balthasar, 1935
Carinaulus jakli Cervenka, 2000
Carinaulus krali Cervenka, 2000
Carinaulus meghalayensis Cervenka, 2000
Carinaulus milaninikodymi Cervenka, 2000
Carinaulus nanxiensis Masumoto, Ki..., 2018
Carinaulus pseudovseteckai Cervenka, 2003
Carinaulus pseudozhanglaensis Cervenka, 2003
Carinaulus pucholti Balthasar, 1961
Carinaulus tryznai Cervenka, 2000
Carinaulus vseteckai Tesar, 1945
Carinaulus wrzecionkoi Cervenka, 2000
Carinaulus zhanglaensis Cervenka, 2000
Cephalocyclus hogei Bates, 1887
Cheleion jendeki Král & Hájek, 2015
Cheleion malayanum Vårdal & For..., 2010
Cheleion watanabei Kakizoe, Mar..., 2019
Chilothorax alexis Frolov, 2001
Chilothorax altaicus Nikolajev, 1984
Chilothorax bistriga Reitter, 1900
Chilothorax chandmanicus Frolov, 2001
Chilothorax comma Reitter, 1892
Chilothorax conspurcatus Linnaeus, 1758
Chilothorax distinctus (Müller, 1776)⇒ Aphodius distinctus O.F. Müller, 1776
Chilothorax dzongensis Petrovitz, 1976
Chilothorax exclamationis Motschulsky, 1849
Chilothorax flavimargo Reitter, 1901
Chilothorax glebi Frolov, 2006
Chilothorax grafi Reitter, 1901
Chilothorax hozuensis Ochi, Tsukam..., 2015
Chilothorax huangyuanensis Cervenka, 1994
Chilothorax jacobsoni Koshantschikov, 1911
Chilothorax kardonensis Huang & Du, 1999
Chilothorax kerzhneri Nikolajev, 1984
Chilothorax kukunorensis Semenov, 1898
Chilothorax lieni Masumoto, Ki..., 2014
Chilothorax melanostictus Schmidt, 1840
Chilothorax melanostictus melanostictus W. Schmidt, 1840
Chilothorax mongolaltaicus Nikolajev, 1984
Chilothorax nigrivittis impubis Koshantschikov, 1913
Chilothorax nigrivittis kongurensis Ochi, Tsukam..., 2015
Chilothorax nigrivittis nigrivittis
Chilothorax ohishii Masumoto, 1975
Chilothorax okadai Nakane, 1951
Chilothorax okadai kiritachiensis Ochi, Kawaha..., 2012
Chilothorax okadai okadai
Chilothorax paykulli Bedel, 1907
Chilothorax potanini Frolov, 2001
Chilothorax punctatus Waterhouse, 1875
Chilothorax shansianus Nakane & Shi..., 1957
Chilothorax sinicus Cervenka, 1994
Chilothorax tanhensis Frolov, 2001
Chiron sulcithorax Perty, 1831
Cnematoplatys benardi A.Schmidt, 1914
Cnematoplatys numensis Stebnicka, 1986
Cnematoplatys sumatrana Balthasar, 1967
Cnemisus kaszabi (Endrödi, 1964)⇒ Cnemisus kaznakovi Semenov, 1903
Cnemisus kaznakovi Semenov, 1903
Cnemisus kozlovi Semenov, 1903
Colobopterus erraticus Linnaeus, 1758Erratic small dung beetle
Colobopterus indagator Mannerheim, 1849
Colobopterus propraetor Balthasar, 1932
Colobopterus quadratus Reiche, 1847
Coprimorphus scrutator Herbst, 1789
Coptochiroides laotianus Cervenka, 2005
Coptochiroides subcostatus Kolbe, 1886
Coptochiroides subcostatus perroti Paulian, 1945
Coptochiroides subcostatus subcostatus
Craterocyphus forticornis Schmidt, 1910
Craterocyphus rhinocerus Reiche, 1847
Cryptoscatomaseter gambrinus Skelley & Go..., 2001Warteye stargazer
Danielssonia minuta Bordat & Howden, 1995
Dialytes foveatus (Schmidt, 1909)⇒ Setylaides foveatus Schmidt, 1909
Dialytes truncatus Melsheimer, 1845
Diastictus vulneratus Sturm, 1805
Didactylia szunyoghyi Endrödi, 1961
Emadiellus rufopustulatus Wiedemann, 1823
Emadiellus rufopustulatus pistor Balthasar, 1945
Emadiellus rufopustulatus rufopustulatus
Eocorythoderus incredibilis Maruyama, 2012
Erytus aequalis Schmidt, 1907
Erytus opacior Koshantschikov, 1894
Erytus pruinosus Reitter, 1892
Esymus merdarius Fabricius, 1775
Esymus pusillus Herbst, 1789
Esymus pusillus pusillus Herbst, 1789
Esymus pusillus roubali Balthasar, 1932
Eudolus quadriguttatus Herbst, 1783
Euheptaulacus carinatus Germar, 1824
