|
|
| | | | Checklist of Subfamily Geotrupinae Latreille, 1802
for Continents
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 2109
116061
1221557 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 2100
115859
1220540 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 1811
113709
1208758 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 928
84435
1006159 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 234
35483
419175 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 189
32132
363198 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 23
3594
50210 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 23
3594
50210 | Families
Genera
Species |
Taxom
| | Family | Geotrupidae
Latreille, 1802
Dor beetles, Dung beetles, Earth-Boring Dung Beetle | 104
1403 | Genera
Species |
Taxom
| | Subfamily | Geotrupinae
Latreille, 1802
| 36
628 | Genera
Species |
Taxom
|
Genera for Continents
36 Genera with 628 Species
| Allotrupes Boucomont, 1896 | | 1 | Species | Taxom | | Anoplotrupes Jekel, 1791 | | 4 | Species | Taxom | | Baraudia Lopez-colon, 1839 | | 2 | Species | Taxom | | Ceratophyus Fischer von ..., 1887 | | 16 | Species | Taxom | | Ceratotrupes Jekel, 1865 | | 5 | Species | Taxom | | Chelotrupes Jekel, 1789 | | 9 | Species | Taxom | | Cnemotrupes Jekel, 1866 | | 24 | Species | Taxom | | Cretogeotrupes Nikolajev, 1990 | | 2 | Species | Taxom | | Enoplotrupes Lucas, 2022 | | 33 | Species | Taxom | | Epigeotrupes Bovo & Zunino, 1891 | | 4 | Species | Taxom | | Frickius Germain, 1897 | | 3 | Species | Taxom | | Geohowdenius Zunino, 1853 | | 2 | Species | Taxom | | Geotrupes Latreille, 0 | Dor Beetle | 98 | Species | Taxom | | Glyptogeotrupes Nikolaev, 2004 | | 3 | Species | Taxom | | Halffterius Zunino, 1866 | | 1 | Species | Taxom | | Haplogeotrupes Nikolaev, 1980 | | 5 | Species | Taxom | | Jekelius Lopez-colon, 1865 | | 30 | Species | Taxom | | Lethrus Scopoli, 2022 | | 159 | Species | Taxom | | Lithogeotrupes Nikolajev, 2008 | | 1 | Species | Taxom | | Megatrupes Zunino, 1967 | | 2 | Species | Taxom | | Mycotrupes LeConte, 1866 | | 5 | Species | Taxom | | Odontotrypes Fairmaire, 0 | | 111 | Species | Taxom | | Onthotrupes Howden, 1865 | | 10 | Species | Taxom | | Parageotrupes Nikolajev & Ren, 2010 | | 1 | Species | Taxom | | Peltotrupes Blanchard, 1955 | | 5 | Species | Taxom | | Phelotrupes Motschulsky, 2010 | | 90 | Species | Taxom | | Pseudotrypocopris MikÜic, 1861 | | 1 | Species | Taxom | | Sericotrupes Zunino, 1802 | | 2 | Species | Taxom | | Silphotrupes Jekel, 1840 | | 6 | Species | Taxom | | Sinogeotrupes Bovo & Zunino, 1905 | | 15 | Species | Taxom | | Taurocerastes Philippi, 1866 | | 1 | Species | Taxom | | Thorectes Mulsant, 1954 | | 56 | Species | Taxom | | Trypocopris Motschulsky, 1918 | | 30 | Species | Taxom | | Trypocoris | | 1 | Species | Taxom | | Typhaeus Leach, 1852 | | 9 | Species | Taxom | | Zuninoeus López-Coló..., 1825 | | 2 | Species | Taxom |
Species list for Continents
628 Species and 92 Subspecies | Allotrupes mandibularis Reitter, 1896 | | | Anoplotrupes balyi Jekel, 1866 | | | Anoplotrupes hornii Blanchard, 1888 | | | Anoplotrupes stercorosus Scriba, 1791 | Dor Beetle | | Baraudia geminata (Gené, 1839) | ⇒ Thorectes geminatus Géné, 1839 | | Ceratophyus alloini Prunier, Tau..., 2016 | | | Ceratophyus amdoensis Král & Jákl, 2020 | | | Ceratophyus dauricus Jekel, 1866 | | | Ceratophyus fischeri (Montrouzier, 1860) | ⇒ Ceratophyus polyceros Pallas, 1771 | | Ceratophyus gopherinus Cartwright, 1966 | Gopher Beetle | | Ceratophyus hoffmannseggi Fairmaire, 1856 | | | Ceratophyus kabaki Nikolajev, 2007 | | | Ceratophyus maghrebinicus Hillert & Kral, 2013 | | | Ceratophyus martinezi Lauffer, 1909 | | | Ceratophyus mesasiaticus Medvedev & N..., 1974 | | | Ceratophyus polyceros Pallas, 1771 | | | Ceratophyus rossii Jekel, 1866 | | | Ceratophyus schaffrathi Hillert & Kral, 2013 | | | Ceratophyus sinicus Zunino, 1973 | | | Ceratophyus sulcicornis Fairmaire, 1887 | | | Ceratotrupes bolivari Halffter & M..., 1962 | Bull klipfish | | Ceratotrupes fronticornis Erichson, 1847 | | | Ceratotrupes sturmi Jekel, 1866 | | | Ceratotrupes sturmii Jekel, 1865 | | | Chelotrupes algarvicus Hillert, KrÃ..., 2012 | | | Chelotrupes brancoi Hillert, KrÃ..., 2012 | | | Chelotrupes feryi Hillert, KrÃ..., 2012 | | | Chelotrupes hendrichi Hillert, KrÃ..., 2012 | | | Chelotrupes hiostius Géné, 1836 | | | Chelotrupes kyliesi Hillert, KrÃ..., 2012 | | | Chelotrupes laevipennis Mulsant & Go..., 1855 | | | Chelotrupes matutinalis Baudi di Selve, 1870 | | | Chelotrupes momus Olivier, 1789 | | | Cnemotrupes blackburni Fabricius, 1781 | | | Cnemotrupes blackburni blackburni blackburni F..., 1781 | | | Cnemotrupes blackburni excrementi Say, 1823 | | | Cnemotrupes blackburnii blackburnii | | | Cnemotrupes blackburnii excrementi Say, 1823 | | | Cnemotrupes cnephosa Howden, 1964 | | | Cnemotrupes cnephosus Howden, 1964 | | | Cnemotrupes guerreroensis (Howden, 1964) | ⇒ Onthotrupes guerreroensis Howden, 1964 | | Cnemotrupes herbeus Jekel, 1866 | | | Cnemotrupes lobatus Howden, 1974 | | | Cnemotrupes nebularum (Howden, 1964) | ⇒ Onthotrupes nebularum Howden, 1964 | | Cnemotrupes occidentalis G. Horn, 1880 | | | Cnemotrupes pecki Howden, 1974 | | | Cnemotrupes pinalonensis Howden, 1994 | | | Cnemotrupes sallaei Jekel, 1866 | | | Cnemotrupes semiopacus Jekel, 1866 | | | Cnemotrupes sobrinus Jekel, 1866 | | | Cnemotrupes splendidus