Harpalus (Latreille 1802) ⇒ Taxonomy and Distribution of Genus

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Arachnids

INSECTOIDGERENGDARK LIGHT

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Taxonomy of Genus Harpalus Latreille, 1802

Class	Insecta Linnaeus, 1758 Insects, True insects	4179 190570 2444470	Families Genera Species
Subclass	Pterygota Lang, 1888 Winged insects	3816 189502 2439455	Families Genera Species
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	3399 186457 2415487	Families Genera Species
Superorder	Holometabola Heider, 1889 Holometabolous Insects	1891 142849 2048615	Families Genera Species
Order	Coleoptera Linnaeus, 1758 Beetles, Coleopterans	592 62779 813516	Families Genera Species
Suborder	Adephaga Schellenberg, 1806 Adephagans, Ground Beetles and Water Beetles	23 6166 123695	Families Genera Species
Superfamily	Caraboidea Latreille, 1802	8 5520 109721	Families Genera Species
Family	Carabidae Latreille, 1802 Carabid beetles, Carabids, Ground Beetles, Tiger Beetles	5446 109537	Genera Species
Subfamily	Harpalinae Bonelli, 1810	1693 52011	Genera Species
Supertribe	Harpalitae Erwin, 1985	217 7410	Genera Species
Tribe	Harpalini Bonelli, 1810	217 7410	Genera Species
Subtribe	Harpalina Bonelli, 1810	90 4097	Genera Species
Genus	Harpalus Latreille, 1802	1753	Species