Euheptaulacus carinatus carinatus Germar, 1824
Euheptaulacus sus Herbst, 1783
Euheptaulacus villosus Gyllenhal, 1806
Euorodalus paracoenosus Balthasar & ..., 1960
Euorodalus stribrnyi Tesar, 1945
Eupleurus antiquus Faldermann, 1835
Eupleurus subterraneus Linnaeus, 1758Subterranean small dung beetle
Eupleurus subterraneus subterraneus Linnaeus, 1758
Gilletianus aganocrossoides Kral, Rakovi..., 2014
Gilletianus asinensis Ochi, Kon & ..., 2016
Gilletianus assamensis Petrovitz, 1976
Gilletianus atsushii Ochi, 1986
Gilletianus banteng Král & Masum..., 2020
Gilletianus bolm Kral, Rakovi..., 2014
Gilletianus cavitensis Ochi, Kon & ..., 2016
Gilletianus comatus Schmidt, 1920
Gilletianus commatoides Balthasar, 1961
Gilletianus costatulus Schmidt, 1908
Gilletianus fukiensis Balthasar, 1952
Gilletianus imasakai Ochi, Kon & ..., 2016
Gilletianus kanchil Král & Masum..., 2020
Gilletianus kinabuhayan kinabuhayan
Gilletianus kinabuhayan lucenaensis Ochi, Kon & ..., 2016
Gilletianus kratochvili Balthasar, 1941
Gilletianus lomsakensis Stebnicka, 1992
Gilletianus luzonensis Ochi, Kon & ..., 2016
Gilletianus matotomensis Ochi, Kon & ..., 2010
Gilletianus miksici Balthasar, 1960
Gilletianus mindorensis Ochi, Kon & ..., 2010
Gilletianus miurai Minkina, 2022
Gilletianus negrosensis Ochi, Kon & ..., 2016
Gilletianus nigrovirgatus Schmidt, 1911
Gilletianus poggii Ochi, Kon & ..., 2014
Gilletianus proclivis Balthasar, 1933
Gilletianus rangoonensis Petrovitz, 1970
Gilletianus reichei Harold, 1859
Gilletianus schoolmeestersi Ochi, Kon & ..., 2016
Gilletianus segmentarius Harold, 1879
Gilletianus segmentaroides Schmidt, 1909
Gilletianus seguyi Paulian, 1945
Gilletianus sundainsulanus Ochi, Kon & ..., 2014
Gilletianus superreichei Ochi, Kon & ..., 2010
Gilletianus takeshii Ochi, Kon & ..., 2010
Gilletianus therondi Balthasar, 1963
Gilletianus timorensis Ochi, Kon & ..., 2016
Gilletianus tonkineus Paulian, 1933
Gilletianus yunnanensis Král, Rakovi..., 2015
Gonaphodius scoparius Harold, 1877
Granulopsammodius mongol Endrödi, 1969
Granulopsammodius plicatuloides Pittino, 1984
Granulopsammodius sinicus Rakovic, 1986
Granulopsammodius tienshanicus Pittino, 1984
Guanyinaphodius kishimotoi Masumoto & K..., 2001
Heptaulacus carinatus (Gebler, 1833)⇒ Euheptaulacus carinatus Germar, 1824
Heptaulacus testudinarius Fabricius, 1775
Koreoxyomus koreanus Kim, 1996
Koshantschikovius chovdgolensis Endrödi, 1983
Labarrus hoabinhensis Balthasar, 1946
Labarrus lividus Olivier, 1789Scarab beetle
Labarrus lividus lividus
Labarrus nabeleki Balthasar, 1936
Labarrus rigidus Balthasar, 1936
Labarrus sublimbatus Motschulsky, 1860
Leiopsammodius chifengleei Kiuchi & Tsai, 2018
Leiopsammodius gestroi Clouët, 1900
Leiopsammodius horaki Rakovic & Kral, 1996
Leiopsammodius indicus Harold, 1877
Leiopsammodius japonicus Harold, 1878
Leiopsammodius laevicollis Klug, 1845
Leiopsammodius liviae Pittino, 1978
Leiopsammodius nomurai Masumoto, 2012
Limarus maculatus Sturm, 1800
Limarus zenkeri Germar, 1813
Liothorax kraatzi Harold, 1868
Liothorax niger Illiger, 1798Beaulieu dung beetle
Liothorax plagiatus Linnaeus, 1767
Loraphodius kashmirensis Sharp, 1878
Loraspis frater Mulsant & Rey, 1870
Megateloides bostrichoides Harold, 1860
Megatelus brahminus Harold, 1879
Melinopterus consputus Creutzer, 1799
Melinopterus longipes Landin, 1949
Melinopterus prodromus Brahm, 1790Spring small dung beetle
Melinopterus punctatosulcatus Sturm, 1805
Melinopterus punctatosulcatus hirtipes Fischer von ..., 1844
Melinopterus reyi Reitter, 1892
Melinopterus scuticollis Semenov, 1898
