Caccobius (Thomson 1859) ⇒ Taxonomy and Distribution of Genus

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Taxonomy of Genus Caccobius Thomson, 1859

Class	Insecta Linnaeus, 1758 Insects, True insects	4179 190570 2444470	Families Genera Species
Subclass	Pterygota Lang, 1888 Winged insects	3816 189502 2439455	Families Genera Species
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	3399 186457 2415487	Families Genera Species
Superorder	Holometabola Heider, 1889 Holometabolous Insects	1891 142849 2048615	Families Genera Species
Order	Coleoptera Linnaeus, 1758 Beetles, Coleopterans	592 62779 813516	Families Genera Species
Suborder	Polyphaga Emery, 1886 Polyphags	274 54416 687671	Families Genera Species
Infraorder	Scarabaeiformia Crowson, 1960	32 5275 90050	Families Genera Species
Superfamily	Scarabaeoidea Latreille, 1802 Bess Beetles, Chafer, Dung Beetles, D´or Beetles	32 5275 90050	Families Genera Species
Family	Scarabaeidae Latreille, 1802 Dung beetles, Flower beetles, Fruit chafers, Rain beetles	4240 75466	Genera Species
Subfamily	Scarabaeinae Latreille, 1802 Dung Beetles, True dung beetles	312 12949	Genera Species
Tribe	Onthophagini Lacordaire, 1856	35 4691	Genera Species
Genus	Caccobius Thomson, 1859	211	Species

Subnodes of Caccobius Thomson, 1859

Subgenus	Caccobius (Caccobius) Thomson, 1859	1	Species
Subgenus	Caccobius (Caccophilus) Jekel, 1872	6	Species
Species	Caccobius amagisanus Matsumura, 1936 ⇒ Caccobius jessoensis
Species	Caccobius amicus Gillet, 1925 ⇒ Onthophagus (Furconthophagus) amicus
Species	Caccobius anomalus Klug, 1855 ⇒ Mimonthophagus anomalus
Species	Caccobius anthracinus Faldermann, 1835
Species	Caccobius anthracites d´Orbigny, 1904
Species	Caccobius asininus Heyden, 1887
Species	Caccobius asper d´Orbigny, 1902 ⇒ Caccophilus discrepans
Species	Caccobius aterrimus Fabricius, 1798
Species	Caccobius atomarius Balthasar, 1935 ⇒ Caccobius brevis
Species	Caccobius atratus Walter & Cambefort, 1977
Species	Caccobius auberti d´Orbigny, 1902
Species	Caccobius balthasari Tesar, 1969 ⇒ Cleptocaccobius balthasarianus
Species	Caccobius balthasarianus Cambefort, 1979
Species	Caccobius bama Walter, 2014
Species	Caccobius bawangensis Ochi & Kon & Kikuta, 1997
Species	Caccobius biceps Orbigny, 1905
Species	Caccobius bicornutus Frey, 1955 ⇒ Mimonthophagus ambiguus
Species	Caccobius bicuspis Wiedemann, 1823
Species	Caccobius bidentatus Boucomont, 1923
Species	Caccobius bimaculatus Mulsant, 1842
Species	Caccobius binodulus Harold, 1877
Species	Caccobius bipustulatus Fiori, 1903 ⇒ Caccobius schreberi
Species	Caccobius biseriatus d´Orbigny, 1902 ⇒ Caccophilus auberti
Species	Caccobius boucomonti Balthasar, 1935
Species	Caccobius brevis Waterhouse, 1875
Species	Caccobius cabellai Walter, 2019
Species	Caccobius callosifrons D Orbigny, 1905
Species	Caccobius castaneus Klug, 1855 ⇒ Caccocnemus castaneus
Species	Caccobius cavatus d´Orbigny, 1908
Species	Caccobius chokakurainus Matsumura, 1938
Species	Caccobius christophi Harold, 1879
Species	Caccobius conifer Frey, 1955 ⇒ Pseudosaproecius validicornis
Species	Caccobius conjunctus Müller, 1902 ⇒ Caccobius schreberi
Species	Caccobius convexifrons Raffray, 1877 ⇒ Caccophilus convexifrons
Species	Caccobius corniceps d´Orbigny, 1913
Species	Caccobius corsicus Schneider, 1902 ⇒ Caccobius schreberi
Species	Caccobius crassoides Boucomont, 1923
Species	Caccobius crassus d´Orbigny, 1902 ⇒ Tomogonus crassus
Species	Caccobius crenatostriatus Balthasar, 1935 ⇒ Caccobius christophi
Species	Caccobius cribrarius Boucomont, 1928
Species	Caccobius croceocinctus d´Orbigny, 1913
Species	Caccobius curvicornis Walter & Cambefort, 1977
Species	Caccobius cuspidiger d´Orbigny, 1913
Species	Caccobius cyclotis Cambefort, 1984
