Checklist for Family Carabidae Latreille, 1802 (Carabid beetles) and region Italy (South Europe)
Class | Insecta Linnaeus, 1758 Insects, True insects | 647 11006 52621 | Families Genera Species | Taxom | Subclass | Pterygota Lang, 1888 Winged insects | 643 10985 52537 | Families Genera Species | Taxom | Infraclass | Neoptera van der Wulp, 1890 Modern Wing-folding insects | 613 10885 52243 | Families Genera Species | Taxom | Superorder | Holometabola Heider, 1889 Holometabolous Insects | 409 8889 45652 | Families Genera Species | Taxom | Order | Coleoptera Linnaeus, 1758 Beetles, Coleopterans | 114 3071 18633 | Families Genera Species | Taxom | Suborder | Adephaga Schellenberg, 1806 Adephagans, Ground Beetles and Water Beetles | 6 342 2879 | Families Genera Species | Taxom | Superfamily | Caraboidea Latreille, 1802
| 1 280 2532 | Family Genera Species | Taxom | | Family | Carabidae Latreille, 1802 Carabid beetles, Carabids, Ground Beetles, Tiger Beetles | 280 2532 | Genera Species | Taxom |
Genera for Italy 280 Genera with 2532 Species #ABCDEGHILMNOPRSTXZ
| Abacetus Dejean, 1828 | | 1 | Species | Taxom | | Abax Bonelli, 1810 | | 20 | Species | Taxom | | Acinopus Dejean, 1821 | | 6 | Species | Taxom | | Acupalpus Latreille, 1829 | | 21 | Species | Taxom | | Agelaea Géné, 1839 | | 1 | Species | Taxom | | Agonoleptus Casey, 1914 | | 2 | Species | Taxom | | Agonum Bonelli, 1810 | | 64 | Species | Taxom | | Agostenus Motschulsky, 1850 | | 1 | Species | Taxom | | Agostinia Jeannel, 1928 | | 2 | Species | Taxom | | Allegrettia Jeannel, 1928 | | 7 | Species | Taxom | | Alpiodytes Jeannel, 1957 | | 2 | Species | Taxom | | Amara Bonelli, 1810 | Seed-Eating Ground Beetles | 109 | Species | Taxom | | Amblystomus Erichson, 1837 | | 14 | Species | Taxom | | Amerizus Chaudoir ver..., 1868 | | 2 | Species | Taxom | | Amphasia Newman, 1838 | | 1 | Species | Taxom | | Anchomenus Bonelli, 1810 | | 2 | Species | Taxom | | Angoleus Villa & Villa, 1833 | | 3 | Species | Taxom | | Anillus Jacquelin du..., 1851 | | 13 | Species | Taxom | | Anisodactylus Dejean, 1829 | | 16 | Species | Taxom | | Anomotarus Chaudoir, 1875 | | 1 | Species | Taxom | | Anophthalmus Sturm, 1844 | | 19 | Species | Taxom | | Anthracus Motschulsky, 1850 | | 6 | Species | Taxom | | Aphoenops Bonvouioir, 1862 | | 1 | Species | Taxom | | Apophua Morley, 1913 | | 4 | Species | Taxom | | Apotomus Illiger, 1807 | | 4 | Species | Taxom | | Apristus Chaudoir, 1846 | | 5 | Species | Taxom | | Aptinus Bonelli, 1810 | | 2 | Species | Taxom | | Argutor Dejean, 1821 | | 2 | Species | Taxom | | Artabas Gozis, 1883 | | 3 | Species | Taxom | | Asaphidion Gozis, 1886 | | 10 | Species | Taxom | | Astigis Rambur, 1838 | | 1 | Species | Taxom | | Atranus LeConte, 1848 | | 2 | Species | Taxom | | Axinopalpus LeConte, 1848 | | 1 | Species | Taxom | | Badister Clairville, 1806 | | 13 | Species | Taxom | | Baudia Ragusa, 1884 | | 4 | Species | Taxom | | Bembidion Latreille, 1802 | | 233 | Species | Taxom | | Bembidium Campa Motsch..., 1844 | | 4 | Species | Taxom | | Binaghites Jeannel, 1937 | | 5 | Species | Taxom | | Blemus Dejean, 1821 | | 1 | Species | Taxom | | Blennidus Motschulsky, 1865 | | 1 | Species | Taxom | | Blethisa Bonelli, 1810 | | 1 | Species | Taxom | | Boldoriella Jeannel, 1928 | | 21 | Species | Taxom | | Bothriopterus Chaudoir, 1838 | | 2 | Species | Taxom | | Brachinus Weber, 1801 | Bombardier Beetle | 31 | Species | Taxom | | Bradybaenus Dejean, 1829 | | 1 | Species | Taxom | | Bradycellus Erichson, 1837 | | 22 | Species | Taxom | | Broscosoma Rosenhauer, 1846 | | 2 | Species | Taxom | | Broscus Panzer, 1813 | | 3 | Species | Taxom | | Calathus Bonelli, 1810 | | 20 | Species | Taxom | | Callisthenes Fischer de W..., 1821 | Beautiful Black Searchers | 1 | Species | Taxom | | Callistus Bonelli, 1809 | | 1 | Species | Taxom | | Calodromius Reitter, 1905 | | 2 | Species | Taxom | | Calomera Motschulsky, 1862 | | 4 | Species | Taxom | | Calosoma Weber, 1801 | Caterpillar hunters | 9 | Species | Taxom | | Campalita Motschulsky, 1865 | | 4 | Species | Taxom | | Carabites Heer, 1852 | | 1 | Species | Taxom | | Carabus Linnaeus, 1758 | Ground beetle | 87 | Species | Taxom | | Cardiomera Bassi, 1834 | | 1 | Species | Taxom | | Carterus Dana, 1851 | | 10 | Species | Taxom | | Cassolaia Wiesner, 1985 | | 1 | Species | Taxom | | Cephalophonus Ganglbauer, 1892 | | 1 | Species | Taxom | | Cephalota Dokhtouroff, 1883 | | 5 | Species | Taxom | | Chlaeniellus Reitter, 1908 | | 6 | Species | Taxom | | Chlaenius Bonelli, 1810 | Vivid Metallic Ground Beetle | 26 | Species | Taxom | | Cicidina | | 1 | Species | Taxom | | Cicindela Linnaeus, 1758 | Common Tiger Beetles | 22 | Species | Taxom | | Cimbrodytes Piva, 2021 | | 1 | Species | Taxom | | Clarencia Sloane, 1917 | | 1 | Species | Taxom | | Clinidium Kirby, 1830 | | 1 | Species | Taxom | | Clivina Latreille, 1802 | | 9 | Species | Taxom | | Cosnania Dejean, 1821 | | 1 | Species | Taxom | | Cryptophonus Brandmayr & ..., 1981 | | 4 | Species | Taxom | | Cychrus Fabricius, 1794 | Rare Snail-eating Beetles | 10 | Species | Taxom | | Cyclotrachelus Chaudoir, 1838 | | 1 | Species | Taxom | | Cylindera Westwood, 1831 | Rounded-thorax Tiger Beetles | 5 | Species | Taxom | | Cymindis Latreille, 1806 | | 23 | Species | Taxom | | Daptus Fischer von ..., 1823 | | 1 | Species | Taxom | | Deltomerus Motschoulsky, 1850 | | 1 | Species | Taxom | | Demetrias Bonelli, 1810 | | 4 | Species | Taxom | | Desera Dejean, 1825 | | 1 | Species | Taxom | | Diachromus Erichson, 1837 | | 1 | Species | Taxom | | Dicaelus Bonelli, 1813 | Notched-mouth Ground Beetles | 2 | Species | Taxom | | Dicheirotrichus Jacquelin du..., 1857 | | 9 | Species | Taxom | | Dimorphoreicheia Magrini, Fan..., 2002 | | 3 | Species | Taxom | | Dinodes Bonet, 1810 | | 1 | Species | Taxom | | Diplocheila Brullé, 1834 | | 1 | Species | Taxom | | Diplous Motschulsky, 1850 | | 1 | Species | Taxom | | Distichus Motschulsky, 1858 | | 1 | Species | Taxom | | Ditomus Dana, 1851 | | 7 | Species | Taxom | | Dixus Billberg, 1820 | | 6 | Species | Taxom | | Doderotrechus Vigna Taglianti, 1968 | | 3 | Species | Taxom | | Dolichus