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Checklist for Subfamily Helophorinae Leach, 1815
and region Continents


Class
 		Insecta
Linnaeus, 1758
Insects, True insects		2110
116133
1222976		Families
Genera
Species		 
Taxom
 

Subclass
 		Pterygota
Lang, 1888
Winged insects		2101
115931
1221959		Families
Genera
Species		 
Taxom
 

Infraclass
 		Neoptera
van der Wulp, 1890
Modern Wing-folding insects		1812
113780
1210168		Families
Genera
Species		 
Taxom
 

Superorder
 		Holometabola
Heider, 1889
Holometabolous Insects		929
84484
1007160		Families
Genera
Species		 
Taxom
 

Order
 		Coleoptera
Linnaeus, 1758
Beetles, Coleopterans		234
35522
419905		Families
Genera
Species		 
Taxom
 

Suborder
 		Polyphaga
Emery, 1886
Polyphags		189
32170
363924		Families
Genera
Species		 
Taxom
 

Infraorder
 		Staphyliniformia
Lameere, 1900
15
5703
91627		Families
Genera
Species		 
Taxom
 

Superfamily
 		Hydrophiloidea
Latreille, 1802
Histerids, Hydriphiloids, Hydrophilids		5
258
4652		Families
Genera
Species		 
Taxom
 
FamilyHydrophilidae
Latreille, 1802
Silver Water Beetles, Water beetles		253
4640		Genera
Species		 
Taxom
 
SubfamilyHelophorinae
Leach, 1815
6
276		Genera
Species		 
Taxom
 


Genera for Continents
6 Genera with 276 Species
Cretotaenia Ponomarenko, 19771SpeciesTaxom
Elophorus Fabricius, 19651SpeciesTaxom
Helophorus Fabricius, 1988282SpeciesTaxom
Hydrophilopsia Ponomarenko, 19871SpeciesTaxom
Laetopsia Fikacek & al., 20105SpeciesTaxom
Zetemenos Bode, 19531SpeciesTaxom


