Species list, Checklist of Supertribe Harpalitae for Kazakhstan (Turkestan)

Class	Insecta Linnaeus, 1758 Insects, True insects	510 3742 11511	Families Genera Species	Taxom
Subclass	Pterygota Lang, 1888 Winged insects	509 3741 11509	Families Genera Species	Taxom
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	473 3671 11418	Families Genera Species	Taxom
Superorder	Holometabola Heider, 1889 Holometabolous Insects	303 2772 9045	Families Genera Species	Taxom
Order	Coleoptera Linnaeus, 1758 Beetles, Coleopterans	73 1123 4192	Families Genera Species	Taxom
Suborder	Adephaga Schellenberg, 1806 Adephagans, Ground Beetles and Water Beetles	6 188 1296	Families Genera Species	Taxom
Superfamily	Caraboidea Latreille, 1802	2 170 1273	Families Genera Species	Taxom
Family	Carabidae Latreille, 1802 Carabid beetles, Carabids, Ground Beetles, Tiger Beetles	164 1263	Genera Species	Taxom
Subfamily	Harpalinae Bonelli, 1810	89 630	Genera Species	Taxom
Supertribe	Harpalitae Erwin, 1985	29 179	Genera Species	Taxom

Genera for Kazakhstan
29 Genera with 179 Species

Acinopus Dejean, 1829		4	Species	Taxom
Acupalpus Latreille, 1850		11	Species	Taxom
Amblystomus Erichson, 1931		3	Species	Taxom
Anisodactylus Dejean, 1796		4	Species	Taxom
Anthracus Motschulsky, 1857		2	Species	Taxom
Bleusea Bedel, 1898		1	Species	Taxom
Bradycellus Erichson, 1846		2	Species	Taxom
Carenochyrus Solsky, 1874		1	Species	Taxom
Cephalophonus Ganglbauer, 1854		1	Species	Taxom
Chilotomus Chaudoir, 1971		1	Species	Taxom
Daptus Fischer von ..., 1823		2	Species	Taxom
Diachromus Erichson, 1758		1	Species	Taxom
Dicheirotrichus Jacquelin du..., 1829		17	Species	Taxom
Ditomus Dana, 1993		2	Species	Taxom
Dixus Billberg, 1872		2	Species	Taxom
Eocarterus Stichel, 1893		2	Species	Taxom
Gynandromorphus Dejean, 1905		3	Species	Taxom
Harpalobrachys Tschitschérine, 1844		1	Species	Taxom
Harpalus Latreille, 1889		114	Species	Taxom
Idiomelas Tschitschérine, 1894		1	Species	Taxom
Liochirus Tschitschérine, 1874		1	Species	Taxom
Loxoncus Schmidt-Gobel, 1858		1	Species	Taxom
Machozetus Chaudoir, 1848		1	Species	Taxom
Microderes Faldermann, 1885		5	Species	Taxom
Neophygas Noonan, 1835		1	Species	Taxom
Ophonus Dejean, 1828		14	Species	Taxom
Parophonus Ganglbauer, 1829		5	Species	Taxom
Pseudoophonus Motschulsky, 1774		3	Species	Taxom
Stenolophus Dejean, 1829	Seedcorn Beetle	6	Species	Taxom

