Taxonomy of Genus Hydrophilus Müller, 1764
Giant Water Scavenger Beetles
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 4179
190570
2444470 | Families
Genera
Species |
Subclass
| Pterygota
Lang, 1888
Winged insects | 3816
189502
2439455 | Families
Genera
Species |
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 3399
186457
2415487 | Families
Genera
Species |
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 1891
142849
2048615 | Families
Genera
Species |
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 592
62779
813516 | Families
Genera
Species |
Suborder
| Polyphaga
Emery, 1886
Polyphags | 274
54416
687671 | Families
Genera
Species |
Infraorder
| Staphyliniformia
Lameere, 1900
| 22
8512
135171 | Families
Genera
Species |
Superfamily
| Hydrophiloidea
Latreille, 1802
Histerids, Hydriphiloids, Hydrophilids | 8
484
7136 | Families
Genera
Species | | Family | Hydrophilidae
Latreille, 1802
Silver Water Beetles, Water beetles | 475
7090 | Genera
Species | | Subfamily | Hydrophilinae
Latreille, 1802
| 82
3621 | Genera
Species | | Tribe | Hydrophilini
Latreille, 1802
| 31
1747 | Genera
Species | | Subtribe | Hydrophilina
Latreille, 1802
| 5
577 | Genera
Species | | Genus | Hydrophilus
Müller, 1764
Giant Water Scavenger Beetles | 291 | Species |
Subnodes of Hydrophilus Müller, 1764 | Subgenus | Hydrophilus (Dibolocelus)
Bedel, 1891
| 2 | Species | | Subgenus | Hydrophilus (Hydrophilus)
Geoffroy, 1762
Giant Water Scavenger Beetles, Serrate ptychomitrium moss | 18 | Species | | Subgenus | Hydrophilus (Temnopterus)
Solier, 1834
| 1 | Species | | Species | Hydrophilus abbreviatus Fabricius, 1801
| | | | Species | Hydrophilus aberrans Kuwert, 1893
| | | | Species | Hydrophilus acherontis Heer, 1852
| | | | Species | Hydrophilus acuminatus Motschulsky, 1853
Dark Diving Beetle | | | | Species | Hydrophilus aegyptiacus Peyron, 1856
| | | | Species | Hydrophilus aeneus De Geer, 1774 ⇒ Helophorus (Helophorus) aequalis
| | | | Species | Hydrophilus affinis Marsham, 1802 ⇒ Helophorus (Helophorus) griseus
| | | | Species | Hydrophilus africanus Kuwert, 1893
| | | | Species | Hydrophilus agilis Randall, 1838
| | | | Species | Hydrophilus analis Paykull, 1798
| | | | Species | Hydrophilus angolensis Erichson, 1843
| | | | Species | Hydrophilus angustatus Villa & Villa, 1838 ⇒ Hydrobius fuscipes
| | | | Species | Hydrophilus angustior Rey, 1885 ⇒ Hydrophilus (Hydrophilus) piceus
| | | | Species | Hydrophilus antiquus ♰ E. Oustalet, 1874
| | | | Species | Hydrophilus armatus Laporte, 1840
| | | | Species | Hydrophilus ater Gmelin, 1790
Cinnamon fern | | | | Species | Hydrophilus atomus Duftschmid, 1805
| | | | Species | Hydrophilus attenuatus Fabricius, 1801
| | | | Species | Hydrophilus auritus F.V.Melsheimer, 1806
| | | | Species | Hydrophilus australis Montrouzier, 1860
| | | | Species | Hydrophilus barbatus Letzner, 1884 ⇒ Hydrophilus (Hydrophilus) aterrimus
| | | | Species | Hydrophilus bedeli Régimbart, 1902
| | | | Species | Hydrophilus bicolor Gyllenhal, 1808 ⇒ Enochrus (Enochrus) melanocephalus
| | | | Species | Hydrophilus bipunctatus Fabricius, 1775
| | | | Species | Hydrophilus bipustulatus Marsham, 1802
| | | | Species | Hydrophilus birmanicus Régimbart, 1902