Melinopterus sphacelatus Panzer, 1798
Mendidius bidens Solsky, 1876
Mendidius bispinifrons Reitter, 1889
Mendidius bivittatus Schmidt, 1922
Mendidius fimbriolatus Mannerheim, 1849
Mendidius gobiensis Endrödi, 1964
Mendidius multiplex Reitter, 1897
Mendidius nelsinae Medvedev, 1968
Mendidius pseudobidens Balthasar, 1941
Mesontoplatys arabicus Harold, 1875
Mesontoplatys sunantae Stebnicka, 1981
Messyrhus brachypterus Rakovic & Kral, 1997
Microtermitodius atomus Pittino, 2006
Mozartius jugosus Lewis, 1895
Mozartius jugosus horii Ochi, Kawaha..., 2006
Mozartius jugosus jugosus
Mozartius jugosus shingoi Ochi, Kawaha..., 2006
Mozartius kyushuensis Ochi & Kawah..., 2002
Mozartius testaceus Nomura & Nakane, 1951
Mozartius testaceus shikokuensis Masumoto, 1984
Mozartius testaceus testaceus
Mozartius uenoi Masumoto, 1984
Mozartius uenoi hadai Kawai, 2003
Mozartius uenoi uenoi
Myrhessus kubotai Ochi, Kawaha..., 2012
Myrhessus nishinoi Ochi, Kawaha..., 2012
Myrhessus samurai Balthasar, 1941
Myrhessus sunadai Ochi, Kawaha..., 2019
Myrhessus yorikoae Ochi, Kawaha..., 2006
Neagolius bilimecki Seidlitz, 1891
Neagolius falcispinis Koshantschikov, 1912
Neagolius haroldi Koshantschikov, 1894
Neagolius hengduanensis Kral, 1997
Neagolius montanus Erichson, 1848
Neagolius montisjuldi Pittino, 1988
Neagolius moronensis Frolov, 2009
Neagolius petri Frolov, 2014
Neagolius takin Kral, 1995
Neagolius zangbo Kral, 1997
Neocalaphodius karolinae Minkina, 2015
Neocalaphodius moestus Fabricius, 1801
Neotrichiorhyssemus boninensis Nakane, 1960
Neotrichiorhyssemus esakii Nomura, 1943
Neotrichiorhyssemus expansicollis Benard, 1930
Neotrichiorhyssemus hauseri Balthasar, 1933
Neotrichiorhyssemus hirsutus Clouët, 1901
Neotrichiorhyssemus kentingensis Masumoto & al., 2013
Neotrichiorhyssemus malabaricus Balthasar, 1963
Neotrichiorhyssemus matthewsi Rakovic, 1983
Neotrichiorhyssemus umbilicatus Petrovitz, 1968
Nettelislasia euparinoides Islas, 1945
Nialus nefandus Reitter, 1907
Nialus varians Duftschmid, 1805
Nimbus contaminatus Herbst, 1783
Nimbus makowskyi Koshantschikov, 1891
Nimbus obliteratus Panzer, 1823
Nipponaphodius gotoi Nomura & Nakane, 1951
Nipponoagoliinus yasutakai Ochi & Kawahara, 2001
Nobius bosaki Tesar, 1945
Nobius serotinus Panzer, 1799
Odochilus bacchusi Rakovic, 1987
Odochilus balthasari Rakovic, 1987
Odochilus borneensis Rakovi� & A..., 2021
Odochilus buruensis Boucomont, 1926
Odochilus convexus Nomura, 1971
Odochilus haroldi Rakovic, 1987
Odochilus jani Bezdek & Krivan, 2001
Odochilus johnsoni Rakovic, 1987
Odochilus kashizakii Ochi, Kon & ..., 2017
Odochilus laotianus Minkina, 2020
Odochilus montanus Masumoto & K..., 2019
Odochilus parajohnsoni Rakovic, 1997
Odochilus parasetosus Rakovic, 1997
Odochilus parasyntheticus Rakovic, 1997
Odochilus philippinensis Rakovic, 1987
Odochilus pittinoi Rakovic, 1987
Odochilus sarawakensis Rakovic, 1997
Odochilus setosus Boucomont, 1914
Odochilus shavrini Rakovi� & A..., 2021
Odochilus sulawesiensis Minkina, Bez..., 2023
Odochilus syntheticus Harold, 1877
Odochilus syntheticus philippinensis (Rakovic, 1987)⇒ Odochilus philippinensis Rakovic, 1987
Odochilus taiwanus Masumoto, La..., 2016
Odochilus thailandicus Rakovic, 1997
Orodaliscus rotundangulus Reitter, 1900
Orodaliscus zangi Schmidt, 1906
Oromus alpinus Scopoli, 1763
Otophorus haemorrhoidalis Linnaeus, 1758Bloody small dung beetle
Oxyomus aciculatus Schmidt, 1909
Oxyomus alligator Ho, Hu & Fik..., 2022
Oxyomus bremeri Stebnicka, 1981
Oxyomus cameratus Schmidt, 1908
Oxyomus debilis Harold, 1877
Oxyomus ishidai Nakane, 1977
