|
|
| | | | Checklist of Tribe Onthophagini Lacordaire, 1856
for Palaearctic (Ecozones)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 409
11833
195881 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 408
11832
195881 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 392
11734
195055 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 321
11427
194050 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 140
8868
175782 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 126
7984
148422 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 14
830
15619 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 14
830
15619 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 671
13345 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 46
1602 | Genera
Species |
Taxom
| | Tribe | Onthophagini
Lacordaire, 1856
| 11
995 | Genera
Species |
Taxom
|
Genera for Palaearctic
11 Genera with 995 Species | Anoctus Sharp, 1942 | | 2 | Species | Taxom | | Caccobius Thomson, 1994 | | 74 | Species | Taxom | | Cassolus Sharp, 1952 | | 5 | Species | Taxom | | Cleptocaccobius Cambefort, 1857 | | 9 | Species | Taxom | | Digitonthophagus Balthasar, 2017 | | 11 | Species | Taxom | | Disphysema Harold, 1873 | | 1 | Species | Taxom | | Euonthophagus Balthasar, 1946 | | 56 | Species | Taxom | | Haroldius Boucomont, 1993 | | 9 | Species | Taxom | | Onthophagus Latreille, 1939 | | 857 | Species | Taxom | | Phalops Erichson, 1883 | | 6 | Species | Taxom | | Proagoderus Lansberge, 1912 | | 6 | Species | Taxom |
Species list for Palaearctic
995 Species and 38 Subspecies | Anoctus foveolatus (Paulian, 1945) | ⇒ Anoctus siamensis Balthasar, 1942 | | Anoctus siamensis Balthasar, 1942 | | | Caccobius (Caccobius) chujoi Paulian, 1942 | | | Caccobius (Caccophilus) tortus Sharp, 1875 | | | Caccobius tortus matsudai Ochi & Araya, 1996 | | | Caccobius tortus tortus | | | Caccobius (Caccophilus) unicornis Fabricius, 1798 | | | Caccobius amagisanus (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccobius anthracinus Faldermann, 1835 | | | Caccobius asininus Heyden, 1887 | | | Caccobius aterrimus Fabricius, 1798 | | | Caccobius atomarius (Balthasar, 1935) | ⇒ Caccobius brevis Waterhouse, 1875 | | Caccobius balthasari (Tesar, 1969) | ⇒ Cleptocaccobius balthasarianus Cambefort, 1979 | | Caccobius bicuspis Wiedemann, 1823 | | | Caccobius bidentatus Boucomont, 1923 | | | Caccobius bimaculatus Mulsant, 1842 | | | Caccobius boucomonti Balthasar, 1935 | | | Caccobius brevis Waterhouse, 1875 | | | Caccobius chokakurainus Matsumura, 1938 | | | Caccobius christophi Harold, 1879 | | | Caccobius corsicus (Schneider, 1902) | ⇒ Caccobius schreberi Linnaeus, 1767 | | Caccobius crenatostriatus (Balthasar, 1935) | ⇒ Caccobius christophi Harold, 1879 | | Caccobius demangei Boucomont, 1919 | | | Caccobius denticollis Harold, 1867 | | | Caccobius diminutivus Walker, 1858 | | | Caccobius excavatus Bai, Zhang &..., 2007 | | | Caccobius flavolimbatus Balthasar, 1942 | | | Caccobius fukiensis Balthasar, 1942 | | | Caccobius gonoderus Fairmaire, 1888 | | | Caccobius haroldi (Jekel, 1872) | ⇒ Caccobius aterrimus Fabricius, 1798 | | Caccobius himalayanus Jekel, 1872 | | | Caccobius hirayamai (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccobius histeroides Ménétriés, 1832 | | | Caccobius imitans Balthasar, 1933 | | | Caccobius indicus Harold, 1867 | | | Caccobius indistinctus Mulsant, 1842 | | | Caccobius inermis (Arrow, 1931) | ⇒ Cleptocaccobius inermis Arrow, 1931 | | Caccobius jessoensis Harold, 1867 | | | Caccobius juvenilis Mulsant, 1842 | | | Caccobius kasuganus Matsumura & ..., 1936 | | | Caccobius kelleri Olsoufieff, 