| | | | Species | Hydrophilus blandus Dejean, 1833
| | | | Species | Hydrophilus bonariensis
| | | | Species | Hydrophilus brasiliensis Laporte de Castelnau, 1840
| | | | Species | Hydrophilus braunii Heer, 1847
| | | | Species | Hydrophilus brodiei Giebel, 1856
| | | | Species | Hydrophilus brunnipalpis Kuwert, 1893
| | | | Species | Hydrophilus caffer Boheman, 1851
| | | | Species | Hydrophilus californicus LeConte & J.L., 1855
| | | | Species | Hydrophilus cambodiensis Régimbart, 1903
| | | | Species | Hydrophilus caraboides Linnaeus, 1758 ⇒ Hydrochara caraboides
Lesser silver water beetle | | | | Species | Hydrophilus carbonarius Heer, 1847
| | | | Species | Hydrophilus carinata Jander, 1927 ⇒ Hydrochara caraboides
| | | | Species | Hydrophilus carinatus Jander, 1927
| | | | Species | Hydrophilus caschmirensis Redtenbacher, 1844
Large Scavenging Water Beetle | | | | Species | Hydrophilus cashmirensis Redtenbacher, 1892
| | | | Species | Hydrophilus castus Say, 1835
| | | | Species | Hydrophilus cavicrus Kuwert, 1893
| | | | Species | Hydrophilus cavisternum Bedel, 1891
| | | | Species | Hydrophilus chalcaspis Eschscholtz, 1818 ⇒ Berosus (Berosus) luridus
| | | | Species | Hydrophilus chalybeatus Curtis, 1844
| | | | Species | Hydrophilus chalybeus Dejean, 1833
| | | | Species | Hydrophilus chrysomelinus Herbst, 1797 ⇒ Helochares (Helochares) lividus
| | | | Species | Hydrophilus cicindeloides Marsham, 1802
| | | | Species | Hydrophilus cinctus F.V.Melsheimer, 1806
| | | | Species | Hydrophilus cinereus Marsham, 1802 ⇒ Helophorus (Empleurus) rufipes
| | | | Species | Hydrophilus coccinelloides Rossi, 1790
| | | | Species | Hydrophilus cognatus Sharp, 1873
| | | | Species | Hydrophilus collaris Fabricius, 1775
| 1 | Subspecies | | Species | Hydrophilus columbinus Kuwert, 1893
| | | | Species | Hydrophilus convexus Laporte, 1840
| | | | Species | Hydrophilus cordiger Herbst, 1784 ⇒ Cercyon (Cercyon) unipunctatus
| | | | Species | Hydrophilus costulata A.P.Semenov, 1900
| | | | Species | Hydrophilus cultrifer Régimbart, 1903
| | | | Species | Hydrophilus dauricus Mannerheim, 1852
| | | | Species | Hydrophilus deperditus Germar, 1839
| | | | Species | Hydrophilus deplanatus Orchymont, 1911
| | | | Species | Hydrophilus dichrous Krynicki, 1832 ⇒ Enochrus (Lumetus) quadripunctatus
| | | | Species | Hydrophilus dorsalis Brullé, 1836
| | | | Species | Hydrophilus duvali M.Hansen, 1999
| | | | Species | Hydrophilus ellipticus LeConte & J.L., 1855
| | | | Species | Hydrophilus elongatus Régimbart, 1902
| | | | Species | Hydrophilus ensifer Brullé, 1837 ⇒ Hydrophilus ater
| 2 | Subspecies | | Species | Hydrophilus erythrocephalus Fabricius, 1792 ⇒ Helochares (Helochares) obscurus
| | | | Species | Hydrophilus exstriatus Say, 1835
| | | | Species | Hydrophilus extricatus Scudder, 1900
| | | | Species | Hydrophilus femoratus É.Blanchard, 1841
| | | | Species | Hydrophilus fimbriatus F.V.Melsheimer, 1806
| | | | Species | Hydrophilus flavicornis Laporte de Castelnau, 1840
| | | | Species | Hydrophilus flavipalpis Boheman, 1851
| | | | Species | Hydrophilus flavipes Steven, 1808 ⇒ Hydrochara flavipes
| | | | Species | Hydrophilus flaviventris Ochs, 1923 ⇒ Hydrochara caraboides
| | | | Species | Hydrophilus foveolata Haworth, 1807 ⇒ Anacaena limbata