Species list for Continents
276 Species and 7 Subspecies
Cretotaenia pallida ♰ A. G. Ponoma..., 1977
Elophorus leechi (McCorkle, 1965)⇒ Helophorus leechi McCorkle, 1965
Helophorus (Atractelophorus) arvernicus Mulsant, 1846
Helophorus (Atractelophorus) brevipalpis Bedel, 1881
Helophorus (Atractelophorus) glacialis A. Villa & G..., 1883
Helophorus (Atractelophorus) nivalis Giraud, 1851
Helophorus (Atracthelophorus) altaicus Ganglbauer, 1901
Helophorus (Atracthelophorus) arvernicus Mulsant, 1846
Helophorus (Atracthelophorus) aspericollis Angus, 1973
Helophorus (Atracthelophorus) auricollis Eschscholtz, 1822
Helophorus (Atracthelophorus) brevipalpis Bedel, 1881
Helophorus (Atracthelophorus) brevitarsis Kuwert, 1886
Helophorus (Atracthelophorus) confrater Kuwert, 1886
Helophorus (Atracthelophorus) glacialis Villa, 1883
Helophorus (Atracthelophorus) montenegrinus Kuwert, 1885
Helophorus (Cyphelophorus) tuberculatus Gyllenhall, 1808
Helophorus (Empleurus) hirsutiventris Angus, 1984
Helophorus (Empleurus) hispanicus Sharp, 1915
Helophorus (Empleurus) nubilis Fabricius, 1776
Helophorus (Empleurus) nubilus Fabricius, 1776Wheat Mud Beetle
Helophorus (Empleurus) porculus Bedel, 1881Turnip mud beetle
Helophorus (Empleurus) rufipes Bosc, 1791Turnip Mud Beetle
Helophorus (Empleurus) schmidti Villa, 1838
Helophorus (Eutrichelophorus) micans Faldermann, 1853
Helophorus (Gephelophorus) auriculatus Sharp, 1915
Helophorus (Gephelophorus) sibiricus Motschulsky, 1860Siberian water scavenger beetle
Helophorus (Helophorus) aequalis C.G. Thomson, 1868
Helophorus (Helophorus) aquaticus Linnaeus, 1758Beetle
Helophorus (Helophorus) asperatus Rey, 1885
Helophorus (Helophorus) bergrothi (J. Sahlberg, 1880)⇒ Helophorus strandi Angus, 1970
Helophorus (Helophorus) croaticus Kuwert, 1886
Helophorus (Helophorus) discrepans Rey, 1885
Helophorus (Helophorus) dorsalis Marsham, 1802
Helophorus (Helophorus) flavipes Fabricius, 1792
Helophorus (Helophorus) fulgidicollis Motschulsky, 1860
Helophorus (Helophorus) grandis Illiger, 1798Dwarf sand verbena
Helophorus (Helophorus) granularis Linnaeus, 1761
Helophorus (Helophorus) griseus Herbst, 1793
Helophorus (Helophorus) hammondi Angus, 1970
Helophorus (Helophorus) illustris Sharp, 1916
Helophorus (Helophorus) khnzoryani Angus, 1970
Helophorus (Helophorus) laticollis C.G. Thomson, 1854
Helophorus (Helophorus) liguricus Angus, 1970
Helophorus (Helophorus) longitarsis Wollaston, 1864
Helophorus (Helophorus) maritimus Rey, 1885
Helophorus (Helophorus) milleri Kuwert, 1886
Helophorus (Helophorus) minutus Fabricius, 1775
Helophorus (Helophorus) nanus Sturm, 1836
Helophorus (Helophorus) niger J. Sahlberg, 1880
Helophorus (Helophorus) obscurus Mulsant, 1844
Helophorus (Helophorus) occidentalis Angus, 1983
Helophorus (Helophorus) pallidipennis
Helophorus (Helophorus) pallidus Gebler, 1830
Helophorus (Helophorus) paraminutus Angus, 1986
Helophorus (Helophorus) pumilio Erichson, 1837
Helophorus (Helophorus) redtenbacheri Kuwert, 1885
Helophorus (Helophorus) strandi Angus, 1970
Helophorus (Helophorus) strigifrons C.G. Thomson, 1868
Helophorus (Helophorus) syriacus Kuwert, 1885
Helophorus (Helophorus) villosus Duftschmid, 1805
Helophorus (Kyphohelophorus) tuberculatus Gyllenhal, 1808
Helophorus (Megahelophorus) aequalis Thomson, C.G., 1868
Helophorus (Megahelophorus) grandis Illiger, 1798
Helophorus (Megalelophorus) aquaticus
Helophorus (Megalelophorus) grandis Illiger, 1798
Helophorus grandis maritimus
Helophorus (Megalelophorus) liguricus Angus, 1970
