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TAXSYNDISISLFRMPIC
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Taxonomy of Genus Copelatus Erichson, 1832


Class
 		Insecta
Linnaeus, 1758
Insects, True insects		4179
190570
2444470		Families
Genera
Species

Subclass
 		Pterygota
Lang, 1888
Winged insects		3816
189502
2439455		Families
Genera
Species

Infraclass
 		Neoptera
van der Wulp, 1890
Modern Wing-folding insects		3399
186457
2415487		Families
Genera
Species

Superorder
 		Holometabola
Heider, 1889
Holometabolous Insects		1891
142849
2048615		Families
Genera
Species

Order
 		Coleoptera
Linnaeus, 1758
Beetles, Coleopterans		592
62779
813516		Families
Genera
Species

Suborder
 		Adephaga
Schellenberg, 1806
Adephagans, Ground Beetles and Water Beetles		23
6166
123695		Families
Genera
Species

Superfamily
 		Dytiscoidea
Leach, 1815
10
495
10977		Families
Genera
Species
FamilyDytiscidae
Leach, 1815
Diving beetles, Predaceous diving beetle, Water tigers		411
10195		Genera
Species
SubfamilyCopelatinae
Van den Branden, 1885
9
1128		Genera
Species
TribeCopelatini
Branden, 1885
9
1128		Genera
Species
GenusCopelatus
Erichson, 1832
673Species
Subnodes of Copelatus Erichson, 1832
SubgenusCopelatus (Copelatus)
Erichson, 1832
49Species
SpeciesCopelatus abditus Balke, Watts, Cooper, Humphreys & Vogler, 2004
SpeciesCopelatus abonnenci Guignot, 1939
Rocky Mountain cliff fern, Rocky Mountain woodsia
SpeciesCopelatus abusus Guignot, 1953
SpeciesCopelatus acamas Guignot, 1955
SpeciesCopelatus aciculatus Guignot, 1959
SpeciesCopelatus acuductus Clark, 1863
SpeciesCopelatus adelbert Megna, Atthakor, Manaono, Hendrich & Balke, 2017
SpeciesCopelatus advena Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus aemulus Bilardo & Rocchi, 1995
SpeciesCopelatus aequabilis Guignot, 1961
SpeciesCopelatus aequatorius Régimbart, 1899
SpeciesCopelatus aethiopicus Régimbart, 1906
SpeciesCopelatus africanus Sharp, 1882  ⇒ Copelatus pulchellus
SpeciesCopelatus agaroensis Balke in Shaverdo, Monaghan, Lees, Ranaivosolo and, 2008
SpeciesCopelatus agilis Fabricius, 1792
SpeciesCopelatus agrias Guignot, 1954
SpeciesCopelatus aldabricus J.Balfour-Browne, 1950
SpeciesCopelatus alternatus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus alutaceus Régimbart, 1900
SpeciesCopelatus amaroides Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus amatolensis Omer-Cooper, 1965
SpeciesCopelatus amazonicus Régimbart, 1889
SpeciesCopelatus ambiguus Bertrand & Legros, 1975
SpeciesCopelatus americanus Balfour-Browne, 1939
SpeciesCopelatus amphibius Ranarilalatiana & Bergsten, 2019
SpeciesCopelatus anastomosans Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus andamanicus Régimbart, 1899  ⇒ Copelatus oblitus
SpeciesCopelatus andobonicus Guignot, 1960
SpeciesCopelatus andreinii Régimbart, 1905
SpeciesCopelatus andrewesi J.Balfour-Browne, 1939
SpeciesCopelatus angolensis Peschet, 1924
SpeciesCopelatus angouei Bilardo & Rocchi, 2011
SpeciesCopelatus angustatus Chevrolat, 1863  ⇒ Copelatus bertrandi
SpeciesCopelatus angustus Gschwendtner, 1932
SpeciesCopelatus ankaratra Ranarilalatiana & Bergsten, 2019
SpeciesCopelatus annobomensis J.Balfour-Browne, 1939
SpeciesCopelatus anthracinus Régimbart, 1895
SpeciesCopelatus antoniorum Hájek & Brancucci, 2011
SpeciesCopelatus apactus
SpeciesCopelatus aphroditae ♰ Balke in Miller & Balke, 2003
SpeciesCopelatus apicinotatus Bilardo & Rocchi, 2018
SpeciesCopelatus apuzzoi Bilardo & Rocchi, 1999
SpeciesCopelatus aruensis J.Balfour-Browne, 1939
SpeciesCopelatus asemus Omer-Cooper, 1931
SpeciesCopelatus assalyi Bilardo & Rocchi, 2010
SpeciesCopelatus assamensis Vazirani, 1970
SpeciesCopelatus assimilis Régimbart, 1895
2Subspecies
SpeciesCopelatus ateles Guignot, 1955
SpeciesCopelatus ater Sharp, 1882  ⇒ Exocelina atra
SpeciesCopelatus atriceps Sharp, 1882  ⇒ Liopterus atriceps
SpeciesCopelatus atripennis Balfour-Browne, 1939
SpeciesCopelatus atrosulcatus Régimbart, 1906
SpeciesCopelatus aubei Montrouzier, 1860  ⇒ Exocelina aubei
SpeciesCopelatus australiae Clark, 1863  ⇒ Exocelina australiae
SpeciesCopelatus australis Clark, 1863  ⇒ Copelatus chevrolati
