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TAXFAMGENSPESUB
Checklist for Family Hybosoridae Erichson, 1847 (Pill scarab beetles)
and region Eurasia (Continents)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
1859
65777
561707
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
1850
65672
561174
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
1595
64213
555201
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
829
47685
461834
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
209
18218
187489
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
167
16090
158751
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
18
1686
21104
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
18
1686
21104
Families
Genera
Species
 
Taxom
 
FamilyHybosoridae
Erichson, 1847
Pill scarab beetles, Scavenger scarab beetle
49
246
Genera
Species
 
Taxom
 


Genera for Eurasia
49 Genera with 246 Species
#ABCEFGHJLMOPS
Anaides Westwood, 1842
0SpeciesTaxom
Astaenomoechus Martínez & ..., 19362SpeciesTaxom
Astenomoechus1SpeciesTaxom
Besuchetostes Paulian, 197211SpeciesTaxom
Brenskea Reitter, 18911SpeciesTaxom
Celaenochrous Kuijten, 19841SpeciesTaxom
Ceratocanthus White, 18871SpeciesTaxom
Coilodes Westwood, 1843
0SpeciesTaxom
Coprologus Heer, 18471SpeciesTaxom
Cretanaides Nikolajev, 19961SpeciesTaxom
Cretanallachius Huang & al., 20151SpeciesTaxom
Cretohybosorus Nikolajev, 19992SpeciesTaxom
Cyphopisthes Gestro, 186010SpeciesTaxom
Ebbrittoniella Martínez, 18832SpeciesTaxom
Eusphaeropeltis Gestro, 198217SpeciesTaxom
Fortishybosorus Yan, Bai & Ren, 20131SpeciesTaxom
Germarostes Paulian, 18394SpeciesTaxom
Hybosorus MacLeay, 18456SpeciesTaxom
Hypseloderus Fairmaire, 20132SpeciesTaxom
Jurahybosorus Nikolajev, 20052SpeciesTaxom
Leptosorus Nikolajev, 20062SpeciesTaxom
Libanochrus Kirejtshuk, ..., 20111SpeciesTaxom
Liparochrus Erichson, 198012SpeciesTaxom
Macrophilharmostes Paulian, 19751SpeciesTaxom
Madrasostes Paulian, 187443SpeciesTaxom
Mesoceratocanthus Nikolajev & al., 20101SpeciesTaxom
Microphaeochroops Pic, 19855SpeciesTaxom
Microphaeolodes Kuijten, 19851SpeciesTaxom
Mimaphodius Nikolajev, 20071SpeciesTaxom
Mimocoelodes Pic, 19301SpeciesTaxom
Oxymorostes Ballerio, 20091SpeciesTaxom
Palaeopycnus Lu, Ballerio..., 20222SpeciesTaxom
Pantolasius Lansberge, 18873SpeciesTaxom
Parolichrus Paulian, 19251SpeciesTaxom
Paulianostes Ballerio, 20003SpeciesTaxom
Paulichrus1SpeciesTaxom
Perignamptus Harold, 18774SpeciesTaxom
Phaeochridius Lansberge, 19252SpeciesTaxom
Phaeochroops Candeze, 198126SpeciesTaxom
Phaeochrous Laporte de C..., 194348SpeciesTaxom
Phaeocroides Péringuey, 19633SpeciesTaxom
Protanaides Nikolajev, 20101SpeciesTaxom
Protohybosorus Nikolajev, 20104SpeciesTaxom
Pterorthochaetes Gestro, 201423SpeciesTaxom
Pulcherhybosorus Yan & al., 20121SpeciesTaxom
Seleucosorus Kuijten, 18612SpeciesTaxom
Silphodes Westwood, 18412SpeciesTaxom
Sinochaetodus Lu, Bai, Ren..., 20181SpeciesTaxom
Synarmostes Germar, 1843
0SpeciesTaxom