Helophorus (Meghelophorus) aequalis C.G. Thomson, 1868
Helophorus (Meghelophorus) aquaticus Linnaeus, 1758
Helophorus (Meghelophorus) grandis Illiger, 1798
Helophorus (Meghelophorus) khnzoriani Angus, 1970
Helophorus (Mesohelophorus) palaeosibiricus ♰ Fikacek & al., 2012
Helophorus (Mesosperchus) costalis ♰ Ponomarenko, 2014
Helophorus (Mesosperchus) gracilis ♰ Prokin & al., 2010
Helophorus (Mesosperchus) inceptivus ♰ Fikacek & al., 2012
Helophorus (Mesosperchus) tarsalis ♰ Ponomarenko, 1977
Helophorus (Mesosperchus) yixianus ♰ Fikacek & al., 2012
Helophorus (Orphelophorus) arcticus Brown, 1937
Helophorus (Orphelophorus) obscurellus Poppius, 1907
Helophorus (Orphelophorus) turca Incekara, 2008
Helophorus (Rhopalohelophorus) alternatus LeConte, 1861
Helophorus (Rhopalohelophorus) angustatus Motschulsky, 1860
Helophorus (Rhopalohelophorus) angusticollis Orchymont, 1945
Helophorus (Rhopalohelophorus) artus Smetana, 1985
Helophorus (Rhopalohelophorus) arvernicus Mulsant, 1846
Helophorus (Rhopalohelophorus) asperatus Rey, 1885
Helophorus (Rhopalohelophorus) asturiensis Kuwert, 1885
Helophorus (Rhopalohelophorus) auricollis Eschscholtz, 1822Toothed pogonatum moss
Helophorus (Rhopalohelophorus) bameuli Angus, 1987
Helophorus (Rhopalohelophorus) barbarae Angus, 1984
Helophorus (Rhopalohelophorus) brevipalpis Bedel, 1881
Helophorus brevipalpis brevipalpis
Helophorus brevipalpis levantinus Angus, 1988
Helophorus (Rhopalohelophorus) brevitarsis Kuwert, 1890
Helophorus (Rhopalohelophorus) browni McCorkle, 1970
Helophorus (Rhopalohelophorus) californicus Smetana, 1985
Helophorus (Rhopalohelophorus) calpensis Angus, 1988
Helophorus (Rhopalohelophorus) carsoni Angus, 1970
Helophorus (Rhopalohelophorus) chamberlaini Smetana, 1985
Helophorus (Rhopalohelophorus) cincticollis Guillebeau, 1893
Helophorus (Rhopalohelophorus) columbianus McCorkle, 1965
Helophorus (Rhopalohelophorus) confrater
Helophorus (Rhopalohelophorus) croaticus Kuwert, 1886
Helophorus (Rhopalohelophorus) cuspifer Smetana, 1985
Helophorus (Rhopalohelophorus) dalmatinus Ganglbauer, 1901
Helophorus (Rhopalohelophorus) discrepans Rey, 1885
Helophorus (Rhopalohelophorus) dorsalis Marsham, 1802
Helophorus (Rhopalohelophorus) eclectus Orchymont, 1945
Helophorus (Rhopalohelophorus) fauveli Ganglbauer, 1901
Helophorus (Rhopalohelophorus) fenderi McCorkle, 1965
Helophorus (Rhopalohelophorus) flavipes Fbricius, 1792
Helophorus (Rhopalohelophorus) fortis LeConte, 1866
Helophorus (Rhopalohelophorus) frater (Orchymont, 1926)⇒ Helophorus furius Smetana, 1987
Helophorus (Rhopalohelophorus) frosti Smetana, 1985
Helophorus (Rhopalohelophorus) fulgidicollis Motschulsky, 1860
Helophorus (Rhopalohelophorus) furius Smetana, 1987
Helophorus (Rhopalohelophorus) glacialis A.Villa & G...., 1833
Helophorus (Rhopalohelophorus) granularis Linnaeus, 1761
Helophorus (Rhopalohelophorus) gratus Angus, 1987
Helophorus (Rhopalohelophorus) griseus Herbst, 1793
Helophorus (Rhopalohelophorus) hatchi McCorkle, 1965
Helophorus (Rhopalohelophorus) illustris Sharp, 1916
Helophorus (Rhopalohelophorus) inflectus McCorkle, 1965
Helophorus (Rhopalohelophorus) jacutus Poppius, 1907
Helophorus (Rhopalohelophorus) jocoteroi Angus & Diaz..., 1991
Helophorus (Rhopalohelophorus) kaszabianus Angus, 1970
Helophorus (Rhopalohelophorus) kervillei Orchymont, 1932
Helophorus (Rhopalohelophorus) kirgisicus Knisch, 1914
Helophorus (Rhopalohelophorus) korotyaevi Angus, 1985
Helophorus (Rhopalohelophorus) kryzanovskii Angus, 1984
Helophorus (Rhopalohelophorus) lacustris LeConte, 1850