SpeciesCopelatus babyrousa Hájek, Hendrich & Balke, 2018
SpeciesCopelatus bacillifer Guignot, 1955
SpeciesCopelatus baculiformis Guignot, 1955
SpeciesCopelatus badeni Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus bagus Balke & Hendrich, 2001
SpeciesCopelatus bakewelli J.Balfour-Browne, 1939
SpeciesCopelatus bangalorensis Vazirani, 1970
SpeciesCopelatus baoulicus Bilardo & Pederzani, 1978
SpeciesCopelatus bapounouensis Bilardo & Rocchi, 2019
SpeciesCopelatus baranensis Hájek, Shaverdo, Hendrich & Balke, 2021
SpeciesCopelatus barbouri Young, 1942
SpeciesCopelatus basalis Boheman, 1848  ⇒ Copelatus pulchellus
SpeciesCopelatus basifasciatus Régimbart, 1895
SpeciesCopelatus basilewskyi Bilardo & Pederzani, 1979
SpeciesCopelatus bechynei Guignot, 1953
SpeciesCopelatus befasicus Guignot, 1956
SpeciesCopelatus bengalensis Guignot, 1955
SpeciesCopelatus bertrandi Nilsson, Bilardo & Rocchi, 1996
SpeciesCopelatus betampona Ranarilalatiana & Bergsten, 2019
SpeciesCopelatus bezdeki Sheth, Ghate & Hájek, 2018
SpeciesCopelatus bibulus Guignot, 1948
SpeciesCopelatus bicolor Fabricius, 1792
SpeciesCopelatus biformis Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus bilunatus Guignot, 1955
SpeciesCopelatus bimaculatus Resende & Vanin, 1991  ⇒ Exocelina bimaculata
SpeciesCopelatus binaghii Bilardo & Pederzani, 1978
SpeciesCopelatus biolleyi Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus birougouensis Bilardo & Rocchi, 2018
SpeciesCopelatus biseriatus J.Balfour-Browne, 1939
SpeciesCopelatus biswasi Mukherjee & Sengupta, 1986
SpeciesCopelatus blancasi Guignot, 1958
SpeciesCopelatus blatchleyi Young, 1953
Ball-head skyrocket, Ballhead gilia, Ballhead ipomopsis
SpeciesCopelatus bolivianus Guignot, 1957
SpeciesCopelatus bombycinus Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus bonvouloiri Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus bottegoi Régimbart, 1895
SpeciesCopelatus bougainvillensis Hájek, Shaverdo, Hendrich & Balke, 2021
SpeciesCopelatus boukali Hendrich & Balke, 1998
SpeciesCopelatus boulevardi Watts, 1978  ⇒ Exocelina boulevardi
SpeciesCopelatus brancuccii Rocchi, 1979
SpeciesCopelatus brasiliensis Zimmermann, 1921
SpeciesCopelatus brendelli Hájek, Hendrich & Balke, 2018
SpeciesCopelatus brevicornis Sharp, 1882
SpeciesCopelatus brevistrigatus Guignot, 1959
SpeciesCopelatus brivioi Rocchi, 1976
SpeciesCopelatus bromeliarum Scott, 1912
SpeciesCopelatus brooksi
SpeciesCopelatus brownei Guignot, 1942  ⇒ Exocelina brownei
SpeciesCopelatus brullei Aubé, 1838
SpeciesCopelatus brunneus J.Balfour-Browne, 1939
SpeciesCopelatus buqueti Aubé, 1838
SpeciesCopelatus burgeoni Gschwendtner, 1930
2Subspecies
SpeciesCopelatus caelatipennis Aubé, 1838
Grand Junction camissonia		4Subspecies
SpeciesCopelatus caelatus Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus caffer J.Balfour-Browne, 1939
SpeciesCopelatus calaquei Bilardo & Rocchi, 2008
SpeciesCopelatus camerunensis Guignot, 1941
2Subspecies
SpeciesCopelatus capensis Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus carayoni Legros, 1949
SpeciesCopelatus carinatus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus celinoides Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus cessaima Caetano, de C. Bena, & Vanin, 2013
SpeciesCopelatus ceylonicus Vazirani, 1969
SpeciesCopelatus cheesmanae J.Balfour-Browne, 1939
SpeciesCopelatus chevrolati Aubé, 1838
Chain fern, Chainfern, Chevrolat´s Dytiscid		2Subspecies
SpeciesCopelatus chibcha Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus chinensis Régimbart, 1899  ⇒ Copelatus japonicus
SpeciesCopelatus chipiriricus Guignot, 1957
SpeciesCopelatus chloroticus Régimbart, 1899
SpeciesCopelatus cinnamomeus Régimbart, 1895
SpeciesCopelatus clarki Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus collarti Gschwendtner, 1932
SpeciesCopelatus collocallosus Falkenström, 1932  ⇒ Copelatus japonicus
SpeciesCopelatus colocallosus
SpeciesCopelatus commatifer Heller, 1916  ⇒ Exocelina commatifera
SpeciesCopelatus compertus Guignot, 1956
SpeciesCopelatus concii Bilardo, 1982