| | | | Species | Hydrophilus foveolatus Régimbart, 1901
| | | | Species | Hydrophilus fraternus C.Heyden, 1859
| | | | Species | Hydrophilus fulvofemoratus Fairmaire, 1869
| | | | Species | Hydrophilus fulvus Fourcroy, 1785 ⇒ Helochares (Helochares) lividus
| | | | Species | Hydrophilus fuscipes O.F. Müller, 1776 ⇒ Hydrobius fuscipes
| | | | Species | Hydrophilus fuscus De Geer, 1774 ⇒ Berosus (Berosus) luridus
| | | | Species | Hydrophilus ganssuensis A. Semenov, 1889
| | | | Species | Hydrophilus gaudini ♰ O. Heer, 1862
| | | | Species | Hydrophilus gayndahensis W. J. MacLeay, 1871 ⇒ Hydrophilus (Hydrophilus) picicornis
| | | | Species | Hydrophilus geticus Protescu, 1938
| | | | Species | Hydrophilus gibbosus Régimbart, 1902
| | | | Species | Hydrophilus gibbus Illiger, 1801
| | | | Species | Hydrophilus giganteus Heer, 1862
| | | | Species | Hydrophilus glaber J.L.LeConte, 1855
| | | | Species | Hydrophilus globosus F.V.Melsheimer, 1806
| | | | Species | Hydrophilus globulus Paykull, 1798
| | | | Species | Hydrophilus grisescens Gyllenhal, 1827
| | | | Species | Hydrophilus griseus Fabricius, 1787 ⇒ Helochares (Helochares) lividus
| | | | Species | Hydrophilus guarani Bachmann, 1966
| | | | Species | Hydrophilus hackeri Orchymont, 1937
| | | | Species | Hydrophilus haemorrhoidalis Fabricius, 1792
| | | | Species | Hydrophilus harpe Short & McIntosh, 2015
| | | | Species | Hydrophilus hastatus Herbst, 1779
| | | | Species | Hydrophilus hemisphaericus Dejean, 1821
| | | | Species | Hydrophilus horni Régimbart, 1902
| | | | Species | Hydrophilus impressus Marsham, 1802
| | | | Species | Hydrophilus indicus Bedel, 1891
| | | | Species | Hydrophilus inermis Küster, 1852 ⇒ Hydrophilus (Hydrophilus) pistaceus
| | | | Species | Hydrophilus infrequens Watts, 1988
| | | | Species | Hydrophilus intermedius Jacquelin du Val, 1857
| | | | Species | Hydrophilus iricolor Régimbart, 1901
| | | | Species | Hydrophilus irinus Brullé, 1837
| | | | Species | Hydrophilus japonicus Sharp, 1873
| | | | Species | Hydrophilus knorrii Heer, 1847
| | | | Species | Hydrophilus kuntzeni ♰ F. E. Zeuner, 1931
| | | | Species | Hydrophilus labiatus F.V.Melsheimer, 1806
| | | | Species | Hydrophilus laevis Sturm, 1826
| | | | Species | Hydrophilus lateralis Fabricius, 1775
| 3 | Subspecies | | Species | Hydrophilus laticollis Régimbart, 1907
| | | | Species | Hydrophilus lepidus Brullé, 1836
| | | | Species | Hydrophilus limbalis LeConte & J.L., 1855
| | | | Species | Hydrophilus limbatus Brullé, 1836
| | | | Species | Hydrophilus lineatus Dejean, 1821
| | | | Species | Hydrophilus lividus Herbst, 1797 ⇒ Helochares (Helochares) lividus
| | | | Species | Hydrophilus loanei Camerano, 1907
| | | | Species | Hydrophilus longipalpis Marsham, 1802
| | | | Species | Hydrophilus longus Bruch, 1915
| | | | Species | Hydrophilus loriai Régimbart, 1902
| | | | Species | Hydrophilus lucidus Dejean, 1833
| | | | Species | Hydrophilus lugubris Motschulsky, 1845 ⇒ Hydrophilus (Hydrophilus) triangularis
| | | | Species | Hydrophilus luridus Gmelin, 1790
| | | | Species | Hydrophilus lutosus Marsham, 1802
| | | | Species | Hydrophilus magnificus ♰ L. Piton, 1939