Subnodes of Harpalus Latreille, 1802

Subgenus	Harpalus (Acardystus) Reitter, 1908	4	Species
Subgenus	Harpalus (Actephilus) Stephens, 1839 ⇒ Harpalus	3	Species
Subgenus	Harpalus (Amblystus) Motschulsky, 1864 ⇒ Harpalus (Harpalus)	6	Species
Subgenus	Harpalus (Artabas) Gozis, 1882	8	Species
Subgenus	Harpalus (Baeticoharpalus) ⇒ Harpalus	1	Species
Subgenus	Harpalus (Brachypangus) Tschitschérine, 1898 ⇒ Harpalus	1	Species
Subgenus	Harpalus (Cephalomorphus) Tschitschérine, 1897	0	Species
Subgenus	Harpalus (Cephalophonus) Ganglbauer, 1892	3	Species
Subgenus	Harpalus (Cephalotypsis) Tschitschérine, 1901 ⇒ Harpalus	1	Species
Subgenus	Harpalus (Cryptophonus) Brandmayr & Zetto Brandmayr, 1981	19	Species
Subgenus	Harpalus (Euharpalops) Casey, 1924	11	Species
Subgenus	Harpalus (Glanodes) Casey, 1914	7	Species
Subgenus	Harpalus (Harpalobius) Reitter, 1900 ⇒ Harpalus (Harpalus)	4	Species
Subgenus	Harpalus (Harpalobrachys) Tschitschérine, 1899	1	Species
Subgenus	Harpalus (Harpaloderus) Reitter, 1900 ⇒ Harpalus	17	Species
Subgenus	Harpalus (Harpalomerus) Casey, 1914 ⇒ Harpalus	5	Species
Subgenus	Harpalus (Harpalophonus) Ganglbauer, 1892	8	Species
Subgenus	Harpalus (Harpalus) Latreille, 1802 Peony	294	Species
Subgenus	Harpalus (Hypsinephus) H.W. Bates, 1878	0	Species
Subgenus	Harpalus (Lasioharpalus) Reitter, 1900 ⇒ Harpalus	11	Species
Subgenus	Harpalus (Licinoderus) St.-Claude Deville, 1905	2	Species
Subgenus	Harpalus (Loboharpalus) Schauberger, 1932	4	Species
Subgenus	Harpalus (Loxophonus) Reitter, 1894 ⇒ Harpalus	1	Species
Subgenus	Harpalus (Megapangus) Casey, 1914	2	Species
Subgenus	Harpalus (Microderes) Faldermann, 1835	0	Species
Subgenus	Harpalus (Microharpalus) Tschitschérine, 1901 ⇒ Harpalus	0	Species
Subgenus	Harpalus (Migadophonus) Tschitschérine, 1897 ⇒ Harpalus	0	Species
Subgenus	Harpalus (Neopangus) Tschitschérine, 1898 ⇒ Harpalus	0	Species
Subgenus	Harpalus (Neophygas) Noonan, 1976	0	Species
Subgenus	Harpalus (Opadius) Casey, 1914	20	Species
Subgenus	Harpalus (Ophonus) Dejean, 1821	1	Species
Subgenus	Harpalus (Pangus) Motschulsky, 1859	0	Species
Subgenus	Harpalus (Pharalus) Casey, 1914 ⇒ Harpalus	2	Species
Subgenus	Harpalus (Pheuginus) Motschulsky, 1844 ⇒ Harpalus	16	Species
Subgenus	Harpalus (Platus) Motschulsky, 1844	0	Species
Subgenus	Harpalus (Plectralidus) Casey, 1914	80	Species
Subgenus	Harpalus (Pseudoophonus) Motschulsky, 1844 Longjaw mudsuckers	53	Species
Subgenus	Harpalus (Semiophonus) Schauberger, 1933	1	Species
Subgenus	Harpalus (Typsiharpalus) Tschitschérine, 1901	0	Species
Subgenus	Harpalus (Zangoharpalus) Huang, 1998	7	Species
Species	Harpalus 4-punctatus Dejean, 1829
Species	Harpalus abasinus Rost, 1892
Species	Harpalus abdominalis Fabricius, 1787
Species	Harpalus abolitus ♰ C. von Heyden, 1866
Species	Harpalus abstrusus Casey, 1914
Species	Harpalus acolpomatus Antoine, 1941 ⇒ Harpalus (Harpalus) dissitus
Species	Harpalus acomanus Casey, 1914
Species	Harpalus acuminatus Motschulsky, 1844 ⇒ Harpalus (Amblystus) latus
Species	Harpalus adamsii Fischer von Waldheim, 1817
Species	Harpalus adanensis Schauberger, 1933
Species	Harpalus adelaidae Castelnau, 1867
Species	Harpalus adenticulatus Huang, 1992
Species	Harpalus admissus Casey, 1924
Species	Harpalus advena LeConte, 1852
Species	Harpalus adventicius Schauberger, 1934 ⇒ Harpalus (Harpalus) praecurrens
Species	Harpalus advolans Nietner, 1856
Species	Harpalus aemulus Wollaston, 1864
Species	Harpalus aeneolus Reitter, 1900
Species	Harpalus aeneonitens MacLeay, 1871
Species	Harpalus aeneopiceus Stephens, 1828
Species	Harpalus aenescens Casey, 1884
Species	Harpalus aeneuoides J. Jacquet, 1942 ⇒
Species	Harpalus aeneus Fabricius, 1775
Species	Harpalus aequabilis Casey, 1914
Species	Harpalus aequilatus Solier, 1849
Species	Harpalus aereus Dejean, 1829 ⇒ Hypharpax aereus
Species	Harpalus aesculoides Jedlicka, 1959
Species	Harpalus aesopus Casey, 1914
Species	Harpalus aestivus Duftschmid, 1812 ⇒
Species	Harpalus afghanus Jedlicka, 1967
Species	Harpalus agilis Péringuey, 1896
Species	Harpalus agitabilis Casey, 1924
Species	Harpalus agonoderus Putzeys, 1878
Species	Harpalus agonus Tschitschérine, 1894
Species	Harpalus agricola Say, 1823
Species	Harpalus aimaki Jedlička, 1968 ⇒ Harpalus (Harpalobius) froelichii
Species	Harpalus ainus Habu & Baba, 1963 ⇒ Harpalus (Amblystus) laevipes
Species	Harpalus akbensis Jedlička, 1958
Species	Harpalus akini Tschitschérine, 1895
Species	Harpalus alacris Chevrolat, 1861 ⇒
Species	Harpalus alajanicus Jedlička, 1957 ⇒ Harpalus (Harpalus) ingenuus ingenuus
Species	Harpalus alaskensis Lindroth, 1968
Species	Harpalus albanensis Schauberger, 1933
Species	Harpalus albarracinus H. Wagner, 1926 ⇒ Ophonus (Hesperophonus) azureus
Species	Harpalus albionicus Mannerheim, 1843 ⇒ Harpalus (Harpalus) cautus
Species	Harpalus albonicus Mannerheim, 1843
Species	Harpalus alluaudi Antoine, 1922 ⇒ Harpalus (Harpalus) madagascariensis
Species	Harpalus alpestris L. Redtenbacher, 1849 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus altaicus Jedlička, 1968 ⇒ Harpalus (Harpalus) uniformis uniformis
Species	Harpalus alternans Castelnau, 1867 ⇒ Gnathaphanus licinoides
Species	Harpalus amaroides Faldermann, 1836 ⇒
Species	Harpalus amasiensis Reitter, 1900 ⇒ Harpalus pulchrinulus
Species	Harpalus amator Reitter, 1900 ⇒
Species	Harpalus ambigenus Reiche, 1853 ⇒
Species	Harpalus amdoensis Semenov, 1889 ⇒ Harpalus (Harpalus) ganssuensis
Species	Harpalus amethystinus Dejean, 1829
Species	Harpalus amiculus Casey, 1924
Species	Harpalus amicus L. Dufour, 1843 ⇒
Species	Harpalus amoenus Heer, 1837 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus amputatoides MlynáÍ, 1979
Species	Harpalus anactus Giebel, 1856
Species	Harpalus anadoluensis Kataev, 1993
Species	Harpalus ancillaris Casey, 1924
Species	Harpalus anderschii Duftschmid, 1812
Species	Harpalus andrewesi Schauberger, 1932 ⇒ Harpalus (Harpalus) amarellus
Species	Harpalus andrewesianus Schauberger, 1932 ⇒ Harpalus (Pseudoophonus) indicus
Species	Harpalus angularis Rey, 1886 ⇒
Species	Harpalus angulatoides Puel, 1935 ⇒ Harpalus cilihumerus
Species	Harpalus angustatus MacLeay, 1871 ⇒ Gnathaphanus melbournensis
Species	Harpalus angusticollis G. Müller, 1921 ⇒ Ophonus (Metophonus) puncticollis
Species	Harpalus angustior J. R. Sahlberg, 1875 ⇒
Species	Harpalus anisodactyliformys Solsky, 1874
Species	Harpalus annulicornis Stephens, 1828 ⇒ Harpalus (Harpaloderus) rufipalpis rufipalpis
Species	Harpalus antarcticus Laporte, 1867
Species	Harpalus anthracinus Fairmaire & Laboulbène, 1854 ⇒
Species	Harpalus antiphon Casey, 1924 ⇒ Harpalus (Harpalus) obnixus
Species	Harpalus antiphonis Csiki, 1932
Species	Harpalus antoinei Schauberger, 1926
Species	Harpalus antoineianus Schauberger, 1929
Species	Harpalus antonrichteri Kirschenhofer, 1992 ⇒ Harpalus (Harpalus) kabakovi
Species	Harpalus apfelbecki Schauberger, 1926
Species	Harpalus apunctatus Puel, 1935 ⇒
Species	Harpalus arcticus Gyllenhal, 1810
Species	Harpalus ardosiacus Lutschnik, 1922 ⇒ Ophonus (Ophonus) ardosiacus
Species	Harpalus arenicola Wesmael, 1872 ⇒ Harpalus (Acardystus) attenuatus
Species	Harpalus argutoroides Bates, 1873
Species	Harpalus armeniacus Chaudoir, 1846 ⇒ Harpalus (Harpalus) subcylindricus
Species	Harpalus armenus K. Daniel, 1904
Species	Harpalus asaphus Antoine, 1941
Species	Harpalus aspromontis Hille, 1914
Species	Harpalus assensus Casey, 1924
Species	Harpalus assimilis Dejean, 1829
Species	Harpalus asturiae Schauberger, 1928 ⇒
Species	Harpalus asturiensis Jedlička, 1957 ⇒
Species	Harpalus ater C.R.Sahlberg, 1827
Species	Harpalus atricornis Stephens, 1835 ⇒ Anisodactylus (Anisodactylus) nemorivagus
Species	Harpalus atrocaeruleus Stephens, 1828 ⇒ Harpalus (Harpaloderus) rufipalpis rufipalpis
Species	Harpalus atroviridis MacLeay, 1871
Species	Harpalus aulicus Dejean, 1829
Species	Harpalus australis Dejean, 1829 ⇒ Hypharpax australis
Species	Harpalus avarus Schauberger, 1930 ⇒ Harpalus (Harpalus) subcylindricus
Species	Harpalus azrouanus Emden & Schauberger, 1932
Species	Harpalus azruensis Antoine, 1925
Species	Harpalus azurescens Gyllenhal, 1827 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus azureus Fabricius, 1775 ⇒ Harpalus (Harpaloderus) sulphuripes sulphuripes
Species	Harpalus babunensis MlynáÍ, 1979
Species	Harpalus bachmayeri MlynáÍ, 1979
Species	Harpalus badakshanus Jedlicka, 1956
Species	Harpalus badius Dejean, 1829
Species	Harpalus baergi Csiki, 1932 ⇒ Harpalus (Amblystus) laevipes
Species	Harpalus ballioni Heyden, 1882 ⇒ Harpalus (Pheuginus) serripes ernsti
Species	Harpalus baltimoriensis Say, 1823
Species	Harpalus barbarus Leder, 1872
Species	Harpalus barthei Antoine, 1920 ⇒ Harpalus (Actephilus) pumilus
Species	Harpalus bartoni Maran, 1935
Species	Harpalus basalis Motschulsky, 1844 ⇒ Harpalus (Amblystus) latus
Species	Harpalus basharicus Schauberger, 1933
Species	Harpalus basilaris Kirby, 1837 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus beatulus Casey, 1924
Species	Harpalus beffai Lutshnik, 1922
Species	Harpalus berckensis Antoine, 1920
Species	Harpalus beta Duftschmid, 1812
Species	Harpalus betschuanus Obst, 1908
Species	Harpalus bicolor Fischer von Waldheim, 1829
Species	Harpalus bifoveolatus Küster, 1846
Species	Harpalus bifrons Gyllenhal, 1810
Species	Harpalus birmanus Andrewes, 1947 ⇒ Harpalus (Zangoharpalus) praticola
Species	Harpalus biroi Jedlicka, 1952
Species	Harpalus bisbipunctatus Puel, 1935 ⇒
Species	Harpalus blanditus Casey, 1924
Species	Harpalus bogemannii Gyllenhal, 1813
Species	Harpalus boisduvalii Castelnau, 1867
Species	Harpalus boluensis Jedlička, 1962 ⇒ Harpalus (Harpalus) smyrnensis smyrnensis
Species	Harpalus bonvouloiri Vuillefroy, 1866
Species	Harpalus boranus G.Müller, 1939
Species	Harpalus borealis Zetterstedt, 1828
Species	Harpalus borysthenicus Krynicki, 1832 ⇒ Harpalus (Harpalomerus) amputatus obtusus
Species	Harpalus bosphoranus Reiche, 1862 ⇒ Harpalus cilihumerus
Species	Harpalus bostockii Castelnau, 1867
Species	Harpalus bouvieri Tschitscherine, 1897
Species	Harpalus bracatus Casey, 1924
Species	Harpalus brachypus Steven, 1809
Species	Harpalus bradycelloides Reitter, 1900 ⇒ Harpalus (Harpalus) pusillus
Species	Harpalus brevicollis Schauberger, 1926
Species	Harpalus breviformis Tschitscherine, 1898
Species	Harpalus bruckmanni ♰ O. Heer, 1862
Species	Harpalus brunnea Gyllenhal, 1810
Species	Harpalus brunneipennis Boheman, 1848
Species	Harpalus brunneus Castelnau, 1867 ⇒ Harpalus (Amblystus) marginellus
Species	Harpalus bucculentus Kraatz, 1874 ⇒ Harpalus (Pheuginus) servus
Species	Harpalus bulgharensis Jedlička, 1958 ⇒ Harpalus (Harpalus) smyrnensis smyrnensis
Species	Harpalus bulirschi Facchini, 2012
Species	Harpalus bungei Chaudoir, 1844 ⇒ Harpalus (Harpalus) bungii
Species	Harpalus burmeisteri Giebel, 1856