Species list for Kazakhstan
179 Species and 35 Subspecies

Acinopus (Acinopus) laevigatus Ménétriés, 1832
Acinopus laevigatus laevigatus Ménétriés..., 1832
Acinopus (Haplacinopus) striolatus Zoubkoff, 1833
Acinopus (Osimus) ammophilus Dejean, 1829
Acupalpus (Acupalpus) dubius Schilsky, 1888
Acupalpus (Acupalpus) elegans Dejean, 1829
Acupalpus (Acupalpus) exiguus Dejean, 1829
Acupalpus (Acupalpus) flavicollis Sturm, 1825
Acupalpus (Acupalpus) limbatus Gebler, 1833
Acupalpus (Acupalpus) luteatus (Duftschmid, 1812)	⇒ Acupalpus dubius Schilsky, 1888
Acupalpus (Acupalpus) maculatus Schaum, 1860
Acupalpus (Acupalpus) meridianus Linnaeus, 1761	Midday harp ground beetle
Acupalpus (Acupalpus) parvulus Sturm, 1825
Acupalpus (Ancylostria) interstitialis Reitter, 1884
Acupalpus (Tachistodes) flaviceps Motschulsky, 1850
Amblystomus metallescens Dejean, 1829
Amblystomus niger (Heer, 1841)	⇒ Amblystomus nigrinus Csiki, 1931
Amblystomus nigrinus Csiki, 1931
Anisodactylus (Anisodactylus) binotatus Fabricius, 1787	Two-marked harp ground beetle
Anisodactylus (Hexatrichus) poeciloides Stephens, 1828	Saltmarsh Short-spur
Anisodactylus poeciloides pseudoaeneus Dejean, 1829
Anisodactylus (Pseudanisodactylus) signatus Panzer, 1796
Anthracus consputus Duftschmid, 1812
Anthracus longicornis Schaum, 1857
Bleusea ammophila Tschitschérine, 1898
Bradycellus (Tachycellus) glabratus Reitter, 1894
Bradycellus (Triliarthrus) caucasicus Chaudoir, 1846
Carenochyrus titanus Solsky, 1874
Cephalophonus cephalotes Fairmaire & ..., 1854
Chilotomus violaceus Kryzhanovski..., 1971
Daptus pictus Fischer von ..., 1823
Daptus vittatus Fischer von ..., 1823
Diachromus germanus Linnaeus, 1758
Dicheirotrichus (Cardiostenus) cymindiformis Reitter, 1901
Dicheirotrichus (Cardiostenus) microderus Solsky, 1874
Dicheirotrichus (Oreoxenus) mannerheimii R.F.Sahlberg, 1844	Bullies
Dicheirotrichus mannerheimii ponojensis J.R. Sahlberg, 1875
Dicheirotrichus (Trichocellus) desertus Motschulsky, 1849
Dicheirotrichus (Trichocellus) discicollis Dejean, 1829
Dicheirotrichus (Trichocellus) discolor Faldermann, 1836
Dicheirotrichus discolor punctidorsis Reitter In T..., 1899
Dicheirotrichus (Trichocellus) externepunctatus Reitter In T..., 1899
Dicheirotrichus (Trichocellus) hauseri Reitter, 1894
Dicheirotrichus (Trichocellus) lacustris L.Redtenbacher, 1858
Dicheirotrichus (Trichocellus) obscuricollis Reitter In T..., 1899
Dicheirotrichus (Trichocellus) stenothorax Kabak & Kataev, 1993
Dicheirotrichus (Trichocellus) subangularis Kataev & Wrase, 2006
Dicheirotrichus (Trichocellus) tscheresovae Komarov, 1995
Dicheirotrichus (Trichocellus) tschitscherini Reitter In T..., 1899
Dicheirotrichus (Trichocellus) ustulatus Dejean, 1829
Ditomus calydonius P. Rossi, 1790
Ditomus calydonius oriens Dvorak, 1993
Dixus eremita Dejean, 1825
Dixus semicylindricus Piochard de ..., 1872
Eocarterus (Eocarterus) chodshenticus Ballion, 1871
Eocarterus (Eocarterus) semenowi Reitter, 1893
Gynandromorphus etruscus Quensel, 1806
Gynandromorphus etruscus etruscus Quensel, 1806
Gynandromorphus etruscus peyroni Carret, 1905
Harpalobrachys leiroides Motschulsky, 1844	Mauve goby
Harpalus (Acardystus) flavescens Piller &Mitt..., 1783
Harpalus (Acardystus) pygmaeus Dejean, 1829
Harpalus (Actephilus) albanicus Reitter, 1900
Harpalus (Actephilus) pumilus Sturm, 1818
Harpalus (Amblystus) latus Linnaeus, 1758
Harpalus (Amblystus) luteicornis Dufischmid, 1812
Harpalus (Amblystus) rubripes Dufischmid, 1812	Bareyed goby
Harpalus (Amblystus) solitaris Dejean, 1829	Checked goby
Harpalus (Artabas) dispar Dejean, 1829
Harpalus dispar splendens Gebler, 1829