Species list for Eurasia
246 Species and 17 Subspecies
Astaenomoechus nevermanni Boucomont, 1936
Astaenomoechus setosus Boucomont, 1936
Astenomoechus solisi
Besuchetostes besucheti Paulian, 1972
Besuchetostes dubium Paulian, 1975
Besuchetostes hindu Paulian, 1975
Besuchetostes howdeni Paulian, 1979
Besuchetostes jaccoudi Paulian, 1977
Besuchetostes keralae Paulian, 1975
Besuchetostes kodaikanalense Paulian, 1975
Besuchetostes loebli Paulian, 1972
Besuchetostes mussardi Paulian, 1972
Besuchetostes peradeniyae Paulian, 1972
Besuchetostes taprobanae Paulian, 1972
Brenskea coronata Reitter, 1891
Celaenochrous sinensis Kuijten, 1984
Ceratocanthus striatulus Lansberge, 1887
Coprologus gracilis Heer, 1847
Cretanaides trogopterus Nikolajev, 1996
Cretanallachius magnificus Huang & al., 2015
Cretohybosorus buryaticus Nikolajev, 1999
Cretohybosorus striatulus Nikolajev, 1999
Cyphopisthes amphicyllis Sharp, 1875
Cyphopisthes crux Sharp, 1875
Cyphopisthes dohertyi Paulian, 1942
Cyphopisthes humeralis (Gestro, 1899)⇒ Cyphopisthes wallacei Pascoe, 1860
Cyphopisthes inexpectatus Paulian, 1981
Cyphopisthes krikkeni Paulian, 1980
Cyphopisthes luzonicus Paulian, 1978
Cyphopisthes minutus Paulian, 1978
Cyphopisthes szentivanyi Paulian, 1973
Cyphopisthes wallacei Pascoe, 1860
Ebbrittoniella gestroi Paulian, 1942
Ebbrittoniella ignita Westwood, 1883
Eusphaeropeltis aureola Gestro, 1899
Eusphaeropeltis aurora Lansberge, 1887
Eusphaeropeltis borneensis Paulian, 1978
Eusphaeropeltis boucomonti Paulian, 1978
Eusphaeropeltis celebicus Paulian, 1978
Eusphaeropeltis chenchaomingi Jiang, Balle..., 2020
Eusphaeropeltis corruscus Gestro, 1899
Eusphaeropeltis drescheri Paulian, 1942
Eusphaeropeltis iris Gestro, 1899
Eusphaeropeltis kedahensis Paulian, 1942
Eusphaeropeltis krikkeni Paulian, 1978
Eusphaeropeltis modiglianii Gestro, 1899
Eusphaeropeltis pulcher Lansberge, 1885
Eusphaeropeltis punctatissimus Lansberge, 1887
Eusphaeropeltis raapi Gestro, 1899
Eusphaeropeltis sabah Paulian, 1989
Eusphaeropeltis scheuerni Paulian, 1982
Fortishybosorus ericeusicus Z. Yan, 2013
Germarostes (Germarostes) aphodioides Illiger, 1800
Germarostes (Germarostes) globosus Say, 1835
Germarostes (Germarostes) rugatus Germar, 1843
Germarostes (Haroldostes) nitens Guérin-Ménev..., 1839
Hybosorus illigeri Reiche, 1853
Hybosorus laportei Westwood, 1845
Hybosorus lividus O. Heer, 1862
Hybosorus ocampoi Bai & Zhang, 2016
Hybosorus orientalis Westwood, 1845
Hybosorus roei (Westwood, 1845)⇒ Hybosorus illigeri Reiche, 1853
Hypseloderus denticollis Fairmaire, 1893
Hypseloderus yupae Masumoto, Ko..., 2013
Jurahybosorus kazakhstanicus Nikolajev, 2008
Jurahybosorus mongolicus G. V. Nikolajev, 2005
Leptosorus fortus D. Ren, 1995
Leptosorus zherikhini Nikolajev, 2006
Libanochrus calvus A. G. Kirejt..., 2011
Liparochrus (Liparochrus) crassicollis Arrow, 1925
Liparochrus (Liparochrus) dolium Heller, 1912