SpeciesCopelatus concolor Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus concolorans J.Balfour-Browne, 1939
SpeciesCopelatus confinis Bilardo & Rocchi, 1999
SpeciesCopelatus congener Guignot, 1941
SpeciesCopelatus congo Gschwendtner, 1938
SpeciesCopelatus consimilis Bilardo & Rocchi, 2002
2Subspecies
SpeciesCopelatus consors Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus convexus ♰ Förster, 1891
SpeciesCopelatus cooperae Bilardo & Pederzani, 1972
SpeciesCopelatus cordovai Megna & Epler, 2012
SpeciesCopelatus cordylinoides J.Balfour-Browne, 1938
SpeciesCopelatus coxalis Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus crassus Régimbart, 1895
SpeciesCopelatus cryptarchoides Régimbart, 1899
SpeciesCopelatus cubaensis Schaeffer, 1908
Barnacles
SpeciesCopelatus cubiensis
SpeciesCopelatus curtistriatus Bilardo & Rocchi, 1995
SpeciesCopelatus curvispinis Bilardo & Rocchi, 2008
SpeciesCopelatus cynthiae Bilardo & Rocchi, 2008
SpeciesCopelatus daemeli Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus danyi Megna & Epler, 2012
SpeciesCopelatus darlingtoni Young, 1942
SpeciesCopelatus davidi Wewalka, 2017
SpeciesCopelatus debilis Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus deccanensis Sheth, Ghate & Hájek, 2018
SpeciesCopelatus decellei Bilardo, 1982
SpeciesCopelatus decemstriatus Aubé, 1838
SpeciesCopelatus decemsulcatus Régimbart, 1895
SpeciesCopelatus deceptor Bilardo & Rocchi, 1995
SpeciesCopelatus dentatipenis Jiang, Hajek & Jia, 2022
SpeciesCopelatus depauperatus Régimbart, 1894
SpeciesCopelatus depressus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus descarpentriesi Bertrand & Legros, 1975
SpeciesCopelatus diffisus Guignot, 1939
Rocky Mountain cliff fern, Rocky Mountain woodsia
SpeciesCopelatus dimorphus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus discoideus Sharp, 1882  ⇒ Copelatus pulchellus
SpeciesCopelatus distinctus Aubé, 1838
SpeciesCopelatus distinguendus Régimbart, 1903
SpeciesCopelatus diversistriatus Jiang, Hajek & Jia, 2022
SpeciesCopelatus divisus Watts, 1978
SpeciesCopelatus djiboutensis Wewalka & Jäch, 2017
SpeciesCopelatus dolosus Guignot, 1956
SpeciesCopelatus doriae Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus doudouensis Bilardo & Rocchi, 2015
SpeciesCopelatus duodecimstriatus Aubé, 1838
SpeciesCopelatus duponti Aubé, 1838
SpeciesCopelatus edax Guignot, 1955
SpeciesCopelatus efoutensis Bilardo & Rocchi, 1995
SpeciesCopelatus ejactus Omer-Cooper, 1965
SpeciesCopelatus ellai Bilardo & Rocchi, 1995
SpeciesCopelatus elongatulus MacLeay, 1871  ⇒ Exocelina elongatula
SpeciesCopelatus elongatus Kolbe, 1883
SpeciesCopelatus elutus Guignot, 1958
SpeciesCopelatus enganensis Guignot, 1940
SpeciesCopelatus epactus Guignot, 1948
SpeciesCopelatus erichsoni Guérin-Méneville, 1849
SpeciesCopelatus erichsonii Guérin-Méneville, 1849
SpeciesCopelatus erraticus Balfour-Browne, 1939
SpeciesCopelatus esteriensis Bilardo & Pederzani, 1978
SpeciesCopelatus eucritus Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus evanidus Bilardo & Rocchi, 1995
SpeciesCopelatus exaratus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus extensus Sharp, 1882
SpeciesCopelatus externus Kirsch, 1873
SpeciesCopelatus fairmairei Zimmermann, 1919
SpeciesCopelatus falciformis Pederzani & Schizzerotto, 2017
SpeciesCopelatus fallax Bilardo & Pederzani, 1978
SpeciesCopelatus fasciatus Bilardo & Rocchi, 1995
SpeciesCopelatus fastidiosus Guignot, 1959
SpeciesCopelatus feae Régimbart, 1888
SpeciesCopelatus felicis Hajek, Jiang & Jia, 2022
SpeciesCopelatus fernandensis Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus ferrugineus Sharp, 1882  ⇒ Exocelina ferruginea
SpeciesCopelatus ferruginicollis Régimbart, 1895
SpeciesCopelatus ferus Guignot, 1953
SpeciesCopelatus festae Régimbart, 1899
SpeciesCopelatus fidschiensis Zimmermann, 1928
Eveningsnow
SpeciesCopelatus fijiensis Guignot, 1955
SpeciesCopelatus filiformis Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus fimbriolatus Dejean, 1836
SpeciesCopelatus fizpaci Bilardo & Rocchi, 2013
SpeciesCopelatus flavicans Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus flavidus Régimbart, 1895