Oxyomus kiuchii Masumoto, 1991
Oxyomus kocoti Minkina, 2018
Oxyomus malgorzatae Minkina, 2016
Oxyomus masumotoi Nomura, 1973
Oxyomus mencli Minkina, 2016
Oxyomus nanxiensis Masumoto, Ki..., 2018
Oxyomus nowaki Minkina, 2016
Oxyomus sylvestris Scopoli, 1763Forest small dung beetle
Oxyomus taipingensis Masumoto, Ki..., 2014
Oxyomus thailandicus Masumoto, 1991
Palnia cambeforti Kral, 1999
Palnia thailandica Bellmann, 2020
Paracrossidius impressiusculus Fairmaire, 1888
Paracrossidius instigator Balthasar, 1932
Paracrossidius kozlovi Koshantschikov, 1916
Paracrossidius laticapitus Zhang, 1992
Paracrossidius melanodiscus Zhang, 1992
Paracrossidius planipennis Petrovitz, 1961
Paracrossidius radekcervenkai Cervenka, 2017
Paracrossidius sichuanensis Zhang, 1992
Paracrossidius transmontanus Balthasar, 1932
Paracrossidius truculentus Balthasar, 1963
Paradidactylia avuncula Balthasar, 1946
Paradidactylia avunculus Balthasar, 1946
Paradidactylia biseriata Schmidt, 1908
Paradidactylia biseriatus Schmidt, 1908
Paradidactylia carinulata Motschulsky, 1863
Paradidactylia carinulatus Motschulsky, 1863
Paradidactylia ovatula Harold, 1861
Paradidactylia ovatulus Harold, 1861
Paradidactylia semiangulus Balthasar, 1952
Paradidactylia shantungensis Balthasar, 1941
Parammoecius corvinus Erichson, 1848
Parammoecius gibbus Germar, 1817
Parammoecius pyrenaeus Jacquelin du..., 1863
Parammoecius yamato (Nakane, 1960)⇒ Ammoecius yamato Nakane, 1960
Paulianellus andrei Minkina & Král, 2018
Paulianellus asahinai Nakane, 1951
Paulianellus laoticus Mencl, Král ..., 2015
Paulianellus maderi Balthasar, 1938
Paulianellus neomaderi Mencl, Král ..., 2015
Paulianellus taiwanensis Mencl, Král ..., 2015
Phaeaphodius costalis Gebler, 1848
Phaeaphodius dauricus Harold, 1863
Phaeaphodius kiulungensis Balthasar, 1932
Phaeaphodius plutenkoi Kral, 2002
Phaeaphodius rectus Motschulsky, 1866
Phalacronothus biguttatus Germar, 1824
Phalacronothus botulus Balthasar, 1945
Phalacronothus pseudoavunculus Masumoto, 1977
Phalacronothus quadrimaculatus Linnaeus, 1761
Phalacronothus semiangulus Balthasar, 1952
Pharaphodius attritus (Balthasar, 1933)⇒ Pharaphodius marginellus Fabricius, 1781
Pharaphodius birmanicus Petrovitz, 1965
Pharaphodius crenatus Harold, 1862
Pharaphodius diadema Wiedemann, 1823
Pharaphodius javanus Schmidt, 1909
Pharaphodius marginellus Fabricius, 1781
Pharaphodius orientalis Harold, 1862
Pharaphodius priscus Motschulsky, 1858
Pharaphodius putearius Reitter, 1895
Pharaphodius rugosostriatus Waterhouse, 1875
Pharaphodius sundaicus Balthasar, 1946
Plagiogonus arenarius Olivier, 1789
Plagiogonus bortkevitschi Koshantschikov, 1916
Plagiogonus burgaltaicus Csiki, 1901
Plagiogonus chan Endrödi, 1967
Plagiogonus culminarius Reitter, 1900
Plagiogonus duporti Paulian, 1945
Plagiogonus inflatus Balthasar, 1938
Plagiogonus krataay Masumoto, 1992
Plagiogonus ohkurai Masumoto, 1992
Plagiogonus palea Balthasar, 1967
Plagiogonus putridus (Geoffroy, 1785)⇒ Plagiogonus arenarius Olivier, 1789
Plagiogonus quadriclypealis Cervenka, 1995
Plagiogonus shirahatai Nakane, 1951
Planolinellus grzegorzi Minkina, 2017
Planolinellus vittatus Say, 1825
Planolinoides aenictus Cooper & Gordon, 1987Benttooths
Planolinoides borealis Gyllenhal, 1827
Planolinus fasciatus Olivier, 1789
Planolinus khaoensis Stebnicka, 1988
Planolinus pratensis Nomura & Nakane, 1951
Platyderides arrowi Schmidt, 1910
Platyderides doiangkhangensis Masumoto, 1991
Platyderides jengi Masumoto, Ki..., 2014
Platyderides klapperichi Balthasar, 1942
Platyderides planipennis Petrovitz, 1961