Species	Harpalus caenus Say, 1823
Species	Harpalus caffer Duftschmid, 1812 ⇒
Species	Harpalus calabrus Schauberger, 1926 ⇒ Ophonus (Ophonus) sabulicola
Species	Harpalus calciatii Della Beffa, 1931
Species	Harpalus calcitrapus Motschulsky, 1844
Species	Harpalus camarguensis Puel, 1937
Species	Harpalus canonicus Casey, 1884
Species	Harpalus captiosus Casey, 1924
Species	Harpalus carbonarius Dejean, 1829
Species	Harpalus carbonatus LeConte, 1860 ⇒ Harpalus (Harpalus) somnulentus
Species	Harpalus cardiaderus Putzeys, 1872 ⇒
Species	Harpalus cardioderus Putzeys, 1872
Species	Harpalus caricus Kataev & Wrase, 1997
Species	Harpalus carolinae Rozen, 1989 ⇒ Harpalus (Euharpalops) spadiceus
Species	Harpalus carteroides Zurcher, 1911
Species	Harpalus cascadiensis Hatch, 1949 ⇒ Harpalus (Amblystus) laevipes
Species	Harpalus castillianus Vuillefroy, 1866 ⇒
Species	Harpalus catharinae Schauberger, 1933 ⇒ Ophonus (Metophonus) subsinuatus
Species	Harpalus caudalis Casey, 1914
Species	Harpalus cayennensis Dejean, 1826
Species	Harpalus celax
Species	Harpalus celioides Ménétriés, 1848 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus celox Casey, 1914 ⇒ Harpalus (Harpalus) somnulentus
Species	Harpalus centralis Schauberger, 1929 ⇒ Cryptophonus tenebrosus
Species	Harpalus cervicis Motschulsky, 1844 ⇒ Harpalus (Harpalus) lumbaris
Species	Harpalus chalusi Jedlicka, 1968
Species	Harpalus changaicus Jedlička, 1968 ⇒
Species	Harpalus chaudoiri Motschulsky, 1850
Species	Harpalus chaudoirianus Lutshnik, 1922
Species	Harpalus chengjiangensis Huang, 1993
Species	Harpalus chevrierii Heer, 1837 ⇒ Harpalus (Amblystus) solitaris
Species	Harpalus chilensis Dejean, 1829
Species	Harpalus chiloschizontus Huang, 1995
Species	Harpalus chinadensis Jedlička, 1932 ⇒ Harpalus (Harpalus) tibeticus
Species	Harpalus chlorizans Bates, 1883 ⇒
Species	Harpalus chlorophanus Panzer, 1799
Species	Harpalus chloropterus Stephens, 1828 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus chloroticus Dejean, 1829
Species	Harpalus chobauti Reitter, 1900 ⇒
Species	Harpalus cilicicus Csiki, 1932
Species	Harpalus ciliciensis MlynáÍ, 1979 ⇒ Harpalus (Harpalus) reflexus anadoluensis
Species	Harpalus cilihumerus Huang; Hu & Sun, 1994
Species	Harpalus cimensis Cockerell, 1922
Species	Harpalus circumfusus Germar, 1823
Species	Harpalus circumpuctatus Chaudour, 1846
Species	Harpalus ciscaucasicus Lutshnik, 1921
Species	Harpalus clandestinus LeConte, 1878 ⇒
Species	Harpalus clarkei Kataev & J.Schmidt, 2020
Species	Harpalus clavipes Boheman, 1848 ⇒ Harpalus (Harpalus) natalensis
Species	Harpalus clientus Casey, 1914 ⇒
Species	Harpalus coeruleipennis Ménétriés, 1832
Species	Harpalus coerulescens Schilsky, 1888 ⇒
Species	Harpalus coeruleus Schilsky, 1888 ⇒
Species	Harpalus cognatus Gyllenhal, 1827
Species	Harpalus collucens Casey, 1914
Species	Harpalus colombensis Nietner, 1857
Species	Harpalus coloradensis Casey, 1914 ⇒ Harpalus (Harpalus) opacipennis
Species	Harpalus columbianus Casey, 1924
Species	Harpalus columbinus Germar, 1817
Species	Harpalus comis Haldeman, 1843
Species	Harpalus complanatus Sturm, 1818 ⇒ Harpalus (Pheuginus) servus
Species	Harpalus complicans Reitter, 1900 ⇒ Harpalus (Harpalus) cyanopterus
Species	Harpalus concinnus Stephens, 1828
Species	Harpalus conditus ♰ S. H. Scudder, 1900
Species	Harpalus confinalis Andrewes, 1932 ⇒ Harpalus (Harpalus) amarellus
Species	Harpalus confinis Stephens, 1828
Species	Harpalus confusus Dejean, 1829
Species	Harpalus congener Péringuey, 1896
Species	Harpalus conradti Schauberger, 1934 ⇒ Harpalus (Harpalus) arcuatus
Species	Harpalus consentaneus Dejean, 1829 ⇒ Harpalus (Acardystus) attenuatus
Species	Harpalus consimile Gyllenhal, 1810
Species	Harpalus consimilis Gyllenhal, 1810
Species	Harpalus consors Tschitscherine, 1897
Species	Harpalus conspectus Casey, 1924
Species	Harpalus constrictus ♰ O. Heer, 1862 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus contactus Casey, 1924
Species	Harpalus contemptoides Puel, 1935 ⇒
Species	Harpalus contemptulus Puel, 1935 ⇒
Species	Harpalus contristatus Duftschmid, 1812 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus contumax Lutshnik, 1933
Species	Harpalus convexicollis Ménétriés, 1832
Species	Harpalus convexiusculus MacLeay, 1871
Species	Harpalus convictor Casey, 1884
Species	Harpalus convivus LeConte, 1865
Species	Harpalus coracinus Sturm, 1818 ⇒
Species	Harpalus cordatoides Schauberger, 1926
Species	Harpalus cordicollis Dejean, 1829
Species	Harpalus correiroi Schatzmayr, 1943
Species	Harpalus corsicus Audinet-Serville, 1821
Species	Harpalus coruscus Tschitscherine, 1895
Species	Harpalus corvus Duftschmid, 1812 ⇒
Species	Harpalus couleensis Hatch, 1949
Species	Harpalus coxii Castelnau, 1867
Species	Harpalus crassipes Duftschmid, 1812 ⇒
Species	Harpalus crassiusculus Fairmaire & Laboulbène, 1854
Species	Harpalus crassus Tschitschérine, 1895 ⇒ Harpalus (Harpalus) major
Species	Harpalus cratognathoides Gerstaecker, 1867
Species	Harpalus crebrior Tschitscherine, 1902
Species	Harpalus crenatellus Casey, 1914
Species	Harpalus creticus MaÍan, 1934 ⇒ Harpalus (Harpaloderus) honestus
Species	Harpalus cribratus Peyron, 1858
Species	Harpalus cribricollis Dejean, 1829 ⇒ Ophonus (Hesperophonus) cribricollis
Species	Harpalus cribripennis Chaudoir, 1842
Species	Harpalus cristatus L.Dufour, 1820
Species	Harpalus csikii Jedlička, 1967 ⇒ Harpalus (Harpalus) viridanus viridanus
Species	Harpalus cuculus Casey, 1924
Species	Harpalus cuncticeps Ball, 1973 ⇒ Harpalus (Glanodes) puncticeps
Species	Harpalus cunctipeps Ball, 1973
Species	Harpalus cuniculinus Duftschmid, 1812 ⇒ Harpalus (Harpaloderus) honestus
Species	Harpalus cupiens Casey, 1924
Species	Harpalus cupreolatus Casey, 1924
Species	Harpalus cupreonitens Dejean, 1829
Species	Harpalus cupreu Dejean, 1829
Species	Harpalus curtatus Mannerheim, 1853 ⇒
Species	Harpalus curticornis Casey, 1914 ⇒
Species	Harpalus curtus Audinet-Serville, 1821
Species	Harpalus cyanellus Kraatz, 1874 ⇒
Species	Harpalus cyanescens Motschulsky, 1844 ⇒ Harpalus (Harpalus) erosus
Species	Harpalus cyaneus Tschitschérine, 1893 ⇒ Harpalus (Harpalus) viridanus viridanus
Species	Harpalus cyclopius Reitter, 1900 ⇒ Harpalus tjanschanicus
Species	Harpalus cyrenaicus Koch, 1939
Species	Harpalus dagliari Jedlička, 1962 ⇒ Harpalus (Harpalus) smyrnensis smyrnensis
Species	Harpalus dampierii Castelnau, 1867
Species	Harpalus dashdorzhi Jedlicka, 1967
Species	Harpalus debdouensis Obenberger, 1918
Species	Harpalus debrunieri Briel, 1964 ⇒ Harpalus (Semiophonus) signaticornis
Species	Harpalus decolor Fairmaire & Laboulbène, 1854 ⇒ Harpalus (Harpaloderus) sulphuripes sulphuripes
Species	Harpalus decoloratus Reitter, 1900 ⇒ Harpalus (Harpalus) oblitus patruelis
Species	Harpalus defixus Walker, 1866
Species	Harpalus dejeani Perty, 1830
Species	Harpalus deletus ♰ E. Oustalet, 1874
Species	Harpalus deludens Casey, 1914
Species	Harpalus denisonensis Castelnau, 1867
Species	Harpalus dentatus Stephens, 1828
Species	Harpalus dentipes Wiedemann, 1823
Species	Harpalus depressicollis Bagnall, 1932 ⇒ Anisodactylus (Anisodactylus) californicus
Species	Harpalus depressus Duftschmid, 1812 ⇒
Species	Harpalus despectus C.R.Sahlberg, 1827
Species	Harpalus deutschii C.R.Sahlberg, 1827
Species	Harpalus deyrollei Castelnau, 1867
Species	Harpalus dichrous Dejean, 1829
Species	Harpalus dierli Jedlicka, 1966
Species	Harpalus difficilis Hope, 1845
Species	Harpalus diffinis Dejean, 1829
Species	Harpalus diffusus Klug, 1833
Species	Harpalus dilatatus ♰ N. Theobald, 1937 ⇒ Harpalus (Acardystus) atratus
Species	Harpalus dingo Castelnau, 1867
Species	Harpalus dinniki Lutshnik, 1933 ⇒ Harpalus (Harpalus) chrysopus chrysopus
Species	Harpalus disaogashimensis Huang In Huang & Zhang, 1995
Species	Harpalus discoideus Erichson, 1837 ⇒ Harpalus (Lasioharpalus) smaragdinus
Species	Harpalus discrepans A.Morawitz, 1862
Species	Harpalus discriminatus J. Jacquet, 1942 ⇒ Harpalus (Lasioharpalus) smaragdinus
Species	Harpalus dispellens Walker, 1859
Species	Harpalus distinctus Rambur, 1838
Species	Harpalus ditomoides Dejean, 1829
Species	Harpalus diversipennis Schatzmayr, 1943
Species	Harpalus diversopunctatus Solsky, 1874
Species	Harpalus diversus Dejean, 1829 ⇒ Harpalus (Harpalus) oblitus oblitus
Species	Harpalus dlabolai Jedlicka, 1968
Species	Harpalus dolens C.R.Sahlberg, 1827
Species	Harpalus dolosus Casey, 1914
Species	Harpalus dorsalis Dejean, 1829
Species	Harpalus dorsiger Klug, 1853
Species	Harpalus drescheri Fischer von Waldheim, 1817
Species	Harpalus duftschmidii Sturm, 1818 ⇒ Harpalus (Lasioharpalus) smaragdinus
Species	Harpalus dulcicollis LaFerté-Sénectère, 1841
Species	Harpalus dulciculus Casey, 1924
Species	Harpalus duponti Coquerel, 1866
Species	Harpalus durescans Casey, 1924 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus dusaultii L.Dufour, 1821
Species	Harpalus dux Casey, 1924
Species	Harpalus dzambuli Jedlička, 1966 ⇒ Harpalus (Harpalus) oblitus oblitus
Species	Harpalus eberti Jedlička, 1966 ⇒ Harpalus (Pseudoophonus) indicus
Species	Harpalus edwardsii Castelnau, 1867
Species	Harpalus effetus Casey, 1914
Species	Harpalus egregius Casey, 1914 ⇒ Harpalus (Amblystus) laevipes
Species	Harpalus eichingeri Jedlička, 1965 ⇒ Harpalus (Harpalus) ganssuensis
Species	Harpalus electus Casey, 1924
Species	Harpalus elegans Preller, 1862
Species	Harpalus elegantulus Ménétriés, 1832
Species	Harpalus ellipticus Ballion, 1878 ⇒
Species	Harpalus emarginatum Gyllenhal, 1827
Species	Harpalus emarginatus Gyllenhal, 1827
Species	Harpalus empiricus Wollaston, 1865
Species	Harpalus enci Jedlička, 1969
Species	Harpalus ephippium Boheman, 1860
Species	Harpalus episcopalis Reiche, 1863
Species	Harpalus episcopus Drapiez, 1819
Species	Harpalus erichsoni Castelnau, 1867
Species	Harpalus erivanus Reitter, 1900 ⇒ Harpalus (Harpalus) smyrnensis raddei
Species	Harpalus ernsti Kataev, 1995
Species	Harpalus erosoides Jedlička, 1953 ⇒ Harpalus (Harpalus) anisodactyliformis
Species	Harpalus errans Tschitschérine, 1897 ⇒ Harpalus (Harpalus) rufomarginatus
Species	Harpalus erratus C.R.Sahlberg, 1827
Species	Harpalus ervini Antoine, 1941
Species	Harpalus erythrocephalus Fabricius, 1787
Species	Harpalus erythrocerus Fischer von Waldheim, 1829
Species	Harpalus escheri Gemminger & Harold, 1868
Species	Harpalus eupatoriae Schauberger, 1932 ⇒ Harpalus (Pheuginus) servus
Species	Harpalus ewaldi Giebel, 1856
Species	Harpalus excavatus ♰ B. Förster, 1891
Species	Harpalus excubans Casey, 1924
Species	Harpalus extensus Walker, 1866 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus externepunctatus Solsky, 1874
Species	Harpalus faber Ménétriés, 1832 ⇒
Species	Harpalus fabichi Maran, 1935 ⇒ Harpalus (Harpaloderus) honestus
Species	Harpalus fabricii Crotch, 1871 ⇒