Fabricius, 1775 | | | Cnemotrupes splendidus miarophagus Say, 1823 | | | Cnemotrupes splendidus splendidus miarophagus Say, 1823 | | | Cnemotrupes truncaticornis Howden, 1964 | | | Cnemotrupes ulkei Blanchard, 1888 | | | Cnemotrupes viridiobscurus Jekel, 1866 | | | Cretogeotrupes convexus ♰ Nikolajev, 1992 | | | Cretogeotrupes nodusus Hong & Wang, 1990 | | | Enoplotrupes (Enoplotrupes) birmanicus Gestro, 1888 | | | Enoplotrupes (Enoplotrupes) chaslei Fairmaire, 1886 | | | Enoplotrupes (Enoplotrupes) crassicornis Boucomont, 1905 | | | Enoplotrupes (Enoplotrupes) largeteaui Oberthür, 1883 | | | Enoplotrupes (Enoplotrupes) sharpi Jordan & Rot..., 1893 | | | Enoplotrupes (Enoplotrupes) sinensis Lucas, 1869 | | | Enoplotrupes (Enoplotrupes) splendens Jordan & Rot..., 1893 | | | Enoplotrupes (Gynocoplotrupes) bieti Oberthur, 1883 | | | Enoplotrupes bieti bieti bieti Oberthur, 1883 | | | Enoplotrupes bieti morishitai Miyake, 1996 | | | Enoplotrupes (Gynocoplotrupes) latus Boucomont, 1909 | | | Enoplotrupes (Gynocoplotrupes) yunnanus Fairmaire, 1888 | | | Enoplotrupes annae Ochi, Kon & ..., 2017 | | | Enoplotrupes apatani Král, Jákl &..., 2020 | | | Enoplotrupes barmanicus Gestro, 1888 | | | Enoplotrupes chaslii Fairmaire, 1886 | | | Enoplotrupes frater Král, Malý &..., 2019 | | | Enoplotrupes gansuensis Král, Malý &..., 2015 | | | Enoplotrupes gaoligong Král, Malý &..., 2015 | | | Enoplotrupes howdeni Král, Malý &..., 2015 | | | Enoplotrupes irregularis Král, Malý &..., 2019 | | | Enoplotrupes kachinensis Král, Malý &..., 2015 | | | Enoplotrupes kubani Král, Malý &..., 2015 | | | Enoplotrupes kumei (Masumoto, 1991) | ⇒ Enoplotrupes sinensis Lucas, 1869 | | Enoplotrupes mishmi Král, Malý &..., 2019 | | | Enoplotrupes monachus Král, Malý &..., 2015 | | | Enoplotrupes rhinoceros Král, Maly &..., 2012 | | | Enoplotrupes schoolmeestersi Ochi, Kon & Bai, 2022 | | | Enoplotrupes tawangensis Gupta, Chand..., 2019 | | | Enoplotrupes tyrannus Král, Maly &..., 2012 | | | Enoplotrupes variicolor (Fairmaire, 1886) | ⇒ Enoplotrupes largeteaui Oberthür, 1883 | | Enoplotrupes xizangensis Ochi, Kon & Bai, 2022 | | | Epigeotrupes castaneipennis Reitter, 1887 | | | Epigeotrupes crenulipennis Fairmaire, 1891 | | | Epigeotrupes infraopacus Fairmaire, 1901 | | | Epigeotrupes kuluensis Bates, 1891 | | | Frickius costulatus Germain, 1897 | | | Frickius variolosus Germain, 1897 | | | Geohowdenius egeriei Germar, 1824 | | | Geohowdenius opacus Haldeman, 1853 | | | Geotrupes (Anoplotrupes) balyi (Jekel, 1865) | ⇒ Anoplotrupes balyi Jekel, 1866 | | Geotrupes (Anoplotrupes) hornii (Blanchard, 1888) | ⇒ Anoplotrupes hornii Blanchard, 1888 | | Geotrupes (Anoplotrupes) stercorosus Scriba, 1791 | | | Geotrupes (Cnemotrupes) blackburnii Fabricius, 1781 | Blackburn'S Earth-Boring Beetle | | Geotrupes blackburnii blackburnii Fabricius, 1781 | | | Geotrupes blackburnii excrementi Say, 1823 | | | Geotrupes (Cnemotrupes) semiopacus (Jekel, 1865) | ⇒ Cnemotrupes semiopacus Jekel, 1866 | | Geotrupes (Cnemotrupes) splendidulus Fabricius, 1775 | | | Geotrupes splendidulus miarophagus Say, 1823 | | | Geotrupes splendidulus splendidulus Fabricius, 1775 | | | Geotrupes (Cnemotrupes) ulkei (Blanchard, 1888) | ⇒ Cnemotrupes ulkei Blanchard, 1888 | | Geotrupes (Geohowdenius) egeriei (Germar, 1824) | ⇒ Geohowdenius egeriei Germar, 1824 | | Geotrupes (Geohowdenius) opacus (Haldeman, 1853) | ⇒ Geohowdenius opacus Haldeman, 1853 | | Geotrupes (Geotrupes) baicalicus Reitter, 1892 | | | Geotrupes (Geotrupes) corinthius Fairmaire, 1886 | | | Geotrupes (Geotrupes) folwarcznyi Cervenka, 2005 | | | Geotrupes (Geotrupes) genestieri Boucomont, 1904 | | | Geotrupes (Geotrupes) ibericus Baraud, 1958 | | | Geotrupes (Geotrupes) jakovlevi Semenov, 1891 | | | Geotrupes (Geotrupes) kashmirensis Sharp, 1878 | | | Geotrupes (Geotrupes) koltzei Reitter, 1893 | | | Geotrupes (Geotrupes) lenardoi Petrovitz, 1973 | | | Geotrupes (Geotrupes) mutator Marsham, 1802 | | | Geotrupes (Geotrupes) olgae Olsoufieff, 1918 | | | Geotrupes (Geotrupes) puncticollis (Malinowsky, 1811) | ⇒ Thorectes puncticollis Lucas, 1845 | | Geotrupes (Geotrupes) spiniger Marsham, 1802 | Dor Beetle | | Geotrupes (Geotrupes) stercorarius (Marsham, 1802) | ⇒ Geotrupes spiniger Marsham, 1802 | | Geotrupes (Glyptogeotrupes) castaneipennis Reitter, 1887 | | | Geotrupes (Glyptogeotrupes) castanipennis Reitter, 1887 | | | Geotrupes (Glyptogeotrupes) crenulipennis Fairmaire, 1891 | | | Geotrupes (Glyptogeotrupes) impressus (Gebler, 1841) | ⇒ Geotrupes jakovlevi Semenov, 1891 | | Geotrupes (Glyptogeotrupes) kuluensis Bates, 1891 | | | Geotrupes (Glyptogeotrupes) molestus Faldermann, 1835 | | | Geotrupes (Megatrupes) cavicollis (Bates, 1887) | ⇒ Megatrupes cavicollis Bates, 1887 | | Geotrupes (Stereopyge) douei Gory, 1841 | | | Geotrupes alpinus (Hagenbach, 1825) | ⇒ Trypocopris alpinus alpinus Sturm & Hage..., 1825 | | Geotrupes amoenus (Jacobson, 1893) | ⇒ Geotrupes koltzei Reitter, 1893 | | Geotrupes armicrus (Fairmaire, 1888) | ⇒ Phelotrupes tenuestriatus Fairmaire, 1887 | | Geotrupes atavus ♰ E. Oustalet, 1874 | | | Geotrupes auratus (Dallatorre, 1879) | ⇒ Phelotrupes auratus auratus auratus Mots..., 1858 | | Geotrupes blackburni (Palisot de B..., 1805) | ⇒ Cnemotrupes splendidus Fabricius, 1775 | | Geotrupes chalybaeus (Mulsant, 1842) | ⇒ Peltotrupes profundus Howden, 1952 | | Geotrupes cnefosa Howden & H.F., 1964 | | | Geotrupes coruscans (Chevrolat, 1840) | ⇒ Trypocopris pyrenaeus pyrenaeus Charpentier, 1825 | | Geotrupes denticulatus (Boucomont, 1905) | ⇒ Phelotrupes denticulatus Boucomont, 1905 | | Geotrupes escorialensis (Jekel, 1866) | ⇒ Thorectes escorialensis Jekel, 1866 | | Geotrupes fisheri (Howden, 1967) | ⇒ Megatrupes fisheri Howden, 1967 | | Geotrupes geminatus (Géné, 1839) | ⇒ Thorectes geminatus | | Geotrupes germari ♰ O. Heer, 1862 | | | Geotrupes guatemalensis (Bates, 1897) | ⇒ Haplogeotrupes guatemalensis Bates, 1887 | | Geotrupes herbea (Jekel, 1865) | ⇒ Onthotrupes herbeus Jekel, 1865 | | Geotrupes hispanus (Reitter, 1893) | ⇒ Thorectes hispanus Reitter, 1892 | | Geotrupes hypocrita (LePeletier &..., 1828) | ⇒ Sericotrupes niger Marsham, 1802 | | Geotrupes inermis (Ménétriés, 1832) | ⇒ Trypocopris inermis Ménétriés, 1832 | | Geotrupes insulana (Howden, 1965) | ⇒ Sinogeotrupes insulanus Howden, 1965 | | Geotrupes intermedius Ferrari, 1852 | | | Geotrupes jakowlewi Semenov, 1891 | | | Geotrupes jekeli (Horn, 1868) | ⇒ Phelotrupes jekeli Harold, 1869 | | Geotrupes jiaoyanshanense ♰ Y. C. Hong, 1983 | | | Geotrupes jiaoyanshanensis Hong, 1983 | | | Geotrupes koiwayai (Masumoto, 1995) | ⇒ Odontotrypes roborowskyi Reitter, 1887 | | Geotrupes laevifrons (Jekel, 1866) | ⇒ Phelotrupes laevifrons Jekel, 1866 | | Geotrupes laevigatus (Mulsant, 1842) | ⇒ Thorectes intermedius Costa, 1839 | | Geotrupes laevis (Haworth, 1807) | ⇒ Trypocopris vernalis vernalis Linnaeus, 1758 | | Geotrupes laevistriatus (Miwa, 1930) | ⇒ Phelotrupes laevistriatus Motschulsky, 1858 | | Geotrupes laevistriatus formosanus (Miwa, 1930) | ⇒ Phelotrupes formosanus Miwa, 1930 | | Geotrupes martensi (Krikken, 1981) | ⇒ Thorectes martensi Krikken, 1981 | | Geotrupes messelensis ♰ F. Meunier, 1921 | | | Geotrupes niger (Marsham, 1802) | ⇒ Sericotrupes niger Marsham, 1802 | | Geotrupes nitidus (Jekel, 1865) | ⇒ Thorectes nitidus Jekel, 1866 | | Geotrupes occidentalis (Horn, 1880) | ⇒ Cnemotrupes occidentalis G. Horn, 1880 | | Geotrupes onitidipes (Bates, 1887) | ⇒ Onthotrupes onitidipes Bates, 1887 | | Geotrupes orichalceus (Fairmaire, 1895) | ⇒ Odontotrypes orichalceus Fairmaire, 1895 | | Geotrupes oshimanus (Fairmaire, 1895) | ⇒ Phelotrupes oshimanus Fairmaire, 1895 | | Geotrupes ovchinnikovi Nikolajev, 2004 | | | Geotrupes proaevus Germar, 1849 | | | Geotrupes purpureus (Küster, 1852) | ⇒ Trypocopris fulgidus Motschulsky, 1845 | | Geotrupes pyrenaeus Charp., 1825 | | | Geotrupes retusus (Le Conte, 1866) | ⇒ Mycotrupes retusus Le Conte, 1866 | | Geotrupes roborovskii | | | Geotrupes rottensis ♰ G. Statz, 1952 | | | Geotrupes ruficornis | | | Geotrupes rugatulus (Jekel, 1865) | ⇒ Thorectes rugatulus Jekel, 1866 | | Geotrupes splendens (Erichson, 1848) | ⇒ Trypocopris pyrenaeus pyrenaeus Charpentier, 1825 | | Geotrupes splendidus Fabricius, 1775 | Golden scarab | | Geotrupes stellosus (Krikken, 1981) | ⇒ Thorectes stellosus stellosus annapurnicus..., 1981 | | Geotrupes substriatellus (Fairmaire, 1897) | ⇒ Sinogeotrupes substriatellus Fairmaire, 1897 | | Geotrupes sylvaticus (Panzer, 1799) | ⇒ Anoplotrupes stercorosus Scriba, 1791 | | Geotrupes uenoi (Masumoto, 1995) | ⇒ Odontotrypes uenoi Masumoto, 1995 | | Geotrupes vernalis Linnaeus, 1758 | | | Geotrupes vetustus ♰ E. F. Germar, 1837 | | | Geotrupes viridiobscurus (Jekel, 1865) | ⇒ Onthotrupes viridiobscurus Jekel, 1865 | | Geotrupes xue | | | Geotrupes yulong | | | Geotrupes yunnanus | | | Glyptogeotrupes impressus Gebler, 1841 | | | Glyptogeotrupes molestus Faldermann, 1835 | | | Glyptogeotrupes ovchinnikovi Nikolajev, 2004 | | | Halffterius rufoclavatus Jekel, 1866 | | | Haplogeotrupes guatemalensis Bates, 1887 | | | Haplogeotrupes guatemalensis guatemalensis guatemalensi..., 1887 | | | Haplogeotrupes guatemalensis tridentatus Howden, 1974 | | | Haplogeotrupes guatemalensis unidentatus Howden, 1974 | | | Haplogeotrupes reddelli Howden, 1980 | | | Jekelius (Jekelius) albarracinus (Wagner, 1928) | ⇒ Thorectes albarracinus Wagner, 1928 | | Jekelius (Jekelius) balearicus López-Coló..., 1984 | | | Jekelius (Jekelius) brullei (Jekel, 1866) | ⇒ Thorectes brullei Jekel, 1866 | | Jekelius brullei africanus (Baraud, 1965) | ⇒ Thorectes brullei africanus Baraud, 1965 | | Jekelius brullei anatolicus (Jekel, 1865) | ⇒ Thorectes brullei anatolicus Jekel, 1865 | | Jekelius brullei brullei | | | Jekelius brullei creticus Fairmaire, 1876 | | | Jekelius brullei syriacus (Jekel, 1865) | ⇒ Thorectes brullei syriacus Jekel, 1865 | | Jekelius (Jekelius) castillanus (López-Coló..., 1985) | ⇒ Thorectes castillanus Lopez-colon, 1985 | | Jekelius (Jekelius) catalonicus López-Coló..., 1991 | | | Jekelius (Jekelius) chalconotus Chevrolat, 1840 | | | Jekelius (Jekelius) chersinus (Delabie, 1954) | ⇒ Thorectes albarracinus Wagner, 1928 | | Jekelius (Jekelius) hernandezi (López-Coló..., 1988) | ⇒ Thorectes hernandezi Lopez-colon, 1988 | | Jekelius (Jekelius) hispanus (Reitter, 1983) | ⇒ Thorectes hispanus Reitter, 1892 | | Jekelius (Jekelius) intermedius (Costa, 1827) | ⇒ Thorectes intermedius Costa, 1839 | | Jekelius (Jekelius) juengueri Romero-Samper, 1996 | | | Jekelius (Jekelius) nitidus (Jekel, 1866) | ⇒ Thorectes nitidus Jekel, 1866 | | Jekelius (Jekelius) punctatolineatus (François, 1904) | ⇒ Thorectes punctatolineatus Francois, 1904 | | Jekelius (Jekelius) sardous (Erichson, 1847) | ⇒ Thorectes sardous Erichson, 1847 | | Jekelius (Jekelius) sericeus (Jekel, 1866) | ⇒ Thorectes sericeus Jekel, 1866 | | Jekelius (Reitterius) punctulatus Jekel, 1866 | | | Jekelius (Rudolfpetrovitzia) marginatus (Poiret, 1787) | ⇒ Thorectes marginatus Poiret, 1787 | | Jekelius bahilloi López-Colón, 2018 | | | Jekelius jeannei Huchet & Sommer, 2020 | | | Jekelius juengeri (Romero-Samper, 1996) | ⇒ Thorectes juengeri Romero-Samper, 1995 | | Jekelius rugatulus Jekel, 1865 | | | Lethrus (Abrognathus) majusculus Semenov, 1899 | | | Lethrus (Abrognathus) pygmaeus Ballion, 1871 | | | Lethrus (Abrognathus) tuberculifrons Ballion, 1871 | | | Lethrus (Autolethrus) raymondi Reitter, 1890 | | | Lethrus (Autolethrus) rotundicollis Fairmaire, 1866 | | | Lethrus (Ceratodirus) aequidentatus Nikolajev, 1968 | | | Lethrus (Ceratodirus) aralicus Nikolajev, 2003 | | | Lethrus (Ceratodirus) borealis Nikolajev, 1973 | | | Lethrus (Ceratodirus) cephalotes (Pallas, 1771) | ⇒ Ceratodirus longimanus Fischer von ..., 1821 | | Lethrus (Ceratodirus) dostojevskii Semenov, 1899 | | | Lethrus (Ceratodirus) gladiator Reitter, 1897 | | | Lethrus (Ceratodirus) karelini Gebler, 1845 | | | Lethrus (Ceratodirus) lamellifer Nikolajev, 1976 | | | Lethrus (Ceratodirus) longimanus Fischer von ..., 1821 | | | Lethrus (Ceratodirus) mugodzharicus Nikolajev, 1987 | | | Lethrus (Ceratodirus) serpentifer Semenov & Me..., 1935 | | | Lethrus (Furcilethrus) antovae Medvedev, 1957 | | | Lethrus (Furcilethrus) banghaasi Reitter, 1893 | | | Lethrus (Furcilethrus) frantsevichi Nikolajev, 1979 | | | Lethrus (Furcilethrus) frater Nikolajev, 1975 | | | Lethrus (Furcilethrus) furcatus Jakovlev, 1890 | | | Lethrus (Furcilethrus) kiritschenkoi Medvedev, 1965 | | | Lethrus (Furcilethrus) legezini Nikolajev, 2001 | | | Lethrus (Furcilethrus) michailovi Nikolajev & ..., 1977 | | | Lethrus (Furcilethrus) mikitovae Nikolajev & ..., 1989 | | | Lethrus (Furcilethrus) nasreddinovi Nikolajev, 1987 | | | Lethrus (Furcilethrus) saryhissoricus Nikolajev, 1987 | | | Lethrus (Furcilethrus) sohrab Nikolajev, 1976 | | | Lethrus staudingeri obsoletus (Semenov, 1891) | ⇒ Lethrus tenuidens obsoletus Semenov, 1894 | | Lethrus staudingeri staudingeri obsoletus Se..., 1894 | | | Lethrus (Furcilethrus) tadzhikorum Medvedev, 1959 | | | Lethrus (Furcilethrus) tenuidens Reitter, 1890 | | | Lethrus tenuidens obsoletus Semenov, 1894 | | | Lethrus tenuidens tenuidens | | | Lethrus (Heteroplistodus) bispinus Jakovlev, 1889 | | | Lethrus (Heteroplistodus) bulbocerus Fischer von ..., 1845 | | | Lethrus bulbocerus bulbocerus bulbocerus F..., 1845 | | | Lethrus bulbocerus tarbagataicus Nikolajev, 2001 | | | Lethrus (Heteroplistodus) cicatricosus Reitter, 1889 | | | Lethrus (Heteroplistodus) ciskungesicus Nikolajev, 2001 | | | Lethrus (Heteroplistodus) crenulatus Gebler, 1845 | | | Lethrus (Heteroplistodus) kentauensis Kral & Olexa, 1996 | | | Lethrus (Heteroplistodus) korshinskii Semenov, 1899 | | | Lethrus (Heteroplistodus) kuldzhensis Lebedev, 1926 | | | Lethrus (Heteroplistodus) lebedevi Semonov & Me..., 1935 | | | Lethrus (Heteroplistodus) lopatini Medvedev, 1959 | | | Lethrus (Heteroplistodus) medvedevi Semenov & Gu..., 1935 | | | Lethrus (Heteroplistodus) obliquus Semenov, 1894 | | | Lethrus (Heteroplistodus) potanini Jakovlev, 1889 | | | Lethrus (Heteroplistodus) serridens Nikolajev, 1971 | | | Lethrus (Heteroplistodus) tekesicus Nikolajev, 2003 | | | Lethrus (Heteroplistodus) tschitsherini Semenov, 1894 | | | Lethrus (Heteroplistodus) uncidens Nikolajev, 1976 | | | Lethrus (Lethrus) acutangulus Ballion, 1871 | | | Lethrus (Lethrus) anisodon Semenov & Gu..., 1935 | | | Lethrus (Lethrus) appendiculatus Jakovlev, 1891 | | | Lethrus (Lethrus) apterus Laxmamann, 1770 | | | Lethrus (Lethrus) ares Král, Rejsek..., 2001 | | | Lethrus (Lethrus) arnoldii Nikolajev, 1987 | | | Lethrus (Lethrus) bajsuntavicus Nikolajev, 2003 | | | Lethrus (Lethrus) brachiicollis Fairmaire, 1855 | | | Lethrus (Lethrus) chistjakovae Nikolajev, 2003 | | | Lethrus (Lethrus) elephas Reitter, 1890 | | | Lethrus (Lethrus) fallax Nikolajev, 1975 | | | Lethrus (Lethrus) geminatus Kraatz, 1882 | | | Lethrus (Lethrus) glaber Medvedev, 1962 | | | Lethrus (Lethrus) karateghinicus Nikolajev, 1976 | | | Lethrus (Lethrus) macrognathus Fairmaire, 1866 | | | Lethrus (Lethrus) marakandicus Nikolajev, 2003 | | | Lethrus (Lethrus) mucronatus Semenov, 1894 | | | Lethrus mucronatus tashkumyricus | | | Lethrus (Lethrus) politus Solsky, 1876 | | | Lethrus (Lethrus) schaumi Reitter, 1890 | | | Lethrus (Lethrus) scoparius Fischer von ..., 1821 | | | Lethrus scoparius laevigatus | | | Lethrus (Lethrus) sieversi Koshantschikov, 1894 | | | Lethrus (Lethrus) sogdianus Semenov, 1894 | | | Lethrus (Lethrus) splendidus Semenov & Me..., 1935 | | | Lethrus (Lethrus) submandibularis Lebedev, 1932 | | | Lethrus (Lethrus) sulcipennis (Kraatz, 1883) | ⇒ Lethrus mucronatus Semenov, 1894 | | Lethrus (Lethrus) superbus Kraatz, 1882 | | | Lethrus (Lethrus) tschatkalensis Protzenko, 1965 | | | Lethrus (Lethrus) vachshianus Nikolajev & ..., 1989 | | | Lethrus (Lethrus) zeravshanicus Nikolajev, 2003 | | | Lethrus (Mesolethrus) elisae Nikolajev, 2001 | | | Lethrus (Mesolethrus) inermis Reitter, 1897 | | | Lethrus (Mesolethrus) microbuccis Ballion, 1871 | | | Lethrus (Mesolethrus) nuratavicus Nikolajev, 1987 | | | Lethrus (Mesolethrus) shakhristanicus Nikolajev, 2003 | | | Lethrus (Neolethrus) arcanus Medvedev, 1971 | | | Lethrus (Paralethrus) bituberculatus