Bonelli, 1810 | | 1 | Species | Taxom | | Dromius Bonelli, 1810 | | 7 | Species | Taxom | | Drypta Latreille, 1796 | | 2 | Species | Taxom | | Duvalius Delarouzee, 1859 | | 109 | Species | Taxom | | Dyschiriodes Jeannel, 1959 | | 27 | Species | Taxom | | Dyschirius Bonelli, 1810 | | 69 | Species | Taxom | | Egadroma Motschulsky, 1855 | | 3 | Species | Taxom | | Elaphropus Motschoulsky, 1839 | | 26 | Species | Taxom | | Elaphrus Fabricius, 1775 | Marsh Ground Beetles | 12 | Species | Taxom | | Ellipsoptera Dokhtouroff, 1883 | Ellipsed-winged Tiger Beetles | 1 | Species | Taxom | | Emphanes Motschulsky, 1850 | | 8 | Species | Taxom | | Epaphius Stephens, 1827 | | 1 | Species | Taxom | | Epomis Bonelli, 1810 | | 2 | Species | Taxom | | Eucarabus Géhin, 1876 | | 1 | Species | Taxom | | Europhilus Chaudoir, 1859 | | 6 | Species | Taxom | | Eurynebria Ganglbauer, 1891 | | 1 | Species | Taxom | | Galerita Fabricius, 1801 | False Bombardier Beetle | 1 | Species | Taxom | | Gastrellarius Casey, 1918 | | 1 | Species | Taxom | | Gigadema J. Thomson, 1859 | | 1 | Species | Taxom | | Glypta Gravenhorst, 1829 | | 40 | Species | Taxom | | Gnathaphanus MacLeay, 1825 | | 1 | Species | Taxom | | Grammognatha Motschulsky, 1850 | | 1 | Species | Taxom | | Graniger Motschulsky, 1864 | | 2 | Species | Taxom | | Gynandromorphus Dejean, 1829 | | 1 | Species | Taxom | | Habroscelimorpha Dokhtouroff, 1883 | Habro Tiger Beetles | 1 | Species | Taxom | | Haptoderus Haptoderus, 1838 | | 6 | Species | Taxom | | Harpalophonus Ganglbauer, 1892 | | 2 | Species | Taxom | | Harpalus Latreille, 1802 | | 84 | Species | Taxom | | Hexagonia Kirby, 1825 | | 1 | Species | Taxom | | Hololeius LaFerté-Séne..., 1851 | | 1 | Species | Taxom | | Hydrium J. LeConte, 1848 | | 1 | Species | Taxom | | Hypharpax MacLeay, 1825 | | 1 | Species | Taxom | | Hypotyphlus Jeannel, 1937 | | 5 | Species | Taxom | | Ichnusodytes Magrini & Fa..., 2019 | | 1 | Species | Taxom | | Italaphaenops Ghidini, 1964 | | 1 | Species | Taxom | | Italodytes Muller, 1938 | | 1 | Species | Taxom | | Lachnocrepis LeConte, 1853 | | 1 | Species | Taxom | | Laemostenus Bonelli, 1810 | | 29 | Species | Taxom | | Lamprias Bonelli, 1810 | | 5 | Species | Taxom | | Lasiotrechus Dejean, 1821 | | 1 | Species | Taxom | | Lebia Latreille, 1802 | colorful foliage ground beetle | 21 | Species | Taxom | | Leistus Frölich, 1799 | | 18 | Species | Taxom | | Lessinodytes Vigna Taglianti, 1982 | | 3 | Species | Taxom | | Licinus Latreille, 1802 | | 10 | Species | Taxom | | Limnaeum Stephens, 1828 | | 2 | Species | Taxom | | Limnastis Ganglbauer, 1892 | | 3 | Species | Taxom | | Limodromus Motschulsky, 1864 | | 3 | Species | Taxom | | Lionychus Wissmann, 1846 | | 7 | Species | Taxom | | Lophoglossus LeConte, 1852 | | 1 | Species | Taxom | | Lophyra Motschulsky, 1859 | | 1 | Species | Taxom | | Lophyridia Jeannel, 1946 | | 3 | Species | Taxom | | Loricera Latreille, 1802 | | 1 | Species | Taxom | | Loxandrus LeConte, 1853 | | 1 | Species | Taxom | | Loxoncus Schmidt-Gobel, 1846 | | 1 | Species | Taxom | | Lymnastis