Helophorus (Rhopalohelophorus) lapponicus C.G. Thomson, 1853
Helophorus (Rhopalohelophorus) laticollis C.G. Thomson, 1853New Forest Mud Beetle
Helophorus (Rhopalohelophorus) latipenis Hilsenhoff, 1995
Helophorus (Rhopalohelophorus) lecontei Knisch, 1924Powell's amaranth
Helophorus (Rhopalohelophorus) ledatus Orchymont, 1945
Helophorus (Rhopalohelophorus) leechi McCorkle, 1965
Helophorus (Rhopalohelophorus) leontis Angus, 1985
Helophorus (Rhopalohelophorus) linearis (LeConte, 1855)⇒ Helophorus nubilus Fabricius, 1776
Helophorus (Rhopalohelophorus) linearoides Orchymont, 1945
Helophorus (Rhopalohelophorus) lineatus Say, 1823
Helophorus (Rhopalohelophorus) longitarsis Wollaston, 1864
Helophorus (Rhopalohelophorus) marginicollis Smetana, 1985
Helophorus (Rhopalohelophorus) minutus Fabricius, 1775
Helophorus (Rhopalohelophorus) montenegrinus (Kuwert, 1885)⇒ Helophorus brevipalpis Bedel, 1881
Helophorus (Rhopalohelophorus) nanus Sturm, 1836
Helophorus (Rhopalohelophorus) nevadensis Sharp, 1916
Helophorus (Rhopalohelophorus) nigricans Poppius, 1907
Helophorus (Rhopalohelophorus) nitiduloides Orchymont, 1945
Helophorus (Rhopalohelophorus) nitidulus LeConte, 1855Calabura
Helophorus (Rhopalohelophorus) nivalis (Thomson, 1853)⇒ Helophorus glacialis A. Villa & G..., 1883
Helophorus (Rhopalohelophorus) oblongus LeConte, 1850
Helophorus (Rhopalohelophorus) obscurus (Mulsant, 1844)⇒ Helophorus lecontei Knisch, 1924
Helophorus (Rhopalohelophorus) orchymonti Smetana, 1985
Helophorus (Rhopalohelophorus) oregonus McCorkle, 1965
Helophorus (Rhopalohelophorus) orientalis Motschulsky, 1860
Helophorus (Rhopalohelophorus) pallidipennis Mulsant & Wa..., 1852
Helophorus (Rhopalohelophorus) pallidus Gebler, 1830
Helophorus (Rhopalohelophorus) parajacutus Angus, 1970
Helophorus (Rhopalohelophorus) paraminutus Angus, 1984
Helophorus (Rhopalohelophorus) paraspelendidus Angus, 1970
Helophorus (Rhopalohelophorus) parasplendidus Angus, 1970
Helophorus parasplendidus parasplendidus Angus, 1970
Helophorus parasplendidus sperryi Smetana, 1985
Helophorus (Rhopalohelophorus) pitcheri Angus, 1970
Helophorus (Rhopalohelophorus) poppii Angus, 1970
Helophorus (Rhopalohelophorus) pumilio Erichson, 1837
Helophorus (Rhopalohelophorus) puncticollis Rey, 1885
Helophorus (Rhopalohelophorus) redtenbacheri Kuwert, 1885
Helophorus (Rhopalohelophorus) rinki Angus, 1985
Helophorus (Rhopalohelophorus) robertsi Smetana, 1985
Helophorus (Rhopalohelophorus) schuhi McCorkle, 1965
Helophorus (Rhopalohelophorus) seidlitzii Kuwert, 1885
Helophorus (Rhopalohelophorus) sempervarians Angus, 1970
Helophorus (Rhopalohelophorus) similes Kuwert, 1887
Helophorus (Rhopalohelophorus) similis Kuwert, 1887
Helophorus (Rhopalohelophorus) singularis Miller, 1881
Helophorus (Rhopalohelophorus) smetanai Hilsenhoff, 1995
Helophorus (Rhopalohelophorus) splendidus Sahlberg, 1880
Helophorus (Rhopalohelophorus) strigifrons Thomson, 1868
Helophorus (Rhopalohelophorus) subarcuatus Rey, 1885
Helophorus (Rhopalohelophorus) tumidus Motschulsky, 1860
Helophorus (Rhopalohelophorus) uvarovi Angus, 1884
Helophorus (Rhopalohelophorus) villosus Duftschmid, 1805
Helophorus (Rhopalohelophorus) wrootae Angus, 1988
Helophorus (Transithelophorus) crinitus Ganglbauer, 1901
Helophorus (Trichelophorus) alternans Géné, 1836
Helophorus (Trichohelophorus) alternans Géné, 1836
Helophorus abeillei Guillebeau, 1896
Helophorus aethiopicus Régimbart, 1907
Helophorus aethiopicus aethiopicus
Helophorus aethiopicus cooperi Orchymont, 1948