| | | | Species | Hydrophilus marginatus Duftschmid, 1805 ⇒ Cymbiodyta marginella
| | | | Species | Hydrophilus marginellus Fabricius, 1792 ⇒ Cercyon (Paracercyon) analis
| | | | Species | Hydrophilus margipallens Marsham, 1802 ⇒ Cymbiodyta marginella
| | | | Species | Hydrophilus masculinus Régimbart, 1901
| | | | Species | Hydrophilus medius Brullé, 1837
| | | | Species | Hydrophilus melanocephalus Fabricius, 1801 ⇒ Enochrus (Lumetus) fuscipennis
| | | | Species | Hydrophilus mesopotamiae Kniñ, 1914
| | | | Species | Hydrophilus minimus Dejean, 1821
| | | | Species | Hydrophilus minutissimus Germar, 1823
| | | | Species | Hydrophilus minutus Gmelin, 1790
| | | | Species | Hydrophilus miserandus Von Heyden, 1859
| | | | Species | Hydrophilus mixtus LeConte & J.L., 1855
| | | | Species | Hydrophilus mollis Marsham, 1802
| | | | Species | Hydrophilus morio Sturm, 1835 ⇒ Hydrophilus (Hydrophilus) aterrimus
| | | | Species | Hydrophilus mundus Boheman, 1851
| | | | Species | Hydrophilus nebulosus Say, 1824
| | | | Species | Hydrophilus neptunus von Heyden & von Heyden, 1866
| | | | Species | Hydrophilus niger G. Eichler, 1876 ⇒ Limnoxenus niger
| | | | Species | Hydrophilus nigricans Zetterstedt, 1838 ⇒ Enochrus (Lumetus) ochropterus
| | | | Species | Hydrophilus nigriceps Fabricius, 1801
| | | | Species | Hydrophilus nigricornis De Geer, 1774 ⇒ Hydrochara caraboides
| | | | Species | Hydrophilus nimbatus Say, 1823
| | | | Species | Hydrophilus nitidulus Brullé, 1837
| | | | Species | Hydrophilus nitidus Marsham, 1802
| | | | Species | Hydrophilus noachicus Heer, 1847
| | | | Species | Hydrophilus novaeguineae Watts, 1988
| | | | Species | Hydrophilus nucleoensis Arce-Pérez & Arriaga-Varela, 2021
| | | | Species | Hydrophilus oberthueri Régimbart, 1902
| | | | Species | Hydrophilus oblongus Olivier, 1792 ⇒ Limnoxenus niger
| | | | Species | Hydrophilus obscurus Fabricius, 1792 ⇒ Helochares (Helochares) obscurus
| | | | Species | Hydrophilus obtusatus Say, 1823 ⇒ Hydrochara obtusata
| | | | Species | Hydrophilus occulatus
| | | | Species | Hydrophilus occultus Orchymont, 1933 ⇒ Hydrochara occulta
| | | | Species | Hydrophilus ochripes Curtis, 1844
| | | | Species | Hydrophilus ochropterus Marsham, 1802
| | | | Species | Hydrophilus olivaceus Fabricius, 1781
| | | | Species | Hydrophilus orbiculare Fabricius, 1775
| | | | Species | Hydrophilus orbicularis Fabricius, 1775
| | | | Species | Hydrophilus orientalis Kuwert, 1893
| | | | Species | Hydrophilus ovalis Ziegler, 1844
| | | | Species | Hydrophilus pallidipalpis W. S. MacLeay, 1825
| | | | Species | Hydrophilus pallidus Rossi, 1792 ⇒ Helochares (Helochares) lividus
| | | | Species | Hydrophilus pallipes É.Blanchard, 1841
| | | | Species | Hydrophilus palpalis Brullé, 1838
| 2 | Subspecies | | Species | Hydrophilus parvulus Herbst, 1797
| | | | Species | Hydrophilus paulinierii Guérin‑Méneville, 1842
| | | | Species | Hydrophilus pergrandis ♰ J. F. Zhang, 1989
| | | | Species | Hydrophilus picaeus Fourcroy & Geoffroy, 1785
| | | | Species | Hydrophilus picinus Marsham, 1802
| | | | Species | Hydrophilus picipes Fabricius, 1787 ⇒ Catops picipes
| | | | Species | Hydrophilus piesbergeni Reitter, 1906
| | | | Species | Hydrophilus pilula O.F. Müller, 1776 ⇒ Coelostoma (Coelostoma) orbiculare
