Cychrus (Fabricius 1794), Rare Snail-eating Beetles ⇒ Taxonomy and Distribution of Genus

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Taxonomy of Genus Cychrus Fabricius, 1794
Rare Snail-eating Beetles

Class	Insecta Linnaeus, 1758 Insects, True insects	4179 190570 2444470	Families Genera Species
Subclass	Pterygota Lang, 1888 Winged insects	3816 189502 2439455	Families Genera Species
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	3399 186457 2415487	Families Genera Species
Superorder	Holometabola Heider, 1889 Holometabolous Insects	1891 142849 2048615	Families Genera Species
Order	Coleoptera Linnaeus, 1758 Beetles, Coleopterans	592 62779 813516	Families Genera Species
Suborder	Adephaga Schellenberg, 1806 Adephagans, Ground Beetles and Water Beetles	23 6166 123695	Families Genera Species
Superfamily	Caraboidea Latreille, 1802	8 5520 109721	Families Genera Species
Family	Carabidae Latreille, 1802 Carabid beetles, Carabids, Ground Beetles, Tiger Beetles	5446 109537	Genera Species
Subfamily	Carabinae Latreille, 1802	155 13828	Genera Species
Tribe	Cychrini Laporte, 1834 Snail-eating ground beetles	12 1144	Genera Species
Subtribe	Cychrina Laporte, 1834	9 1110	Genera Species
Genus	Cychrus Fabricius, 1794 Rare Snail-eating Beetles	730	Species

Subnodes of Cychrus Fabricius, 1794

Subgenus	Cychrus (Cychropsis) Boileau, 1901	69	Species
Subgenus	Cychrus (Cychrus)	339	Species
Subgenus	Cychrus (Kryptocychrus) Cavazzuti, 1997 ⇒ Cychrus	1	Species
Subgenus	Cychrus (Shuocychropsis) Imura, 2002	11	Species
Species	Cychrus abuthumensis Deuve & J.Schmidt, 2005
Species	Cychrus acuticollis L. Dufour, 1857 ⇒ Cychrus (Cychrus) dufouri
Species	Cychrus aeneicollis Beutenmüller, 1903
Species	Cychrus akcakocae Breuning, 1969 ⇒ Cychrus (Cychrus) anatolicus ponticus
Species	Cychrus albanicus Hopp, 1929
Species	Cychrus alternatus Motschulsky, 1859
Species	Cychrus altitudinum Deuve & Mourzine, 1998
Species	Cychrus altomontensis Branes, 1987 ⇒ Cychrus (Cychrus) spinicollis
Species	Cychrus alutaceus Sturm, 1826
Species	Cychrus ambulans Deuve, 1991
Species	Cychrus andrewsii T.W.Harris, 1839
Species	Cychrus angelli Beutenmüller, 1918
Species	Cychrus angulatus T.W.Harris, 1839
Species	Cychrus angustemarginatus Mandl, 1977
Species	Cychrus angusticollis Mannerheim, 1823
Species	Cychrus angustithorax Deuve, 1991 ⇒ Cychrus (Cychrus) cordicollis
Species	Cychrus anomalognathus Imura & Cavazzuti, 2018
Species	Cychrus artenuatus Fabricius, 1792
Species	Cychrus atratus Venet, 1926
Species	Cychrus baiburtensis Breuning, 1975
Species	Cychrus baimaicus Deuve, 2009
Species	Cychrus balcanicus Hopffgarten, 1881 ⇒ Cychrus (Cychrus) anatolicus occidentalis
Species	Cychrus ballionis Retowski, 1888 ⇒ Cychrus (Cychrus) aeneus starcki
Species	Cychrus baojiensis Imura, 1993
Species	Cychrus barkamensis Deuve, 1991	6	Subspecies
Species	Cychrus basumtsoensis Deuve, 2001
Species	Cychrus becvari Deuve, 2018
Species	Cychrus belasicensis Mandl, 1989 ⇒ Cychrus (Cychrus) semigranosus balcanicus
Species	Cychrus belousovi O.Berlov, 1997
Species	Cychrus bicarinatus LeConte, 1853