Helophorus aethiops J.Balfour-Br..., 1954
Helophorus algiricus Motschulsky, 1860
Helophorus altosichuanensis Angus, 2019
Helophorus angusi Hebauer, 1999
Helophorus antiquus Giebel, 1856
Helophorus apfelbecki (Kniñ, 1910)⇒ Helophorus nivalis Giraud, 1851
Helophorus aquila Angus, Fengl..., 2014
Helophorus armeniacus Ganglbauer, 1901
Helophorus atlantis Angus & Aouad, 2009
Helophorus banghaasi Kniñ, 1919
Helophorus beibienkoi Angus, 1984
Helophorus biltoni Angus, Mahdi..., 2005
Helophorus brodiei Giebel, 1856
Helophorus coopei J.V.Matthews, 1976
Helophorus costulatus Kuwert, 1887
Helophorus daedalus d´Orchymont, 1932
Helophorus difficilis Angus, 1988
Helophorus discretus d´Orchymont, 1925
Helophorus dixoni Angus, 1987
Helophorus dracomontanus Angus, 2017
Helophorus elizabethae Angus, 1987
Helophorus erzurumicus Ä°ncekara, M..., 2005
Helophorus exilis Heer, 1862
Helophorus faustianus Sharp, 1916
Helophorus fennicus (Paykull, 1798)⇒ Helophorus rufipes Bosc, 1791
Helophorus fosteri Angus, 1985
Helophorus gurjevae Angus, 1985
Helophorus guttulus Motschulsky, 1860
Helophorus helenae Angus, 1998
Helophorus hilaris Sharp, 1916
Helophorus imaensis d´Orchymont, 1926
Helophorus inexpectatus Angus, 1995
Helophorus inquinatus (Mannerheim, 1852)⇒ Helophorus auricollis Eschscholtz, 1822
Helophorus jaechi Angus, 1995
Helophorus karatavicus Angus & Lito..., 2018
Helophorus kayae Angus, 1985
Helophorus kazakhstanicus Angus, 1995
Helophorus kerimi Ganglbauer, 1901
Helophorus kozlovi Zaitzev, 1908
Helophorus kryzhanovskii Angus, 1985
Helophorus laferi Angus, 1995
Helophorus lamicola Zaitzev, 1908
Helophorus lewisi Angus, 1985
Helophorus limbatus Motschulsky, 1860
Helophorus longipennis Ganglbauer, 1901
Helophorus maculatus Motschulsky, 1860
Helophorus magnus Heer, 1862
Helophorus manchuricus Sharp, 1915
Helophorus matsumurai Nakane, 1963
Helophorus meighenensis J.V.Matthews, 1976
Helophorus mervensis Semenov, 1900
Helophorus mongoliensis Angus, 1970
Helophorus montanus d´Orchymont, 1926
Helophorus nivicola Sainte‑..., 1926
Helophorus oregonensis
Helophorus oscillator Sharp, 1915
Helophorus oxygonus Bedel, 1881
Helophorus paramontanus Angus, 1985
Helophorus peyerimhoffi d´Orchymont, 1926
Helophorus pici Guillebeau, 1893
Helophorus pictus Ganglbauer, 1901
Helophorus ponticus Angus, 1988
Helophorus praenanus Lomnicki, 1894
Helophorus regularis Sharp, 1882
Helophorus richterae Angus, 1985
Helophorus rigescens ♰ Scudder, 1890
Helophorus robustus Angus, Litov..., 2019
Helophorus salinus Angus & Lito..., 2018
Helophorus schoedli Angus, 2007
Helophorus sempevarians Angus, 1970
Helophorus ser Zaitzev, 1908
Helophorus shatrovskyi Angus, 1985
Helophorus sinoglacialis Angus, Rynde..., 2017
Helophorus subcarinatus Angus, 1985
Helophorus terminassianae Angus, 1984
Helophorus thauma Angus & Toledo, 2010
Helophorus theryi d´Orchymont, 1925
Helophorus timidus Motschulsky, 1860
Helophorus trisulcatus Guillebeau, 1896
Helophorus vereschaginae Angus, Litov..., 2019
Helophorus walkeri (Sharp, 1916)⇒ Helophorus obscurus Mulsant, 1844
Helophorus yammounensis Angus, 1988
Helophorus yangae Angus, FikáÄ..., 2016
Helophorus zagrosicus Angus, 1988
Hydrophilopsia longitarsalis ♰ A. G. Ponoma..., 1987
Laetopsia baissensis ♰ Ponomarenko, 1987
Laetopsia bontsaganica ♰ A. A. Prokin, 2009
Laetopsia hydraenoides ♰ A. Prokin, 2010
Laetopsia mongolica ♰ A. G. Ponoma..., 1987
Laetopsia shatrovskiyi ♰ A. Prokin, 2010
Zetemenos sexlineatus ♰ A. Bode, 1953


Links, sources and further information
Helophorinae at gbif.org