Taxonomy of Genus Malthinus Latreille, 1806
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 4179
190570
2444470 | Families
Genera
Species |
Subclass
| Pterygota
Lang, 1888
Winged insects | 3816
189502
2439455 | Families
Genera
Species |
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 3399
186457
2415487 | Families
Genera
Species |
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 1891
142849
2048615 | Families
Genera
Species |
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 592
62779
813516 | Families
Genera
Species |
Suborder
| Polyphaga
Emery, 1886
Polyphags | 274
54416
687671 | Families
Genera
Species |
Infraorder
| Elateriformia
Crowson, 1960
| 52
4609
79150 | Families
Genera
Species |
Superfamily
| Elateroidea
Leach, 1815
Click Beetles, Firefly Beetles, Skipjack Beetles | 23
2729
37099 | Families
Genera
Species | | Family | Cantharidae
Imhoff, 1856
Cantharids, Leatherwings, Sailor beetles, Soldier beetle | 294
7560 | Genera
Species | | Subfamily | Malthininae
Kiesenwetter, 1852
| 26
1650 | Genera
Species | | Tribe | Malthinini
Kiesenwetter, 1852
| 8
1512 | Genera
Species | | Genus | Malthinus
Latreille, 1806
| 531 | Species |
Subnodes of Malthinus Latreille, 1806 | Subgenus | Malthinus (Indomalthinus)
Brancucci in Wittmer & Brancucci, 1978
| 8 | Species | | Subgenus | Malthinus (Malthinus)
Latreille, 1806
| 205 | Species | | Species | Malthinus abdelkader Marseul, 1868
| | | | Species | Malthinus aegyptiacus Ferrante, 1908
| | | | Species | Malthinus alexanderi Fender, 1966 ⇒ Caccodes alexanderi
| | | | Species | Malthinus algericus Pic, 1900
| | | | Species | Malthinus angusticollis Pic, 1898
| | | | Species | Malthinus angustipennis Pic, 1908
| | | | Species | Malthinus armipes Kiesenwetter, 1871
| | | | Species | Malthinus atlasicus Wittmer, 1940
| | | | Species | Malthinus atriceps Pic, 1900
| | | | Species | Malthinus atripennis LeConte, 1881 ⇒ Athemellus atripennis
| | | | Species | Malthinus axillaris Kiesenwetter, 1852
| | | | Species | Malthinus axillaroides Wittmer, 1936
| | | | Species | Malthinus balteatus Suffrian, 1851
| | | | Species | Malthinus baroniurbanii Wittmer, 1974
| | | | Species | Malthinus basimarginatus Wittmer, 1966
| | | | Species | Malthinus batnensis Pic, 1900
| | | | Species | Malthinus bicolor LeConte, 1884 ⇒ Caccodes bicolor
Salmon gill-maggot | | | | Species | Malthinus biglianii Fiori, 1915
| | | | Species | Malthinus biguttatus Linnaeus, 1758
| | | | Species | Malthinus biguttulus Paykull, 1800 ⇒ Malthinus biguttatus
| | | | Species | Malthinus biinterruptus Pic, 1903
| | | | Species | Malthinus bilineatus Kiesenwetter, 1852
| | | | Species | Malthinus binotaticollis Pic, 1913 ⇒ Malthinus seriepunctatus
| | | | Species | Malthinus biskrensis Pic, 1900
| | | | Species | Malthinus bordei Pic, 1911
| | | | Species | Malthinus brevenotaticeps Pic, 1923
| | | | Species | Malthinus brevereductus Pic, 1953
| | | | Species | Malthinus brevior Pic, 1900
| | | | Species | Malthinus brevipennis ⇒ Caccodes brevipennis
| | | | Species | Malthinus brunneosulcata Pic, 1940
| | | | Species | Malthinus caifensis Pic, 1900
| | | | Species | Malthinus calaber Pic, 1909
| | | | Species | Malthinus capito
| | | | Species | Malthinus carbonarius Waltl, 1838 ⇒ Malthodes (Malthodes) maurus
| | | | Species | Malthinus centrocarinatus Wittmer, 1969