Harpalus (Cephalophonus) cephalotes (Fairmaire & ..., 1854)	⇒ Harpalus capito A.Morawitz, 1862
Harpalus cephalotes cephalotes
Harpalus (Cryptophonus) tenebrosus (Dejean, 1829)	⇒
Harpalus (Euharpalops) fraternus LeConte, 1852
Harpalus (Harpalobius) autumnalis Duftschmid, 1812
Harpalus (Harpalobius) froelichii Sturm, 1818	Brush-thighed Seed-eater
Harpalus (Harpalobius) fuscipalpis Sturm, 1818
Harpalus (Harpalobius) hirtipes Panzer, 1796
Harpalus neglectus neglectus Audinet-Serv..., 1821
Harpalus (Harpalophonus) circumpunctatus Chaudoir, 1846
Harpalus (Harpalophonus) hospes Sturm, 1818
Harpalus hospes ciscaucasicus Lutshnik, 1921
Harpalus hospes hospes Sturm, 1818
Harpalus (Harpalophonus) stevenii Dejean, 1829
Harpalus (Harpalophonus) terrestris Motschulsky, 1844
Harpalus (Harpalus) affinis (Schrank, 1781)	⇒ Harpalus serripes ernsti Kataev, 1995
Harpalus (Harpalus) akinini Tschitschérine, 1895
Harpalus (Harpalus) alajensis Tschitschérine, 1898
Harpalus (Harpalus) alexeevi Kataev, 1990
Harpalus (Harpalus) brevicornis Germar, 1824
Harpalus (Harpalus) caeruleatus Bates, 1878
Harpalus (Harpalus) cisteloides Motschulsky, 1844
Harpalus cisteloides cisteloides Motschulsky, 1844
Harpalus (Harpalus) cyanopterus Tschitschérine, 1897
Harpalus (Harpalus) erosus Mannerheim, 1825
Harpalus (Harpalus) ferghanensis Tschitschérine, 1899
Harpalus (Harpalus) glasunovi Kataev, 1987
Harpalus glasunovi glasunovi Kataev, 1987
Harpalus (Harpalus) inexspectatus Kataev, 1989
Harpalus (Harpalus) kabakianus Kataev, 1988
Harpalus (Harpalus) kadyrbekovi Kataev, 1988
Harpalus (Harpalus) kiritshenkoi Kataev, 1990
Harpalus (Harpalus) kryzhanovskii Kataev, 1988
Harpalus (Harpalus) lumbaris Mannerheim, 1825
Harpalus (Harpalus) macronotus Tschitschérine, 1893
Harpalus (Harpalus) masoreoides Bates, 1878
Harpalus (Harpalus) medvedevi Kataev, 2006
Harpalus (Harpalus) oblitus (Dejean, 1829)	⇒ Harpalus fraternus LeConte, 1852
Harpalus oblitus oblitus Dejean, 1829
Harpalus (Harpalus) opacipennis Haldemann, 1843
Harpalus (Harpalus) pseudoserripes Reitter, 1900
Harpalus (Harpalus) pusillus Motschulsky, 1850
Harpalus (Harpalus) remboides Solsky, 1874
Harpalus (Harpalus) saxicola Dejean, 1829
Harpalus (Harpalus) subcylindricus Dejean, 1829
Harpalus (Harpalus) tianschanicus Semenov, 1889
Harpalus (Harpalus) uniformis Motschulsky, 1844
Harpalus uniformis staudingeri Jedlicka, 1953
Harpalus uniformis uniformis Motschulsky, 1844
Harpalus (Harpalus) viridanus Motschulsky, 1844
Harpalus viridanus viridanus Motschulsky, 1844
Harpalus (Harpalus) vittatus Gebler, 1833	Frillfin goby
Harpalus vittatus vittatus Gebler, 1833
Harpalus (Harpalus) xanthopus Gemminger & ..., 1868
Harpalus xanthopus winkleri Schauberger, 1923
Harpalus xanthopus xanthopus Gemminger & ..., 1868
Harpalus (Harpalus) zabroides Dejean, 1829
Harpalus (Harpalus) zhdankoi Kataev, 1990
Harpalus (Lasioharpalus) dimidiatus P. Rossi, 1790
Harpalus (Lasioharpalus) distinguendus Duftschmid, 1812
Harpalus distinguendus distinguendus Duftschmid, 1812
Harpalus (Lasioharpalus) smaragdinus Duftschmid, 1812
Harpalus (Opadius) laevipes Zetterstedt, 1828
Harpalus (Opadius) nigritarsis C.R. Sahlberg, 1827	Highspine goby
Harpalus (Pheuginus) anxius Duftschmid, 1812
Harpalus (Pheuginus) politus Dejean, 1829
Harpalus politus politus Dejean, 1829
Harpalus (Pheuginus) serripes Quensel, 1806
Harpalus serripes ernsti Kataev, 1995
Harpalus serripes serripes Quensel, 1806
Harpalus (Pheuginus) servus Duftschmid, 1812
Harpalus (Pheuginus) tardus Panzer, 1796
Harpalus (Plectralidus) amplicollis Ménétriés, 1848
Harpalus (Plectralidus) angulatus Putzeys, 1878