Motschulsky, 1862 | | 5 | Species | Taxom | | Masoreus Dejean, 1821 | | 4 | Species | Taxom | | Megacephala Latreille, 1802 | | 1 | Species | Taxom | | Metadromius Bedel, 1907 | | 2 | Species | Taxom | | Metallina Motschulsky, 1850 | | 6 | Species | Taxom | | Metrius Eschscholtz, 1829 | | 1 | Species | Taxom | | Microderes Faldermann, 1836 | | 1 | Species | Taxom | | Microlestes Schmidt-Goebel, 1846 | | 13 | Species | Taxom | | Microtyphlus Linder, 1863 | | 1 | Species | Taxom | | Miscodera Eschscholtz, 1830 | | 1 | Species | Taxom | | Molops Bonelli, 1810 | | 12 | Species | Taxom | | Myas Sturm, 1826 | | 3 | Species | Taxom | | Myriochila Motschulsky, 1858 | | 1 | Species | Taxom | | Myriochile Motschulsky, 1862 | | 1 | Species | Taxom | | Nebria Latreille, 1802 | Gazelle Beetles | 57 | Species | Taxom | | Neocolpodes Jeannel, 1948 | | 1 | Species | Taxom | | Nomius Laporte, 1835 | | 1 | Species | Taxom | | Notaphus Dejean, 1821 | | 5 | Species | Taxom | | Notiobia Perty, 1830 | | 2 | Species | Taxom | | Notiophilus Dumeril, 1806 | | 13 | Species | Taxom | | Ocydromus Latreille, 1802 | | 74 | Species | Taxom | | Ocys Stephens, 1828 | | 12 | Species | Taxom | | Odacantha Paykull, 1798 | | 1 | Species | Taxom | | Odontium Latreille, 1802 | | 8 | Species | Taxom | | Oedesis Motschulsky, 1850 | | 2 | Species | Taxom | | Olisthopus Dejean, 1828 | | 7 | Species | Taxom | | Omaseus Mannerheim, 1824 | | 2 | Species | Taxom | | Omoglymmius Ganglbauer, 1892 | | 2 | Species | Taxom | | Omophron Latreille, 1802 | Round Sand Beetles | 6 | Species | Taxom | | Omus Eschscholtz, 1829 | Night-stalking Tiger Beetles | 1 | Species | Taxom | | Oodes Bonelli, 1810 | | 2 | Species | Taxom | | Ophonus Dejean, 1821 | | 34 | Species | Taxom | | Oreonebria K. Daniel, 1903 | | 7 | Species | Taxom | | Orotrechus J. Muller, 1913 | | 35 | Species | Taxom | | Orthomus Chaudoir, 1838 | | 3 | Species | Taxom | | Oxypselaphus Chaudoir, 1843 | | 1 | Species | Taxom | | Panagaeus Latreille, 1802 | | 2 | Species | Taxom | | Pangus Dejean, 1821 | | 1 | Species | Taxom | | Paradromius Fowler, 1887 | | 2 | Species | Taxom | | Parallelomorphus Motschulsky, 1850 | | 2 | Species | Taxom | | Paranchus Lindroth, 1974 | | 1 | Species | Taxom | | Paratachys Casey, 1918 | | 3 | Species | Taxom | | Parazuphium Jeannel, 1942 | | 5 | Species | Taxom | | Parophonus Ganglbauer, 1891 | | 8 | Species | Taxom | | Patrobus Dejean, 1821 | | 5 | Species | Taxom | | Paussus L., 1775 | | 2 | Species | Taxom | | Pedius Motschulsky, 1850 | | 2 | Species | Taxom | | Peliocypas Schmidt-Gobel, 1846 | | 1 | Species | Taxom | | Pentagonica Schmidt-Göbel, 1846 | | 1 | Species | Taxom | | Percus Bonelli, 1810 | | 15 | Species | Taxom | | Pericompsus LeConte, 1852 | | 1 | Species | Taxom | | Perigona Laporte, 1835 | | 1 | Species | Taxom | | Perileptus Schaum, 1860 | | 1 | Species | Taxom | | Philochthus Stephens, 1828 | | 10 | Species | Taxom | | Philorhizus Hope, 1838 | | 11 | Species | Taxom | | Phonias Gozis, 1886 | | 3 | Species | Taxom | | Phrator Semenov, 1922 | | 1 | Species | Taxom | | Phyla