| | | | Species | Malthinus chelifer Kiesenwetter, 1851
| | | | Species | Malthinus chiosensis
| | | | Species | Malthinus chisosensis Fender, 1963 ⇒ Caccodes chisosensis
| | | | Species | Malthinus chobauti Bourgeois, 1897
| | | | Species | Malthinus cilicius Svihla, 2002
| | | | Species | Malthinus cincticollis Kiesenwetter, 1866
| | | | Species | Malthinus cognatus Stephens, 1830 ⇒ Malthodes (Malthodes) minimus
| | | | Species | Malthinus collaris J. R. Sahlberg, 1903 ⇒ Malthinus flaveolus
| | | | Species | Malthinus concolor Stephens, 1830 ⇒ Malthinus (Malthinus) frontalis
| | | | Species | Malthinus conspicuus Kiesenwetter, 1852
| 2 | Subspecies | | Species | Malthinus corcyreus Pic, 1901
| | | | Species | Malthinus crassicornis Maeklin, 1846
| | | | Species | Malthinus crassipes Pic, 1899
| | | | Species | Malthinus crenulatomimus Wittmer, 1984 ⇒ Malthinellus crenulatomimus
| | | | Species | Malthinus crenulatus Wittmer, 1979 ⇒ Malthinellus crenulatus
| | | | Species | Malthinus croceicollis Wollaston, 1862
| | | | Species | Malthinus curtenotatus Pic, 1931
| | | | Species | Malthinus curzensis
| | | | Species | Malthinus dalmatinus Pic, 1902
| | | | Species | Malthinus damascenus Wittmer, 1936
| | | | Species | Malthinus danieli ♰ A. Kuśka, 2010
| | | | Species | Malthinus deguchii Takahashi, 2018
| | | | Species | Malthinus delagrangei Pic, 1900
| | | | Species | Malthinus delicatulus Bourgeois, 1899 ⇒ Malthinus facialis
| | | | Species | Malthinus depauperatus Wollaston, 1862
| | | | Species | Malthinus difficilis LeConte, 1851
| | | | Species | Malthinus diffusus Kiesenwetter, 1866
| | | | Species | Malthinus dimidiaticollis Rosenhauer, 1847
| | | | Species | Malthinus dimorphus Wittmer, 1935
| | | | Species | Malthinus dispar Germar, 1823
| | | | Species | Malthinus distinctipes Pic, 1901 ⇒ Malthinus seriepunctatus
| | | | Species | Malthinus diversicollis Pic, 1906
| | | | Species | Malthinus diversicolor Pic, 1905
| | | | Species | Malthinus diversithorax Pic, 1901 ⇒ Malthinus scriptus
| | | | Species | Malthinus dromioides Bourgeois, 1885
| | | | Species | Malthinus drurei Pic, 1904
| | | | Species | Malthinus dryocoetes Rottenberg, 1871
| | | | Species | Malthinus dubius Motschulsky, 1853
| | | | Species | Malthinus duplolineatus Champion, 1926
| | | | Species | Malthinus elongaticollis Pic, 1907
| | | | Species | Malthinus euboeicus Wittmer, 1974
| | | | Species | Malthinus excisipes Pic, 1899
| | | | Species | Malthinus externus Pic, 1900
| | | | Species | Malthinus facialis C. G. Thomson, 1864
| | | | Species | Malthinus fasciatus A. G. Olivier, 1790 ⇒ Malthinus balteatus
| | | | Species | Malthinus femoralis Pic, 1900
| | | | Species | Malthinus filicornis Kiesenwetter, 1852 ⇒ Malthinus scriptus
| | | | Species | Malthinus fjellandi Fender, 1963 ⇒ Caccodes fjellandi
| | | | Species | Malthinus flammeicollis Wollaston, 1865
| | | | Species | Malthinus flaveolus Herbst, 1784
| | | | Species | Malthinus flavicans Bourgeois, 1903
| | | | Species | Malthinus flavicollis Pic, 1904
| | | | Species | Malthinus flavipennis Pic, 1903
| | | | Species | Malthinus flavus Latreille, 1806 ⇒ Malthinus flaveolus
| | | | Species | Malthinus forcipipher Kiesenwetter, 1852
| | | | Species | Malthinus fulvicollis Stephens, 1830 ⇒ Malthodes (Malthodes) minimus