Species	Harpalus fallax Le Conte, 1859 ⇒ Harpalus (Harpalus) somnulentus
Species	Harpalus fanaticus Casey, 1924 ⇒
Species	Harpalus fanensis Kataev & Wrase, 1993
Species	Harpalus fastiditus Dejean, 1829 ⇒ Harpalus (Harpalus) oblitus patruelis
Species	Harpalus fastuosus Faldermann, 1836
Species	Harpalus fatuus LeConte, 1863
Species	Harpalus femoralis Chaudoir, 1846 ⇒
Species	Harpalus femoratus Dejean, 1829
Species	Harpalus fenisex Casey, 1914
Species	Harpalus feroculus Casey, 1924
Species	Harpalus ferrugatus Reitter, 1902
Species	Harpalus ferrugineus Fabricius, 1775
Species	Harpalus festivus K. Daniel, 1904 ⇒ Harpalus (Harpalophonus) stevenii
Species	Harpalus finitimus Schauberger, 1934 ⇒ Harpalus (Zangoharpalus) praticola
Species	Harpalus flavipalpis MacLeay, 1864
Species	Harpalus flavipennis Reitter, 1900
Species	Harpalus flavitarsis Dejean, 1829 ⇒ Harpalus (Pheuginus) modestus
Species	Harpalus flaviventris Sturm, 1818 ⇒
Species	Harpalus flavolimbatus Heer, 1837
Species	Harpalus fleischeri Eichler, 1924 ⇒ Harpalus (Actephilus) pumilus
Species	Harpalus flindersii Castelnau, 1867
Species	Harpalus foliarius Gistel, 1857
Species	Harpalus forsstroemi Gyllenhal, 1810
Species	Harpalus forsstromii Gyllenhal, 1810
Species	Harpalus forstroemii Gyllenhal, 1810
Species	Harpalus fortnumi Castelnau, 1867
Species	Harpalus foveicollis Motschulsky, 1844 ⇒
Species	Harpalus fractus Casey, 1924 ⇒ Harpalus (Harpalus) ellipsis
Species	Harpalus freudei Jedlicka, 1965
Species	Harpalus friulanus J. Müller, 1926 ⇒ Harpalus (Pheuginus) anxius
Species	Harpalus froelichi Sturm, 1818
Species	Harpalus frolichii Sturm, 1818
Species	Harpalus frontalis Fischer von Waldheim, 1829
Species	Harpalus fugax Faldermann, 1836 ⇒ Harpalus cilihumerus
Species	Harpalus fugitans Casey, 1914
Species	Harpalus fulgidicollis L.Dufour, 1820
Species	Harpalus fuliginosus Duftschmid, 1812 ⇒
Species	Harpalus fulvipes Della Beffa, 1909 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus funerarius Rightmyer, 2004 ⇒ Harpalus (Euharpalops) reversus
Species	Harpalus funestus Audinet‑Serville, 1821 ⇒ Harpalus (Actephilus) pumilus
Species	Harpalus furtivus LeConte, 1865 ⇒
Species	Harpalus furviculus Casey, 1924 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus furvus C.R.Sahlberg, 1827
Species	Harpalus fuscipalpoides Schauberger, 1926 ⇒ Harpalus (Harpalus) viridanus viridanus
Species	Harpalus fuscus Gmelin, 1790
Species	Harpalus futilis Casey, 1914 ⇒ Harpalus (Harpalus) somnulentus
Species	Harpalus galiberti Puel, 1925
Species	Harpalus gammeli Schauberger, 1933
Species	Harpalus gaudionis Reiche, 1862 ⇒ Harpalus (Harpalophonus) hospes hospes
Species	Harpalus gayndahensis MacLeay, 1871
Species	Harpalus gebieni Schauberger, 1930 ⇒ Harpalus (Pseudoophonus) pastor niigatanus
Species	Harpalus gemmeus Casey, 1914
Species	Harpalus geniculatus
Species	Harpalus germanicus Reitter, 1900 ⇒ Harpalus (Amblystus) solitaris
Species	Harpalus germanus Chaudoir, 1876
Species	Harpalus germari Castelnau, 1867
Species	Harpalus ginfushanus Jedlička, 1958 ⇒ Harpalus (Pseudoophonus) aenigma
Species	Harpalus glaberellus Sturm, 1818 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus glaberrimus Motschulsky, 1848 ⇒ Harpalus (Harpalomerus) amputatus obtusus
Species	Harpalus glamorgani Lequillon, 1841
Species	Harpalus glebalis Coquerel, 1859
Species	Harpalus gondocoroensis
Species	Harpalus gory Gory, 1833
Species	Harpalus goryi Boisduval, 1835
Species	Harpalus goudotii Dejean, 1829
Species	Harpalus grancanariensis Emden, 1928 ⇒ Harpalus (Cryptophonus) schaumii
Species	Harpalus grandiceps Boheman, 1860
Species	Harpalus grandicollis Reiche, 1862 ⇒
Species	Harpalus gregarius Fauvel, 1897
Species	Harpalus grimmi Sturm, 1826
Species	Harpalus griseoides Reitter, 1900
Species	Harpalus grottagliensis Schauberger, 1927 ⇒ Ophonus (Hesperophonus) azureus
Species	Harpalus habroterus Antoine, 1941 ⇒ Harpalus (Acardystus) attenuatus
Species	Harpalus haldemani Casey, 1914
Species	Harpalus hamasiensis J. Müller, 1947 ⇒ Harpalus (Harpalus) impressus
Species	Harpalus hassani Antoine, 1941
Species	Harpalus hauseri Schauberger, 1928 ⇒ Harpalus (Harpalomerus) amputatus obtusus
Species	Harpalus hebescens Bates, 1882
Species	Harpalus heeri Giebel, 1856
Species	Harpalus helopioides Faldermann, 1836 ⇒
Species	Harpalus hespericus Rosenhauer, 1856 ⇒ Harpalus (Acardystus) attenuatus
Species	Harpalus heterostictus Tschitscherine, 1898
Species	Harpalus himalayicus Jedlička, 1966 ⇒ Harpalus (Harpalus) indicola indicola
Species	Harpalus hirsutulus Dejean, 1829
Species	Harpalus hirsutus Ménétriés, 1843
Species	Harpalus hispanicus Schauberger, 1926 ⇒ Ophonus (Ophonus) sabulicola
Species	Harpalus hispanus Rambur, 1838
Species	Harpalus hissariensis Lutshnik, 1922
Species	Harpalus hoffmanni Schauberger, 1928
Species	Harpalus hokkaidensis Habu & Baba, 1963 ⇒
Species	Harpalus honestoides Reitter, 1900 ⇒ Harpalus (Harpaloderus) honestus
Species	Harpalus hoppingi Stevens, 1919 ⇒ Harpalus (Euharpalops) fraternus
Species	Harpalus horni Jedlička, 1928
Species	Harpalus hottentota Duftschmid, 1812 ⇒ Harpalus (Acardystus) atratus
Species	Harpalus hottentotta Duftschmid, 1812 ⇒ Harpalus (Acardystus) atratus
Species	Harpalus hsifanicus Schauberger, 1932
Species	Harpalus huachua Watson, 1927 ⇒ Harpalus (Glanodes) huachuca
Species	Harpalus hurkai Divoký, Pulpán & Rébl, 1990
Species	Harpalus hweisinensis Schauberger, 1929 ⇒ Harpalus (Pseudoophonus) pastor pastor
Species	Harpalus hylacis Say, 1823
Species	Harpalus hyperboreus Motschulsky, 1844 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus hypocrita Dejean, 1829 ⇒
Species	Harpalus hypogeomysis Huang, 1993
Species	Harpalus iberiae Schauberger, 1929 ⇒ Ophonus (Metophonus) cordatus
Species	Harpalus ibericus Pantel, 1888 ⇒ Harpalus (Artabas) dispar dispar
Species	Harpalus ignavus Duftschmid, 1812 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus ignobilis Boheman, 1858
Species	Harpalus ihai Habu, 1968 ⇒ Harpalus (Zangoharpalus) tinctulus luteicornoides
Species	Harpalus ilgasdensis Jedlička, 1962 ⇒ Harpalus (Harpalus) smyrnensis smyrnensis
Species	Harpalus illawarrensis Castelnau, 1867
Species	Harpalus illectus Casey, 1914
Species	Harpalus illeticus D. L. Crawford, 1909 ⇒ Harpalus (Opadius) desertus
Species	Harpalus imitans Schauberger, 1926
Species	Harpalus immaturus Motschulsky, 1860
Species	Harpalus immixtus
Species	Harpalus impiger Duftschmid, 1812 ⇒ Harpalus (Harpalobius) autumnalis
Species	Harpalus impotens LeConte, 1858
Species	Harpalus impressipennis Dejean, 1829 ⇒ Harpalus (Plectralidus) decipiens decipiens
Species	Harpalus impuncticeps Reitter, 1900 ⇒ Harpalus (Semiophonus) signaticornis
Species	Harpalus impunctus Wiedemann, 1825 ⇒
Species	Harpalus inaequalipennis Castelnau, 1867
Species	Harpalus inaequalis Reitter, 1900
Species	Harpalus incertus Dejean, 1829 ⇒ Harpalus (Harpalus) oblitus oblitus
Species	Harpalus incisus Dejean, 1829
Species	Harpalus incrassatus Boheman, 1848
Species	Harpalus inderiensis Motschulsky, 1848 ⇒ Harpalus (Harpalophonus) terrestris
Species	Harpalus ineditus Dejean, 1829
Species	Harpalus infelix Castelnau, 1867
Species	Harpalus infrequens Lutshnik, 1921
Species	Harpalus inmixtus Casey, 1924
Species	Harpalus inornatus Germar, 1848
Species	Harpalus inschanicus Breit, 1914
Species	Harpalus insignis Gautier Des Cottes, 1872 ⇒ Harpalus (Harpaloderus) honestus
Species	Harpalus instructus Casey, 1924 ⇒ Harpalus (Amblystus) laevipes
Species	Harpalus insularis Ragusa, 1886 ⇒ Harpalophonus italus
Species	Harpalus intactus Casey, 1924 ⇒ Harpalus (Harpalus) somnulentus
Species	Harpalus integer Fabricius, 1798
Species	Harpalus interioris Castelnau, 1867
Species	Harpalus intermedius Desbrochers des Loges, 1865 ⇒ Harpalus (Acardystus) attenuatus
Species	Harpalus intermittens Solsky, 1874
Species	Harpalus interpunctatus Kirby, 1837
Species	Harpalus interruptus Fischer von Waldheim, 1829
Species	Harpalus interstinctus Motschulsky, 1844 ⇒ Harpalus (Artabas) dispar splendens
Species	Harpalus interstitialis Gredler, 1863
Species	Harpalus inunctus Sturm, 1818 ⇒ Harpalus (Harpalobius) autumnalis
Species	Harpalus irinus Reiche, 1843
Species	Harpalus iripennis Schaeffer, 1910
Species	Harpalus iris Alluaud, 1918
Species	Harpalus irvinei Andrewes, 1930
Species	Harpalus iskerensis Jedlička, 1964 ⇒ Harpalus (Harpalus) melaneus melaneus
Species	Harpalus israelita Brullé, 1875
Species	Harpalus itoshimanus Habu, 1954
Species	Harpalus jacqueti Puel, 1936 ⇒
Species	Harpalus jailensis Schauberger, 1926
Species	Harpalus janetscheki Jedlička, 1970 ⇒ Harpalus (Pseudoophonus) indicus
Species	Harpalus janthinus Gautier Des Cottes, 1872 ⇒ Harpalus (Harpaloderus) honestus
Species	Harpalus janus Fairmaire, 1856 ⇒ Harpalus (Harpaloderus) honestus
Species	Harpalus japonicus A. Morawitz, 1862 ⇒ Harpalus (Pseudoophonus) sinicus
Species	Harpalus jeanelli
Species	Harpalus jedlickai Schauberger, 1929
Species	Harpalus jianyangensis Huang, Lei, Yan & Hu, 1996 ⇒ Harpalus (Harpalus) pallidipennis
Species	Harpalus judaeus Brullé, 1875
Species	Harpalus kabylianus Reiche, 1862
Species	Harpalus kailiensis Huang, 1992
Species	Harpalus karakorum Jedlička, 1958
Species	Harpalus karennius Bates, 1892
Species	Harpalus kashmirensis Bates, 1889
Species	Harpalus kaszabi Jedlicka, 1956
Species	Harpalus katavensis Jedlicka, 1957
Species	Harpalus kiautschouensis Schauberger, 1930 ⇒ Harpalus (Pseudoophonus) sinicus
Species	Harpalus kidanicus Kataev, 1989
Species	Harpalus kidanikus Kataev, 1989
Species	Harpalus kinfushanicus Schauberger, 1933 ⇒ Harpalus (Pseudoophonus) aenigma
Species	Harpalus kingii Castelnau, 1867 ⇒ Hypharpax kingii
Species	Harpalus kiselevi Kataev & Shilenkov, 1990
Species	Harpalus klementzae Kataev, 1984
Species	Harpalus knauthi Ganglbauer, 1901
Species	Harpalus knorri Giebel, 1856
Species	Harpalus kohistanicus Kirschenhofer, 1992 ⇒ Harpalus (Harpalus) melaneus melaneus
Species	Harpalus kolenatii Lutshnik, 1922 ⇒ Harpalus (Harpalus) breviusculus
Species	Harpalus koniensis Schauberger, 1927
Species	Harpalus korbi Schauberger, 1933
Species	Harpalus kreftii Castelnau, 1867
Species	Harpalus krueperi Apfelbeck, 1902
Species	Harpalus kuangcensis MlynáÍ, 1974 ⇒ Harpalus (Pseudoophonus) aenigma
Species	Harpalus kubanicus Jedlička, 1957
Species	Harpalus lacustris Casey, 1914
Species	Harpalus laesus LeConte, 1858
Species	Harpalus laeviceps Ménétriés, 1832
Species	Harpalus laevicollis Duftschmid, 1812
Species	Harpalus laevigatus L.Dufour, 1820
Species	Harpalus laevis Curtis, 1839
Species	Harpalus laevistriatus Sturm, 1818
Species	Harpalus laminatus Fairmaire, 1859
Species	Harpalus lamottei
Species	Harpalus langloisii Peyron, 1858
Species	Harpalus lapeyrousei Castelnau, 1867
Species	Harpalus lapponica C.R. Sahlberg, 1827