Liparochrus (Liparochrus) dux Arrow, 1909
Liparochrus (Liparochrus) fossulatus Westwood, 1852
Liparochrus (Liparochrus) krikkeni Paulian, 1980
Liparochrus (Liparochrus) papuus Lansberge, 1885
Liparochrus (Liparochrus) sculptilis Westwood, 1852
Liparochrus (Liparochrus) silphoides Harold, 1874
Liparochrus (Parolichrus) ingens (Felsch, 1909)⇒ Parolichrus ingens Felsche, 1909
Liparochrus (Parolichrus) lugubris (Arrow, 1925)⇒ Parolichrus lugubris Arrow, 1925
Liparochrus (Ranidichrus) irianae (Paulian, 1980)⇒ Paulichrus irianae Paulian, 1980
Liparochrus (Ranidichrus) modestus Paulian, 1980
Macrophilharmostes major Paulian, 1975
Madrasostes agostii Paulian, 1993
Madrasostes bartolozzii Ballerio, 2013
Madrasostes boucomonti Paulian, 1978
Madrasostes burckhardti Paulian, 1989
Madrasostes chungi Ballerio, 2018
Madrasostes clypeale Paulian, 1993
Madrasostes clypealis Paulian, 1993
Madrasostes deharvengi Gao, 2009
Madrasostes depressum Paulian, 1992
Madrasostes feae Gestro, 1899
Madrasostes franzi Paulian, 1978
Madrasostes granulatum Paulian, 1975
Madrasostes hashimi Ballerio & M..., 2010
Madrasostes inaequale Paulian, 1992
Madrasostes ivani Ballerio, 2018
Madrasostes kazumai Ochi, Johki ..., 1990
Madrasostes kazumai hisamatsui Ochi, 1990
Madrasostes kazumai kazumai hisamatsui Ochi, 1990
Madrasostes malayanum Paulian, 1979
Madrasostes malayanus Paulian, 1979
Madrasostes masumotoi Paulian, 1987
Madrasostes matangense Ballerio, 2018
Madrasostes mirificum Ballerio & M..., 2010
Madrasostes nigrum Paulian, 1975
Madrasostes orousseti Paulian, 1981
Madrasostes parcepunctatum Paulian, 1989
Madrasostes punctatum Paulian, 1989
Madrasostes rafflesi Paulian, 1979
Madrasostes renaudpauliani Ballerio, 2018
Madrasostes reticulatum Lansberge, 1887
Madrasostes sabah Paulian, 1989
Madrasostes sculpturatum Paulian, 1989
Madrasostes simplex Paulian, 1989
Madrasostes sumatranum Paulian, 1992
Madrasostes suzukii Ochi, Tsai &..., 2005
Madrasostes taiwanense Ochi, Tsai &..., 2005
Madrasostes tamil Paulian, 1975
Madrasostes thai Paulian, 1987
Madrasostes thoracicum Paulian, 1989Spinybacked orbweaver
Madrasostes tonkinense Paulian, 1945
Madrasostes tonkinensis Paulian, 1945
Madrasostes variolosum Harold, 1874
Madrasostes variolosus Harold, 1874
Mesoceratocanthus tuberculifrons G. V. Nikolajev, 2010
Microphaeochroops hirsutus Pic, 1930
Microphaeochroops laetus Arrow, 1942
Microphaeochroops nigrosetosus Kuijten, 1985
Microphaeochroops peninsularis Arrow, 1942
Microphaeochroops varius Kuijten, 1985
Microphaeolodes mulumontis Kuijten, 1985
Mimaphodius pusillus Nikolajev, 2007
Mimocoelodes minutus Pic, 1930
Oxymorostes riedeli Ballerio, 2009
Palaeopycnus circus Lu, Ballerio..., 2022
Palaeopycnus fushengii Lu, Ballerio..., 2022
Pantolasius bandaharae Kuijten, 1985
Pantolasius kerincii Keith, 2020
Pantolasius vethi Lansberge, 1887
Parolichrus lugubris Arrow, 1925
Paulianostes acromialis Pascoe, 1860