SpeciesCopelatus fluviaticus Guignot, 1957
SpeciesCopelatus fontanus J.Balfour-Browne, 1950
SpeciesCopelatus fonticola Guignot, 1955
SpeciesCopelatus fossilis ♰ Říha, 1974
SpeciesCopelatus fractistriatus Bilardo & Rocchi, 1995
SpeciesCopelatus fractus Guignot, 1953
SpeciesCopelatus fragilis Sharp, 1882  ⇒ Copelatus regimbarti
SpeciesCopelatus freudei Guignot, 1955
SpeciesCopelatus fryi J.Balfour-Browne, 1939
SpeciesCopelatus fulviceps J.Balfour-Browne, 1938
SpeciesCopelatus fulvus Guignot, 1952
SpeciesCopelatus fuscipennis Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus fuscomaculatus Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus gabonicus Bilardo & Pederzani, 1978
SpeciesCopelatus galapagoensis G.R.Waterhouse, 1845
SpeciesCopelatus gapa Watts, 1978  ⇒ Exocelina gapa
SpeciesCopelatus garambanus Guignot, 1955
SpeciesCopelatus gardineri Scott, 1912
SpeciesCopelatus geayi Régimbart, 1904
SpeciesCopelatus geniculatus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus gibsoni Vazirani, 1974
SpeciesCopelatus gigas Bilardo & Rocchi, 2019
SpeciesCopelatus glyphicus Say, 1823  ⇒ Copelatus vitraci
SpeciesCopelatus glyptus Guignot, 1954  ⇒ Exocelina glypta
SpeciesCopelatus gracilis Sharp, 1882  ⇒ Exocelina gracilis
SpeciesCopelatus griffinii Régimbart, 1899
SpeciesCopelatus gschwendtneri Guignot, 1939
Rocky Mountain cliff fern, Rocky Mountain woodsia
SpeciesCopelatus guadelupensis Legros, 1948
SpeciesCopelatus guérini Aubé, 1838
SpeciesCopelatus guerini Aubé, 1838
SpeciesCopelatus guineensis Guignot, 1955
SpeciesCopelatus haemorrhoidalis Fabricius, 1787  ⇒ Copelatus aruensis
SpeciesCopelatus haemorroidalis
SpeciesCopelatus hararensis Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus hardenbergi J.Balfour-Browne, 1950
SpeciesCopelatus harrarensis Guignot, 1961
SpeciesCopelatus hasegawai Sato, 1988  ⇒ Copelatus nakamurai
SpeciesCopelatus hebeter Guignot, 1953
SpeciesCopelatus heterogynus Régimbart, 1899
SpeciesCopelatus hisamatsui Sato, 1961  ⇒ Copelatus tenebrosus
SpeciesCopelatus horni Régimbart, 1902
SpeciesCopelatus hova Alluaud, 1900
SpeciesCopelatus hydroporoides Murray, 1859  ⇒ Exocelina novaecaledoniae
SpeciesCopelatus ibrahimi Zalat, Saleh, Angus & Kaschef, 2000
SpeciesCopelatus idanus Guignot, 1955
SpeciesCopelatus iguelaensis Bilardo & Rocchi, 2002
SpeciesCopelatus ilybioides Régimbart, 1895
SpeciesCopelatus imasakai Matsui & Kitayama, 2000
SpeciesCopelatus imitator Bilardo & Rocchi, 2002
SpeciesCopelatus imperfectus Guignot, 1952
SpeciesCopelatus impressicollis Sharp, 1882  ⇒ Copelatus distinctus
SpeciesCopelatus inaequalis Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus incertus Omer-Cooper, 1957
SpeciesCopelatus incisus Guignot, 1954
SpeciesCopelatus incognitus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus indicus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus inopinatus Bilardo & Rocchi, 1995
SpeciesCopelatus insidiosus Bilardo & Rocchi, 1995
SpeciesCopelatus insolitus Chevrolat, 1863
SpeciesCopelatus instabilis Régimbart, 1897
SpeciesCopelatus instriatus Zimmermann, 1921
SpeciesCopelatus insuetus Guignot, 1941
SpeciesCopelatus insulanus Guignot, 1939
Rocky Mountain cliff fern, Rocky Mountain woodsia
SpeciesCopelatus insularis Balfour-Browne, 1950
SpeciesCopelatus integer Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus internus Régimbart, 1904
SpeciesCopelatus interruptus Perroud & Montrouzier, 1864
SpeciesCopelatus interstriatus J.Balfour-Browne, 1939
SpeciesCopelatus inuber Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus ipiformis Régimbart, 1895
SpeciesCopelatus irinus Régimbart, 1899
SpeciesCopelatus ischius Guignot, 1956
SpeciesCopelatus jactator Guignot, 1955
SpeciesCopelatus jaechi Bilardo & Rocchi, 2016
SpeciesCopelatus jamaicensis Young, 1942
SpeciesCopelatus japonicus Sharp, 1884
SpeciesCopelatus jarrigei Legros, 1954
SpeciesCopelatus javanus Régimbart, 1883
SpeciesCopelatus jocosus Guignot, 1955
SpeciesCopelatus johannis Guignot, 1954
SpeciesCopelatus kainantuensis Balke, 2001
SpeciesCopelatus kalaharii Gschwendtner, 1935