Species	Cychrus biceratops Cavazzuti, 1999
Species	Cychrus bilobus Say, 1823
Species	Cychrus bovelinii Heer, 1837
Species	Cychrus breuningi Schweiger, 1964 ⇒ Cychrus (Cychrus) anatolicus ponticus
Species	Cychrus businskyianus Imura, 2000
Species	Cychrus capito Lewis, 1881
Species	Cychrus carniolicus Motschulsky, 1866 ⇒ Cychrus (Cychrus) attenuatus attenuatus
Species	Cychrus carsicola Depoli, 1929 ⇒ Cychrus (Cychrus) attenuatus attenuatus
Species	Cychrus cavazzutii Deuve, 2002
Species	Cychrus chuandongziensis Deuve, 2017
Species	Cychrus coadunatus DeGeer, 1774 ⇒ Cychrus (Cychrus) caraboides caraboides
Species	Cychrus colasi Deuve, 1993
Species	Cychrus constrictus LeConte, 1853
Species	Cychrus convexus A. Morawitz, 1863 ⇒ Cychrus (Cychrus) morawitzi morawitzi
Species	Cychrus coptolabroides Imura & Cavazzuti, 2017
Species	Cychrus cordatus LeConte, 1853
Species	Cychrus coreicus Breuning, 1950
Species	Cychrus corvus Fall, 1910
Species	Cychrus costai Emery, 1872
Species	Cychrus crenatus Motschulsky, 1859
Species	Cychrus cristatus T.W.Harris, 1839
Species	Cychrus cyanicolor Deuve, 2006
Species	Cychrus dabashanensis Imura, 1995
Species	Cychrus dakogicus Cavazzuti, 2003
Species	Cychrus daliangshanus Imura, 1998
Species	Cychrus dandoensis Deuve, 2007
Species	Cychrus darcangensis Deuve, 1995
Species	Cychrus deoraliensis Deuve, 2013
Species	Cychrus depressus Eidam, 1925 ⇒ Cychrus (Cychrus) caraboides caraboides
Species	Cychrus dilatatus LeConte, 1853
Species	Cychrus dinaricus Mandl, 1989
Species	Cychrus discoidicollis Deuve, 2017
Species	Cychrus dissolutus Schaum, 1863
Species	Cychrus diversithorax Pic, 1916
Species	Cychrus drannadaghensis Breuning, 1967 ⇒ Cychrus (Cychrus) anatolicus ilgazdaghensis
Species	Cychrus dreuxi Deuve, 2013
Species	Cychrus elevatus Knoch, 1801
Species	Cychrus elongatus Audinet‑Serville, 1821 ⇒ Cychrus (Cychrus) caraboides caraboides
Species	Cychrus equiphilus Deuve, 1998
Species	Cychrus eremicola Deuve & J.Schmidt, 2005
Species	Cychrus erlangshanensis Kleinfeld, 2000
Species	Cychrus erwanrozoi Deuve, 2012
Species	Cychrus fabriciusi Burmeister, 1939
Species	Cychrus fopinganus Cavazzuti, 1999
Species	Cychrus forceps Imura & Cavazzuti, 2017
Species	Cychrus forestae Deuve, 1996
Species	Cychrus frivaldszkyi Roeschke, 1907
Species	Cychrus fuchsianus Rivers, 1890
Species	Cychrus fugongensis Imura & Cavazzuti, 2018
Species	Cychrus fulleri G.Horn, 1879
Species	Cychrus fulvus Letzner, 1849
Species	Cychrus gansuicola Deuve, 2007
Species	Cychrus germari Chaudoir, 1861
Species	Cychrus gigantea Deuve, 1992
Species	Cychrus glabripennis Imura, 1998
Species	Cychrus gnathochaetus Deuve, 1998
Species	Cychrus goergneri Korell & Kleinfeld, 1988
Species	Cychrus gorodinskiianus Deuve, 2011
Species	Cychrus gorodinskiorum Deuve, 2013
Species	Cychrus gracilis Géhin, 1885
Species	Cychrus graecus Breuning, 1967
Species	Cychrus granulatus Motschulsky, 1850 ⇒ Cychrus (Cychrus) aeneus aeneus
Species	Cychrus grososae Deuve, 1995
Species	Cychrus grummulifer Deuve, 1994
Species	Cychrus guyotii LeConte, 1863