Ballion, 1871 | | | Lethrus bituberculatus bituberculatus bituberculat..., 1871 | | | Lethrus bituberculatus impressifrons Ballion, 1870 | | | Lethrus bituberculatus macrodon Nikolajev, 1969 | | | Lethrus (Paralethrus) crassus Hillert, 2004 | | | Lethrus (Paralethrus) kabaki Nikolajev, 1998 | | | Lethrus (Paralethrus) karatavicus (Nikolajev & ..., 1971) | ⇒ Paralethrus karatavicus Nikolajev & ..., 1971 | | Lethrus (Paralethrus) turkestanicus Ballion, 1871 | | | Lethrus (Scelolethrus) aktavicus Nikolajev, 2003 | | | Lethrus (Scelolethrus) andrejewae Nikolajev, 1987 | | | Lethrus (Scelolethrus) auriculatus Semenov, 1894 | | | Lethrus (Scelolethrus) bactrianus Semenov, 1894 | | | Lethrus (Scelolethrus) baglanicus Nikolajev, 2003 | | | Lethrus (Scelolethrus) carinatus Nikolajev, 1976 | | | Lethrus (Scelolethrus) chorassanicus Semenov & Me..., 1935 | | | Lethrus (Scelolethrus) costatus Semenov, 1894 | | | Lethrus (Scelolethrus) crypticus Kral & Olexa, 1996 | | | Lethrus (Scelolethrus) gissaricus Nikolajev, 1976 | | | Lethrus (Scelolethrus) kattaghanicus Nikolajev, 1976 | | | Lethrus (Scelolethrus) koshantschikovi Semenov, 1891 | | | Lethrus (Scelolethrus) kryzhanovskii Medvedev, 1971 | | | Lethrus (Scelolethrus) mediocris Fairmaire, 1892 | | | Lethrus (Scelolethrus) mithras Reitter, 1904 | | | Lethrus (Scelolethrus) nikolajevi Medvedev, 1971 | | | Lethrus (Scelolethrus) obtritus Nikolajev, 1976 | | | Lethrus (Scelolethrus) rosmarus (Ballion, 1871) | ⇒ Paralethrus turkestanicus Ballion, 1870 | | Lethrus (Scelolethrus) spinimanus Jakovlev, 1892 | | | Lethrus (Scelolethrus) substriatus Kraatz, 1883 | | | Lethrus (Scelolethrus) sulcatus Kraatz, 1883 | | | Lethrus (Scelolethrus) tekkensis Kral & Olexa, 1996 | | | Lethrus (Scelolethrus) tenuepunctus Semenov & Me..., 1935 | | | Lethrus (Scelolethrus) tenuestriatus Nikolajev, 1976 | | | Lethrus (Scelolethrus) valentinae Nikolajev, 2003 | | | Lethrus aenescens (Fairmaire, 1892) | ⇒ Paralethrus turkestanicus Ballion, 1870 | | Lethrus ahriman Pak & Gubin, 2020 | | | Lethrus asmodeus Pak & Gubin, 2020 | | | Lethrus baiocchii Hillert & Sechi, 2014 | | | Lethrus bituberculifrons (Lebedev, 1926) | ⇒ Paralethrus bituberculatus Ballion, 1870 | | Lethrus dinotherium Wilkins, 1885 | | | Lethrus dostojewskii Semenov, 1899 | | | Lethrus fedtschenkoi (Semenov, 1894) | ⇒ Lethrus superbus Kraatz, 1882 | | Lethrus golestanicus Král & Hillert, 2021 | | | Lethrus halkidikiensis Kral & Hillert, 2013 | | | Lethrus jacobsoni (Semenov & Me..., 1935) | ⇒ Paralethrus bituberculatus Ballion, 1870 | | Lethrus jakovlevi (Semenov, 1891) | ⇒ Heteroplistodus bulbocerus Fischer von ..., 1845 | | Lethrus kiritshenkoi Medvedev, 1965 | | | Lethrus klimenkoi Bagaturov, 2018 | | | Lethrus korzhinskii (Semenov, 1899) | ⇒ Lethrus korshinskii Semenov, 1899 | | Lethrus koshantschikowi Semenov, 1892 | | | Lethrus kozhantchikovi | | | Lethrus kozhantschikovi (Semenov, 1891) | ⇒ Lethrus koshantschikovi Semenov, 1891 | | Lethrus laevigatus (Ballion, 1871) | ⇒ Lethrus scoparius Fischer von ..., 1821 | | Lethrus litvinovi Semenov, 1900 | | | Lethrus liviae Pittino, 2011 | | | Lethrus muskinabadensis Nikolajev & ..., 2014 | | | Lethrus nazarovi Gusakov, 2017 | | | Lethrus paki Ivanova, 2012 | | | Lethrus percutens Gusakov & Kl..., 2011 | | | Lethrus perun Kral & Hillert, 2013 | | | Lethrus pigmaeus | | | Lethrus rustam Nikolajev & ..., 2014 | | | Lethrus schneideri Kral & Hillert, 2013 | | | Lethrus strymonensis Kral & Hillert, 2013 | | | Lethrus thracicus Král & Hillert, 2018 | | | Lethrus toropovi Shapovalov, 2022 | | | Lithogeotrupes confusus Nikolajev, 2008 | | | Megatrupes cavicollis Bates, 1887 | | | Megatrupes fisheri Howden, 1967 | | | Mycotrupes cartwrighti Olson & Hubbell, 1954 | | | Mycotrupes gaigei Olson & Hubbell, 1954 | | | Mycotrupes lethroides Westwood, 1837 | | | Mycotrupes pedester Howden, 1954 | Scrub Island burrowing scarab | | Mycotrupes retusus Le Conte, 1866 | Sandhills Earth Boring Scarab Beetle | | Odontotrypes abaensis Ochi, Kon & Bai, 2018 | | | Odontotrypes anthracinus Král & Lu, 2020 | | | Odontotrypes arnaudi Howden, 2006 | | | Odontotrypes baimaensis Nikolajev & ..., 2014 | | | Odontotrypes balangensis Ochi, Kon & Bai, 2019 | | | Odontotrypes balthasari MikÜic, 1958 | | | Odontotrypes bhutan Král, Maly &..., 2001 | | | Odontotrypes biconiferus Fairmaire, 1887 | | | Odontotrypes bimaculatus Cervenka, 2006 | | | Odontotrypes cariosus Fairmaire, 1886 | | | Odontotrypes cavazzutii (Král, Maly &..., 2001) | ⇒ Odontotrypes zhongdianensis cavazzutii Kral & Maly ..., 2001 | | Odontotrypes caverniferus Nikolajev, 2014 | | | Odontotrypes cheni Ochi, Kon & Bai, 2017 | | | Odontotrypes cicatricosus Nikolajev, 1977 | | | Odontotrypes cobaltipennis Ochi, Kon & Bai, 2019 | | | Odontotrypes cribripennis Fairmaire, 1887 | | | Odontotrypes daochengicus Ochi, Kon & Bai, 2017 | | | Odontotrypes davidiani Nikolajev, 2009 | | | Odontotrypes densopunctatus Nikolajev & ..., 2014 | | | Odontotrypes duplopunctatus Nikolajev & ..., 2014 | | | Odontotrypes emei Král, Maly &..., 2001 | | | Odontotrypes erchushanus Ochi, Kon & Bai, 2018 | | | Odontotrypes farkaci Král, Maly &..., 2001 | | | Odontotrypes farkaci farkaci | | | Odontotrypes farkaci habaensis Ochi, Kon & Bai, 2018 | | | Odontotrypes fujiokai Ochi, Kon & ..., 2010 | | | Odontotrypes gaoersishanus Ochi, Kon & Bai, 2017 | | | Odontotrypes glaber Nikolajev, 1998 | | | Odontotrypes