Platyderides truculentus Balthasar, 1963
Platyderides vietnamicus Cervenka, 1995
Platytomus mongolicus Medvedev, 1974
Platytomus variolosus Kolenati, 1846
Platytomus yadai Ochi, Kawaha..., 2006
Pleuraphodius berlandi Paulian, 1945
Pleuraphodius javanus Balthasar, 1945
Pleuraphodius lewisi Waterhouse, 1875
Pleuraphodius luzonensis Petrovitz, 1972
Pleuraphodius malangensis Balthasar, 1960
Pleuraphodius tectoformibus Schmidt, 1909
Pleuraphodius viscaensis Stebnicka, 1993
Pleurophorus apicipennis Reitter, 1892
Pleurophorus banksi Pittino & Ma..., 1986
Pleurophorus beccarii Pittino & Ma..., 1986
Pleurophorus caesus Creutzer, 1796Cut small dung beetle
Pleurophorus cambeforti Pittino & Ma..., 1986
Pleurophorus dubius
Pleurophorus formosanus Pittino & Ma..., 1986
Pleurophorus impressicollis Boheman, 1858
Pleurophorus laticollis Pittino & Ma..., 1986
Pleurophorus pannonicus Petrovitz, 1961
Pleurophorus setosus Pittino & Ma..., 1986
Pleurophorus thailandicus Pittino & Ma..., 1986
Pleurophorus tonkinensis Balthasar, 1933
Proctophanes distinguendus Schmidt, 1910
Psammodius asper (Fabricius, 1775)⇒ Psammodius sulcicollis Illiger, 1802
Psammodius convexus Waterhouse, 1875
Psammodius coreanus (Kim, 1980)⇒ Rakovicius coreanus Kim, 1980
Psammodius flavolittoralis Kim, 1980
Psammodius generosus (Reitter, 1892)⇒ Psammodius nocturnus Reitter, 1892
Psammodius hangangnensis Kim, 1980
Psammodius inscitus (Walker, 1858)⇒ Rhyssemus inscitus Walker, 1858
Psammodius japonicus (Harold, 1878)⇒ Leiopsammodius japonicus Harold, 1878
Psammodius kobayashii Nomura, 1973
Psammodius kondoi (Masumoto, 1984)⇒ Psammodius kobayashii Nomura, 1973
Psammodius laevicollis (Klug, 1845)⇒ Leiopsammodius laevicollis Klug, 1845
Psammodius maruyamai Ochi, Kawaha..., 2011
Psammodius nocturnus Reitter, 1892
Psammodius pierottii Pittino, 1979Mud blenny
Psammodius sculpticollis (Fairmaire, 1897)⇒ Leiopsammodius indicus Harold, 1877
Psammodius sungshinarum Kim, 1980
Psammoporus abdita Nikritin, 1975
Psammoporus comis Lewis, 1895
Psammoporus friebi Balthasar, 1935
Psammoporus hybridus Reitter, 1892
Psammoporus kamtschaticus Motschulsky, 1860
Psammoporus latipunctus Gredler, 1866
Psammoporus mimica Pittino, 2006
Psammoporus nakanei (Masumoto, 1986)⇒ Aegialia kamtschaticus Motschulsky, 1860
Psammoporus nakanei horii Ochi & Kawahara, 2012
Psammoporus nakanei nakanei
Psammoporus tsukamotoi Masumoto, 1986
Pseudacrossus absconditus Balthasar, 1932
Pseudacrossus grebenschikovi Balthasar, 1961
Pseudacrossus grebenscikovi Balthasar, 1961
Pseudacrossus grombczewskyi Koshantschikov, 1891
Pseudacrossus juxtus Petrovitz, 1972
Pseudacrossus kalabi Kral, 1997
Pseudacrossus nasutus Reitter, 1887
Pseudacrossus przewalskyi Reitter, 1887
Pseudacrossus qinghaiensis Kral, 1997
Pseudacrossus serrimargo Koshantschikov, 1913
Pseudacrossus smetanai Kral, 2011
Pseudacrossus subsericeus Ballion, 1878
Pseudacrossus tenebricosus Schmidt, 1922
Pseudoahermodontus benesi Dellacasa & ..., 1997
Pseudoheptaulacus azumai Ochi, Kon & ..., 2014
Pseudoheptaulacus iniquus Schmidt, 1909
Pseudoheptaulacus reikoae Ochi, Kon & ..., 2014
Pseudoheptaulacus riekoae Ochi, Kon & ..., 2014
Pseudostereomera mirabilis Bordat & Howden, 1995
Pterobius itiokai Maruyama, 2010
Pubinus tomentosus O.F.Muller, 1776
Qingaphodius nikodymi Kral, 1997
Qingaphodius ochotonarum Kral, 1997
Rakovicius ainu Lewis, 1895
Rakovicius coreanus Kim, 1980
Rakovicius kawaii (Pittino, 2006)⇒ Rakovicius subopacus Nomura, 1973
Rakovicius kyotoensis Ochi, Kawaha..., 2019
Rakovicius lanae Masumoto, 2012
Rakovicius subopacus Nomura, 1973