Paulianostes georyssoides Gestro, 1899
Paulianostes panggoling Ballerio, 2000
Paulichrus irianae Paulian, 1980
Perignamptus carinipennis Gestro, 1899
Perignamptus loriae Gestro, 1899
Perignamptus rossi Paulian, 1978
Perignamptus sharpi Harold, 1877
Phaeochridius derasus Harold, 1880
Phaeochridius uniformis Arrow, 1925
Phaeochroops acuticollis Arrow, 1907
Phaeochroops angulatus Benderitter, 1923
Phaeochroops colopacilis Kuijten, 1981
Phaeochroops curtulus Schmidt, 1912
Phaeochroops freenae Kuijten, 1981
Phaeochroops gigas Arrow, 1907
Phaeochroops gilleti Benderitter, 1923
Phaeochroops hisaeae Nishikawa, 1989
Phaeochroops indicus Arrow, 1907
Phaeochroops lakhonicus Kuijten, 1981
Phaeochroops lansbergei Candeze, 1876
Phaeochroops laotianus Paulian, 1945
Phaeochroops longisetosus Kuijten, 1981
Phaeochroops maruyamai Nishikawa, 1989
Phaeochroops masumotoi Nishikawa, 1996
Phaeochroops meghalayicus Keith, 2001
Phaeochroops myanmarensis Keith, 2000
Phaeochroops ninbin Kuijten, 1981
Phaeochroops opacicollis Arrow, 1909
Phaeochroops peninsularis Arrow, 1909
Phaeochroops rattus Arrow, 1909
Phaeochroops seres Kuijten, 1981
Phaeochroops silphoides Fairmaire, 1898
Phaeochroops taiwanus Nomura, 1973
Phaeochroops vulpecula Arrow, 1909
Phaeochroops vulturius Kuijten, 1981
Phaeochrous alternatus (Fairmaire, 1879)⇒ Phaeochrous emarginatus Laporte, 1840
Phaeochrous asiaticus (Lewis, 1896)⇒ Phaeochrous emarginatus Laporte, 1840
Phaeochrous beccarii Harold, 1871
Phaeochrous borealis Kuijten, 1984
Phaeochrous celebensis (Pic, 1928)⇒ Phaeochrous emarginatus Laporte, 1840
Phaeochrous compactus Kuijten, 1978
Phaeochrous davaonis Kuijten, 1981
Phaeochrous dispar lujai Schouteden, 1918
Phaeochrous dissimilis Arrow, 1909
Phaeochrous dissimilis dissimilis
Phaeochrous dissimilis vietnamicola Kuijten, 1978
Phaeochrous diversipes Pic, 1928
Phaeochrous dubius Westwood, 1846
Phaeochrous elevatus Kuijten, 1978
Phaeochrous emarginatus Laporte, 1840
Phaeochrous emarginatus benderitteri Pic, 1928
Phaeochrous emarginatus buruensis Kuijten, 1978
Phaeochrous emarginatus davidis Fairmaire, 1886
Phaeochrous emarginatus emarginatus suturalis La..., 1885
Phaeochrous emarginatus suturalis Lansberge, 1885
Phaeochrous emarginatus thilliezi Keith, 1999
Phaeochrous enigmaticus Kuijten, 1978
Phaeochrous enigmaticus enigmaticus
Phaeochrous enigmaticus ovchinnikovi Nikolajev, 2008
Phaeochrous gracilis (Petrovitz, 1975)⇒ Phaeochrous emarginatus Laporte, 1840
Phaeochrous hainanensis Zhang-Youwei, 1990
Phaeochrous indicus Westwood, 1846
Phaeochrous intermedius Pic, 1928
Phaeochrous intermedius intermedius occidentalis..., 1978
Phaeochrous intermedius occidentalis Kuijten, 1978
Phaeochrous lobatus Kuijten, 1978
Phaeochrous madagascariensis Westwood, 1846
Phaeochrous madrassicus Kuijten, 1978
Phaeochrous orbachi Keith & Ball..., 2019
Phaeochrous pallidus (Arrow, 1909)⇒ Phaeochrous emarginatus Laporte, 1840