Harpalus angulatus scytha Tschitschér..., 1899
Harpalus (Plectralidus) brevis Motschiulsky, 1844
Harpalus (Plectralidus) calathoides Motschulsky, 1844
Harpalus (Plectralidus) compressus Motschulsky, 1844
Harpalus (Plectralidus) cyclogonus Chaudoir, 1844
Harpalus cyclogonus cyclogonus Chaudoir, 1844
Harpalus (Plectralidus) foveiger Tschitschérine, 1895
Harpalus (Plectralidus) fuscicornis Ménétriés, 1832
Harpalus (Plectralidus) kirgisicus Motschulsky, 1844
Harpalus (Plectralidus) lutshniki Schauberger, 1932
Harpalus (Plectralidus) mitridati Pliginskiy, 1915
Harpalus (Plectralidus) nigritarsis C.R. Sahlberg, 1827
Harpalus (Plectralidus) oodioides Dejean, 1829
Harpalus (Plectralidus) optabilis Dejean, 1829
Harpalus (Plectralidus) pulvinatus Ménétriés, 1848
Harpalus pulvinatus lubricus Reitter, 1900
Harpalus pulvinatus pulvinatus Ménétriés..., 1848
Harpalus (Plectralidus) rufiscapus Gebler, 1833
Harpalus (Plectralidus) salinus Dejean, 1829
Harpalus salinus salinus Dejean, 1829
Harpalus (Plectralidus) tarsalis Mannenheim, 1825
Harpalus (Plectralidus) tichonis Jakobson, 1907
Harpalus (Plectralidus) torridoides Reitter, 1900
Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)	⇒ Pseudoophonus calceatus Duftschmid, 1812
Harpalus (Pseudoophonus) capito A.Morawitz, 1862
Harpalus (Pseudoophonus) griseus (Panzer, 1796)	⇒ Pseudoophonus griseus Panzer, 1797
Harpalus (Pseudoophonus) rufipes (DeGeer, 1774)	⇒
Harpalus (Semiophonus) signaticornis Duftschmid, 1812
Harpalus alaskensis Lindroth, 1968
Harpalus tjanschanicus Semenov, 1889
Idiomelas (Idiomelas) nigripes Reitter, 1894
Liochirus cycloderus Solsky, 1874
Loxoncus (Loxoncus) procerus Schaum, 1858
Machozetus lehmanni Ménétriés, 1848
Microderes (Microderes) brachypus Steven, 1809
Microderes (Microderes) undulatus Gebler, 1841
Microderes (Neopangus) diversopunctatus Solsky, 1874
Microderes diversopunctatus heterostictus Tschitschérine, 1898
Microderes (Neopangus) namanganensis Heyden, 1885
Neophygas microcephalus Faldermann, 1835
Ophonus (Hesperophonus) azureus Fabricius, 1775
Ophonus (Hesperophonus) chlorizans Solsky, 1874
Ophonus (Hesperophonus) convexicollis Ménétriés, 1832
Ophonus (Hesperophonus) minimus Motschulsky, 1845
Ophonus (Metophonus) cordatus Duftschmid, 1812
Ophonus (Metophonus) hystrix Reitter, 1894
Ophonus hystrix dissors Tschitschérine, 1895
Ophonus hystrix kamenskii Kataev & Kabak, 2001
Ophonus (Metophonus) laticollis (Mannerheim, 1825)	⇒ Ophonus puncticollis Paykull, 1798
Ophonus lystrix kamenskii Kataev & Kabak, 2001
Ophonus (Metophonus) puncticollis Paykull, 1798
Ophonus (Metophonus) rufibarbis Fabricius, 1792
Ophonus (Metophonus) transversus Motschulsky, 1844
Ophonus (Ophonus) stictus Stephens, 1828	Oolite Downy-back
Parophonus (Ophonomimus) hirsutulus Dejean, 1829
Parophonus (Ophonomimus) interstitialis Reitter, 1889
Parophonus (Parophonus) laeviceps Ménétriés, 1832
Parophonus (Parophonus) mendax P.Rossi, 1790
Parophonus (Parophonus) planicollis (Dejean, 1829)	⇒ Parophonus laeviceps Ménétriés, 1832
Pseudoophonus (Platus) calceatus Duftschmid, 1812
Pseudoophonus (Pseudoophonus) griseus Panzer, 1797
Pseudoophonus (Pseudoophonus) rufipes De Geer, 1774
Stenolophus (Egadroma) marginatus Dejean, 1829
Stenolophus (Stenolophus) abdominalis Géné, 1836
Stenolophus abdominalis persicus Mannerheim, 1844
Stenolophus (Stenolophus) discophorus Fischer von ..., 1823
Stenolophus (Stenolophus) mixtus Herbst, 1784
Stenolophus (Stenolophus) proximus Dejean, 1829

Subnodes of Harpalitae with occurrences in Kazakhstan

Tribe

Harpalini
Bonelli, 1810

29
179

Genera
Species

Taxom