Rakovicius takashii Ochi, Kawaha..., 2019
Rakovicius thailandicus Balthasar, 1965
Renaudius tesari Paulian, 1942
Renaudius thailandicus (Masumoto, 1991)⇒ Oxyomus thailandicus Masumoto, 1991
Rhinocerotopsis nakasei Maruyama, 2009
Rhodaphodius foetens Fabricius, 1787
Rhyparus accessoricostatus Mencl, Rakov..., 2013
Rhyparus adebratti Bordat, 1996
Rhyparus amamianus (Nakane, 1956)⇒ Rhyparus helophoroides Fairmaire, 1893
Rhyparus annamariae Minkina, 2017
Rhyparus anneae Stebnicka, 1998
Rhyparus apoensis Ochi & Kakizoe, 2021
Rhyparus apomontanus Ochi & Kakizoe, 2021
Rhyparus approximans Fairmaire, 1893
Rhyparus australiae (Lea, 1923)⇒ Rhyparus helophoroides Fairmaire, 1893
Rhyparus azumai Nakane, 1956
Rhyparus azumai loebli Paulian, 1983
Rhyparus bacanensis Minkina, Ani..., 2022
Rhyparus baracatanensis Ochi & Kakizoe, 2021
Rhyparus barclayi Masumoto, Oc..., 2019
Rhyparus basilanensis Ochi & Kakizoe, 2021
Rhyparus birmanicus Fairmaire, 1897
Rhyparus breviceps Paulian, 1984
Rhyparus burckhardti Paulian, 1989
Rhyparus buruensis Minkina, Ani..., 2022
Rhyparus chinensis Balthasar, 1952
Rhyparus danielssoni Bordat, 1996
Rhyparus dentatus Fairmaire, 1896
Rhyparus denticollis Fairmaire, 1893
Rhyparus edieae Stebnicka, 1998
Rhyparus flavohirtus Anichtchenko..., 2021
Rhyparus gracilis Arrow, 1905
Rhyparus helephoroides (Fairmaire, 1893)⇒ Rhyparus helophoroides Fairmaire, 1893
Rhyparus helophoroides Fairmaire, 1893
Rhyparus huaphanensis Ochi, Kon & ..., 2019
Rhyparus ifugaoensis Ochi & Kakizoe, 2021
Rhyparus jakli Minkina & An..., 2022
Rhyparus kinabalu Paulian, 1989
Rhyparus kitanoi Miyake, 1982
Rhyparus kitanoi kitanoi
Rhyparus kitanoi taiwanus Ochi, 2001
Rhyparus klapperichorum Paulian, 1983
Rhyparus laosensis Ochi, Kon & ..., 2018
Rhyparus loebli (Paulian, 1983)⇒ Rhyparus azumai loebli Paulian, 1983
Rhyparus luzonensis Ochi & Kakizoe, 2021
Rhyparus magnus Schmidt, 1911
Rhyparus merkli Minkina & An..., 2022
Rhyparus micros Bordat, 1996
Rhyparus mindanaoensis Anichtchenko..., 2021
Rhyparus minor Paulian, 1989
Rhyparus multipunctatus Paulian, 1984
Rhyparus myslenickorum Minkina, 2019
Rhyparus nahangensis Minkina & Král, 2022
Rhyparus negrosensis Ochi & Kakizoe, 2021
Rhyparus obsoletus Fairmaire, 1893
Rhyparus orousseti (Paulian, 1981)⇒ Rhyparus helophoroides Fairmaire, 1893
Rhyparus paraloebli Ochi, Kon & ..., 2018
Rhyparus paraphilippinensis Ochi & Kakizoe, 2021
Rhyparus peninsularis Arrow, 1905
Rhyparus philippinensis Arrow, 1905
Rhyparus pseudominor Bordat, 1996
Rhyparus punctithoracicus Ochi, Kon & Bai, 2019
Rhyparus risbeci (Paulian, 1934)⇒ Rhyparus helophoroides Fairmaire, 1893
Rhyparus saundersi Pascoe, 1866
Rhyparus schoolmeestersi Ochi, Kon & ..., 2018
Rhyparus semikitanoi Ochi, Kon & ..., 2018
Rhyparus stebnickae Minkina, Ani..., 2022
Rhyparus striatus Arrow, 1935
Rhyparus sumatrensis Fairmaire, 1893
Rhyparus thailandicus Masumoto & Ochi, 2018
Rhyparus verrucosus Schmidt, 1916
Rhyparus vietnamicus Ochi, Kusui ..., 2018
Rhyparus wushensis Ochi & Masumoto, 2018
Rhyparus xanthi Frivaldszky, 1883
Rhyparus yangi Ochi, Kon & Bai, 2019
Rhyssemodes bouvieri Clouët, 1901
Rhyssemodes malyi Rakovic, 1982
Rhyssemodes ningxia Rakovi�, Kr..., 2017
Rhyssemodes orientalis Mulsant & Go..., 1874
Rhyssemodes sindicus Pittino, 1984
Rhyssemodes taklamakan Rakovi�, Kr..., 2017
Rhyssemus australis (Petrovitz, 1963)⇒ Rhyssemus inscitus Walker, 1858
Rhyssemus birmensis Clouët, 1901
Rhyssemus chinensis Pittino, 1996
Rhyssemus feae Clouët, 1901
Rhyssemus germanus Linnaeus, 1767Germany small dung beetle