| | | | Species | Malthinus fuscescens Blanchard, 1846
| | | | Species | Malthinus fuscipes Bourgeois, 1897
| | | | Species | Malthinus fuscus Waltl, 1838
| | | | Species | Malthinus gafsaensis Wittmer, 1969
| | | | Species | Malthinus gardneri Pic, 1929
| | | | Species | Malthinus garganicus Fiori, 1915
| | | | Species | Malthinus geigei Wittmer, 1971
| | | | Species | Malthinus geniculatus Kiesenwetter, 1859
| | | | Species | Malthinus georgiensis Fender, 1966
| | | | Species | Malthinus glabellus Kiesenwetter, 1852
| | | | Species | Malthinus gomerensis
| | | | Species | Malthinus grandiceps Pic, 1901
| | | | Species | Malthinus granicollis McKey-Fender, 1951 ⇒ Caccodes granicollis
| | | | Species | Malthinus gratiosus Pic, 1901
| | | | Species | Malthinus griseipennis Pic, 1913 ⇒ Malthinus flaveolus
| | | | Species | Malthinus griseonotata Pic, 1951
| | | | Species | Malthinus henrici Pic, 1901
| | | | Species | Malthinus horaki Svihla, 1980 ⇒ Malthinus (Malthinus) planulus
| | | | Species | Malthinus huachucae Fender, 1951 ⇒ Caccodes huachucae
| | | | Species | Malthinus humeralis Kiesenwetter, 1874
| | | | Species | Malthinus immaculatus Stephens, 1830 ⇒ Malthinus (Malthinus) frontalis
| | | | Species | Malthinus immunis Marsham, 1802
| | | | Species | Malthinus impressicollis Fairmaire, 1884
| | | | Species | Malthinus inaequalithorax Pic, 1933
| | | | Species | Malthinus incanus Kiesenwetter, 1852
| | | | Species | Malthinus incrassicornis Pic, 1900
| | | | Species | Malthinus incrassipes Wittmer, 1956
| | | | Species | Malthinus indisjuncta Pic, 1910
| | | | Species | Malthinus indistincta Pic, 1910
| | | | Species | Malthinus inflavus Marseul, 1878
| | | | Species | Malthinus ingens Champion, 1915
| | | | Species | Malthinus insignipes Pic, 1907
| | | | Species | Malthinus insularis Pic, 1915
| | | | Species | Malthinus ionicus Pic, 1901
| | | | Species | Malthinus japonicus Ohbayashi, 1949
| | | | Species | Malthinus kamengensis Wittmer, 1966
| | | | Species | Malthinus karenpankowskiae Pankowski & Fanti, 2023
| | | | Species | Malthinus kiesenwetteri C. N. F. Brisout de Barneville, 1863
| | | | Species | Malthinus klapperichi Wittmer, 1951
| | | | Species | Malthinus knulli Fender, 1963 ⇒ Caccodes knulli
| | | | Species | Malthinus kobensis Lewis, 1895
| | | | Species | Malthinus kocheri Pic, 1953
| | | | Species | Malthinus kraatzi Pic, 1900
| | | | Species | Malthinus krueperi Pic, 1901 ⇒ Malthinus balteatus
| | | | Species | Malthinus kruperi Pic, 1901
| | | | Species | Malthinus lacteifrons Marseul, 1878
| | | | Species | Malthinus laesus Kiesenwetter, 1871
| | | | Species | Malthinus laetus Waltl, 1838 ⇒ Malthodes neglectus
| | | | Species | Malthinus laevicollis Kiesenwetter, 1859
| | | | Species | Malthinus lateflavicans Pic, 1910
| | | | Species | Malthinus lateralis Motschulsky, 1853 ⇒ Malthinus balteatus
| | | | Species | Malthinus laterufus Pic, 1900
| | | | Species | Malthinus laticeps Fiori, 1915 ⇒ Caccodes laticeps
| | | | Species | Malthinus laticollis Pic, 1900
| | | | Species | Malthinus latipennis Germar, 1823
| | | | Species | Malthinus latirufus Pic, 1901
| | | | Species | Malthinus limbatus Pic, 1900
| | | | Species | Malthinus lineata Pic, 1899
| | | | Species | Malthinus lineatocollis Champion, 1920
| | | | Species | Malthinus lituratus Motschulsky, 1853