gnom Nikolajev, 2009 | | | Odontotrypes gogona Král, Maly &..., 2001 | | | Odontotrypes gongga Král, Maly &..., 2001 | | | Odontotrypes guoluoshanus Ochi, Kon & Bai, 2018 | | | Odontotrypes guzanensis Ochi, Kon & Bai, 2019 | | | Odontotrypes haba Král, Maly &..., 2001 | | | Odontotrypes hamai Ochi, Kon & ..., 2017 | | | Odontotrypes hayekae Miksic, 1961 | | | Odontotrypes hayekae adamiki Cervenka, 2007 | | | Odontotrypes hayekae hayekae | | | Odontotrypes hayeki MikÜic, 1961 | | | Odontotrypes howdeni Král, Maly &..., 2001 | | | Odontotrypes hubeicus Král, Malý &..., 2014 | | | Odontotrypes impressiusculus Fairmaire, 1887 | | | Odontotrypes jiuding Král, Maly &..., 2001 | | | Odontotrypes kabaki Nikolajev, 2006 | | | Odontotrypes kabakovi Nikolajev, 2007 | | | Odontotrypes kalabi Král, Maly &..., 2001 | | | Odontotrypes karnali Král, Maly &..., 2001 | | | Odontotrypes korolevi Nikolajev, 2013 | | | Odontotrypes kryzhanovskii Nikolajev, 1998 | | | Odontotrypes kucerai Král, Maly &..., 2001 | | | Odontotrypes lama Král, Maly &..., 2001 | | | Odontotrypes lassallei Král, Maly &..., 2001 | | | Odontotrypes lazikouensis Ochi, Kon & Bai, 2018 | | | Odontotrypes leptopterus Nikolajev & ..., 2014 | | | Odontotrypes liupan Král & Lu, 2020 | | | Odontotrypes luguensis Nikolajev, 2013 | | | Odontotrypes maedai Howden, 2006 | | | Odontotrypes marcoberrai Ochi, Kon & Bai, 2019 | | | Odontotrypes medogensis Ochi, Kon & Bai, 2018 | | | Odontotrypes medvedevi Nikolajev, 1998 | | | Odontotrypes meigu Shokhin, 2008 | | | Odontotrypes meyomintang Král, Maly &..., 2001 | | | Odontotrypes minshanensis Ochi, Kon & Bai, 2021 | | | Odontotrypes mirek Král, Maly &..., 2001 | | | Odontotrypes mursini Nikolajev, 1998 | | | Odontotrypes nepalensis Ochi, Kon & ..., 2017 | | | Odontotrypes nikodymi (Král, Maly &..., 2001) | ⇒ Odontotrypes semenowi Reitter, 1887 | | Odontotrypes opacipennis Ochi, Kon & Bai, 2021 | | | Odontotrypes orichalceus Fairmaire, 1895 | | | Odontotrypes paulusi Král, Maly &..., 2001 | | | Odontotrypes pauma (Král, Maly &..., 2001) | ⇒ Odontotrypes radiosus Fairmaire, 1895 | | Odontotrypes purpureiaeneus Nikolaev, 1976 | | | Odontotrypes purpureipunctatus Boucomont, 1905 | | | Odontotrypes qinghaiensis Ochi, Kon & Bai, 2020 | | | Odontotrypes qinling Král, Maly &..., 2001 | | | Odontotrypes radiosus Fairmaire, 1895 | | | Odontotrypes roborowskyi Reitter, 1887 | | | Odontotrypes rufipes Boucomont, 1905 | | | Odontotrypes rugososulcatus Nikolajev, 2009 | | | Odontotrypes sabde Král, Maly &..., 2001 | | | Odontotrypes satanas Král, Maly &..., 2001 | | | Odontotrypes schoolmeestersi Ochi, Kon & Bai, 2017 | | | Odontotrypes semenowi Reitter, 1887 | | | Odontotrypes semenowi nikodymi Král, Malý &..., 2001 | | | Odontotrypes semenowi semenowi | | | Odontotrypes semicribrosus Fairmaire, 1891 | | | Odontotrypes semirugosus Fairmaire, 1887 | | | Odontotrypes semiscribrosus Fairmaire, 1891 | | | Odontotrypes shokhini Nikolajev, 2009 | | | Odontotrypes sichuanensis Nikolajev, 2013 | | | Odontotrypes slavek Král, Maly &..., 2001 | | | Odontotrypes szetshwanus Nikolaev, 1977 | | | Odontotrypes taurus Boucomont, 1905 | | | Odontotrypes tawangensis Gupta, Chand..., 2016 | | | Odontotrypes tibetanus Nikolaev, 1976 | | | Odontotrypes tryznai Král, Maly &..., 2001 | | | Odontotrypes turnai Král, Maly &..., 2001 | | | Odontotrypes uenoi Masumoto, 1995 | | | Odontotrypes viridipennis Ochi, Kon & Bai, 2021 | | | Odontotrypes weidengensis Ochi, Kon & Bai, 2018 | | | Odontotrypes weiweii Ochi, Kon & Bai, 2017 | | | Odontotrypes xinluhaiensis Ochi, Kon & Bai, 2018 | | | Odontotrypes xizangensis Ochi, Kon & Bai, 2019 | | | Odontotrypes xue Král, Maly &..., 2001 | | | Odontotrypes yangi Ochi, Kon & Bai, 2017 | | | Odontotrypes yulong Král, Maly &..., 2001 | | | Odontotrypes zeduoensis Ochi, Kon & Bai, 2018 | | | Odontotrypes zhongdianensis Král, Maly &..., 2001 | | | Odontotrypes zhongdianensis cavazzutii Kral & Maly ..., 2001 | | | Odontotrypes zhongdianensis zhongdianensis | | | Onthotrupes guerreroensis Howden, 1964 | | | Onthotrupes herbeus Jekel, 1865 | | | Onthotrupes lobatus Howden, 1974 | | | Onthotrupes nebularum Howden, 1964 | | | Onthotrupes nuntiatus Kohlmann & A..., 2020 | | | Onthotrupes onitidipes Bates, 1887 | | | Onthotrupes pecki Howden, 1974 | | | Onthotrupes sobrinus Jekel, 1865 | | | Onthotrupes viridiobscurus Jekel, 1865 | | | Parageotrupes incanus ♰ G. V. Nikolajev, 2010 | | | Peltotrupes profundus Howden, 1952 | Crested weedfish | | Peltotrupes profundus dubius Howden, 1955 | | | Peltotrupes profundus profundus dubius Howden, 1955 | | | Peltotrupes youngi Howden, 1955 | Ocala burrowing scarab | | Phelotrupes (Chromogeotrupes) auratus Motschulsky, 1858 | | | Phelotrupes auratus auratus auratus Mots..., 1858 | | | Phelotrupes auratus yaku Tsukamoto, 1958 | | | Phelotrupes (Chromogeotrupes) bicolor Fairmaire, 1888 | | | Phelotrupes (Chromogeotrupes) immarginatus (Boucomont, 1904) | ⇒ Chromogeotrupes immarginatus Boucomont, 1904 | | Phelotrupes (Chromogeotrupes) scutellatus Fairmaire, 1887 | | | Phelotrupes (Eogeotrupes) amethystinus Jekel, 1866 | | | Phelotrupes amethystinus amethystinus | | | Phelotrupes amethystinus chinmontanus Ochi, Kon & ..., 2017 | | | Phelotrupes (Eogeotrupes) cambeforti Král, Maly &..., 2001 | | | Phelotrupes (Eogeotrupes) deuvei Král, Maly &..., 2001 | | | Phelotrupes (Eogeotrupes) formosanus Miwa, 1930 | | | Phelotrupes (Eogeotrupes) laevifrons (Jekel, 1866) | ⇒ Eogeotrupes laevifrons Jekel, 1865 | | Phelotrupes (Eogeotrupes) laevistriatus Motschulsky, 1858 | | | Phelotrupes (Eogeotrupes) oshimanus Fairmaire, 1895 | | | Phelotrupes oshimanus oshimanus | | | Phelotrupes oshimanus yukiae Ochi, Toguch..., 2022 | | | Phelotrupes (Eogeotrupes) tenuestriatus (Fairmaire, 1887) | ⇒ Eogeotrupes tenuestriatus Fairmaire, 1887 | | Phelotrupes (Phelotrupes) businskyorum Král, Maly &..., 2001 | | | Phelotrupes businskyorum businskyorum | | | Phelotrupes businskyorum kachinmontanus Ochi, Kon & ..., 2017 | | | Phelotrupes (Phelotrupes) davidis Deyrolle, 1878 | | | Phelotrupes (Phelotrupes) denticulatus Boucomont, 1905 | | | Phelotrupes (Phelotrupes) holzschuhi Král, Maly &..., 2001 | | | Phelotrupes holzschuhi holzschuhi | | | Phelotrupes holzschuhi myanmarensis Ochi, Kon & ..., 2017 | | | Phelotrupes (Phelotrupes) imurai Masumoto, 1995 | | | Phelotrupes (Phelotrupes) indicus Boucomont, 1905 | | | Phelotrupes (Phelotrupes) jekeli Harold, 1869 | | | Phelotrupes (Phelotrupes) kubani Král, Maly &..., 2001 | | | Phelotrupes (Phelotrupes) metallescens Fairmaire, 1894 | | | Phelotrupes (Phelotrupes) obscuratus Fairmaire, 1889 | | | Phelotrupes (Phelotrupes) orientalis Westwood, 1839 | | | Phelotrupes orientalis burmanicus Miksic, 1960 | | | Phelotrupes orientalis orientalis | | | Phelotrupes (Phelotrupes) wrzecionkoi Král, Maly &..., 2001 | | | Phelotrupes (Phelotrupes) yunnanensis Král, Maly &..., 2001 | | | Phelotrupes (Sinogeotrupes) insulanus Howden, 1965 | | | Phelotrupes (Sinogeotrupes) taiwanus Miyake & Yamaya, 1995 | | | Phelotrupes abei Ochi & Kon, 2015 | | | Phelotrupes annamiticus Král, 2016 | | | Phelotrupes armatus Boucomont, 1905 | | | Phelotrupes belousovi Nikolajev, 2007 | | | Phelotrupes beuteli Ochi, Kon & Bai, 2017 | | | Phelotrupes bolm (Kral & Maly ..., 2001) | ⇒ Sinogeotrupes bolm Král, Maly &..., 2001 | | Phelotrupes brunipennis Ochi & Kon, 2015 | | | Phelotrupes caii Ochi, Kon & Bai, 2017 | | | Phelotrupes cheni Ochi, Kon & Bai, 2010 | | | Phelotrupes chenwenlongi Ochi, Masumo..., 2017 | | | Phelotrupes compressidens Fairmaire, 1891 | | | Phelotrupes haroldi (Boucomont, 1905) | ⇒ Phelotrupes jekeli Harold, 1869 | | Phelotrupes hunanensis (Kral & Maly ..., 2001) | ⇒ Sinogeotrupes hunanensis Král, Maly &..., 2001 | | Phelotrupes jendeki (Kral & Maly ..., 2001) | ⇒ Sinogeotrupes jendeki Král, Maly &..., 2001 | | Phelotrupes kachinensis Ochi & Kon, 2015 | | | Phelotrupes keitai Ochi, Kon & ..., 2018 | | | Phelotrupes kerzhneri Shokhin, 2008 | | | Phelotrupes lama Cervenka, 2009 | | | Phelotrupes linzhiensis Ochi, Kon & Bai, 2021 | | | Phelotrupes luhuoensis Ochi, Kon & Bai, 2019 | | | Phelotrupes masudai Ochi, Kon & ..., 2010 | | | Phelotrupes masumotoi (Kral & Maly ..., 2001) | ⇒ Sinogeotrupes masumotoi Král, Maly &..., 2001 | | Phelotrupes motuoensis Ochi, Kon & Bai, 2020 | | | Phelotrupes nikolajevi (Kral & Maly ..., 2001) | ⇒ Sinogeotrupes nikolajevi Král, Maly &..., 2001 | | Phelotrupes oberthuri (Boucomont, 1905) | ⇒ Sinogeotrupes oberthuri Boucomont, 1905 | | Phelotrupes reni daxueshanus Ochi, Kon & Bai, 2018 | | | Phelotrupes reni gansuensis Ochi, Kon & Bai, 2017 | | | Phelotrupes reni reni | | | Phelotrupes schoolmeestersi Ochi, Kon & ..., 2010 | | | Phelotrupes schoolmeestersi gaoligongicus Ochi, Kon & Bai, 2018 | | | Phelotrupes schoolmeestersi schoolmeestersi | | | Phelotrupes semiannamiticus Ochi, Kon & ..., 2020 | | | Phelotrupes semibicolor Ochi, Kon & Bai, 2017 | | | Phelotrupes semidenticulatus Ochi, Kon & Bai, 2017 | | | Phelotrupes shangrilensis Ochi, Kon & Bai, 2017 | | | Phelotrupes smetanai (Kral & Maly ..., 2001) | ⇒ Sinogeotrupes smetanai Král, Maly &..., 2001 | | Phelotrupes strnadi (Kral & Maly ..., 2001) | ⇒ Sinogeotrupes strnadi Král, Maly &..., 2001 | | Phelotrupes strnadi axanensis Ochi, Kon & ..., 2018 | | | Phelotrupes strnadi strnadi | | | Phelotrupes subaeneus Ochi, Kon & Bai, 2010 | | | Phelotrupes substriatellus Fairmaire, 1897 | | | Phelotrupes todai serai Ochi, Kon & ..., 2020 | | | Phelotrupes todai todai | | | Phelotrupes tsukamotoi Ochi, Kon & ..., 2010 | | | Phelotrupes typhoeus Linnaeus, 1758 | | | Phelotrupes variolicollis Boucomont, 1904 | | | Phelotrupes vietnamicus Ochi, Kon & ..., 2017 | | | Phelotrupes weiweii Ochi, Kon & Bai, 2017 | | | Phelotrupes yangi Ochi, Kon & Bai, 2010 | | | Phelotrupes zhangi Ochi, Kon & Bai, 2010 | | | Pseudotrypocopris amedei Fairmaire, 1861 | | | Sericotrupes niger Marsham, 1802 | | | Silphotrupes escorialensis (Jekel, 1866) | ⇒ Thorectes escorialensis Jekel, 1866 | | Silphotrupes escorialensis escorialensis | | | Silphotrupes escorialensis opaculus (Reitter, 1893) | ⇒ Thorectes escorialensis opaculus Reitter, 1893 | | Silphotrupes orocantabricus Verdú & Gal..., 2000 | | | Silphotrupes punctatissimus (Chevrolat, 1840) | ⇒ Thorectes punctatissimus Chevrolat, 1840 | | Sinogeotrupes armatus Boucomont, 1905 | | | Sinogeotrupes belousovi Nikolajev, 2007 | | | Sinogeotrupes bolm Král, Maly &..., 2001 | | | Sinogeotrupes compressidens Fairmaire, 1891 | | | Sinogeotrupes hunanensis Král, Maly &..., 2001 | | | Sinogeotrupes insulanus Howden, 1965 | | | Sinogeotrupes jendeki Král, Maly &..., 2001 | | | Sinogeotrupes masumotoi Král, Maly &..., 2001 | | | Sinogeotrupes nikolajevi Král, Maly &..., 2001 | | | Sinogeotrupes oberthuri Boucomont, 1905 | | | Sinogeotrupes