Species	Harpalus lapponicus Zetterstedt, 1828 ⇒ Harpalus (Amblystus) solitaris
Species	Harpalus lascivus Casey, 1914
Species	Harpalus latebricola Casey, 1914
Species	Harpalus latescans Casey, 1924
Species	Harpalus latianus Schauberger, 1923
Species	Harpalus laticollis Sturm, 1825 ⇒ Masoreus (Masoreus) wetterhallii wetterhallii
Species	Harpalus latitibia Rey, 1886 ⇒ Harpalus (Acardystus) attenuatus
Species	Harpalus latiusculus Kolbe, 1883
Species	Harpalus lecontei Casey, 1914 ⇒ Harpalus (Euharpalops) fraternus
Species	Harpalus leirodes Motschulsky, 1844
Species	Harpalus lentus Sturm, 1818 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus lepcha Andrewes, 1930 ⇒ Harpalus (Zangoharpalus) praticola
Species	Harpalus leptopus Bates, 1883
Species	Harpalus lerati Antoine, 1920 ⇒ Harpalus (Harpalus) subcylindricus
Species	Harpalus leucops Wiedemann, 1823
Species	Harpalus leviceps Casey, 1924
Species	Harpalus lewisi Linsley, 1945 ⇒ Harpalus (Opadius) lewisii
Species	Harpalus lheritieri ♰ L. Piton, 1940
Species	Harpalus liasinus Giebel, 1856
Species	Harpalus limbatus Duftschmid, 1812 ⇒ Harpalus (Amblystus) latus
Species	Harpalus limbopunctatus Fuss, 1858
Species	Harpalus liodes Bates, 1878 ⇒ Harpalus (Harpalus) brevicornis
Species	Harpalus liolus Bates, 1882
Species	Harpalus littoralis Rambur, 1838 ⇒ Harpalus (Cryptophonus) fulvus
Species	Harpalus livens Gyllenhal, 1810
Species	Harpalus lividulus Casey, 1914
Species	Harpalus lobipes Tschitschérine, 1898 ⇒ Harpalus (Loboharpalus) rubefactus rubefactus
Species	Harpalus loeffleri Jedlicka, 1966
Species	Harpalus lokayi Jedlička, 1929 ⇒
Species	Harpalus longihornus Lei & Huang, 1997
Species	Harpalus longior Kirby, 1837
Species	Harpalus loxophonoides Kataev & Wrase, 1993
Species	Harpalus lubricus Reitter, 1900
Species	Harpalus lucidicollis Dejean, 1829
Species	Harpalus lucidipennis Jedlička, 1958 ⇒ Harpalus (Harpalus) caiphus
Species	Harpalus lucidulus Boheman, 1848
Species	Harpalus lucidus LeConte, 1865
Species	Harpalus luridipennis Germar, 1822
Species	Harpalus lusitanicus Schatzmayr, 1943
Species	Harpalus lustralis Casey, 1884
Species	Harpalus lustrans Casey, 1884
Species	Harpalus lutarius Gistel, 1857
Species	Harpalus luteicornoides Breit, 1913
Species	Harpalus lutshnikianus Kataev, 1987 ⇒ Harpalus (Harpalus) kandaharensis
Species	Harpalus luxatus Audinet-Serville, 1821
Species	Harpalus lycaon Linder, 1860 ⇒ Acupalpus (Acupalpus) exiguus
Species	Harpalus lycius Kataev & Wrase, 1997
Species	Harpalus lydius Kataev & Wrase, 1997
Species	Harpalus lyratus Motschulsky, 1844 ⇒ Harpalus (Harpalus) vittatus vittatus
Species	Harpalus maceratus ♰ H. F. Wickham, 1911
Species	Harpalus machadoi Jeanne, 1971
Species	Harpalus macilentus Casey, 1914 ⇒ Harpalus (Harpalus) innocuus
Species	Harpalus maculicornis Stephens, 1835 ⇒ Harpalus (Harpaloderus) rufipalpis rufipalpis
Species	Harpalus maculifrons Ménétriés, 1848
Species	Harpalus maculiventris
Species	Harpalus magnodentatus Schauberger, 1929 ⇒ Harpalus (Pseudoophonus) tridens
Species	Harpalus malacus Casey, 1914 ⇒
Species	Harpalus mandibularis Castelnau, 1867
Species	Harpalus manhattanis Casey, 1884
Species	Harpalus mansuetus Casey, 1914 ⇒
Species	Harpalus marginatus Dalla Torre, 1877
Species	Harpalus marginepunctatus Dejean, 1829
Species	Harpalus marginicollis Castelnau, 1867 ⇒ Gnathaphanus melbournensis
Species	Harpalus maritimus O. Schneider, 1898 ⇒ Harpalus (Pheuginus) servus
Species	Harpalus massageta Silfverberg, 1977 ⇒
Species	Harpalus maurus Tschitschérine, 1897 ⇒ Harpalus (Harpalus) cisteloides cisteloides
Species	Harpalus maxillosus Dejean, 1829 ⇒ Harpalus (Acardystus) attenuatus
Species	Harpalus maxwellensis Csiki, 1932
Species	Harpalus maxwelli Casey, 1924 ⇒
Species	Harpalus mayeti Verdier, Quezel & Rioux, 1951
Species	Harpalus medicus Kataev, 1993
Species	Harpalus meirai Mateu, 1958
Species	Harpalus melaenus Lutshnik, 1922
Species	Harpalus melampus Duftschmid, 1812 ⇒
Species	Harpalus melanarius Dejean, 1829 ⇒ Notiobia (Anisotarsus) melanaria
Species	Harpalus melanopus Haldeman, 1843
Species	Harpalus melanotus Jacquet, 1942
Species	Harpalus melbournensis Castelnau, 1867
Species	Harpalus melleti Heer, 1837
Species	Harpalus melletii Heer, 1837
Species	Harpalus meridionalis Dejean, 1829
Species	Harpalus merula Germar, 1823
Species	Harpalus merus Tschitschérine, 1895 ⇒ Harpalus (Harpalus) akinini
Species	Harpalus metallescens Rye, 1874 ⇒ Harpalus (Amblystus) latus
Species	Harpalus meticulosus Coquerel, 1866
Species	Harpalus metuens Casey, 1924
Species	Harpalus mexicanus Dejean, 1829
Species	Harpalus micans Klug, 1833
Species	Harpalus microcephalus Faldermann, 1835
Species	Harpalus mihatschi Reitter, 1900 ⇒ Harpalus (Harpalus) reflexus reflexus
Species	Harpalus minor Reitter, 1884 ⇒
Species	Harpalus minorita Gistel, 1857
Species	Harpalus minutissimus Facchini, 2015
Species	Harpalus minutus Dejean, 1829 ⇒ Harpalus (Actephilus) pumilus
Species	Harpalus misellus Tschitschérine, 1897 ⇒ Harpalus (Harpalus) bungii
Species	Harpalus miser Tschitscherine, 1897
Species	Harpalus mithridatis Csiki, 1932
Species	Harpalus miwai Jedlicka, 1946
Species	Harpalus mobilis Casey, 1914 ⇒ Harpalus (Harpalus) opacipennis
Species	Harpalus modulatus Casey, 1924
Species	Harpalus moestus Dejean, 1829 ⇒ Hypharpax moestus
Species	Harpalus mongolicus Jedlicka, 1966
Species	Harpalus monilicornis Latreille, 1805
Species	Harpalus montanellus Mateu, 1953
Species	Harpalus montanus Sturm, 1818 ⇒ Trichotichnus (Trichotichnus) laevicollis
Species	Harpalus montezumae Csiki, 1932
Species	Harpalus monticola Dejean, 1829 ⇒
Species	Harpalus montivagus Reitter, 1900 ⇒ Harpalus (Amblystus) laevipes
Species	Harpalus montuosus Csiki, 1932
Species	Harpalus mormonicus Casey, 1914
Species	Harpalus motschoulskyanus Schauberger, 1928 ⇒ Harpalus (Amblystus) laevipes
Species	Harpalus motschulskyi Gemminger & Harold, 1868 ⇒ Harpalus (Harpalobius) hirtipes
Species	Harpalus muchei Jedlicka, 1961
Species	Harpalus muciulus Huang, 1992
Species	Harpalus muellerianus Schauberger, 1933
Species	Harpalus mulsanti Marseul, 1866 ⇒
Species	Harpalus multisetosus C. G. Thomson, 1884 ⇒ Harpalus (Actephilus) picipennis
Species	Harpalus munganasti Reitter, 1908 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus murzini Kataev, 2008
Species	Harpalus mutabilis Haldeman, 1843
Species	Harpalus nactus Casey, 1914
Species	Harpalus nama Obst, 1908
Species	Harpalus namanganensis Heyden, 1885
Species	Harpalus namanus
Species	Harpalus nanulus Tschitschérine, 1898
Species	Harpalus nanus Dejean, 1829 ⇒ Harpalus (Harpalus) fuscoaeneus
Species	Harpalus narynensis Jedlička, 1958 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus neglectoides Jedlička, 1960 ⇒ Harpalus (Harpaloderus) neglectus neglectus
Species	Harpalus nepalensis Jedlička, 1966 ⇒ Harpalus (Zangoharpalus) praticola
Species	Harpalus nero ♰ E. Oustalet, 1874
Species	Harpalus niger J.Frivaldszky, 1889
Species	Harpalus nigerrimus Dejean, 1831
Species	Harpalus nigricans Schilsky, 1888 ⇒
Species	Harpalus nigricornis Stephens, 1835
Species	Harpalus nigrinus Schilsky, 1888 ⇒ Harpalus (Harpalus) kabakianus
Species	Harpalus nigripalpis Stephens, 1835
Species	Harpalus nigripennis Motschulsky, 1844 ⇒ Harpalus (Actephilus) pumilus
Species	Harpalus nigripes J. R. Sahlberg, 1913 ⇒
Species	Harpalus nigritarsus C.R.Sahlberg, 1827
Species	Harpalus nigrocaeruleus Stephens, 1828
Species	Harpalus nigrocoeruleus Stephens, 1828
Species	Harpalus nigrosericeus Goeze, 1777
Species	Harpalus niigatanus Schauberger, 1929
Species	Harpalus nimius Casey, 1924
Species	Harpalus nindae
Species	Harpalus niponensis H. W. Bates, 1883
Species	Harpalus nitens ♰ O. Heer, 1870 ⇒ Trichotichnus (Trichotichnus) nitens
Species	Harpalus nitescans Casey, 1914
Species	Harpalus nitescens Stannard, 1958 ⇒ Harpalus (Opadius) desertus
Species	Harpalus nitidulus Motschulsky, 1844 ⇒
Species	Harpalus nitidus Sturm, 1818 ⇒ Harpalus (Harpaloderus) honestus
Species	Harpalus nivalis Casey, 1924
Species	Harpalus nobilitatus Faldermann, 1836 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus nonsignatus Krynicki, 1832
Species	Harpalus nossibianus Brancsik, 1893
Species	Harpalus notatus Stephens, 1828 ⇒ Harpalus (Harpaloderus) rufipalpis rufipalpis
Species	Harpalus novaehollandiae Castelnau, 1867
Species	Harpalus novaezelandiae Laporte, 1867
Species	Harpalus nugax Casey, 1914 ⇒
Species	Harpalus nuperus ♰ S. H. Scudder, 1900
Species	Harpalus nuristanus Jedlicka, 1956
Species	Harpalus obenbergeri Jedlička, 1928 ⇒ Harpalus (Harpalus) aequicollis
Species	Harpalus obesulus LeConte, 1852 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus obesus A. Morawitz, 1862 ⇒ Harpalus (Harpalus) major
Species	Harpalus oblongiusculus Castelnau, 1867
Species	Harpalus oblongus Casey, 1914 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus obscuricornis Motschulsky, 1844 ⇒ Harpalus (Harpaloderus) rufipalpis rufipalpis
Species	Harpalus obscuripennis Dejean, 1829
Species	Harpalus obscurus Sturm, 1818 ⇒ Harpalus (Harpaloderus) sulphuripes sulphuripes
Species	Harpalus obsoletus Dejean, 1829
Species	Harpalus obtusangulus Faldermann, 1835
Species	Harpalus obtusicollis Putzeys, 1878 ⇒
Species	Harpalus obtusus ♰ B. Förster, 1891
Species	Harpalus occidentalis Chaudoir, 1868
Species	Harpalus ochreatus Reiche & Saulcy, 1855 ⇒ Harpalus (Acardystus) pygmaeus
Species	Harpalus ocreatus Say, 1830
Species	Harpalus octopunctatus Dejean, 1829
Species	Harpalus offusus Förster, 1891
Species	Harpalus olenini Poppius, 1906
Species	Harpalus opacus Dejean, 1829
Species	Harpalus opalinus LeConte, 1863
Species	Harpalus ophonoides Jedlicka, 1958
Species	Harpalus opicus Casey, 1914 ⇒
Species	Harpalus oppositus Casey, 1914
Species	Harpalus oreas Bates, 1891
Species	Harpalus oregonensis Casey, 1914
Species	Harpalus orienticola Tschitscherine, 1902
Species	Harpalus orientis Schauberger, 1926 ⇒ Ophonus (Metophonus) puncticollis
Species	Harpalus otini Antoine, 1941 ⇒ Harpalus (Harpalus) angustitarsis
Species	Harpalus oustaleti ♰ F. Meunier, 1915
Species	Harpalus ovalis Motschulsky, 1844 ⇒ Cryptophonus melancholicus reichei
Species	Harpalus ovatus Chaudoir, 1844 ⇒
Species	Harpalus pacificus Fall, 1926
Species	Harpalus paganettii Flach, 1907
Species	Harpalus pakistanus Jedlička, 1964
Species	Harpalus palaestinus Jedlička, 1958 ⇒ Harpalus (Harpalus) caiphus
Species	Harpalus palangoides Reiche, 1868
Species	Harpalus pallidipes Reitter, 1918
Species	Harpalus pallidus Dejean, 1829
Species	Harpalus palumbinus Lutshnik, 1922 ⇒ Harpalus (Harpalus) caeruleatus
Species	Harpalus papagonalis Casey, 1914
Species	Harpalus paradoxus Haldeman, 1843
Species	Harpalus parallelus Dejean, 1829
Species	Harpalus paroensis Castelnau, 1867
Species	Harpalus parowanus Casey, 1924 ⇒
Species	Harpalus parumpunctatus W. Kolbe, 1924