| | | | Species | Malthinus longevittatus Pic, 1953
| | | | Species | Malthinus longicornis Kiesenwetter, 1866
| | | | Species | Malthinus longipennis Motschulsky, 1853 ⇒ Malthinus (Malthinus) frontalis
| | | | Species | Malthinus longithorax Pic, 1899
| | | | Species | Malthinus luteola Trella, 1925 ⇒ Malthinus facialis
| | | | Species | Malthinus luteolus Stephens, 1830
| | | | Species | Malthinus luteus Pic, 1935
| | | | Species | Malthinus maculaticeps Pic, 1920
| | | | Species | Malthinus maculipennis Pic, 1900
| | | | Species | Malthinus maculithorax Pic, 1898
| | | | Species | Malthinus maculiventris Fairmaire, 1884 ⇒ Malthinus scriptus
| | | | Species | Malthinus marginatus Latreille, 1806
| | | | Species | Malthinus marginicollis Ganglbauer, 1906
| | | | Species | Malthinus marginipennis Bourgeois, 1888 ⇒ Malthinus scriptus
| | | | Species | Malthinus maritimus Pic, 1899 ⇒ Malthinus facialis
| | | | Species | Malthinus masoni M.G.Pankowski & Fanti, 2022
| | | | Species | Malthinus mathieui Pic, 1900
| | | | Species | Malthinus maurus Laporte de Castelnau, 1840
| | | | Species | Malthinus maximiceps Pic, 1917
| | | | Species | Malthinus mediocrenotata Pic, 1914 ⇒ Malthinus seriepunctatus
| | | | Species | Malthinus mediocrenotatus Pic, 1914
| | | | Species | Malthinus melanocerus
| | | | Species | Malthinus merkli J. Weise, 1895 ⇒ Malthinus balteatus
| | | | Species | Malthinus mexensis Wittmer, 1969
| | | | Species | Malthinus mimica Pic, 1906 ⇒ Malthinus seriepunctatus
| | | | Species | Malthinus montandoni Pic, 1907
| | | | Species | Malthinus moricei Pic, 1930
| | | | Species | Malthinus mucoreus Kiesenwetter, 1879
| | | | Species | Malthinus multonotata Pic, 1910 ⇒ Malthinus seriepunctatus
| | | | Species | Malthinus mutabilis Wollaston, 1862
| | | | Species | Malthinus mutata Pic, 1910
| | | | Species | Malthinus mutilatus Kiesenwetter, 1879
| | | | Species | Malthinus nakanei Wittmer, 1955
| | | | Species | Malthinus neapolitanus Pic, 1905
| | | | Species | Malthinus nepalensis Wittmer, 1966
| | | | Species | Malthinus nigribuccis Marseul, 1878
| | | | Species | Malthinus nigricollis Stephens, 1830 ⇒ Malthodes (Malthodes) marginatus
| | | | Species | Malthinus nigrina Wittmer, 1936
| | | | Species | Malthinus nigripes Bourgeois, 1897
| | | | Species | Malthinus nigrithorax Wittmer, 1936
| | | | Species | Malthinus nigronotatus Pic, 1908
| | | | Species | Malthinus nitidithorax Pic, 1905
| | | | Species | Malthinus normandi Pic, 1900
| | | | Species | Malthinus notaticeps Wittmer, 1956
| | | | Species | Malthinus notaticollis Pic, 1900
| | | | Species | Malthinus novaki Reitter, 1889
| | | | Species | Malthinus obscuriceps Pic, 1900 ⇒ Malthinus balteatus
| | | | Species | Malthinus obscuricollis Fairmaire, 1883
| | | | Species | Malthinus obscuricolor Pic, 1920
| | | | Species | Malthinus obscurior Bourgeois, 1897
| | | | Species | Malthinus obscuripennis Pic, 1898
| | | | Species | Malthinus obscuripes Kiesenwetter, 1866
| | | | Species | Malthinus obscurithorax Pic, 1900
| | | | Species | Malthinus occipitalis LeConte, 1851
| 3 | Subspecies | | Species | Malthinus okinawanus Wittmer, 1970
| | | | Species | Malthinus olcesei Pic, 1898
| | | | Species | Malthinus orbiculatus N.Takahashi, 0
| | | | Species | Malthinus ornatus Rosenhauer, 1856