Species	Caccobius demangei Boucomont, 1919
Species	Caccobius denticollis Harold, 1867	3	Subspecies
Species	Caccobius devagiriensis Nithya & Sabu, 2013
Species	Caccobius dickinsoni Masumoto, Ochi & Higurashi, 2022
Species	Caccobius diminutivus Walker, 1858
Species	Caccobius discrepans Péringuey, 1901
Species	Caccobius dorsalis Harold, 1867 ⇒ Caccophilus dorsalis
Species	Caccobius dybowskii d´Orbigny, 1902
Species	Caccobius elephantinus Balthasar, 1967
Species	Caccobius excavatus Bai, Zhang & Yang, 2007
Species	Caccobius ferrugineus Fahraeus, 1857
Species	Caccobius flavipennis Balthasar, 1963 ⇒ Caccobius denticollis
Species	Caccobius flavolimbatus Balthasar, 1942
Species	Caccobius foveolatus Frey, 1955
Species	Caccobius fukiensis Balthasar, 1942
Species	Caccobius fuliginosus Roth, 1851 ⇒ Caccophilus reticuliger
Species	Caccobius gallinus Arrow, 1907
Species	Caccobius gananensis d´Orbigny, 1904
Species	Caccobius gemma Sharp, 1875 ⇒ Onthophagus gemma
Species	Caccobius genierorum Walter, 2014
Species	Caccobius gibbosulus d´Orbigny, 1915
Species	Caccobius gilleti Balthasar, 1933
Species	Caccobius globaticeps d´Orbigny, 1905
Species	Caccobius gonoderus Fairmaire, 1888	1	Subspecies
Species	Caccobius grossegranosus Balthasar, 1963
Species	Caccobius haroldi Jekel, 1872 ⇒ Caccobius aterrimus
Species	Caccobius himalayanus Jekel, 1872
Species	Caccobius hirayamai Matsumura, 1936 ⇒ Caccobius jessoensis
Species	Caccobius hirsutus Frey, 1958
Species	Caccobius histerinus Fahraeus, 1857
Species	Caccobius histeroides Ménétriés, 1832
Species	Caccobius histrio Balthasar, 1967
Species	Caccobius humilis Orbigny, 1902
Species	Caccobius imitans Balthasar, 1933
Species	Caccobius imitatus
Species	Caccobius imperfectus Müller, 1902 ⇒ Caccobius schreberi
Species	Caccobius incertus Péringuey, 1901 ⇒ Caccophilus fuliginosus
Species	Caccobius inconspicuus Fahraeus, 1857 ⇒ Caccophilus fuliginosus
Species	Caccobius indicus Harold, 1867
Species	Caccobius indistinctus Mulsant, 1842
Species	Caccobius inermis Arrow, 1931 ⇒ Cleptocaccobius inermis
Species	Caccobius infuscatus Müller, 1902 ⇒ Caccobius schreberi
Species	Caccobius inops Péringuey, 1901
Species	Caccobius ivorensis Cambefort, 1984
Species	Caccobius jessoensis Harold, 1867
Species	Caccobius jononis Matsumura, 1936 ⇒ Gibbonthophagus viduus
Species	Caccobius jossoi Walter, 2014
Species	Caccobius juvenilis Mulsant, 1842
Species	Caccobius kasuganus Matsumura & Yohena, 1936
Species	Caccobius kelleri Olsoufieff, 1907
Species	Caccobius koichii Matsumura, 1934
Species	Caccobius koltzei Reitter, 1892 ⇒ Caccobius sordidus
Species	Caccobius krikkeni Cambefort, 1980
Species	Caccobius laetus Balthasar, 1963 ⇒ Caccobius gonoderus
Species	Caccobius lamottei Cambefort, 1983 ⇒ Cleptocaccobius lamottei
Species	Caccobius lateralis d´Orbigny, 1905
Species	Caccobius leleupi Balthasar, 1963
Species	Caccobius litigiosus Péringuey, 1901 ⇒ Caccophilus nigritulus
Species	Caccobius longipennis d´Orbigny, 1904
Species	Caccobius maruyamai Masumoto, Ochi & Sakchoowong, 2012
Species	Caccobius mastrucatus Péringuey, 1901 ⇒ Caccophilus obtusus
Species	Caccobius masumotoi Cambefort, 1990
Species	Caccobius matsudai Ochi & Araya, 1996
Species	Caccobius matsumotoi Cambefort, 1990
Species	Caccobius matsuoi Matsumura & Yohena, 1937
Species	Caccobius medioniger Ancey, 1883 ⇒ Caccocnemus castaneus
Species	Caccobius megaponerae Brauns, 1914 ⇒ Megaponerophilus megaponerae
Species	Caccobius meridionalis Boucomont, 1914
Species	Caccobius microcephalus Harold, 1877 ⇒ Caccobius jessoensis
Species	Caccobius minimus d´Orbigny, 1898 ⇒ Caccophilus viridicollis
Species	Caccobius minor Walter & Cambefort, 1977 ⇒ Cleptocaccobius minor