Species	Cychrus habaensis Deuve & Mourzine, 1998
Species	Cychrus haeckeli Deuve, 1997
Species	Cychrus heinzi Deuve, 2013
Species	Cychrus heishuiensis Deuve, 2004
Species	Cychrus helicophilus Deuve & Mourzine, 2009
Species	Cychrus hemphili G Horn, 1879 ⇒ Cychrus (Cychrus) hemphillii
Species	Cychrus hopaensis Deuve, 2002
Species	Cychrus hoppei Ganglbauer, 1891 ⇒ Cychrus (Cychrus) caraboides caraboides
Species	Cychrus huangi Deuve, 1992
Species	Cychrus hungi Kleinfeld, 2000
Species	Cychrus huxianensis Imura, 1993
Species	Cychrus huyaensis Imura & Cavazzuti, 2018
Species	Cychrus ibericus Jeanne, 1976 ⇒ Cychrus (Cychrus) spinicollis
Species	Cychrus idahoensis Webb, 1901
Species	Cychrus ilgazdaghensis MaÍan, 1932
Species	Cychrus indianae Blatchley, 1910
Species	Cychrus inessae O.Berlov, 1997	2	Subspecies
Species	Cychrus inops Deuve, 1997
Species	Cychrus intermedius Hampe, 1850 ⇒ Cychrus (Cychrus) caraboides caraboides
Species	Cychrus interruptus Ménétriés, 1843
Species	Cychrus irregularis Beutenmüller, 1903
Species	Cychrus jeanimarcilhaci Deuve, 2003
Species	Cychrus jiaolieshanus Imura & Cavazzuti, 2019
Species	Cychrus jinpingshanus Imura & Cavazzuti, 2017
Species	Cychrus jiudingensis Häckel & Sehnal, 2014
Species	Cychrus jiulongensis Deuve, 1994	2	Subspecies
Species	Cychrus jollyi Deuve & Mourzine, 1997
Species	Cychrus kabakianus Imura & Cavazzuti, 2017
Species	Cychrus kalabianus Deuve, 2019
Species	Cychrus kangdingensis Deuve, 1991 ⇒ Cychrus (Cychrus) cordicollis
Species	Cychrus kangensis Deuve & Tian, 2013
Species	Cychrus karalianus Deuve, 1996
Species	Cychrus karamani Mandl, 1989 ⇒ Cychrus (Cychrus) semigranosus semigranosus
Species	Cychrus kazantsevi Deuve, 2000
Species	Cychrus kelloggi Dury, 1912
Species	Cychrus krali Deuve, 1991
Species	Cychrus kucerai Deuve, 1997
Species	Cychrus lapponicus Géhin, 1885 ⇒ Cychrus (Cychrus) caraboides caraboides
Species	Cychrus latialis Luigioni, 1922
Species	Cychrus laticollis Jeanne, 1972
Species	Cychrus lativentris Motschulsky, 1850
Species	Cychrus lauzonensis Schatzmayr, 1939 ⇒ Cychrus (Cychrus) grajus
Species	Cychrus lazikouensis Deuve, 1997
Species	Cychrus lecontei Dejean, 1826
Species	Cychrus leonardii T.W.Harris, 1839
Species	Cychrus lianghensis Cavazzuti, 2008
Species	Cychrus liguricus Straneo, 1933 ⇒ Cychrus (Cychrus) attenuatus attenuatus
Species	Cychrus lingbaoensis Korell & Kleinfeld, 1999
Species	Cychrus liuyei Deuve & Tian, 2013
Species	Cychrus luolaluokuanus Imura & Cavazzuti, 2019
Species	Cychrus maerkangensis Cavazzuti, 2008
Species	Cychrus makalu Deuve, 2003
Species	Cychrus makarovi Berlov, 1997
Species	Cychrus manasluensis Deuve & J.Schmidt, 2005
Species	Cychrus manningensis Deuve & Mourzine, 2009
Species	Cychrus maoerganicus Deuve & Tian, 2007
Species	Cychrus maoniuensis Deuve, 1996
Species	Cychrus marginatus Fischer von Waldheim, 1820
Species	Cychrus markamensis Korell & Kleinfeld, 1999
Species	Cychrus mauxi Deuve, 1991
Species	Cychrus medicimons Imura, 1999
Species	Cychrus medodiyensis Breuning, 1969
Species	Cychrus mengenensis Breuning, 1975 ⇒ Cychrus (Cychrus) anatolicus ponticus