SpeciesCopelatus kammuriensis Tamu & Tsukamoto, 1955
SpeciesCopelatus karnatakus Holmen & Vazirani, 1990
SpeciesCopelatus katangensis Guignot, 1952
SpeciesCopelatus kely Ranarilalatiana & Bergsten, 2019
SpeciesCopelatus kietensis Hájek, Shaverdo, Hendrich & Balke, 2021
SpeciesCopelatus kilimandjarensis Bilardo & Pederzani, 1972
SpeciesCopelatus kindianus Guignot, 1954
SpeciesCopelatus kongouensis Bilardo & Rocchi, 1999
SpeciesCopelatus koreanus Mori, 1932
SpeciesCopelatus labratus Sharp, 1882
SpeciesCopelatus laccophilinus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus laeticulus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus laevipennis Hájek, Shaverdo, Hendrich & Balke, 2021
SpeciesCopelatus lamottei Legros, 1954
SpeciesCopelatus lanzai Bilardo & Rocchi, 1995
SpeciesCopelatus laraensis Bilardo & Rocchi, 1995
SpeciesCopelatus lasckonyi Bilardo & Rocchi, 1995
2Subspecies
SpeciesCopelatus latens Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus laticollis Régimbart, 1899
SpeciesCopelatus latifasciatus Bilardo & Rocchi, 1999
SpeciesCopelatus latipes Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus latus J.Balfour-Browne, 1950
SpeciesCopelatus leonardii Bilardo & Rocchi, 1999
SpeciesCopelatus leonensis Legros, 1954
SpeciesCopelatus leotensi
SpeciesCopelatus lepersonneae Gschwendtner, 1943
SpeciesCopelatus levicollis Guignot, 1952  ⇒ Copelatus polystrigus
SpeciesCopelatus lignosus Guignot, 1955
SpeciesCopelatus lineatipennis Guignot, 1955
SpeciesCopelatus lineatus Guérin-Méneville, 1838
SpeciesCopelatus lividus Sharp, 1882  ⇒ Copelatus chloroticus
SpeciesCopelatus longicornis Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus lootensi Guignot, 1955
SpeciesCopelatus louayensis Bilardo & Rocchi, 2004
SpeciesCopelatus luctuosus Guignot, 1939
Rocky Mountain cliff fern, Rocky Mountain woodsia
SpeciesCopelatus luridescens Régimbart, 1889
SpeciesCopelatus luteocinctus Guignot, 1955
SpeciesCopelatus luzonicus Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus macellus Guignot, 1950
Miller Canyon woodlandsnail
SpeciesCopelatus maculatus Sharp, 1882
SpeciesCopelatus madangensis Balke, 2001
SpeciesCopelatus madoni Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus mahajanga Pederzani & Hájek, 2005
SpeciesCopelatus mahleri Holmen & Vazirani, 1990
SpeciesCopelatus makokouensis Bilardo & Rocchi, 1995
SpeciesCopelatus malaisei Guignot, 1954
SpeciesCopelatus malavergnorum Manuel & Ramahandrison, 2022
SpeciesCopelatus mancus Sharp, 1887
Diamond Valley suncup
SpeciesCopelatus marginalis Gschwendtner, 1932  ⇒ Copelatus polystrigus
SpeciesCopelatus marginatus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus marginipennis Laporte, 1835
SpeciesCopelatus martinbaehri Hendrich, Shaverdo, Hájek & Balke, 2019
SpeciesCopelatus martini Manuel, Deler-Hernández, Megna & Hájek, 2018
SpeciesCopelatus masculinus Régimbart, 1899
SpeciesCopelatus massaicus Guignot, 1941
SpeciesCopelatus mathani Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus mauiensis Blackburn, 1884  ⇒ Exocelina parvula
SpeciesCopelatus maushomi Sheth, Ghate & Hájek, 2018
SpeciesCopelatus mbokoensis Bilardo & Rocchi, 2006
SpeciesCopelatus melanarius Sharp, 1882  ⇒ Exocelina melanaria
SpeciesCopelatus melanogrammus Régimbart, 1883
SpeciesCopelatus mendax Bilardo & Rocchi, 2011
SpeciesCopelatus miaowangi Hajek, Jiang & Jia, 2022
SpeciesCopelatus mimetes Guignot, 1957
SpeciesCopelatus minimus Bilardo & Rocchi, 1995
SpeciesCopelatus minor Bilardo & Pederzani, 1978
2Subspecies
SpeciesCopelatus minutissimus J.Balfour-Browne, 1939
SpeciesCopelatus mocquerysi Régimbart, 1895
SpeciesCopelatus mohelicus Guignot, 1959
SpeciesCopelatus monticola Guignot, 1951
SpeciesCopelatus montivagus Young, 1942
SpeciesCopelatus mopanshanensis Jiang, Zhao & Hajek, 2022
SpeciesCopelatus mulangensis Bameul, 2003
SpeciesCopelatus multistriatus Zimmermann, 1921
SpeciesCopelatus multistriolatus Guignot, 1956  ⇒ Copelatus teranishii
SpeciesCopelatus mundus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus mutabilis Bilardo & Rocchi, 1999
SpeciesCopelatus mvoungensis Bilardo & Rocchi, 2004