| | | | Species | Hydrophilus plicifer Bedel, 1891
| | | | Species | Hydrophilus politus Dejean, 1821
| | | | Species | Hydrophilus pollens Sharp, 1887
| | | | Species | Hydrophilus praedator Dejean, 1833
| | | | Species | Hydrophilus profanifuga Semenow, 1900 ⇒ Hydrochara flavipes
| | | | Species | Hydrophilus pseudovatus Arce-Pérez & Arriaga-Varela, 2021
| | | | Species | Hydrophilus punctatus Fourcroy & Geoffroy, 1785
| | | | Species | Hydrophilus punctiformis Sturm, 1826
| | | | Species | Hydrophilus purpurascens Régimbart, 1901
| | | | Species | Hydrophilus pusillus Müller, 1776
| | | | Species | Hydrophilus pygmaeus Fabricius, 1792
| 1 | Subspecies | | Species | Hydrophilus quadripunctatus Herbst, 1797
| | | | Species | Hydrophilus quadristriatus Horn, 1871
| | | | Species | Hydrophilus regimbarti Zaitzev, 1908
| | | | Species | Hydrophilus rehmanni Heer, 1847
| | | | Species | Hydrophilus resplendens Kuwert, 1893
| | | | Species | Hydrophilus rottensis ♰ G. Statz, 1939
| | | | Species | Hydrophilus rotundus Say, 1825
| | | | Species | Hydrophilus roubali Vavra, 1934 ⇒ Hydrochara flavipes
| | | | Species | Hydrophilus ruficornis Boisduval, 1835 ⇒ Hydrophilus (Hydrophilus) picicornis
| | | | Species | Hydrophilus rufipes Fabricius, 1792
| | | | Species | Hydrophilus rufocinctus Bedel, 1891
| | | | Species | Hydrophilus rufomarginatus M.Hansen, 1999
| | | | Species | Hydrophilus ruminianus Heer, 1865
| | | | Species | Hydrophilus sabellifer Fairmaire, 1879
| | | | Species | Hydrophilus sabelliferus Fairmaire, 1879 ⇒ Hydrophilus (Hydrophilus) picicornis
| | | | Species | Hydrophilus sartus A.P.Semenov, 1900
| | | | Species | Hydrophilus scissipalpis Blackburn, 1901
| | | | Species | Hydrophilus scrobiculata Panzer, 1799 ⇒ Hydrochara caraboides
| | | | Species | Hydrophilus scrobiculatus Panzer, 1799
| | | | Species | Hydrophilus scutellaris Dejean, 1836
| | | | Species | Hydrophilus semenovi Zaitzev, 1908
| | | | Species | Hydrophilus semicylindricus Eschscholtz, 1822
| | | | Species | Hydrophilus seminulum Paykull, 1798 ⇒ Chaetarthria seminulum
| | | | Species | Hydrophilus senegalensis Percheron, 1835
| | | | Species | Hydrophilus signaticollis T.Charpentier, 1825
| | | | Species | Hydrophilus similis Orchymont, 1919
| | | | Species | Hydrophilus simulator Bedel, 1891
| | | | Species | Hydrophilus smaragdina Bach, 1856 ⇒ Hydrochara caraboides
| | | | Species | Hydrophilus sordidus Marsham, 1802
| | | | Species | Hydrophilus spectabilis ♰ O. Heer, 1862
| | | | Species | Hydrophilus spinicollis Eschscholtz, 1822
| | | | Species | Hydrophilus spinipennis Gory, 1834
| | | | Species | Hydrophilus spinosus Steven, 1808
| | | | Species | Hydrophilus stagnalis Marsham, 1802 ⇒ Helophorus (Helophorus) grandis
| | | | Species | Hydrophilus stenopterus O. Heer, 1862
| | | | Species | Hydrophilus sternitalis Reitter, 1906
| | | | Species | Hydrophilus striatulus Fabricius, 1801
| | | | Species | Hydrophilus striatus Turton, 1802
| | | | Species | Hydrophilus striolatus LeConte & J.L., 1855
| | | | Species | Hydrophilus subaenea Motschulsky, 1845 ⇒ Hydrochara caraboides
| | | | Species | Hydrophilus subaeneus Motschulsky, 1845
| | | | Species | Hydrophilus subcupreus Say, 1825