| | | | Species | Malthinus oujdaensis Wittmer, 1969
| | | | Species | Malthinus pallicolor Fairmaire, 1883
| | | | Species | Malthinus pallidipennis Pic, 1900
| | | | Species | Malthinus pallidipes Fairmaire, 1884
| | | | Species | Malthinus pallidithorax Pic, 1899 ⇒ Malthinus seriepunctatus
| | | | Species | Malthinus paradoxus Dohrn, 1882
| | | | Species | Malthinus pardoi Wittmer, 1969
| | | | Species | Malthinus pauljohnsoni Pankowski & Fanti, 2023
| | | | Species | Malthinus persicus Pic, 1901
| | | | Species | Malthinus phrygius Svihla, 2002
| | | | Species | Malthinus piceonotatus Pic, 1916
| | | | Species | Malthinus pici Wittmer, 1936
| | | | Species | Malthinus pinicola Stephens, 1830 ⇒ Malthinus (Malthinus) frontalis
| | | | Species | Malthinus pithanoides Escalera, 1913
| | | | Species | Malthinus praetextus Bourgeois, 1899 ⇒ Malthinus balteatus
| | | | Species | Malthinus pubescens Pic, 1911
| | | | Species | Malthinus pumilus Brebisson, 1835 ⇒ Malthodes (Malthodes) pumilus
| | | | Species | Malthinus punctatus Geoffroy, 1785 ⇒ Malthinus flaveolus
| | | | Species | Malthinus pyrrhoderus Fairmaire, 1883 ⇒ Malthinus rubricollis
| | | | Species | Malthinus quadrimaculatus Fender, 1951 ⇒ Caccodes quadrimaculatus
| | | | Species | Malthinus quadrinodosus Champion, 1926
| | | | Species | Malthinus quedenfeldti Pic, 1910
| | | | Species | Malthinus raffrayi Pic, 1910 ⇒ Malthinus seriepunctatus
| | | | Species | Malthinus raveli Pic, 1906 ⇒ Malthinus seriepunctatus
| | | | Species | Malthinus recurvus Baudi di Selve, 1872
| | | | Species | Malthinus rhaphidiceps Kiesenwetter, 1852
| | | | Species | Malthinus rifbjergi Fanti & Damgaard, 2018
| | | | Species | Malthinus robustus Motschulsky, 1853
| | | | Species | Malthinus rosinae Pic, 1900
| | | | Species | Malthinus rubricollis Baudi di Selve, 1859
| | | | Species | Malthinus rufescens Pic, 1901
| | | | Species | Malthinus ruficeps Pic, 1899
| | | | Species | Malthinus ruficollis Latreille, 1806 ⇒ Malthodes (Malthodes) minimus
| | | | Species | Malthinus rufifrons Motschulsky, 1860
| | | | Species | Malthinus rufithorax Pic, 1903
| | | | Species | Malthinus sabauda Pic, 1901
| | | | Species | Malthinus sabaudus Pic, 1901 ⇒ Malthinus facialis
| | | | Species | Malthinus sagamiensis Takahashi, 1992
| | | | Species | Malthinus sahlbergi Pic, 1909
| | | | Species | Malthinus sanguinicollis Fallen, 1807
| | | | Species | Malthinus sauteri Pic, 1934
| | | | Species | Malthinus schatzmayri Pic, 1951
| | | | Species | Malthinus schaubi Wittmer, 1969
| | | | Species | Malthinus scriptipennis Pic, 1900
| | | | Species | Malthinus scriptus Kiesenwetter, 1852
| | | | Species | Malthinus scutellaris Rosenhauer, 1856
| | | | Species | Malthinus sellatus Blanchard, 1846
| | | | Species | Malthinus semiobscurus Pic, 1900
| | | | Species | Malthinus sericellus Marseul, 1868
| | | | Species | Malthinus seriepunctatus Kiesenwetter, 1852
| | | | Species | Malthinus serrulatus N.Takahashi, 0
| | | | Species | Malthinus sibilleae Wittmer, 1969
| | | | Species | Malthinus sicanus Kiesenwetter, 1871
| | | | Species | Malthinus sikkimensis Pic, 1924
| | | | Species | Malthinus simoni Pic, 1913
| | | | Species | Malthinus simplicipes Pic, 1899
| | | | Species | Malthinus sinuatipes Pic, 1908 ⇒ Malthinus balteatus