Species	Harpalus pastus Ménétriés, 1848 ⇒
Species	Harpalus patrona Casey, 1914
Species	Harpalus patruelis Dejean, 1829
Species	Harpalus patrueloides Castelnau, 1867
Species	Harpalus paululus Casey, 1914 ⇒ Harpalus (Harpalus) innocuus
Species	Harpalus pauper Tschitscherine, 1897
Species	Harpalus pecirkai Jedlička, 1928 ⇒ Harpalus (Pseudoophonus) tridens
Species	Harpalus pelagicus Wollaston, 1865
Species	Harpalus pellax Casey, 1914 ⇒ Harpalus (Harpalus) somnulentus
Species	Harpalus peneaui Antoine, 1922
Species	Harpalus penglainus Huang; Hu & Sun, 1994
Species	Harpalus pennsylvanicus Say, 1823 ⇒ Harpalus (Pseudoophonus) pensylvanicus
Species	Harpalus perezi Vuillefroy, 1868 ⇒
Species	Harpalus periglabellus Huang, 1992
Species	Harpalus peringueyi Kuntzen, 1919
Species	Harpalus peritus Casey, 1914 ⇒ Harpalus (Harpalus) somnulentus
Species	Harpalus perlucidus Schauberger, 1929 ⇒ Harpalus (Harpalus) oblitus patruelis
Species	Harpalus peronii Castelnau, 1867
Species	Harpalus perplexus Gyllenhal, 1827 ⇒ Harpalus (Lasioharpalus) smaragdinus
Species	Harpalus persianus Jedlička, 1958 ⇒
Species	Harpalus persolus Casey, 1914
Species	Harpalus perspicuus Casey, 1914
Species	Harpalus peruvianus Dejean, 1829
Species	Harpalus perversus Roubal, 1917 ⇒ Harpalus (Amblystus) latus
Species	Harpalus petisii Duftschmid, 1812 ⇒
Species	Harpalus petraeus Motschulsky, 1844
Species	Harpalus petreus Motschulsky, 1844
Species	Harpalus pewtzowi Tschitschérine, 1897 ⇒ Harpalus (Harpalus) caeruleatus
Species	Harpalus pexus Ménétriés, 1848 ⇒ Harpalus (Harpalophonus) circumpunctatus
Species	Harpalus pharisaeus Reiche & Saulcy, 1855
Species	Harpalus phoenix Tschitscherine, 1902
Species	Harpalus phoeopterus
Species	Harpalus piceus Ménétriés, 1843
Species	Harpalus picicornis Zetterstedt, 1828
Species	Harpalus picilabris Stephens, 1835 ⇒ Harpalus (Acardystus) attenuatus
Species	Harpalus picipes MacLeay, 1864 ⇒
Species	Harpalus piger Duftschmid, 1812 ⇒
Species	Harpalus pilosus Huang, 1995
Species	Harpalus pimalicus Casey, 1914 ⇒ Harpalus (Amblystus) laevipes
Species	Harpalus piperi Casey, 1924
Species	Harpalus placidus Casey, 1884
Species	Harpalus planatus Motschulsky, 1844 ⇒ Harpalus (Harpalus) viridanus viridanus
Species	Harpalus planicollis Dejean, 1829
Species	Harpalus planipennis MacLeay, 1871
Species	Harpalus planiusculus Kraatz, 1873
Species	Harpalus planoimpressus Castelnau, 1867
Species	Harpalus platyderus H. W. Bates, 1881
Species	Harpalus platypterus Fischer von Waldheim, 1829
Species	Harpalus plebejus Tschitschérine, 1897 ⇒
Species	Harpalus pleistocenicus ♰ A. M. Lomnicki, 1894
Species	Harpalus pleuriticus Kirby, 1837 ⇒ Harpalus (Harpalus) somnulentus
Species	Harpalus plustschewskyi Tschitschérine, 1895 ⇒
Species	Harpalus poeciloides Stephens, 1828
Species	Harpalus pohnerti Jedlička, 1929 ⇒ Harpalus (Harpalus) nigrans
Species	Harpalus polychromus Tschitschérine, 1897 ⇒ Harpalus (Harpalus) quadratus
Species	Harpalus polychrous Gistel, 1857
Species	Harpalus ponticus Schauberger, 1926 ⇒ Ophonus (Ophonus) sabulicola
Species	Harpalus porosus Germar, 1817
Species	Harpalus posticus Dejean, 1829
Species	Harpalus potisii Stephens, 1835 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus praedatorius Gistel, 1857
Species	Harpalus praestans Casey, 1924
Species	Harpalus praetermissus Lutshnik, 1921 ⇒ Harpalus (Harpalophonus) terrestris
Species	Harpalus precarius Péringuey, 1896
Species	Harpalus probatus Casey, 1914 ⇒
Species	Harpalus promissus Reiche, 1872
Species	Harpalus promptus Boheman, 1848 ⇒ Harpalus (Harpalus) natalensis
Species	Harpalus propinquus Ballion, 1871
Species	Harpalus proteus Paykull, 1790
Species	Harpalus provensis Casey, 1924 ⇒ Harpalus (Harpalus) ellipsis
Species	Harpalus proximus LeConte, 1848 ⇒
Species	Harpalus przewalskyi Tschitschérine, 1894 ⇒ Harpalus (Harpalus) viridanus viridanus
Species	Harpalus pseudanisodactylus
Species	Harpalus pseudoaeneolus Schauberger, 1930
Species	Harpalus pseudochalybaeus Schauberger, 1926
Species	Harpalus pseudocorporosus Schauberger, 1930 ⇒
Species	Harpalus pseudodimidiatus Schauberger, 1928 ⇒
Species	Harpalus pseudokrueperi Schauberger, 1933 ⇒ Ophonus (Hesperophonus) similis
Species	Harpalus pseudoodes A. Fleischer, 1909 ⇒ Harpalus (Harpalus) davidianus davidianus
Species	Harpalus pseudoparallelus Schauberger, 1926
Species	Harpalus pseudoquadricollis Schauberger, 1932 ⇒ Ophonus (Ophonus) ardosiacus
Species	Harpalus psittaceus Geoffroy, 1785 ⇒
Species	Harpalus psittacinus Reitter, 1900 ⇒
Species	Harpalus pubescens O. F. Müller, 1776
Species	Harpalus pubiger L.Dufour, 1820
Species	Harpalus pubipennis Küster, 1849
Species	Harpalus pubitarsis Casey, 1914
Species	Harpalus pueli Antoine, 1920 ⇒
Species	Harpalus pulcher Dejean, 1829
Species	Harpalus pulchrinulus Reitter, 1900
Species	Harpalus pullus Gyllenhal, 1827
Species	Harpalus pulverulentus P. Rossi, 1790
Species	Harpalus pumilio Casey, 1914
Species	Harpalus punctatellus Reiche, 1850
Species	Harpalus punctatipennis Rambur, 1838
Species	Harpalus punctatulus Duftschmid, 1812
Species	Harpalus punctibasis Poppius, 1906 ⇒ Harpalus (Harpalus) nigrans
Species	Harpalus puncticollis Paykull, 1798
Species	Harpalus punctifer Laporte, 1867
Species	Harpalus punctiferus Castelnau, 1867
Species	Harpalus punctiger Stephens, 1828 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus punctilabris W.S.MacLeay, 1825
Species	Harpalus punctulatus W.S.Macleay, 1825
Species	Harpalus putridus Zetterstedt, 1828
Species	Harpalus pygmaeolus Reitter, 1908
Species	Harpalus quadraticollis Castelnau, 1867
Species	Harpalus quadricollis L. Redtenbacher, 1844 ⇒ Harpalus (Cryptophonus) idiotus
Species	Harpalus quasianxius Kataev, 1989
Species	Harpalus quenselii Gyllenhal, 1810
Species	Harpalus raddei Tschitschérine, 1897
Species	Harpalus ragusae J. Müller, 1924 ⇒
Species	Harpalus rajah Wiedemann, 1824
Species	Harpalus ramburi Rosenhauer, 1856 ⇒ Harpalus (Plectralidus) decipiens decipiens
Species	Harpalus rana Castelnau, 1867 ⇒ Hypharpax moestus
Species	Harpalus ranula Castelnau, 1867
Species	Harpalus rayei Linder, 1864
Species	Harpalus rebellus Schauberger, 1926
Species	Harpalus recensus Casey, 1914 ⇒
Species	Harpalus recisus Casey, 1914
Species	Harpalus rectangulus Casey, 1914
Species	Harpalus redikortzewi Lutshnik, 1934
Species	Harpalus redikorzevi Lutshnik, 1934
Species	Harpalus redivivus ♰ H. F. Wickham, 1917
Species	Harpalus regeli Ballion, 1878
Species	Harpalus regularis Motschulsky, 1844 ⇒ Harpalus (Harpalobius) froelichii
Species	Harpalus reichei Desbrochers des Loges, 1867
Species	Harpalus reinigi Schauberger, 1934 ⇒ Harpalus (Lasioharpalus) smaragdinus
Species	Harpalus reitteri Wrase & Kataev, 2011
Species	Harpalus relator Reitter, 1900
Species	Harpalus relucens Bates, 1873
Species	Harpalus renoicus Casey, 1914 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus retowskianus Reitter, 1887 ⇒ Harpalus (Harpalus) chrysopus chrysopus
Species	Harpalus retowskii Heyden, 1883 ⇒ Harpalus (Harpalus) chrysopus chrysopus
Species	Harpalus retratus Cresson, 1876 ⇒ Harpalus (Plectralidus) retractus
Species	Harpalus rewolinskii Noonan, 1991 ⇒ Harpalus (Harpalus) somnulentus
Species	Harpalus rhemboides Solsky, 1874
Species	Harpalus rhodopus Schauberger, 1930 ⇒
Species	Harpalus rivularis Motschulsky, 1860 ⇒ Harpalus (Harpalobius) froelichii
Species	Harpalus roseni Brancsik, 1899
Species	Harpalus rossii Ponza, 1805
Species	Harpalus rotudangulus Jedlička, 1957 ⇒ Harpalus (Harpalus) erosus
Species	Harpalus rotundatus Chaudoir, 1844 ⇒ Harpalus (Harpalomerus) amputatus obtusus
Species	Harpalus rotundicollis Kolenati, 1845 ⇒ Harpalus (Harpalus) breviusculus
Species	Harpalus roubali Schauberger, 1928 ⇒
Species	Harpalus rubrosuturatus Kuntzen, 1919
Species	Harpalus rudis LeConte, 1863
Species	Harpalus rufescens Dejean, 1829
Species	Harpalus rufibarbis Fabricius, 1792 ⇒ Ophonus (Metophonus) rufibarbis
Species	Harpalus ruficeps Curtis, 1833 ⇒ Harpalus (Amblystus) latus
Species	Harpalus ruficornis Fabricius, 1775 ⇒ Pseudoophonus (Pseudoophonus) rufipes
Species	Harpalus ruficrus Ménétriés, 1832
Species	Harpalus rufimanus LeConte, 1848 ⇒
Species	Harpalus rufitarsis Duftschmid, 1812 ⇒ Harpalus (Harpaloderus) rufipalpis rufipalpis
Species	Harpalus rufithorax C.R.Sahlberg, 1827
Species	Harpalus rufocincta C.R. Sahlberg, 1827 ⇒ Amara (Celia) praetermissa
Species	Harpalus rufocinctus Chaudoir, 1843 ⇒ Harpalus (Harpalus) rufomarginatus
Species	Harpalus rufopiceus Casey, 1914
Species	Harpalus rufus Brüggemann, 1873 ⇒ Harpalus (Harpalobius) autumnalis
Species	Harpalus rugicollis Motschulsky, 1860 ⇒ Harpalus (Pseudoophonus) sinicus
Species	Harpalus rugosipennis Castelnau, 1867
Species	Harpalus rugosus Nietner, 1857
Species	Harpalus rugulosus Heer, 1837 ⇒ Harpalus (Amblystus) latus
Species	Harpalus rupicola Sturm, 1818 ⇒ Ophonus (Metophonus) rupicola
Species	Harpalus rusticus Say, 1823
Species	Harpalus sabulicola Panzer, 1796
Species	Harpalus sabulosus Audinet-Serville, 1821
Species	Harpalus sachalinensis Matsumura, 1911 ⇒ Harpalus (Amblystus) laevipes
Species	Harpalus sadoensis Schauberger, 1929 ⇒ Harpalus (Pseudoophonus) ussuriensis ussuriensis
Species	Harpalus sagowskii Lutshnik, 1909
Species	Harpalus sahlbergi Zetterstedt, 1837
Species	Harpalus sahlbergianus Lutshnik, 1922
Species	Harpalus salinator Pantel, 1888 ⇒
Species	Harpalus salzmanni Germar, 1823
Species	Harpalus sareptanus K. Daniel, 1904 ⇒ Harpalus (Harpalophonus) circumpunctatus
Species	Harpalus sarmaticus Motschulsky, 1850 ⇒
Species	Harpalus sarmatus Schauberger, 1933
Species	Harpalus satanas Reitter, 1900 ⇒
Species	Harpalus satyrus Sturm, 1818
Species	Harpalus scabricollis Quensel, 1806
Species	Harpalus scabripectus Huang; Hu & Sun, 1994
Species	Harpalus scaritides Sturm, 1818 ⇒ Pangus scaritides
Species	Harpalus schaubergeri Csiki, 1932
Species	Harpalus schaubergerianus Puel, 1937
Species	Harpalus schaubergianus
Species	Harpalus schlotheimi Giebel, 1856
Species	Harpalus schouberti Tschitschérine, 1898
Species	Harpalus schreibersii Duftschmid, 1812 ⇒
Species	Harpalus scipio Reitter, 1900 ⇒
Species	Harpalus sculptipennis Castelnau, 1867
Species	Harpalus sculpturalis Castelnau, 1867
Species	Harpalus scytha Tschitschérine, 1899
Species	Harpalus seclusus Casey, 1914 ⇒
Species	Harpalus segnis Dejean, 1829 ⇒ Harpalus (Harpalobius) froelichii
Species	Harpalus sejunctus Casey, 1914 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus seladon Schauberger, 1926 ⇒ Ophonus (Metophonus) rufibarbis
Species	Harpalus semicastaneus Reitter, 1900 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus semiviolaceus Dejean, 1829 ⇒
Species	Harpalus senilis Nietner, 1857
Species	Harpalus serbicus Matits, 1910
Species	Harpalus serdicanus Apfelbeck, 1904 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus seriatus Chaudoir, 1846 ⇒ Harpalus (Plectralidus) metallinus metallinus