SpeciesCopelatus mysorensis Vazirani, 1970
SpeciesCopelatus nakamurai Guéorguiev, 1970
SpeciesCopelatus nangaensis Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus nanlingensis Hajek, Jiang & Jia, 2022
SpeciesCopelatus nauclerus Guignot, 1939
Rocky Mountain cliff fern, Rocky Mountain woodsia
SpeciesCopelatus neavei J.Balfour-Browne, 1950
SpeciesCopelatus neelumae Vazirani, 1973
SpeciesCopelatus neglectus Sharp, 1882
SpeciesCopelatus neogestroi Balke in Shaverdo, Monaghan, Lees, Ranaivosolo and, 2008
SpeciesCopelatus neoguineensis Zimmermann, 1919
Florida Mountain woodlandsnail
SpeciesCopelatus nguemai Bilardo & Rocchi, 2011
SpeciesCopelatus niger Gschwendtner, 1932
SpeciesCopelatus nigricans Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus nigricollis Paulino de Oliveira, 1880
SpeciesCopelatus nigritulus Sharp, 1883  ⇒ Exocelina australiae
SpeciesCopelatus nigrolineatus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus nigropennis Zimmermann, 1927
SpeciesCopelatus nigrostriatus Régimbart, 1895
SpeciesCopelatus nilgiricus Balfour-Browne, 1939
SpeciesCopelatus nilssoni Wewalka, 1982
SpeciesCopelatus nimbaensis Legros, 1958
SpeciesCopelatus nitens Bilardo & Rocchi, 1999
SpeciesCopelatus nitidus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus nodieri Régimbart, 1895
SpeciesCopelatus normalis Erichson, 1847
SpeciesCopelatus notius Omer-Cooper, 1965
SpeciesCopelatus novaecaledoniae Balfour-Browne, 1939
SpeciesCopelatus nzei Bilardo & Rocchi, 1999
SpeciesCopelatus oberthuri Sharp, 1882
SpeciesCopelatus oblitus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus oblongus Illiger, 1801
SpeciesCopelatus obscurus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus obtusus Boheman, 1848  ⇒ Copelatus pulchellus
SpeciesCopelatus occultus Bilardo & Rocchi, 1995
SpeciesCopelatus ogasawarensis Kamiya, 1932
SpeciesCopelatus onorei Pederzani & Rocchi, 1982
SpeciesCopelatus ornatipennis Zimmermann, 1926
SpeciesCopelatus owas Régimbart, 1895
SpeciesCopelatus pachys Guignot, 1955
SpeciesCopelatus pallidus Régimbart, 1895
SpeciesCopelatus paludorum Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus pandanorum Scott, 1912
SpeciesCopelatus pantosi Guignot, 1958
SpeciesCopelatus parabaptus Guignot, 1955
SpeciesCopelatus parallelipipedus Régimbart, 1895
SpeciesCopelatus parallelus Zimmermann, 1920
SpeciesCopelatus pardii Rocchi, 1990
SpeciesCopelatus parisii Guignot, 1959
SpeciesCopelatus parumstriatus Gschwendtner, 1934
SpeciesCopelatus parvulus Boisduval, 1835  ⇒ Exocelina parvula
SpeciesCopelatus paryphes Guignot, 1955
SpeciesCopelatus patrizii Peschet, 1922
SpeciesCopelatus patrizzii Guignot, 1961
SpeciesCopelatus pederzanii Bilardo & Rocchi, 1995
SpeciesCopelatus pentheres Guignot, 1953
SpeciesCopelatus pereirai Guignot, 1955
SpeciesCopelatus perfectus Sharp, 1882  ⇒ Exocelina perfecta
SpeciesCopelatus peridinus Guignot, 1955
SpeciesCopelatus peruanus Erichson, 1847
SpeciesCopelatus pinnifer Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus piriensis Omer-Cooper, 1965
SpeciesCopelatus platynotus Régimbart, 1895
SpeciesCopelatus polystrigus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus ponomarenkoi ♰ Říha, 1974
SpeciesCopelatus portitor
SpeciesCopelatus posticatus Fabricius, 1801
SpeciesCopelatus poungai Bilardo & Rocchi, 2006
SpeciesCopelatus predaveterus ♰ Miller in Miller & Balke, 2003
SpeciesCopelatus prolixus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus prolongatus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus propino Guignot, 1960
SpeciesCopelatus propinquus Régimbart, 1895
SpeciesCopelatus proximus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus pseudostriatus Ranarilalatiana & Bergsten, 2019
SpeciesCopelatus pulchellus Klug, 1834
SpeciesCopelatus pulicarius Régimbart, 1895
SpeciesCopelatus pumilus Régimbart, 1895
SpeciesCopelatus punctatus Bilardo & Rocchi, 1995
SpeciesCopelatus punctipennis Lea, 1899  ⇒ Exocelina punctipennis
SpeciesCopelatus punctulatus Aubé, 1838  ⇒ Copelatus sahlbergi
SpeciesCopelatus pusillus Sharp, 1882  ⇒ Copelatus tenebrosus
SpeciesCopelatus puzhelongi Jiang, Hajek & Jia, 2022