| | | | Species | Malthinus smyrnensis Pic, 1915
| | | | Species | Malthinus spissicornis Champion, 1926
| | | | Species | Malthinus stigmatias Kiesenwetter, 1866
| | | | Species | Malthinus stigmatus Kiesenwetter, 1866
| | | | Species | Malthinus striatulus Mulsant, 1862 ⇒ Malthinus biguttatus
| | | | Species | Malthinus subcostatus Schaeffer, 1908 ⇒ Caccodes subcostatus
Barnacles | | | | Species | Malthinus subfusca Pic, 1906 ⇒ Malthinus flaveolus
| | | | Species | Malthinus subgeniculatus Pic, 1930
| | | | Species | Malthinus subopaca Pic, 1900 ⇒ Malthinus seriepunctatus
| | | | Species | Malthinus subopaciceps Pic, 1915
| | | | Species | Malthinus subopacus Pic, 1900
| | | | Species | Malthinus subulatus Champion, 1915
| | | | Species | Malthinus sulcicollis Marseul, 1878
| | | | Species | Malthinus syriacus Marseul, 1868
| | | | Species | Malthinus taforaltensis Wittmer, 1969
| | | | Species | Malthinus tamarisci Pic, 1900
| | | | Species | Malthinus tangerianus Pic, 1902
| | | | Species | Malthinus tauri Pic, 1907
| | | | Species | Malthinus taygetanus Pic, 1902 ⇒ Malthinus balteatus
| | | | Species | Malthinus tenietensis Pic, 1900
| | | | Species | Malthinus terrestris Pic, 1905
| | | | Species | Malthinus theresae Pic, 1899
| | | | Species | Malthinus theryi Pic, 1920
| | | | Species | Malthinus tibialis Wittmer, 1966
| | | | Species | Malthinus tibiellus Pic, 1918
| | | | Species | Malthinus tiburtina Baudi di Selve, 1893
| | | | Species | Malthinus torretassoi Pic, 1934
| | | | Species | Malthinus tortiscelis Pic, 1898
| | | | Species | Malthinus tricolor Fender, 1951 ⇒ Caccodes tricolor
| | | | Species | Malthinus triggiber Marseul, 1878
| | | | Species | Malthinus trinotaticeps Pic, 1951
| | | | Species | Malthinus tuniseus Fairmaire, 1875
| | | | Species | Malthinus turcicus Pic, 1899
| | | | Species | Malthinus unicolor Dufour, 1851 ⇒ Malthodes (Malthodes) maurus
| | | | Species | Malthinus unionis Pic, 1909
| | | | Species | Malthinus validiceps Pic, 1928
| | | | Species | Malthinus vandykei Wittmer, 1956
| | | | Species | Malthinus variegatipennis Blanchard, 1846
| | | | Species | Malthinus varus Bourgeois, 1900
| | | | Species | Malthinus vaucheri Pic, 1901
| | | | Species | Malthinus vaulogeri Pic, 1915
| | | | Species | Malthinus venezuelensis
| | | | Species | Malthinus versatilis Delkeskamp, 1939
| | | | Species | Malthinus verticalis Pic, 1930
| | | | Species | Malthinus vitalei Pic, 1916
| | | | Species | Malthinus vitellinus Kiesenwetter, 1866
| | | | Species | Malthinus wewalkai Wittmer, 1980
| | |
Synonyms
Distribution of Malthinus Latreille, 1806 Algeria, Austria, Azerbaijan, Belarus, Belgium, Bhutan, Bulgaria, Canada, China, Cyprus, Czech Republic, Denmark, El Salvador, Estonia, Finland, France, Georgia, Germany, Greece, India, Iran, Ireland, Italy, Japan, Jordan, Kyrgyzstan, Latvia, Lithuania, Luxembourg, Mexico, Montenegro, Morocco, Nepal, Netherlands, Norway, Pakistan, Panama, Poland, Portugal, Republic South Africa, Russia, Serbia, Slovakia, South Korea, Spain, Sweden, Switzerland, Syria, Taiwan, Tajikistan, Thailand, Tunisia, Turkey, Turkmenistan, United Kingdom, USA, Venezuela, Vietnam
Found in fossile formations Green River Formation, Much Wenlock Limestone Formation
Links, sources and further information
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