Motschulsky, 1844 | | 2 | Species | Taxom | | Plataphus Motschulsky, 1864 | | 1 | Species | Taxom | | Platycarabus Morawitz, 1886 | | 1 | Species | Taxom | | Platyderus Stephens, 1827 | | 11 | Species | Taxom | | Platynus Bonelli, 1810 | | 22 | Species | Taxom | | Platypatrobus Darlington, 1938 | | 1 | Species | Taxom | | Platytarus Fairmaire, 1850 | | 3 | Species | Taxom | | Plochionus Dejean, 1825 | | 1 | Species | Taxom | | Plocionus Latreille & ..., 1824 | | 1 | Species | Taxom | | Poecilus Bonelli, 1810 | | 20 | Species | Taxom | | Pogonistes Chaudoir, 1871 | | 2 | Species | Taxom | | Pogonus Dejean, 1821 | | 7 | Species | Taxom | | Polistichus Bonelli, 1810 | | 2 | Species | Taxom | | Polyderis Motschulsky, 1862 | | 2 | Species | Taxom | | Porotachys Netolitzky, 1914 | | 1 | Species | Taxom | | Princidium Motschulsky, 1864 | | 5 | Species | Taxom | | Pseudanophthalmus Jeannel, 1920 | | 1 | Species | Taxom | | Pseudolimnaeum Kraatz, 1888 | | 2 | Species | Taxom | | Pseudomasoreus Desbrochers ..., 1904 | | 1 | Species | Taxom | | Pseudoophonus Motschulsky, 1844 | | 3 | Species | Taxom | | Pseudophonus Motschoulsky, 1848 | | 3 | Species | Taxom | | Pseudosteropus Chaudoir, 1838 | | 1 | Species | Taxom | | Pseudotetracha Fleutiaux, 1894 | | 1 | Species | Taxom | | Pterostichus Bonelli, 1810 | Woodland Ground Beetle | 122 | Species | Taxom | | Reicheia Saulcy, 1862 | | 2 | Species | Taxom | | Reicheiodes Ganglbauer, 1892 | | 3 | Species | Taxom | | Rhegmatobius Jeannel, 1937 | | 11 | Species | Taxom | | Rhipocarabus Reitter, 1896 | | 0 | Species | Taxom | | Rhysodes Dalman, 1823 | | 2 | Species | Taxom | | Sardaphaenops Cerruti & He..., 1956 | | 2 | Species | Taxom | | Scaphinotus Dejean, 1826 | Snail-Eating Beetles | 3 | Species | Taxom | | Scarites Fabricius, 1775 | Scarites Ground Beetle | 10 | Species | Taxom | | Schizogenius Putzeys, 1846 | | 4 | Species | Taxom | | Scotodipnus Schaum, 1860 | | 8 | Species | Taxom | | Scybalicus Schaum, 1862 | | 1 | Species | Taxom | | Selenophorus Dejean, 1829 | | 2 | Species | Taxom | | Semiardistomis Kult, 1950 | | 1 | Species | Taxom | | Semiophonus Schauberger, 1933 | | 1 | Species | Taxom | | Sericoda Kirby, 1837 | | 2 | Species | Taxom | | Siagona Latreille, 1804 | | 1 | Species | Taxom | | Sikelioreicheia Magrini & Fa..., 2019 | | 1 | Species | Taxom | | Sinechostictus Motschulsky, 1864 | | 13 | Species | Taxom | | Sirdenus Dejean, 1828 | | 2 | Species | Taxom | | Somotrichus Seidlitz, 1887 | | 2 | Species | Taxom | | Speluncarius Reitter, 1886 | | 3 | Species | Taxom | | Speomolops Patrizi, 1955 | | 1 | Species | Taxom | | Speotrechus Jeannel, 1922 | | 0 | Species | Taxom | | Sphaeroderus Dejean, 1831 | small snail eating beetle | 1 | Species | Taxom | | Sphodropsis Seidlitz, 1887 | | 1 | Species | Taxom | | Sphodrus Clairville, 1806 | | 2 | Species | Taxom | | Stenolophus Dejean, 1821 | Seedcorn Beetle | 16 | Species | Taxom | | Stomis Clairville, 1806 | | 6 | Species | Taxom | | Synechostictus Motschulsky, 1864 | | 8 | Species | Taxom | | Syntomus Hope, 1838 | | 10 | Species | Taxom | | Synuchus Gyllenhal, 1810 | | 2 | Species | Taxom | | Syrdenus Dejean, 1828 | | 2 | Species | Taxom | | Tachys Dejean, 1821 | | 15 | Species | Taxom | | Tachyta Kirby, 1837 | | 3 | Species | Taxom | | Tachyura Motschulsky, 1862 | | 8 | Species | Taxom | | Tanythrix Schaum, 1858 | | 5 | Species | Taxom | | Tapinopterus Schaum, 1858 | | 1 | Species | Taxom | | Teleutaea Förster, 1869 | | 1 | Species | Taxom | | Tetragonoderus Dejean, 1829 | | 1 | Species | Taxom | | Thalassophilus Wollaston, 1854 | | 1 | Species | Taxom | | Therates Latreille, 1816 | | 1 | Species | Taxom | | Trechicus LeConte, 1853 | | 1 | Species | Taxom | | Trechoblemus Ganglbauer, 1891 | | 1 | Species | Taxom | | Trechus Clairville, 1806 | | 108 | Species | Taxom | | Trepanes Motschulsky, 1864 | | 2 | Species | Taxom | | Trichocellus Ganglbauer, 1892 | | 2 | Species | Taxom | | Trichotichnus A. Morawitz, 1863 | | 7 | Species | Taxom | | Tschitscherinellus Csiki, 1906 | | 1 | Species | Taxom | | Typhlochoromus Moczarski, 1913 | | 2 | Species | Taxom | | Typhloreicheia Holdhaus, 1924 | | 84 | Species | Taxom | | Typhlotrechus J. Muhler, 1913 | | 2 | Species | Taxom | | Xestonotus LeConte, 1853 | | 1 | Species | Taxom | | Zabrus Clairville, 1806 | | 6 | Species | Taxom | | Zuphium Latreille, 1806 | | 2 | Species | Taxom |
Subnodes of Carabidae with occurrences in Italy | Subfamily | Apotominae LeConte, 1853
| 1 4 | Genus Species | Taxom
| | Subfamily | Brachininae Bonelli, 1810
| 2 33 | Genera Species | Taxom
| | Subfamily | Broscinae Hope, 1838
| 3 6 | Genera Species | Taxom
| | Subfamily | Carabinae Latreille, 1802
| 10 117 | Genera Species | Taxom
| | Subfamily | Cicindelinae Latreille, 1802 Tiger beetles | 15 47 | Genera Species | Taxom
| | Subfamily | Elaphrinae Latreille, 1802
| 2 13 | Genera Species | Taxom
| | Subfamily | Harpalinae Bonelli, 1810
| 144 1108 | Genera Species | Taxom
| | Subfamily | Loricerinae Bonelli, 1810
| 1 1 | Genus Species | Taxom
| | Subfamily | Nebriinae Laporte, 1834
| 5 96 | Genera Species | Taxom
| | Subfamily | Omophroninae Bonelli, 1810 Round sand beetles | 2 7 | Genera Species | Taxom
| | Subfamily | Patrobinae Kirby, 1837
| 4 8 | Genera Species | Taxom
| | Subfamily | Paussinae Latreille, 1807 Ant nest beetles, Flanged bombardier beetles, Paussines | 2 3 | Genera Species | Taxom
| | Subfamily | Psydrinae LeConte, 1853
| 1 1 | Genus Species | Taxom
| | Subfamily | Rhysodinae Castelnau, 1840 Wrinkled bark beetles | 3 5 | Genera Species | Taxom
| | Subfamily | Scaritinae Bonelli, 1810 Pedunculate ground beetles | 14 218 | Genera Species | Taxom
| | Subfamily | Siagoninae Bonelli, 1813
| 1 1 | Genus Species | Taxom
| | Subfamily | Trechinae Bonelli, 1810
| 60 849 | Genera Species | Taxom
|
 | | Carabidae (Latreille 1802) |
Links, sources and further information Carabidae at carabidae.org Carabidae at carabidfauna2.net Carabidae at artfakta.se Carabidae at dyntaxa.se Carabidae at gbif.org Carabidae at irmng.org Carabidae at itis.gov Carabidae at marinespecies.org
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