smetanai Král, Maly &..., 2001 | | | Sinogeotrupes strnadi Král, Maly &..., 2001 | | | Sinogeotrupes substriatellus Fairmaire, 1897 | | | Sinogeotrupes taiwanus Miyake & Yamaya, 1995 | | | Sinogeotrupes variolicollis Boucomont, 1905 | | | Taurocerastes patagonicus Philippi, 1866 | | | Thorectes (Baraudia) geminatus | | | Thorectes (Jekelius) albarracinus Wagner, 1928 | | | Thorectes (Jekelius) brullei Jekel, 1866 | | | Thorectes brullei africanus Baraud, 1965 | | | Thorectes brullei anatolicus Jekel, 1865 | | | Thorectes brullei brullei Jekel, 1866 | | | Thorectes brullei creticus Fairmaire, 1876 | | | Thorectes brullei syriacus Jekel, 1865 | | | Thorectes (Jekelius) castillanus Lopez-colon, 1985 | | | Thorectes (Jekelius) chalconotus (Chevrolat, 1840) | ⇒ Thorectes nitidus Jekel, 1866 | | Thorectes (Jekelius) hernandezi Lopez-colon, 1988 | | | Thorectes (Jekelius) hispanus Reitter, 1892 | | | Thorectes (Jekelius) intermedius Costa, 1839 | | | Thorectes intermedius intermedius | | | Thorectes (Jekelius) juengeri Romero-Samper, 1995 | | | Thorectes (Jekelius) marginatus Poiret, 1787 | | | Thorectes (Jekelius) nitidus Jekel, 1866 | | | Thorectes (Jekelius) punctatolineatus Francois, 1904 | | | Thorectes (Jekelius) sardous Erichson, 1847 | | | Thorectes (Jekelius) sericeus Jekel, 1866 | | | Thorectes (Silphotrupes) escorialensis Jekel, 1866 | | | Thorectes escorialensis opaculus Reitter, 1893 | | | Thorectes (Silphotrupes) geminatus Géné, 1839 | | | Thorectes (Silphotrupes) orocantabricus Verdú & Gal..., 2000 | | | Thorectes (Silphotrupes) punctatissimus Chevrolat, 1840 | | | Thorectes (Thorectes) armifrons Reitter, 1892 | | | Thorectes (Thorectes) asperifrons Fairmaire, 1866 | | | Thorectes (Thorectes) banghaasi Reitter, 1892 | | | Thorectes (Thorectes) baraudi Lopez-colon, 1981 | | | Thorectes (Thorectes) coiffaiti Baraud, 1969 | | | Thorectes (Thorectes) coloni Ruiz, 1998 | | | Thorectes (Thorectes) demoflysi Baraud, 1965 | | | Thorectes (Thorectes) distinctus Marseul, 1878 | | | Thorectes (Thorectes) ferreri Lopez-colon, 1983 | | | Thorectes (Thorectes) laevigatus (Fabricius, 1798) | ⇒ Thorectes intermedius Costa, 1839 | | Thorectes (Thorectes) latus Sturm, 1826 | | | Thorectes (Thorectes) lusitanicus Jekel, 1866 | | | Thorectes (Thorectes) martensi Krikken, 1981 | | | Thorectes (Thorectes) nepalensis Baraud, 1974 | | | Thorectes (Thorectes) puncticollis Lucas, 1845 | | | Thorectes (Thorectes) punctulatus Jekel, 1866 | | | Thorectes (Thorectes) reflexus Jekel, 1866 | | | Thorectes (Thorectes) rugatulus Jekel, 1866 | | | Thorectes (Thorectes) shankara Carpaneto & ..., 1999 | | | Thorectes (Thorectes) stellosus Krikken, 1981 | | | Thorectes stellosus annapurnicus Krikken, 1981 | | | Thorectes stellosus stellosus annapurnicus..., 1981 | | | Thorectes (Thorectes) trituberculatus Reitter, 1892 | | | Thorectes (Thorectes) valencianus Baraud, 1966 | | | Thorectes (Thorectes) variolipennis Marseul, 1876 | | | Thorectes (Zuninoeus) hoppei Sturm & Hage..., 1825 | | | Thorectes balearicus (Lopez Colon, 1984) | ⇒ Thorectes albarracinus Wagner, 1928 | | Thorectes catalonicus (Lopez Colon, 1990) | ⇒ Thorectes albarracinus Wagner, 1928 | | Thorectes chersinus (Delabie, 1954) | ⇒ Thorectes albarracinus Wagner, 1928 | | Trypocopris (Pseudotrypocopris) amedei Fairmaire, 1861 | | | Trypocopris (Trypocopris) alpinus Sturm & Hage..., 1825 | | | Trypocopris alpinus alpinus Sturm & Hage..., 1825 | | | Trypocopris alpinus balcanicola Miksic, 1954 | | | Trypocopris alpinus marianii Mariani, 1958 | | | Trypocopris alpinus opacus (Mariani, 1958) | ⇒ Trypocopris alpinus marianii Mariani, 1958 | | Trypocopris (Trypocopris) fulgidus Motschulsky, 1845 | | | Trypocopris (Trypocopris) pyrenaeus Charpentier, 1825 | | | Trypocopris pyrenaeus bartolii Dellacasa, 1974 | | | Trypocopris pyrenaeus coruscans Chevrolat, 1840 | | | Trypocopris pyrenaeus cyanicolor Capra, 1930 | | | Trypocopris pyrenaeus pyrenaeus Charpentier, 1825 | | | Trypocopris pyrenaeus splendens Heer, 1841 | | | Trypocopris (Trypocopris) vernalis (Linnaeus, 1758) | ⇒ Geotrupes vernalis Linnaeus, 1758 | | Trypocopris vernalis apenninicus Mariani, 1958 | | | Trypocopris vernalis caspius Motschulsky, 1845 | | | Trypocopris vernalis fausti Petrovitz, 1962 | | | Trypocopris vernalis fauveli (Bedel, 1911) | ⇒ Trypocopris vernalis obscurus Mulsant, 1842 | | Trypocopris vernalis insularis Schneider, 1878 | | | Trypocopris vernalis manifestus Reitter, 1893 | | | Trypocopris vernalis obscurus Mulsant, 1842 | | | Trypocopris vernalis rambouseki Tesar, 1935 | | | Trypocopris vernalis vernalis Linnaeus, 1758 | | | Trypocopris caspius Motschulsky, 1845 | | | Trypocopris fausti Reitter, 1890 | | | Trypocopris inermis Ménétriés, 1832 | | | Trypocopris zaitzevi Olsoufieff, 1918 | | | Trypocoris vernalis | | | Typhaeus (Chelotrupes) hiostius (Géné, 1836) | ⇒ Chelotrupes hiostius Géné, 1836 | | Typhaeus (Chelotrupes) momus (Olivier, 1789) | ⇒ Chelotrupes momus Olivier, 1789 | | Typhaeus (Typhaeus) fossor Waltl, 1838 | | | Typhaeus (Typhaeus) lateridens Guérin-Ménev..., 1838 | | | Typhaeus (Typhaeus) typhoeus Linnaeus, 1758 | Minotaur Beetle | | Typhaeus typhoeoides Fairmaire, 1852 | | | Zuninoeus hoppei (Sturm & Hage..., 1825) | ⇒ Thorectes hoppei Sturm & Hage..., 1825 |
Subnodes of Geotrupinae with occurrences in Continents
Links, sources and further information
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