Rhyssemus hamatus Petrovitz, 1963
Rhyssemus histrio Balthasar, 1961
Rhyssemus inscitus Walker, 1858
Rhyssemus insignicollis (Lea, 1923)⇒ Rhyssemus inscitus Walker, 1858
Rhyssemus koreanus Stebnicka, 1980
Rhyssemus laoticus Pittino, Rak..., 2013
Rhyssemus limbolarius Petrovitz, 1963
Rhyssemus malasiacus (Lansberg, 1886)⇒ Rhyssemus inscitus Walker, 1858
Rhyssemus mongol (Endrödi, 1969)⇒ Granulopsammodius mongol Endrödi, 1969
Rhyssemus mussardi (Petrovitz, 1975)⇒ Rhyssemus inscitus Walker, 1858
Rhyssemus nanshanchicus Masumoto, 1977
Rhyssemus papuanus (Petrovitz, 1965)⇒ Rhyssemus inscitus Walker, 1858
Rhyssemus philippineus (Masumoto, 1980)⇒ Rhyssemus inscitus Walker, 1858
Rhyssemus samurai (Balthasar, 1941)⇒ Myrhessus samurai Balthasar, 1941
Rhyssemus sculptilipennis Schmidt, 1922
Rhyssemus sculptilipennis sculptilipennis
Rhyssemus tarsalis (Waterhouse, 1876)⇒ Rhyssemus inscitus Walker, 1858
Rhyssemus thailandicus Pittino, 1996
Rhyssemus tonkineus (Balthasar, 1945)⇒ Rhyssemus feae Clouët, 1901
Rhyssemus tuberculatus (Nomura & Nakane, 1951)⇒ Myrhessus samurai Balthasar, 1941
Rhyssemus uncispinis Pittino, 1984
Saprosites armatus Pittino, 2008
Saprosites bangueyensis Paulian, 1944
Saprosites cariniceps Pittino, 2008
Saprosites celebicus Paulian, 1944
Saprosites crockerensis Stebnicka, 1991
Saprosites cycloporum Paulian, 1944
Saprosites difficilis Harold, 1877
Saprosites dudichi Endrödi, 1951
Saprosites enarotadii Stebnicka, 1998
Saprosites fodori Endrödi, 1951
Saprosites formosensis (Nomura, 1939)⇒ Saprosites japonicus Waterhouse, 1875
Saprosites implicatus Schmidt, 1922
Saprosites japonicus Waterhouse, 1875
Saprosites javanus Balthasar, 1941
Saprosites kapitensis Petrovitz, 1973
Saprosites kinabalu Pittino, 2008
Saprosites komumi Stebnicka, 1998
Saprosites laeviceps Harold, 1877
Saprosites laticollis Pittino, 2008
Saprosites longethorax Paulian, 1944
Saprosites mahavus Stebnicka, 2012
Saprosites marchionalis (Harold, 1877)⇒ Saprosites cycloporum Paulian, 1944
Saprosites matangi Stebnicka, 2012
Saprosites merkli Pittino, 2008
Saprosites mjoebergi Gillet, 1924
Saprosites narae Lewis, 1895
Saprosites pahangensis Stebnicka, 2012
Saprosites penrisseni Stebnicka, 2012
Saprosites pygmaeus Harold, 1877Birthworts
Saprosites raffrayi Paulian, 1944
Saprosites rougemonti Pittino, 2008
Saprosites rufopolitus Gillet, 1924
Saprosites rufus Paulian, 1944
Saprosites titschacki Balthasar, 1941
Saprosites topali Pittino, 2008
Saprosites verecundus Schmidt, 1909
Saprosites wauensis Stebnicka, 1984
Saprosites yanoi Nomura, 1939
Saprosites yasutakai Ochi & Kawahara, 2019
Saprovisca leytensis Stebnicka, 1993
Saprovisca sarangay Skelley, 2015
Sariangus sariangi Rakovic & Mencl, 2011
Scabrostomus tanytarsus Skelley & Wo..., 1991Masked stargazer
Setylaides canescens Balthasar, 1933
Setylaides chiangmai Stebnicka, 2008
Setylaides foveatus Schmidt, 1909
Setylaides monstrosus Harold, 1877
Setylaides nakanoi Ochi & Kawahara, 2019
Setylaides tsujii Kakizoe, Kaw..., 2021
Siamaphodius naafon Masumoto, 1991
Sigorus porcus Fabricius, 1792Mahogany Ridgeback scarab
Silluvia elongata Landin, 1949
Silluvia gansuensis Frolov, 2008
Silluvia igori Frolov, 2008
Silluvia kabaki Frolov, 2002
Silluvia shashi Stebnicka, 1977
Silluvia sinica Stebnicka, 1977
Silluvia wassuensis Petrovitz, 1963
Silluvia yunnanica Frolov, 2008
Sinaphodius natalicus Mencl, Rakov..., 2013
Sinaphodius oharai Ochi & Kon, 2008
Sinaphodius philippinensis Cervenka, 1994
Sinaphodius sinuatus Harold, 1860
Sinaphodius sulawesiensis Ochi & Kon, 2004
Sinaphodius yunnanus Cervenka, 1994