Species	Caccobius mirabilepunctatus Cambefort, 1971
Species	Caccobius mixtus Mulsant, 1842
Species	Caccobius montreuilli Walter, 2019
Species	Caccobius morettoi Walter, 2014
Species	Caccobius multifidius
Species	Caccobius multifidus Orbigny, 1908
Species	Caccobius mundus Ménétriés, 1838
Species	Caccobius narashinensis Matsumura, 1936 ⇒ Gibbonthophagus viduus
Species	Caccobius niger Fiori, 1903 ⇒ Caccobius schreberi
Species	Caccobius nigra Tesar, 1969 ⇒ Caccobius denticollis
Species	Caccobius nigritulus Klug, 1855
Species	Caccobius nikkoensis Lewis, 1895
Species	Caccobius nitidiceps Fairmaire, 1893
Species	Caccobius nutans Sharp, 1875
Species	Caccobius obscurus Mulsant, 1842
Species	Caccobius obtusus Fahraeus, 1857
Species	Caccobius ocellipennis d´Orbigny, 1913
Species	Caccobius pantherinus Arrow, 1931
Species	Caccobius parvicollis d´Orbigny, 1913 ⇒ Pycnopanelus parvicollis
Species	Caccobius pentagonus d´Orbigny, 1908
Species	Caccobius picipennis d´Orbigny, 1902 ⇒ Caccophilus convexifrons
Species	Caccobius pilosus Frey, 1958
Species	Caccobius pluridens Orbigny, 1905
Species	Caccobius polygonus d´Orbigny, 1911
Species	Caccobius postlulatus
Species	Caccobius postlutatus d´Orbigny, 1905
Species	Caccobius pseudocorniceps Josso, 2018
Species	Caccobius pseudolaevis d´Orbigny, 1908
Species	Caccobius pudens Péringuey, 1901 ⇒ Caccophilus convexifrons
Species	Caccobius pulicarius Harold, 1875
Species	Caccobius pullus Jekel, 1872
Species	Caccobius punctatissimus Harold, 1867
Species	Caccobius quadridens Orbigny, 1908
Species	Caccobius reticuliger d´Orbigny, 1904
Species	Caccobius rufipennis Motschulsky, 1858
Species	Caccobius sappoensis Matsumura, 1936
Species	Caccobius sapporensis Matsumura, 1936 ⇒ Caccobius jessoensis
Species	Caccobius schaedlei d´Orbigny, 1902
Species	Caccobius scheuerni Cambefort, 1990
Species	Caccobius schreberi Linnaeus, 1767	6	Subspecies
Species	Caccobius semiaeneus d´Orbigny, 1905
Species	Caccobius semicoeruleus d´Orbigny, 1905 ⇒ Caccophilus viridicollis
Species	Caccobius semiluteus d´Orbigny, 1905
Species	Caccobius sericeus Frey, 1958
Species	Caccobius sericoides Balthasar, 1966
Species	Caccobius setifer d´Orbigny, 1905
Species	Caccobius setulosus Motschulsky, 1864
Species	Caccobius shansicus Nakane, 1951
Species	Caccobius sibircus
Species	Caccobius sibiricus Balthasar, 1935 ⇒ Caccobius kelleri
Species	Caccobius signaticollis Orbigny, 1902
Species	Caccobius signatipennis Harold, 1867
Species	Caccobius simplex Boucomont, 1923
Species	Caccobius sordidus Harold, 1886
Species	Caccobius stipes
Species	Caccobius suzukii Matsumura, 1936 ⇒ Caccobius boucomonti
Species	Caccobius tai Cambefort, 1984
Species	Caccobius tigreanus Harold, 1859 ⇒ Caccophilus fuliginosus
Species	Caccobius torticornis Arrow, 1931
Species	Caccobius tsunoellus Matsumara, 1938 ⇒ Caccophilus tortus
Species	Caccobius tuberculifer d´Orbigny, 1902 ⇒ Caccophilus auberti
Species	Caccobius tuberifrons d´Orbigny, 1902
Species	Caccobius ugandicus Frey, 1975
Species	Caccobius ultor Sharp, 1875
Species	Caccobius uniseries Orbigny, 1905
Species	Caccobius upembanus Frey, 1958
Species	Caccobius vacerrosus Lewis, 1895
Species	Caccobius villiersi Cambefort, 1975
Species	Caccobius viridicollis d´Orbigny, 1902 ⇒ Caccophilus convexifrons	2	Subspecies
Species	Caccobius vulcanus Fabricius, 1801
Species	Caccobius yamauchii Matsumura, 1936 ⇒ Caccobius (Caccophilus) unicornis
Species	Caccobius youngai Balthasar, 1967
Species	Caccobius yubariensis Matsumura, 1936 ⇒ Caccobius jessoensis
Species	Caccobius yunnanicus Kabakov, 1994 ⇒ Caccobius masumotoi
Species	Caccobius zairensis Walter & Cambefort, 1977
Species	Caccobius zambiensis Walter, 2014
Species	Caccobius zebra Balthasar, 1963 ⇒ Caccobius denticollis