Checklist for Tribe Helluonini Hope, 1838 (Flat-horned Ground Beetles)
and region Continents
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 2110
116133
1222976 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 2101
115931
1221959 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 1812
113780
1210168 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 929
84484
1007160 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 234
35522
419905 | Families
Genera
Species |
Taxom
|
Suborder
| Adephaga
Schellenberg, 1806
Adephagans, Ground Beetles and Water Beetles | 16
3006
54917 | Families
Genera
Species |
Taxom
|
Superfamily
| Caraboidea
Latreille, 1802
| 2
2636
47670 | Families
Genera
Species |
Taxom
| | Family | Carabidae
Latreille, 1802
Carabid beetles, Carabids, Ground Beetles, Tiger Beetles | 2600
47579 | Genera
Species |
Taxom
| | Subfamily | Harpalinae
Bonelli, 1810
| 1322
25865 | Genera
Species |
Taxom
| | Supertribe | Anthiitae
Bonelli, 1813
| 40
403 | Genera
Species |
Taxom
| | Tribe | Helluonini
Hope, 1838
Flat-horned Ground Beetles | 26
206 | Genera
Species |
Taxom
|
Genera for Continents
26 Genera with 206 Species #ACDEGHMNOPT
| Ametroglossus Sloane, 1887 | | 1 | Species | Taxom | | Colfax Andrewes, 1920 | | 2 | Species | Taxom | | Creagris Nietner, 1867 | | 12 | Species | Taxom | | Dailodontus Reiche, 1834 | | 2 | Species | Taxom | | Dicranoglossus Chaudoir, 1867 | | 1 | Species | Taxom | | Epimicodema Sloane, 1871 | | 1 | Species | Taxom | | Erephognathus Alluaud, 2016 | | 3 | Species | Taxom | | Gigadema J. Thomson, 1864 | | 15 | Species | Taxom | | Helluapterus Sloane, 1914 | | 1 | Species | Taxom | | Helluarchus Sloane, 1914 | | 2 | Species | Taxom | | Helluo Bonelli, 1890 | | 2 | Species | Taxom | | Helluobrochus Reichardt, 1871 | | 26 | Species | Taxom | | Helluodema Castelnau, 1842 | | 2 | Species | Taxom | | Helluomorpha Castelnau de..., 1862 | | 5 | Species | Taxom | | Helluomorphoides Ball, 1834 | Flat-horned Ground Beetles | 26 | Species | Taxom | | Helluonidius Chaudoir, 1968 | | 9 | Species | Taxom | | Helluopapa Darlington, 1968 | | 2 | Species | Taxom | | Helluopapua Darlington, 1968 | | 2 | Species | Taxom | | Helluosoma Castelnau, 1888 | | 7 | Species | Taxom | | Macrocheilus Hope, 1920 | | 64 | Species | Taxom | | Meladroma Motschoulsky, 1801 | | 4 | Species | Taxom | | Neohelluo Sloane, 1914 | | 1 | Species | Taxom | | Omphra Dejean, 1834 | | 7 | Species | Taxom | | Platyhelluo Baehr, 2005 | | 1 | Species | Taxom | | Pleuracanthus Gray, 1974 | | 4 | Species | Taxom | | Triaenogenius Chaudoir, 1943 | | 19 | Species | Taxom |
Species list for Continents
206 Species and 15 Subspecies | Ametroglossus ater W. McLeay, 1887 | | | Colfax stevensi Andrewes, 1920 | | | Colfax stewensi Andrewes, 1920 | | | Creagris bigemmis Andrewes, 1931 | | | Creagris binoculus Bates, 1892 | | | Creagris bisignata Landin, 1955 | | | Creagris distacta Wiedemann, 1823 | | | Creagris hamaticollis Bates, 1892 | | | Creagris horikawae (Nakahara, 1913) | ⇒ Pseudoformicaleo nubecula Gerstäcker, 1885 | | Creagris labrosa Nietner, 1857 | | | Creagris lineola Andrewes, 1926 | | | Creagris matsuokae (Okamoto, 1910) | ⇒ Pseudoformicaleo nubecula Gerstäcker, 1885 | | Creagris nubecula (Gerstaecker, , 1885) | ⇒ Pseudoformicaleo nubecula Gerstäcker, 1885 | | Creagris rubrothorax Louwerens, 1949 | | | Creagris wilsonii Castelnau, 1867 | | | Dailodontus cayennensis Dejean, 1826 | | | Dailodontus clandestinus Klug, 1834 | | | Dicranoglossus resplendens Castelnau, 1867 | | | Epimicodema mastersii W.J.Macleay, 1871 | | | Erephognathus coerulescens Fairmaire, 1903 | | | Erephognathus margarithrix Alluaud, 1936 | | | Erephognathus ranomafanae Kavanaugh & ..., 2016 | | | Gigadema biordinatum Sloane, 1914 | | | Gigadema bostocki Castelnau, 1867 | | | Gigadema bostockii Laporte, 1867 | | | Gigadema dux Blackburn, 1901 | | | Gigadema froggatti W.J.Macleay, 1888 | | | Gigadema grande MacLeay, 1864 | | | Gigadema gulare Sloane, 1914 | | | Gigadema longipenne Germar, 1848 | | | Gigadema mandibulare Blackburn, 1892 | | | Gigadema maxillare Sloane, 1914 | | | Gigadema minuta (Castelnau, 1867) | ⇒ Gigadema sulcatum W.J.Macleay, 1864 | | Gigadema nocte Newman, 1842 | | | Gigadema obscurum Sloane, 1914 | | | Gigadema rugaticolle Blackburn, 1901 | | | Gigadema sulcatum W.J.Macleay, 1864 | | | Helluapterus niger Sloane, 1914 | | | Helluarchus robustus Sloane, 1914 | | | Helluarchus whitei Lea, 1914 | | | Helluo costatus Bonelli, 1813 | | | Helluo insignis Sloane, 1890 | | | Helluobrochus anthracinus Klug, 1834 | | | Helluobrochus ares Reichardt, 1974 | | | Helluobrochus argus Reichardt, 1974 | | | Helluobrochus bacchus Reichardt, 1972 | | | Helluobrochus bechynei Reichardt, 1974 | | | Helluobrochus birai Reichardt, 1974 | | | Helluobrochus brasiliensis Dejean, 1825 | | | Helluobrochus brevicollis Dejean, 1831 | | | Helluobrochus bucki Reichardt, 1974 | | | Helluobrochus capixaba Reichardt, 1974 | | | Helluobrochus collaris Chaudoir, 1877 | | | Helluobrochus cribratus Reiche, 1843 | | | Helluobrochus cribricollis Chaudoir, 1872 | | | Helluobrochus darlingtoni Reichardt, 1974 | | | Helluobrochus horqueta Reichardt, 1974 | | | Helluobrochus inconspicuus Chaudoir, 1848 | | | Helluobrochus lacordairei Dejean, 1831 | | | Helluobrochus linearis Bates, 1871 | | | Helluobrochus luctuosus Chaudoir, 1872 | | | Helluobrochus negrei Reichardt, 1974 | | | Helluobrochus oculatus Reichardt, 1974 | | | Helluobrochus oopselaphus Reichardt, 1974 | | | Helluobrochus petrus Reichardt, 1974 | | | Helluobrochus pilipalpis Reichardt, 1974 | | | Helluobrochus sanguinolentus Klug, 1834 | | | Helluobrochus subrostratus Bates, 1871 | | | Helluodema brunneum Sloane, 1917 | | | Helluodema unicolor Hope, 1842 | | | Helluomorpha araujoi Reichardt, 1974 | | | Helluomorpha eulinae Reichardt, 1974 | | | Helluomorpha heros Gory, 1833 | | | Helluomorpha macroptera Chaudoir, 1850 | | | Helluomorpha protogaea ♰ C. Giebel, 1862 | | | Helluomorphoides balli Reichardt, 1974 | | | Helluomorphoides brunneus Putzeys, 1846 | | | Helluomorphoides clairvillei Dejean, 1831 | | | Helluomorphoides diana Reichardt, 1974 | | | Helluomorphoides femoratus Dejean, 1831 | | | Helluomorphoides ferrugineus LeConte, 1853 | | | Helluomorphoides gigantops Reichardt, 1974 | | | Helluomorphoides glabratus Bates, 1871 | | | Helluomorphoides io Reichardt, 1974 | | | Helluomorphoides juno Reichardt, 1974 | | | Helluomorphoides latitarsis Casey, 1913 | | | Helluomorphoides longicollis Bates, 1883 | | | Helluomorphoides mexicanus Chaudoir, 1872 | | | Helluomorphoides nigerrimus Klug, 1834 | | | Helluomorphoides nigripennis Dejean, 1831 | | | Helluomorphoides oculeus Bates, 1871 | | | Helluomorphoides papago Casey, 1913 | | | Helluomorphoides praeustus Dejean, 1825 | Bluespot goby | | Helluomorphoides praeustus bicolor T.W.Harris, 1828 | | | Helluomorphoides praeustus floridanus Ball, 1956 | | | Helluomorphoides praeustus praeustus Dejean, 1825 | | | Helluomorphoides ritae Reichardt, 1974 | | | Helluomorphoides rubricollis Schaum, 1863 | | | Helluomorphoides squiresi Chaudoir, 1872 | | | Helluomorphoides texanus LeConte, 1853 | | | Helluomorphoides unicolor Brullé, 1834 | | | Helluonidius aterrimus W.J.Macleay, 1873 | | | Helluonidius chrysocomes Maindron, 1908 | | | Helluonidius chrysocomus Maindron, 1908 | | | Helluonidius cyaneus Castelnau, 1867 | | | Helluonidius cyanipennis Hope, 1842 | | | Helluonidius laevifrons Darlington, 1968 | | | Helluonidius latipennis W.J.Macleay, 1887 | | | Helluonidius latipes Darlington, 1968 | | | Helluonidius politus Darlington, 1968 | | | Helluopapa cheesmani Darlington, 1971 | | | Helluopapa toxopei Darlington, 1968 | | | Helluopapua cheesmani Darlington, 1971 | | | Helluopapua toxopei Darlington, 1968 | | | Helluosoma atrum Laporte, 1867 | | | Helluosoma bisetosum Baehr, 2017 | | | Helluosoma bouchardi Baehr, 2005 | | | Helluosoma crenulicolle Baehr, 2017 | | | Helluosoma hangayi Baehr, 2005 | | | Helluosoma kalumburu Baehr, 2017 | | | Helluosoma longicolle MacLeay, 1888 | | | Macrocheilus allardi Basilewsky, 1957 | | | Macrocheilus alluaudi Burgeon, 1937 | | | Macrocheilus angustatus Basilewsky, 1949 | | | Macrocheilus asteriscus White, 1844 | | | Macrocheilus bandipurensis Akhil & Divy..., 2019 | | | Macrocheilus basilewskyi A.Serrano, 2000 | | | Macrocheilus bensoni Hope, 1838 | | | Macrocheilus bicolor Andrewes, 1920 | | | Macrocheilus biguttatus Gory, 1832 | | | Macrocheilus bimaculatus (Dejean, 1831) | ⇒ Macrocheilus bensoni Hope, 1838 | | Macrocheilus binotatus Andrewes, 1931 | | | Macrocheilus biplagiatus Boheman, 1848 | | | Macrocheilus burgeoni Basilewsky, 1967 | | | Macrocheilus chaudoiri Andrewes, 1919 | | | Macrocheilus cheni D.Y. Zhao & ..., 2010 | | | Macrocheilus chinnarensis Akhil & Divy..., 2019 | | | Macrocheilus clasispilus Basilewsky, 1967 | | | Macrocheilus crampeli Alluaud, 1916 | | | Macrocheilus cribrarius Fairmaire, 1901 | | | Macrocheilus cruciatus Marc, 1840 | | | Macrocheilus deuvei Zhao & Tian, 2012 | | | Macrocheilus deuvie (Zhao & Tian, 2012) | ⇒ Macrocheilus deuvei Zhao & Tian, 2012 | | Macrocheilus devagiriensis Shiu & Sabu, 2012 | | | Macrocheilus diplospilus Basilewsky, 1967 | | | Macrocheilus dorsalis Klug, 1834 | | | Macrocheilus dorsiger Chaudoir, 1876 | | | Macrocheilus elegantulus Burgeon, 1937 | | | Macrocheilus ferruginipes Fairmaire, 1892 | | | Macrocheilus fuscipennis Zhao & Tian, 2010 | | | Macrocheilus gigas Zhao & Tian, 2010 | | | Macrocheilus hybridus Péringuey, 1896 | | | Macrocheilus immanis Andrewes, 1920 | | | Macrocheilus impictus Wiedemann, 1823 | | | Macrocheilus labrosus Dejean, 1831 | | | Macrocheilus longicollis Péringuey, 1904 | | | Macrocheilus macromaculatus Louwerens, 1949 | | | Macrocheilus madagascariensis Basilewsky, 