Species	Harpalus sericeus Duftschmid, 1812 ⇒
Species	Harpalus sericinus Gemminger & Harold, 1868
Species	Harpalus seriepunctatus Sturm, 1818 ⇒ Harpalus (Harpalobius) autumnalis
Species	Harpalus setipes Motschulsky, 1848
Species	Harpalus sevastopolitanus Schauberger, 1926
Species	Harpalus sibircus Csiki, 1932
Species	Harpalus sibiricus Csiki, 1932 ⇒
Species	Harpalus silicicola Antoine, 1925
Species	Harpalus similatus Gyllenhal, 1810
Species	Harpalus similis Dejean, 1829 ⇒ Ophonus (Hesperophonus) similis
Species	Harpalus simplex Duftschmid, 1812 ⇒
Species	Harpalus simulans J.Sahlberg, 1880
Species	Harpalus sinensis Jedlička, 1957 ⇒ Harpalus (Harpalus) ganssuensis
Species	Harpalus sinis ♰ Heer, 1847
Species	Harpalus sinuaticollis Facchini, 2015
Species	Harpalus skutariensis Reitter, 1900 ⇒ Harpalus (Harpalus) anatolicus anatolicus
Species	Harpalus sobrinus Dejean, 1829 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus socors Casey, 1914 ⇒
Species	Harpalus sodalis Csiki, 1932 ⇒ Harpalus pulchrinulus
Species	Harpalus soginoides J. R. Sahlberg, 1903 ⇒ Harpalus (Harpalus) alajensis
Species	Harpalus solaris Gistel, 1857
Species	Harpalus solieri Dejean, 1831 ⇒ Cryptophonus tenebrosus
Species	Harpalus solutus Casey, 1914
Species	Harpalus somcheticus Schauberger, 1933
Species	Harpalus spaldingi Casey, 1924
Species	Harpalus spec
Species	Harpalus splendens Gebler, 1829 ⇒ Harpalus (Harpalus) colchicus
Species	Harpalus staudingeri Jedlička, 1953
Species	Harpalus staudingerianus Schauberger, 1932 ⇒ Harpalus (Harpalus) viridanus viridanus
Species	Harpalus stepheni Priesner, 1921 ⇒ Harpalus (Glanodes) stephani
Species	Harpalus stephensii Kirby, 1837
Species	Harpalus steveni Dejean, 1829
Species	Harpalus stictus Freude, 1974 ⇒ Ophonus (Ophonus) ardosiacus
Species	Harpalus stierlini ♰ O. Heer, 1862 ⇒ Harpalus (Artabas) dispar splendens
Species	Harpalus stigmatophorus Puel, 1935 ⇒
Species	Harpalus stigmosus Germar, 1823
Species	Harpalus stocktonensis Casey, 1924 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus stoetzneri Schauberger, 1933
Species	Harpalus stolidus Walker, 1858
Species	Harpalus strasseri Reitter, 1900 ⇒
Species	Harpalus stupidus LeConte, 1859 ⇒ Harpalus (Harpalus) somnulentus
Species	Harpalus sturmii Dejean, 1829 ⇒ Harpalus (Harpalophonus) hospes hospes
Species	Harpalus stygius ♰ O. Heer, 1862 ⇒
Species	Harpalus subalbanicus Puel, 1938 ⇒ Harpalus (Harpalus) subcylindricus
Species	Harpalus subangulatus Reitter, 1900
Species	Harpalus subcaeruleus Stephens, 1828
Species	Harpalus subchalibaeus Reitter, 1900 ⇒
Species	Harpalus subchalybaeus Reitter, 1900 ⇒
Species	Harpalus subcontemptus Puel, 1935 ⇒
Species	Harpalus subcordatus Poppius, 1908 ⇒ Harpalus (Harpalomerus) amputatus obtusus
Species	Harpalus subcostatus Dejean, 1829
Species	Harpalus subenormis Casey, 1924 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus sublaevigatus Tschitschérine, 1898 ⇒ Harpalus (Plectralidus) pulvinatus pulvinatus
Species	Harpalus sublaevis Poppius, 1908 ⇒ Harpalus (Artabas) dispar splendens
Species	Harpalus subolivaceus W.S.MacLeay, 1825
Species	Harpalus subpunctatus Stephens, 1828
Species	Harpalus subquadratus Dejean, 1829
Species	Harpalus subsericeus W.J.MacLeay, 1888
Species	Harpalus subsimilis Chaudoir, 1846 ⇒ Harpalus (Harpalophonus) circumpunctatus
Species	Harpalus subsinuatus Duftschmid, 1812 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus subsulcatus Motschulsky, 1848 ⇒ Harpalus (Harpalomerus) amputatus obtusus
Species	Harpalus subtilis Tschitscherine, 1898
Species	Harpalus subvirens Chaudoir, 1846 ⇒
Species	Harpalus suchebaatori Jedlička, 1967 ⇒
Species	Harpalus sulcatulus Faldermann, 1836 ⇒ Harpalus (Harpalus) xanthopus winkleri
Species	Harpalus sulcicollis Paykull, 1798
Species	Harpalus sulcifer Tschitscherine, 1902
Species	Harpalus sulcipennis Faldermann, 1836
Species	Harpalus supremus Schauberger, 1927 ⇒ Ophonus (Hesperophonus) azureus
Species	Harpalus surinamensis Fabricius, 1792 ⇒ Harpalus (Amblystus) latus
Species	Harpalus sushkini Lutshnik, 1934 ⇒
Species	Harpalus sutschanensis Schauberger, 1929 ⇒ Harpalus (Pseudoophonus) pastor pastor
Species	Harpalus suturalis Chaudoir, 1846
Species	Harpalus syriacus Dejean, 1829
Species	Harpalus syriensis Jedlička, 1958 ⇒ Harpalus (Harpalus) caiphus
Species	Harpalus szetschuanensis Schauberger, 1930 ⇒ Harpalus (Pseudoophonus) pastor pastor
Species	Harpalus szetschuanus Schauberger, 1930 ⇒ Harpalus (Pseudoophonus) singularis
Species	Harpalus tabidus ♰ O. Heer, 1847
Species	Harpalus tadocrus Ball, 1972
Species	Harpalus taphrioides Motschulsky, 1844 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus tardigradus ♰ O. Heer, 1862
Species	Harpalus tardoides Jedlička, 1965 ⇒
Species	Harpalus tarsatus Zetterstedt, 1828
Species	Harpalus tarsicus Jedlička, 1958 ⇒
Species	Harpalus taschketensis Jedlicka, 1958
Species	Harpalus tasmanicus Castelnau, 1867 ⇒ Hypharpax moestus
Species	Harpalus tauricus Motschulsky, 1850 ⇒ Harpalus (Harpalophonus) stevenii
Species	Harpalus temperatus H. Kolbe, 1886 ⇒ Harpalus (Harpalus) pallidipennis
Species	Harpalus tenebrioides Waltl, 1838
Species	Harpalus teneriffae Wollaston, 1864 ⇒ Harpalus (Cryptophonus) schaumii
Species	Harpalus tenuelimbatus Reitter, 1900 ⇒
Species	Harpalus tenuestriatus Jeannel, 1948	2	Subspecies
Species	Harpalus testaceipes Roubal, 1926
Species	Harpalus testaceus Haldeman, 1843 ⇒
Species	Harpalus tetradicus Puel, 1935 ⇒
Species	Harpalus thoracicus Motschulsky, 1844 ⇒ Harpalus (Harpalus) pallidipennis
Species	Harpalus thoracinus Casey, 1914
Species	Harpalus thouzeti Castelnau, 1867
Species	Harpalus tibialis Audinet‑Serville, 1821 ⇒
Species	Harpalus tinctus Quensel, 1806
Species	Harpalus tingens Reitter, 1900
Species	Harpalus tingitanus Fairmaire, 1876 ⇒ Harpalus (Harpalus) oblitus patruelis
Species	Harpalus tjanschanicus Semenov, 1889
Species	Harpalus torretassoi Schatzmayr, 1932
Species	Harpalus torridiformis Poppius, 1906 ⇒
Species	Harpalus torridus Motschulsky, 1844 ⇒
Species	Harpalus torta W.S.MacLeay, 1825
Species	Harpalus transcaspicus Tschitschérine, 1898
Species	Harpalus transparens Motschulsky, 1844
Species	Harpalus transversus Casey, 1914
Species	Harpalus trebinjensis A.Fleischer, 1921
Species	Harpalus trechoides Hope, 1845
Species	Harpalus trentinus Csiki, 1932 ⇒ Ophonus (Metophonus) puncticollis
Species	Harpalus tricolor Guérin-Méneville, 1844
Species	Harpalus tridentatus Fischer von Waldheim, 1821
Species	Harpalus tristis Lutshnik, 1922 ⇒
Species	Harpalus trivialis Tschitschérine, 1898 ⇒
Species	Harpalus truncatus Rosenhauer, 1842 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus tschiliensis Schauberger, 1929 ⇒ Harpalus (Pseudoophonus) pastor pastor
Species	Harpalus tsushimanus Habu & Baba, 1968 ⇒ Harpalus (Zangoharpalus) tinctulus luteicornoides
Species	Harpalus tucumanus Dejean, 1831
Species	Harpalus turculus Bates, 1878 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus turkestanicus Csiki, 1932 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus typhonius Gistel, 1857
Species	Harpalus uenci Jedlicka, 1969
Species	Harpalus uintanus Casey, 1924
Species	Harpalus ulamaeformis
Species	Harpalus ulomaeformis ♰ H. F. Wickham, 1917
Species	Harpalus undulatus Gebler, 1841
Species	Harpalus unicolor Motschulsky, 1844 ⇒ Harpalus (Harpalus) viridanus viridanus
Species	Harpalus uriensis Schauberger, 1933
Species	Harpalus ustulatus Dejean, 1829
Species	Harpalus ustus Quensel, 1806
Species	Harpalus uteanus Casey, 1914 ⇒ Harpalus (Harpalus) somnulentus
Species	Harpalus vacivus Casey, 1914
Species	Harpalus validior Rey, 1886 ⇒ Harpalus (Plectralidus) metallinus metallinus
Species	Harpalus vandiemenensis Castelnau, 1867
Species	Harpalus varicornis LeConte, 1848 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus variipes Bates, 1883 ⇒ Harpalus (Harpalus) bungii
Species	Harpalus vasilinini Lutshnik, 1916
Species	Harpalus velox Castelnau, 1867 ⇒ Haplaner velox
Species	Harpalus veluchianus G.Müller, 1931
Species	Harpalus velutinus Dejean, 1829
Species	Harpalus ventricosus Casey, 1924 ⇒ Harpalus (Harpalobius) fuscipalpis
Species	Harpalus vernalis Duftschmid, 1812 ⇒ Harpalus (Actephilus) pumilus
Species	Harpalus versicolor Reitter, 1900 ⇒ Harpalus (Harpalus) viridanus viridanus
Species	Harpalus versutulus Casey, 1924
Species	Harpalus verticalis Erichson, 1842
Species	Harpalus vespertinus Casey, 1884 ⇒ Harpalus (Harpalus) ellipsis
Species	Harpalus vestigialis Erichson, 1842
Species	Harpalus veterum ♰ T. D. A. Cockerell, 1921
Species	Harpalus vicarius Harold, 1878
Species	Harpalus vicinus Dejean, 1829 ⇒
Species	Harpalus viduus LeConte, 1859 ⇒ Harpalus (Euharpalops) providens
Species	Harpalus villosulus Reiche, 1860
Species	Harpalus violaceus Chaudoir, 1844 ⇒ Harpalus (Harpalus) erosus
Species	Harpalus virens Schilsky, 1888 ⇒
Species	Harpalus virescens Faldermann, 1836 ⇒ Harpalus (Plectralidus) metallinus metallinus
Species	Harpalus viridellus Csiki, 1932
Species	Harpalus viridiaeneus Palisot de Beauvois, 1811
Species	Harpalus viridis Say, 1823 ⇒ Harpalus (Harpaloderus) sulphuripes sulphuripes
Species	Harpalus viridulus Geoffroy, 1785 ⇒ Harpalus (Amblystus) rubripes
Species	Harpalus vivesi Jeanne, 1971 ⇒ Harpalus (Harpaloderus) rufipalpis wagneri
Species	Harpalus vividus Dejean, 1829
Species	Harpalus volaki Kult, 1944 ⇒ Ophonus (Metophonus) subsinuatus
Species	Harpalus voltae Basilewsky, 1947
Species	Harpalus vulpeculus Say, 1823
Species	Harpalus vulpinus K. Daniel, 1904 ⇒ Harpalus (Harpalophonus) stevenii
Species	Harpalus wadei Casey, 1924
Species	Harpalus wagneri Schauberger, 1926
Species	Harpalus washingtoniensis Van Dyke, 1926
Species	Harpalus waterhousii Castelnau, 1867
Species	Harpalus weiratheri J. Müller, 1931
Species	Harpalus wellensii Drapiez, 1819
Species	Harpalus wetterhallii Gyllenhal, 1813
Species	Harpalus whitfieldi Scudder, 1900
Species	Harpalus whitfieldii ♰ S. H. Scudder, 1900
Species	Harpalus wilcoxii Castelnau, 1867
Species	Harpalus wilkensii Chaudoir, 1837
Species	Harpalus winkleri Schauberger, 1923
Species	Harpalus wollastoni Dawson, 1854
Species	Harpalus xanthocnemis Alluaud, 1897
Species	Harpalus xanthorhaphus Wiedemann, 1825
Species	Harpalus xaxarsi Schauberger, 1928
Species	Harpalus xinjiangensis Huang; Hu & Sun, 1994
Species	Harpalus yadongensis Huang, 1998 ⇒ Harpalus (Zangoharpalus) praticola
Species	Harpalus yarrae Castelnau, 1867 ⇒ Hypharpax moestus
Species	Harpalus yunnanus Schauberger, 1930 ⇒ Harpalus (Pseudoophonus) sinicus
Species	Harpalus zhicharevi Silfverberg, 1977 ⇒ Harpalus (Harpalophonus) terrestris
Species	Harpalus zicharevi Csiki, 1932
Species	Harpalus zigzag Schauberger, 1926
Species	Harpalus zouhari Jedlička, 1961