| | | | Species | Hydrophilus sublaevis LeConte & J.L., 1855
| | | | Species | Hydrophilus subrotundus Fabricius, 1792
| | | | Species | Hydrophilus substriata Sturm, 1843 ⇒ Hydrochara caraboides
| | | | Species | Hydrophilus substriatus Sturm, 1843
| | | | Species | Hydrophilus subvariolosa A.P.Semenov, 1900
| | | | Species | Hydrophilus sumatrae Kuwert, 1893
| | | | Species | Hydrophilus temnopteroides Orchymont, 1913
| | | | Species | Hydrophilus testaceus Fabricius, 1801
| | | | Species | Hydrophilus torquatus Marsham, 1802
| | | | Species | Hydrophilus tristis Motschulsky, 1859
| | | | Species | Hydrophilus truncatellus Thunberg, 1794
| | | | Species | Hydrophilus turkestanus Kuwert, 1893
| | | | Species | Hydrophilus undatus Fabricius, 1792
| | | | Species | Hydrophilus unguicularis Régimbart, 1902
| | | | Species | Hydrophilus variegatus Herbst, 1797 ⇒ Helochares (Helochares) obscurus
| | | | Species | Hydrophilus variolosus Dejean, 1833
| | | | Species | Hydrophilus verrucosus Marsham, 1802
| | | | Species | Hydrophilus vexatorius ♰ O. Heer, 1847
| | | | Species | Hydrophilus violaceonitens Jacquelin du Val, 1857
| | | | Species | Hydrophilus viridicollis L. Redtenbacher, 1844 ⇒ Hydrophilus (Hydrophilus) piceus
| | | | Species | Hydrophilus viridis Watts, 1988
| | | | Species | Hydrophilus vitalisi Orchymont, 1919
| | | | Species | Hydrophilus wehnckei Régimbart, 1902
| | | | Species | Hydrophilus wencki Paulino d'Oliveira, 1880
| | | | Species | Hydrophilus westwoodi Giebel, 1856
| | |
Synonyms Native, vernacular names and spellings of Hydrophilus | English | Giant Water Scavenger Beetles | | Dutch | Spinnende watertorren |
Distribution of Hydrophilus Müller, 1764 Albania, Algeria, Angola, Antigua and Barbuda, Argentina, Armenia, Australia, Austria, Azerbaijan, Bahamas, Barbados, Belarus, Belgium, Belize, Benin, Bhutan, Bolivia, Bosnia and Herzegovina, Botswana, Brazil, Bulgaria, Burundi, Cambodia, Cameroon, Canada, Cape Verde, Central African Republic, Chile, China, Colombia, Costa Rica, Cuba, Czech Republic, Democratic Republic of the Congo, Denmark, Dominican Republic, Ecuador, Egypt, Estonia, Ethiopia, Fiji, Finland, France, Gambia, Georgia, Germany, Ghana, Greece, Guatemala, Guinea-Bissau, Guyana, Haiti, Honduras, Hungary, India, Indonesia, Iran, Iraq, Israel, Italy, Japan, Kazakhstan, Kenya, Laos, Latvia, Lesotho, Liberia, Lithuania, Luxembourg, Madagascar, Malaysia, Mali, Mauritania, Mauritius, Mexico, Mongolia, Morocco, Mozambique, Myanmar, Namibia, Nepal, Netherlands, Nicaragua, Niger, Nigeria, North Korea, Norway, Oman, Pakistan, Panama, Papua New Guinea, Paraguay, Peru, Philippines, Poland, Portugal, Republic South Africa, Romania, Russia, Saint Lucia, Samoa, Saudi Arabia, Senegal, Serbia, Sierra Leone, Slovakia, Somalia, South Korea, Spain, Sri Lanka, Sudan, Suriname, Sweden, Switzerland, Syria, Taiwan, Tanzania, Thailand, Togo, Trinidad and Tobago, Tunisia, Turkey, Turkmenistan, Uganda, Ukraine, United Kingdom, Uruguay, USA, Uzbekistan, Vanuatu, Venezuela, Vietnam, Yemen, Zambia, Zimbabwe
Found in fossile formations Florissant Fossil Beds, Green River Formation, Rott Formation
Links, sources and further information
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