1953 | | | Macrocheilus moraisi A.Serrano, 2000 | | | Macrocheilus niger Andrewes, 1920 | | | Macrocheilus nigrotibialis Heller, 1900 | | | Macrocheilus ocellatus Basilewsky, 1953 | | | Macrocheilus overlaeti Burgeon, 1937 | | | Macrocheilus parvimaculatus Zhao & Tian, 2010 | | | Macrocheilus perrieri Fairmaire, 1899 | | | Macrocheilus perrieri perrieri Fairmaire, 1899 | | | Macrocheilus perrieri unicolor Alluaud, 1935 | | | Macrocheilus persimilis Basilewsky, 1970 | | | Macrocheilus proximus Péringuey, 1896 | | | Macrocheilus quadratus Zhao & Tian, 2010 | | | Macrocheilus quadrinotatus Burgeon, 1937 | | | Macrocheilus saulcyi Chevrolat, 1854 | | | Macrocheilus scapularis Reiche, 1843 | | | Macrocheilus sinuatilabris Zhao & Tian, 2010 | | | Macrocheilus solidipalpis Zhao & Tian, 2010 | | | Macrocheilus spectandus Péringuey, 1904 | | | Macrocheilus taedatus Basilewsky, 1960 | | | Macrocheilus tripustulatus Dejean, 1825 | | | Macrocheilus vanharteni Felix & Muil..., 2007 | | | Macrocheilus varians Péringuey, 1904 | | | Macrocheilus viduatus Péringuey, 1899 | | | Macrocheilus viduatus katanganus Basilewsky, 1960 | | | Macrocheilus viduatus viduatus Péringuey, 1899 | | | Macrocheilus vinctus Basilewsky, 1960 | | | Macrocheilus vitalisi Andrewes, 1920 | | | Meladroma angustata Basilewsky, 1970 | | | Meladroma informicollis Liebke, 1928 | | | Meladroma katangensis Burgeon, 1937 | | | Meladroma umbraculata Fabricius, 1801 | | | Neohelluo angulicollis Sloane, 1914 | | | Omphra atrata Klug, 1834 | | | Omphra complanata Reiche, 1843 | | | Omphra drumonti Raj, Sabu & ..., 2012 | | | Omphra hirta Fabricius, 1801 | | | Omphra pilosa Klug, 1834 | | | Omphra rotundicollis Chaudoir, 1872 | | | Omphra rufipes Klug, 1834 | | | Platyhelluo weiri Baehr, 2005 | | | Pleuracanthus inca Reichardt, 1974 | | | Pleuracanthus psittacus Reichardt, 1974 | | | Pleuracanthus sulcipennis Gray, 1832 | | | Pleuracanthus tridens Reichardt, 1974 | | | Triaenogenius (Androchaetus) lugubris Schaum, 1863 | | | Triaenogenius (Triaenogenius) arabicus Gestro, 1889 | | | Triaenogenius (Triaenogenius) carinulatus Fairmaire, 1887 | | | Triaenogenius carinulatus agnatus Péringuey, 1896 | | | Triaenogenius carinulatus carinulatus Fairmaire, 1887 | | | Triaenogenius carinulatus vicinus Péringuey, 1896 | | | Triaenogenius (Triaenogenius) congoensis Basilewsky, 1959 | | | Triaenogenius (Triaenogenius) corpulentus Chaudoir, 1877 | | | Triaenogenius (Triaenogenius) denticulatus Basilewsky, 1959 | | | Triaenogenius (Triaenogenius) ferox Erichson, 1843 | | | Triaenogenius (Triaenogenius) gerstaeckeri Chaudoir, 1877 | | | Triaenogenius (Triaenogenius) kochi Basilewsky, 1964 | | | Triaenogenius (Triaenogenius) lugubrinus Boheman, 1860 | | | Triaenogenius (Triaenogenius) sculpturatus Gerstaecker, 1867 | | | Triaenogenius sculpturatus infuscatus Basilewsky, 1943 | | | Triaenogenius sculpturatus patrizii Alluaud, 1923 | | | Triaenogenius sculpturatus rhodesianus Basilewsky, 1959 | | | Triaenogenius sculpturatus sculpturatus Gerstaecker, 1867 | | | Triaenogenius sculpturatus subcostatus G.Müller, 1943 | |
Subnodes of Helluonini with occurrences in Continents
Links, sources and further information Helluonini at itis.gov
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