SpeciesCopelatus quadrisignatus Régimbart, 1877
SpeciesCopelatus quadristriatus Pederzani & Schizzerotto, 2017
SpeciesCopelatus racenisi Guignot, 1951
SpeciesCopelatus ragazzii Régimbart, 1887
SpeciesCopelatus rasilis Lea, 1899  ⇒ Exocelina rasilis
SpeciesCopelatus regimbarti Branden, 1885
SpeciesCopelatus regimbartii Fairmaire, 1898
SpeciesCopelatus restrictus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus rimosus Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus rivalis Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus rocchii Bilardo, 1982
SpeciesCopelatus romani Zimmermann, 1924
Eveningsnow
SpeciesCopelatus rosulae Hajek, Jiang & Jia, 2022
SpeciesCopelatus royi Legros, 1958
SpeciesCopelatus rubiginosus Guignot, 1954
SpeciesCopelatus rufibasalis Gschwendtner, 1938
SpeciesCopelatus ruficapillus Régimbart, 1895
SpeciesCopelatus ruficollis Schaller, 1783
SpeciesCopelatus ruteri Legros, 1954
SpeciesCopelatus ryukyuensis Sato, 1961  ⇒ Copelatus oblitus
SpeciesCopelatus saegeri Guignot, 1955
SpeciesCopelatus safiotra Ranarilalatiana & Bergsten, 2019
SpeciesCopelatus sahlbergi J.Balfour-Browne, 1939
Chain fern, Chainfern		2Subspecies
SpeciesCopelatus sallaei Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus sanfilippoi Bilardo & Pederzani, 1972
SpeciesCopelatus saverii Pederzani & Schizzerotto, 2017
SpeciesCopelatus scalptus Guignot, 1954
SpeciesCopelatus scaphites Guignot, 1955
SpeciesCopelatus schaefferi Young, 1942
SpeciesCopelatus schalleri Gmelin, 1790
SpeciesCopelatus schereri Wewalka, 1981
SpeciesCopelatus schuhi Hendrich & Balke, 1998
SpeciesCopelatus scutatus Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus scytalotus Guignot, 1956
SpeciesCopelatus sectilis Guignot, 1941
SpeciesCopelatus semiformis Guignot, 1954
SpeciesCopelatus senegalensis Lagar & Hernando, 1991
SpeciesCopelatus sexstriatus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus seydeli Guignot, 1958
SpeciesCopelatus sharpi Branden, 1885  ⇒ Exocelina australis
SpeciesCopelatus shizong Balke & Bergsten, 2003
SpeciesCopelatus sibelaemontis Hájek, Hendrich, Hawlitschek & Balke, 2010
SpeciesCopelatus signatus Sharp, 1873
SpeciesCopelatus silvestrii Régimbart, 1903
SpeciesCopelatus simoni Régimbart, 1889
SpeciesCopelatus simplex Clark, 1863  ⇒ Copelatus aldabricus
SpeciesCopelatus singularis Bilardo & Rocchi, 1995
SpeciesCopelatus sociennus J.Balfour-Browne, 1952
1Subspecies
SpeciesCopelatus solitarius Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus somalicus Guignot, 1952
SpeciesCopelatus sordidipennis Régimbart, 1895
SpeciesCopelatus spangleri Vazirani, 1974  ⇒ Copelatus wewalkai
SpeciesCopelatus spoliatus Guignot, 1955
SpeciesCopelatus stavropolitanus ♰ Říha, 1974
SpeciesCopelatus stillicidicola Bilardo & Rocchi, 1995
SpeciesCopelatus striatellus Boheman, 1848
SpeciesCopelatus striaticollis Lucas, 1857
SpeciesCopelatus striatipennis Dejean, 1836  ⇒ Copelatus sulcipennis
SpeciesCopelatus striatopterus Aubé, 1838
SpeciesCopelatus striatulus Aubé, 1838
SpeciesCopelatus strigipennis Laporte, 1835
SpeciesCopelatus strigosulus Fairmaire, 1878
SpeciesCopelatus strigulifer Régimbart, 1903
SpeciesCopelatus strigulosus Sharp, 1882  ⇒ Copelatus pulchellus
SpeciesCopelatus strinatii Guignot, 1958
SpeciesCopelatus striolatus Peschet, 1917
SpeciesCopelatus stygis Guignot, 1958
SpeciesCopelatus subconcolor Guignot, 1953
SpeciesCopelatus subdeficiens Régimbart, 1902
SpeciesCopelatus subfasciatus A. Zimmermann, 1919  ⇒ Copelatus oblitus
SpeciesCopelatus subjectus Sharp, 1882
SpeciesCopelatus subsimilis Guignot, 1958
SpeciesCopelatus substriatus Kirsch, 1873
SpeciesCopelatus subterraneus Guéorguiev, 1978
SpeciesCopelatus sudrei Bameul, 2003
SpeciesCopelatus sulcatus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus sulcipennis Laporte, 1835
SpeciesCopelatus sumbawensis Régimbart, 1899
SpeciesCopelatus suppar Guignot, 1956
SpeciesCopelatus supplementaris Régimbart, 1895
SpeciesCopelatus sylvaticoides Bilardo & Rocchi, 2013
SpeciesCopelatus sylvaticus Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus takakurai Satô, 1985