Species	Cychrus meridionalis Chaudoir, 1861 ⇒ Cychrus (Cychrus) italicus
Species	Cychrus merkelii G.Horn, 1890
Species	Cychrus mesodiyensis Breuning, 1964
Species	Cychrus mewaensis Deuve, 1997
Species	Cychrus mexicanus Bates, 1882
Species	Cychrus micangshanensis Imura & Okamoto, 2003
Species	Cychrus microthorax Deuve, 1992
Species	Cychrus milyaevi Berlov, 1997
Species	Cychrus mimus G.Horn, 1874
Species	Cychrus minjiangicus Deuve, 2009
Species	Cychrus minor ♰ G. H. Horn, 1876
Species	Cychrus minshanicus Deuve, 1992
Species	Cychrus modestior Deuve, 1991
Species	Cychrus montanus Winkler, 1939
Species	Cychrus montenegrinus Apfelbeck, 1904
Species	Cychrus morvanianus Deuve, 2003
Species	Cychrus mosorensis Winkler, 1939 ⇒ Cychrus (Cychrus) rugicollis
Species	Cychrus motianling Imura & Cavazzuti, 2018
Species	Cychrus mulinensis Deuve, 1992
Species	Cychrus nigricans Letzner, 1849
Species	Cychrus nixiensis Imura & Cavazzuti, 2018
Species	Cychrus obliquus LeConte, 1868
Species	Cychrus obsidianus Imura & Cavazzuti, 2020
Species	Cychrus occidentalis Mandl, 1977
Species	Cychrus olympicola Breuning, 1969 ⇒ Cychrus (Cychrus) semigranosus graecus
Species	Cychrus oreophilus Rivers, 1890
Species	Cychrus ovalis Motschulsky, 1859
Species	Cychrus pamensis Deuve, 1992
Species	Cychrus paramontanus Sehnal & Häckel, 2006
Species	Cychrus paraspinosus Korell & Kleinfeld, 1999
Species	Cychrus pehri SokoláÍ, 1907 ⇒ Cychrus (Cychrus) caraboides caraboides
Species	Cychrus peristericus Roubal, 1930 ⇒ Cychrus (Cychrus) semigranosus albanicus
Species	Cychrus picipes Herbst, 1806 ⇒ Cychrus (Cychrus) attenuatus attenuatus
Species	Cychrus pindicus Mandl, 1989 ⇒ Cychrus (Cychrus) semigranosus graecus
Species	Cychrus ponticus Schweiger, 1962
Species	Cychrus proboscideus Olivier, 1795 ⇒ Cychrus (Cychrus) attenuatus attenuatus
Species	Cychrus prolapsus Cavazzuti, 1999
Species	Cychrus prymnaeus Fischer von Waldheim, 1828 ⇒ Cychrus (Cychrus) caraboides caraboides
Species	Cychrus pseudobruggei Deuve, 1991
Species	Cychrus pseudokoiwayai Deuve & Mourzine, 1998
Species	Cychrus puchneri Kleinfeld, 2016
Species	Cychrus punctatus LeConte, 1859
Species	Cychrus punctipennis Reitter, 1901 ⇒ Cychrus (Cychrus) rugicollis
Species	Cychrus purpuratus Beutenmüller, 1918
Species	Cychrus pustulosus Casey, 1905
Species	Cychrus pygmaeus Chaudoir, 1837 ⇒ Cychrus (Cychrus) caraboides caraboides
Species	Cychrus pyrenaeus Kraatz, 1879 ⇒ Cychrus (Cychrus) caraboides caraboides
Species	Cychrus reductior Deuve, 2010
Species	Cychrus reflexus Fabricius, 1801
Species	Cychrus regularis LeConte, 1884
Species	Cychrus reiseri Apfelbeck, 1904 ⇒ Cychrus (Cychrus) rugicollis
Species	Cychrus relictus G.Horn, 1881
Species	Cychrus reni Deuve & Tian, 2013
Species	Cychrus reticulatus Motschulsky, 1850
Species	Cychrus reuteri Kleinfeld, 2018
Species	Cychrus rhilensis Roeschke, 1907
Species	Cychrus rickseckeri LeConte, 1884 ⇒ Cychrus (Cychrus) hemphillii
Species	Cychrus ridingsii Bland, 1863
Species	Cychrus rossehnali Deuve, 2013