Sinodiapterna hammondi Dellacasa, 1986
Sinodiapterna songrini Stebnicka & ..., 1992
Sinodiapterna troitzkyi Jacobson, 1897
Sinodiapterna walamiensis Masumoto, Wa..., 2017
Sinodiapterna yushana Li & Wang, 2017
Sinodiapterna zeni Ochi, 1991
Stenotothorax hibernalis Nakane & Tsu..., 1956
Stenotothorax hibernalis hibernalis
Stereomera masumotoi Maruyama & N..., 2011
Stereomera pusilla Arrow, 1905
Subrinus sturmi Harold, 1870
Sybacodes borneensis Minkina, Men..., 2018
Sybacodes liliae Dellacasa, 1978
Sybacodes lutulentus Fairmaire, 1896
Sybacodes lutulentus alternatus Fairmaire, 1897
Sybacodes lutulentus lutulentus
Sybacodes simplicicollis Fairmaire, 1896
Sybacodes simplicicollis aureopilosus Schmidt, 1909
Sybacodes simplicicollis simplicicollis
Sybax impressicollis Boheman, 1857
Termitodiellus besucheti Paulian, 1983
Termitodiellus hainanensis Jiang & Wang, 2020
Termitodiellus hammondi Krikken & Hu..., 1987
Termitodiellus interruptus Krikken & Hu..., 1987
Termitodiellus luzonensis Howden, 1995
Termitodiellus mindanaoensis Minkina & Ka..., 2020
Termitodiellus monticola Krikken & Hu..., 1987
Termitopisthes termiticola Gestro, 1891
Termitopisthes wasmanni A.Schmidt, 1911
Termitotrox cupido Maruyama, 2012
Termitotrox icarus Kakizoe, Lia..., 2020
Termitotrox venus Kakizoe & Ma..., 2015
Teuchestes analis Fabricius, 1787
Teuchestes brachysomus Solsky, 1874
Teuchestes fossor Linnaeus, 1758Digger small dung beetle
Teuchestes guangdong Rakovic & Mencl, 2012
Teuchestes hongson Rakovic & Mencl, 2012
Teuchestes sinofraternus Dellacasa & ..., 1983
Teuchestes uenoi Ochi, Kawaha..., 2006
Trichiorhyssemus asperulus Waterhouse, 1875
Trichiorhyssemus boninensis (Nakane, 1960)⇒ Neotrichiorhyssemus esakii Nomura, 1943
Trichiorhyssemus fokiensis Petrovitz, 1968
Trichiorhyssemus hatakei Ochi, Kawaha..., 2019
Trichiorhyssemus hauseri (Balthasar, 1933)⇒ Neotrichiorhyssemus hauseri Balthasar, 1933
Trichiorhyssemus immaturus Minkina, 2022
Trichiorhyssemus kitayamai Ochi & Kawah..., 2001
Trichiorhyssemus lasionotus Clouët, 1901
Trichiorhyssemus microtrichius Ochi, Kawaha..., 2011
Trichiorhyssemus nakoshii Ochi, Kawaha..., 2021
Trichiorhyssemus nishinoi Ochi, Kawaha..., 2020
Trichiorhyssemus pseudoinscitus Pittino, 1984
Trichiorhyssemus taiwanus Ochi, Masumo..., 2015
Trichiorhyssemus yumikoae Pittino & Kawai, 2007
Trichonotulus amdoensis Král, Rakovi..., 2015
Trichonotulus cechovskyi Minkina & Mencl, 2018
Trichonotulus dzamosanicus Stebnicka, 1973
Trichonotulus khonensis Stebnicka, 1981
Trichonotulus scrofa Fabricius, 1787Sow small dung beetle
Trichonotulus shantungensis Balthasar, 1941
Tristaphodius asperulus Cervenka, 1994
Tristaphodius mortuorum Balthasar, 1932
Turanella latevittis Reitter, 1887
Volinus sticticus Panzer, 1798

Subnodes of Aphodiinae with occurrences in Far East
TribeAegialiini
Laporte, 1840
4
27
Genera
Species

Taxom

TribeAphodiini
Leach, 1815
40
329
Genera
Species

Taxom

TribeAulonocnemini
Janssens, 1946
1
2
Genus
Species

Taxom

TribeChironini
Harold, 1867
1
1
Genus
Species

Taxom

TribeCorythoderini
Schmidt, 1910
2
3
Genera
Species

Taxom

TribeEupariini
Schmidt, 1910
8
96
Genera
Species

Taxom

TribeOdochilini
Rakovič, 1987
1
23
Genus
Species

Taxom

TribePsammodiini
Mulsant, 1842
15
113
Genera
Species

Taxom

TribeRhyparini
Schmidt, 1910
5
78
Genera
Species

Taxom

TribeStereomerini
Howden & Storey, 1992
7
11
Genera
Species

Taxom

TribeTermitotrogini
Wasmann, 1918
1
3
Genus
Species

Taxom



Links, sources and further information
Aphodiinae at artfakta.se
Aphodiinae at dyntaxa.se
Aphodiinae at itis.gov