Phaeochrous perroudi (Pic, 1943)⇒ Phaeochrous emarginatus Laporte, 1840
Phaeochrous philippinensis Westwood, 1846
Phaeochrous pletus Kuijten, 1978
Phaeochrous portuum Kuijten, 1978
Phaeochrous pseudintermedius Kuijten, 1978
Phaeochrous rudis Kuijten, 1984
Phaeochrous ruficollis Fairmaire, 1893
Phaeochrous rufus (Pic, 1928)⇒ Phaeochrous madagascariensis Westwood, 1846
Phaeochrous separabilis Zhang-Youwei, 1990
Phaeochrous sulawesi Kuijten, 1978
Phaeochrous tertiarium J. V. Deichm..., 1881
Phaeochrous tokaraensis Nomura, 1961
Phaeochrous tonkineus Pic, 1943
Phaeocroides mapellii (Petrovitz, 1972)⇒ Phaeocroides nanus Arrow, 1942
Phaeocroides nanus Arrow, 1942
Phaeocroides orientalis Petrovitz, 1963
Protanaides sibiricus G. V. Nikolajev, 2010
Protohybosorus grandissimus G. V. Nikolajev, 2010
Protohybosorus karatavicus G. V. Nikolajev, 2010
Protohybosorus mesasiaticus Nikolajev, 2010
Protohybosorus mesoasiaticus G. V. Nikolajev, 2010
Pterorthochaetes andamanus Paulian, 1937
Pterorthochaetes armatus Paulian, 1945
Pterorthochaetes brevis Sharp, 1875
Pterorthochaetes brevisetosus Gestro, 1899
Pterorthochaetes coomani Paulian, 1945
Pterorthochaetes cribricollis Gestro, 1899
Pterorthochaetes dembickyi Ballerio, 2014
Pterorthochaetes gestroi Harold, 1874
Pterorthochaetes gestroi gestroi longisetosus..., 1899
Pterorthochaetes gestroi longisetosus Gestro, 1899
Pterorthochaetes haroldi Sharp, 1875
Pterorthochaetes hirtus Gestro, 1899
Pterorthochaetes incertus Gestro, 1899
Pterorthochaetes insularis Gestro, 1899
Pterorthochaetes latus Sharp, 1875
Pterorthochaetes laxepunctatus Paulian, 1973
Pterorthochaetes montanus Ballerio, 1999
Pterorthochaetes picinus Sharp, 1875
Pterorthochaetes puncticollis Sharp, 1875
Pterorthochaetes septemtrionalis Ballerio, 1999
Pterorthochaetes sulawesii Paulian, 1987
Pterorthochaetes tsurui Ballerio & M..., 2010
Pterorthochaetes yunnanensis Ballerio, 2014
Pulcherhybosorus tridentatus Z. Yan, 2012
Seleucosorus punctatissimus Reiche, 1861
Seleucosorus punctatissumus Reiche, 1861
Silphodes hirtipes (MacLeay, 1864)⇒ Phaeochrous emarginatus Laporte, 1840
Silphodes sumatrensis (Westwood, 1841)⇒ Phaeochrous emarginatus Laporte, 1840
Sinochaetodus tridentatus Lu, Bai, Ren..., 2018

Subnodes of Hybosoridae with occurrences in Eurasia
SubfamilyAnaidinae
Nikolajev, 1996
3
2
Genera
Species

Taxom

SubfamilyCeratocanthinae
A. Martínez, 1968
Pill Scarab Beetles
15
119
Genera
Species

Taxom

SubfamilyHybosorinae
Erichson, 1847
Scavenger Scarab Beetles
17
94
Genera
Species

Taxom

SubfamilyLiparochrinae
Ocampo, 2006
1
12
Genus
Species

Taxom

SubfamilyMimaphodiinae
Nikolajev, 2007
1
1
Genus
Species

Taxom

SubfamilyPachyplectrinae
Ocampo, 2006
Thick Scavenger Scarab Beetles
1
1
Genus
Species

Taxom



Links, sources and further information
Hybosoridae at gbif.org
Hybosoridae at irmng.org
Hybosoridae at itis.gov