Species	Cychrus rosti Roeschke, 1907 ⇒ Cychrus (Cychrus) aeneus starcki	8	Subspecies
Species	Cychrus rostislavi Deuve, 2009
Species	Cychrus rostratus Linnaeus, 1758 ⇒ Cychrus (Cychrus) caraboides caraboides
Species	Cychrus rotundatus Letzner, 1849
Species	Cychrus rotundicollis Jan, 1835
Species	Cychrus rugiceps G.Horn, 1872
Species	Cychrus rugipennis Motschulsky, 1861
Species	Cychrus rugocephalus Deuve, 2014
Species	Cychrus ruspolii Breuning, 1964
Species	Cychrus sabaudus Fauvel, 1882 ⇒ Cychrus (Cychrus) caraboides caraboides
Species	Cychrus sabdensis Deuve, 1994	2	Subspecies
Species	Cychrus sapporensis
Species	Cychrus schmidtii Chaudoir, 1837
Species	Cychrus sertarensis Deuve, 1995
Species	Cychrus setosulus Deuve, 1990 ⇒ Cychrus (Cychrus) stoetzneri stoetzneri
Species	Cychrus shadeicus Deuve, 2007
Species	Cychrus shamaevi Deuve, 1999
Species	Cychrus shangrilanus Imura & Cavazzuti, 2017
Species	Cychrus shankoucola Deuve, 1994	2	Subspecies
Species	Cychrus sharovae O. Berlov, 1997
Species	Cychrus shennongjia Imura, 1996
Species	Cychrus shimenicus Cavazzuti, 2007
Species	Cychrus siguniangensis Deuve, 1994
Species	Cychrus simillimus Deuve, 1997
Species	Cychrus snowi LeConte, 1881
Species	Cychrus soazigae Deuve, 1997
Species	Cychrus sola Cavazzuti, 1997
Species	Cychrus sorini Deuve & Mourzine, 1997
Species	Cychrus starcki Reitter, 1888	2	Subspecies
Species	Cychrus stenostomus Knoch, 1801
Species	Cychrus striatopunctatus Chaudoir, 1844
Species	Cychrus striatus LeConte, 1859
Species	Cychrus subkazantsevi Deuve, 2001
Species	Cychrus subtilis Schaum, 1863
Species	Cychrus tenuithorax Deuve, 2013
Species	Cychrus tenuoritarsalis Deuve & J.Schmidt, 2005
Species	Cychrus testeus Scudder, 1878
Species	Cychrus thamensis Deuve, 2013
Species	Cychrus tillyi Berlov, 1997
Species	Cychrus torretassoi Schatzmayr, 1928
Species	Cychrus torulosus Fischer von Waldheim, 1823 ⇒ Cychrus (Cychrus) semigranosus semigranosus
Species	Cychrus trabzonensis Deuve, 2002
Species	Cychrus travnikanus Apfelbeck, 1904
Species	Cychrus tridens Cavazzuti, 2007
Species	Cychrus tridentifer Cavazzuti, 2010
Species	Cychrus tubaemons Imura & Cavazzuti, 2017	2	Subspecies
Species	Cychrus velutinus Ménétriés, 1843
Species	Cychrus ventricosus Dejean, 1831
Species	Cychrus viduus Dejean, 1826
Species	Cychrus vignai Cavazzuti, 2011
Species	Cychrus violaceus LeConte, 1863
Species	Cychrus volkovi O.Berlov, 1997 ⇒ Cychrus (Cychrus) aeneus aeneus
Species	Cychrus weiperti Deuve & J.Schmidt, 2010
Species	Cychrus wenxianensis Deuve & Mourzine, 1997
Species	Cychrus wheatleyi ♰ G. H. Horn, 1876
Species	Cychrus wuduensis Deuve, 1994
Species	Cychrus xilingicus Cavazzuti, 2001
Species	Cychrus yajiangensis Deuve & Mourzine, 1998
Species	Cychrus yeliguanensis Cavazzuti, 2007
Species	Cychrus yuae Deuve, 1989
Species	Cychrus yulogxuicus Deuve, 1991
Species	Cychrus zamotailovi Berlov, 1997
Species	Cychrus zariquieyi Born, 1924
Species	Cychrus zheguensis Cavazzuti, 2003
Species	Cychrus zhimahensis Imura & Cavazzuti, 2019
Species	Cychrus zhongdianus Deuve, 2005
Species	Cychrus zuopengensis Imura & Cavazzuti, 2019