SpeciesCopelatus tanaus Guignot, 1953
SpeciesCopelatus taprobanicus Wewalka & Vazirani, 1985
SpeciesCopelatus tarapotensis Zimmermann, 1920
SpeciesCopelatus tarsalis Balfour-Browne, 1939
SpeciesCopelatus tenebrosis Régimbart, 1880
SpeciesCopelatus tenebrosus Régimbart, 1880
SpeciesCopelatus tengchongensis Hajek, Jiang & Jia, 2022
SpeciesCopelatus teranishii Kamiya, 1938
SpeciesCopelatus terminalis Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus ternatensis Régimbart, 1899
SpeciesCopelatus terzanii Bilardo & Saverio, 2016
SpeciesCopelatus thiriouxi Peschet, 1917
SpeciesCopelatus thrasys Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus tibialis Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus tinctor Guignot, 1954
SpeciesCopelatus togoensis Régimbart, 1895
SpeciesCopelatus tokaraensis Nakane, 1963  ⇒ Copelatus oblitus
SpeciesCopelatus tomokunii Satô, 1985
SpeciesCopelatus tondangoyei Bilardo & Rocchi, 2013
SpeciesCopelatus torcutus Guignot, 1954
SpeciesCopelatus tostus J.Balfour-Browne, 1950
SpeciesCopelatus trifilis Guignot, 1958
SpeciesCopelatus triglyphus Guignot, 1955
SpeciesCopelatus trilobatus Régimbart, 1895
SpeciesCopelatus tschaga Bilardo & Pederzani, 1972
SpeciesCopelatus tucuchiensis J.Balfour-Browne, 1939
SpeciesCopelatus uludanuensis Hendrich & Balke, 1995
SpeciesCopelatus undecimstriatus Aubé, 1838
SpeciesCopelatus unguicularis Régimbart, 1903
SpeciesCopelatus urceolus Hájek, Shaverdo, Hendrich & Balke, 2021
SpeciesCopelatus usagarensis Zimmermann, 1926
SpeciesCopelatus vagatus Guignot, 1952
Chain fern, Chainfern
SpeciesCopelatus vagus Bilardo & Rocchi, 1995
SpeciesCopelatus validus Sharp, 1882
Grand Junction camissonia
SpeciesCopelatus vancli Hájek, Hendrich & Balke, 2018
SpeciesCopelatus vanninii Bilardo & Rocchi, 1999
SpeciesCopelatus variegatus Régimbart, 1895
SpeciesCopelatus variistriatus Hájek, Shaverdo, Hendrich & Balke, 2021
SpeciesCopelatus venustus Bilardo & Rocchi, 1995
SpeciesCopelatus victoriae Clark, 1863
SpeciesCopelatus vigintistriatus Fairmaire, 1869
SpeciesCopelatus vigintisulcatus Régimbart, 1895
SpeciesCopelatus villiersi Guignot, 1950
Miller Canyon woodlandsnail
SpeciesCopelatus virungaensis Bilardo, 1982
SpeciesCopelatus vitraci Legros, 1948
SpeciesCopelatus vivax Guignot, 1953
SpeciesCopelatus vokoka Ranarilalatiana & Bergsten, 2019
SpeciesCopelatus wallacei J.Balfour-Browne, 1939
SpeciesCopelatus waltoni J.Balfour-Browne, 1950
SpeciesCopelatus wayanadensis Manivannan & Madani, 2012
SpeciesCopelatus waymarni
SpeciesCopelatus wewalkai Holmen & Vazirani, 1990
SpeciesCopelatus weyersi Régimbart, 1900  ⇒ Copelatus irinus
SpeciesCopelatus weymarni J. Balfour-Browne, 1946
SpeciesCopelatus xanthogrammus Régimbart, 1900
SpeciesCopelatus xiniudong Jiang, Hajek & Jia, 2022
SpeciesCopelatus yacumensis Guignot, 1957
SpeciesCopelatus yaguarete Scheers, 2018
SpeciesCopelatus yombensis Bilardo & Rocchi, 2015
SpeciesCopelatus zadiensis Bilardo & Rocchi, 1995
SpeciesCopelatus zanabato Manuel & Ramahandrison, 2022
SpeciesCopelatus zanatanensis Ranarilalatiana & Bergsten, 2019
SpeciesCopelatus zimmermanni Gschwendtner, 1934

Synonyms
Exocelina (Broun, 1886)
Liopterus (Stephens, 1835)
Pelocatus (Zaitzev, 1908)


Distribution of Copelatus Erichson, 1832
Argentina, Australia, Austria, Belgium, Belize, Benin, Bhutan, Bolivia, Brazil, Cameroon, Canada, Chile, China, Colombia, Costa Rica, Cuba, Denmark, Dominican Republic, East Timor, Ecuador, Egypt, Estonia, Ethiopia, Finland, France, Gabon, Gambia, Georgia, Germany, Greece, Guatemala, Guinea-Bissau, Guyana, Honduras, India, Indonesia, Ireland, Italy, Jamaica, Japan, Kenya, Latvia, Liberia, Lithuania, Luxembourg, Madagascar, Mexico, Mozambique, Namibia, Nepal, Netherlands, Nicaragua, Norway, Panama, Papua New Guinea, Peru, Poland, Republic South Africa, Russia, Senegal, Seychelles, Sierra Leone, Singapore, Slovakia, South Korea, Spain, Sri Lanka, Suriname, Sweden, Switzerland, Taiwan, Tanzania, Thailand, USA, Venezuela, Zimbabwe


Links, sources and further information
Copelatus at gbif.org
Copelatus at gbif.org
Copelatus at irmng.org
Copelatus at itis.gov