Synonyms

Acardystus (Reitter, 1908)

Actephilus (Stephens, 1833)

Acthephilus (Reitter, 1900)

Actophilus (Agassiz, 1846)

Amblystus (Motschulsky, 1864)

Anisochirus (Jeannel, 1946)

Artabas (Gozis, 1882)

Asmerinx (Reitter, 1899)

Baeticoharpalus (Serrano & Lencina, 2009)

Bioderus (Motschoulsky, 1848)

Brachyophonus (Sciaky, 1986)

Brachypangus (Tschitschérine, 1898)

Cephalomorphus (Tschitscherin, 1897)

Cephalophonus (Ganglbauer, 1892)

Cephalotypsis (Tschitscherin, 1901)

Conicus (Motschoulsky, 1844)

Cordoharpalus (Hatch, 1949)

Crytophonus (Brandmayr & Zetto-Brandmayr, 1982)

Empeirus (Motschoulsky, 1844)

Epharpalus (Jacobson, 1907)

Epiharpalus (Reitter, 1900)

Erpeinus (Motschoulsky, 1844)

Euharpalops (Casey, 1924)

Glanodes (Casey, 1914)

Haploharpalus (Schauberger, 1926)

Harpalellus (Lindroth, 1968)

Harpalobius (Reitter, 1900)

Harpalobrachys (Tschitschérine, 1899)

Harpaloderus (Reitter, 1900)

Harpalomerus (Casey, 1914)

Harpalophonus (Ganglbauer, 1892)

Harpaloxis (Reitter, 1900)

Harpaloxys (Reitter, 1900)

Harpalus (Actephilus) (Stephens, 1839)

Harpalus (Baeticoharpalus) (Serrano & Lencina, 2009)

Harpalus (Brachypangus) (Tschitschérine, 1898)

Harpalus (Cephalotypsis) (Tschitschérine, 1901)

Harpalus (Harpaloderus) (Reitter, 1900)

Harpalus (Harpalomerus) (Casey, 1914)

Harpalus (Lasioharpalus) (Reitter, 1900)

Harpalus (Loxophonus) (Reitter, 1894)

Harpalus (Microharpalus) (Tschitschérine, 1901)

Harpalus (Migadophonus) (Tschitschérine, 1897)

Harpalus (Neopangus) (Tschitschérine, 1898)

Harpalus (Pharalus) (Casey, 1914)

Harpalus (Pheuginus) (Motschulsky, 1844)

Hesperophonus (Antoine, 1959)

Hypsinephus (Bates, 1878)

Incisophonus (Sciaky, 1986)

Lasiarpalus (Jacobson, 1907)

Lasioharpalus (Reitter, 1900)

Licinoderus (Sainte-Claire-Deville, 1905)

Loboharpalus (Schauberger, 1932)

Loxophonus (Reitter, 1894)

Macrophonus (Tschitscherin, 1901)

Megapangus (Casey, 1914)

Metophonus (Bedel, 1897)

Micrarpalus (Jacobson, 1907)

Microderes (Faldermann, 1835)

Microderus (Motschoulsky, 1844)

Microharpalus (Tschitscherin, 1901)

Migadophonus (Tschitschérine, 1897)

Neoharpalus (Mateu, 1954)

Neopangus (Tschitscherin, 1898)

Neopardileus (Habu, 1954)

Neophygas (Noonan, 1976)

Nephoharpalus (Huang, Lei, Yan & Hu, 1996)

Ooistus (Motschoulsky, 1864)

Opadius (Casey, 1914)

Ophonus (Dejean, 1821)

Ophonus (Brachyophonus) (Sciaky, 1987)

Ophonus (Hesperophonus) (Antoine, 1959)

Ophonus (Incisophonus) (Sciaky, 1987)

Ophonus (Macrophonus) (Tschitschérine, 1901)

Pangus (Motschulsky, 1859)

Pardileus (des Gozis, 1882)

Parhalus (Jeannel, 1946)

Pharalus (Casey, 1914)

Pheuginus (Motschoulsky, 1845)

Phygas (Motschoulsky, 1848)

Platus (Motschoulsky, 1844)

Plectralidus (Casey, 1914)

Pseudoaphonus (Chaudoir, 1863)

Pseudoophonus (Motschulsky, 1844)

Pseudoophonus (Platus) (Motschulsky, 1844)

Pseudophonus (Motschoulsky in Ménétriés, 1848)

Rapahlus (Lu?nik, 1922)

Semiophonus (Schauberger, 1933)

Sulcophonus (Schauberger, 1933)

Typsarpalus (Jacobson, 1907)

Native, vernacular names and spellings of Harpalus

German

Schnelläufer, Schnellläufer

Distribution of Harpalus Latreille, 1802

Afghanistan

Albania

Algeria

Andorra

Angola

Argentina

Armenia

Australia
27 Species

Austria
64 Species

Azerbaijan

Belarus

Belgium

Bhutan

Bolivia

Bosnia and Herzegovina

Botswana

Brazil
5 Species

Bulgaria

Cambodia

Canada
82 Species

Cape Verde

Chile

China
163 Species

Colombia

Comoros

Costa Rica

Croatia

Cyprus

Czech Republic

Democratic Republic of the Congo
1 Species

Denmark

Ecuador

Egypt

Eritrea

Estonia

Ethiopia

Finland

France
70 Species

Georgia
55 Species

Germany
76 Species

Greece
68 Species

Guatemala

Honduras

Hungary

Iceland

India
23 Species

Indonesia
6 Species

Iran
44 Species

Iraq

Ireland

Israel

Italy
83 Species

Jamaica

Japan
43 Species

Jordan

Kazakhstan
92 Species

Kenya

Kyrgyzstan

Laos

Latvia

Lebanon

Lesotho

Liberia

Libya

Liechtenstein

Lithuania

Luxembourg

Madagascar
13 Species

Malawi

Malta

Mauritius

Mexico
22 Species

Moldova

Mongolia

Morocco

Mozambique

Myanmar
2 Species

Namibia

Nepal
11 Species

Netherlands

New Zealand

North Korea

Norway

Oman

Pakistan

Palestine

Panama

Peru

Philippines

Poland

Portugal

Republic South Africa
54 Species

Romania

Russia
155 Species

Saudi Arabia

Senegal

Serbia

Sierra Leone

Slovakia

Slovenia

Somalia

South Korea

Spain
63 Species

Sudan

Sweden

Switzerland

Syria

Taiwan

Tajikistan

Tanzania
4 Species

Thailand

Togo

Tunisia

Turkey
83 Species

Turkmenistan

Uganda

Ukraine

United Kingdom
68 Species

USA
152 Species

Uzbekistan

Vietnam
5 Species

Yemen

Zambia

Zimbabwe

Distribution on islands

Orkney Islands

Found in fossile formations

Florissant Fossil Beds, Green River Formation, Rott Formation, Sarka Formation

Links, sources and further information

Harpalus at artfakta.se
Harpalus at gbif.org
Harpalus at irmng.org
Harpalus at itis.gov