1907 | | | Caccobius koichii Matsumura, 1934 | | | Caccobius koltzei (Reitter, 1892) | ⇒ Caccobius sordidus Harold, 1886 | | Caccobius masumotoi Cambefort, 1990 | | | Caccobius matsudai Ochi & Araya, 1996 | | | Caccobius matsuoi Matsumura & ..., 1937 | | | Caccobius meridionalis Boucomont, 1914 | | | Caccobius microcephalus (Harold, 1877) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccobius minimus (d´Orbigny, 1898) | ⇒ Caccophilus viridicollis Fahraeus, 1857 | | Caccobius mixtus Mulsant, 1842 | | | Caccobius mundus Ménétriés, 1838 | | | Caccobius nikkoensis Lewis, 1895 | | | Caccobius nitidiceps Fairmaire, 1893 | | | Caccobius nutans Sharp, 1875 | | | Caccobius obscurus Mulsant, 1842 | | | Caccobius pantherinus Arrow, 1931 | | | Caccobius pulicarius Harold, 1875 | | | Caccobius pullus Jekel, 1872 | | | Caccobius sapporensis (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccobius scheuerni Cambefort, 1990 | | | Caccobius schreberi Linnaeus, 1767 | | | Caccobius semicoeruleus (d´Orbigny, 1905) | ⇒ Caccophilus viridicollis Fahraeus, 1857 | | Caccobius setulosus Motschulsky, 1864 | | | Caccobius sibiricus (Balthasar, 1935) | ⇒ Caccobius kelleri Olsoufieff, 1907 | | Caccobius sordidus Harold, 1886 | | | Caccobius suzukii (Matsumura, 1936) | ⇒ Caccobius boucomonti Balthasar, 1935 | | Caccobius torticornis Arrow, 1931 | | | Caccobius tsunoellus (Matsumara, 1938) | ⇒ Caccophilus tortus Sharp, 1875 | | Caccobius ultor Sharp, 1875 | | | Caccobius vacerrosus Lewis, 1895 | | | Caccobius viridicollis (d´Orbigny, 1902) | ⇒ Caccophilus convexifrons Raffray, 1877 | | Caccobius vulcanus Fabricius, 1801 | | | Caccobius yamauchii (Matsumura, 1936) | ⇒ Caccobius unicornis Fabricius, 1798 | | Caccobius yubariensis (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccobius yunnanicus (Kabakov, 1994) | ⇒ Caccobius masumotoi Cambefort, 1990 | | Cassolus fukiensis Balthasar, 1960 | | | Cassolus gotoi Masumoto, 1986 | | | Cassolus humeralis Arrow, 1907 | | | Cassolus nudus Sharp, 1875 | | | Cassolus quadridentatus Balthasar, 1952 | | | Cleptocaccobius balthasarianus Cambefort, 1979 | | | Cleptocaccobius convexifrons Raffray, 1877 | | | Cleptocaccobius inermis Arrow, 1931 | | | Cleptocaccobius scrofa Balthasar, 1942 | | | Cleptocaccobius shansicus Nakane, 1951 | | | Cleptocaccobius simplex Boucomont, 1923 | | | Cleptocaccobius simplex simplex | | | Cleptocaccobius viridicollis Fahraeus, 1857 | | | Digitonthophagus aksumensis Génier, 2017 | | | Digitonthophagus antilope Fabricius, 1798 | | | Digitonthophagus bonasus Fabricius, 1775 | | | Digitonthophagus catta Fabricius, 1787 | Beetle | | Digitonthophagus dorcas A. G. Olivier, 1789 | | | Digitonthophagus eucatta Génier, 2017 | | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Digitonthophagus intermedius Reiche, 1840 | | | Digitonthophagus metallicus Fabricius, 1798 | | | Digitonthophagus uks Génier, 2017 | | | Disphysema candezei Harold, 1873 | | | Euonthophagus aeneobrunneus Kabakov, 1977 | | | Euonthophagus alces Fabricius, 1792 | | | Euonthophagus amyntas Olivier, 1789 | | | Euonthophagus amyntas alces Fabricius, 1792 | | | Euonthophagus amyntas amyntas Olivier, 1789 | | | Euonthophagus amyntas auchenia Redtenbacher, 1850 | | | Euonthophagus amyntas subviolaceus Ménétries, 1832 | | | Euonthophagus aspadanaensis Petrovitz, 1965 | | | Euonthophagus atramentarius Ménétriés, 1832 | | | Euonthophagus auchenia L. Redtenbacher, 1850 | | | Euonthophagus bedeli Reitter, 1891 | | | Euonthophagus carbonarius Klug, 1855 | | | Euonthophagus clementi Baraud, 1960 | | | Euonthophagus conterminus Petrovitz, 1971 | | | Euonthophagus crocatus Mulsant & Go..., 1870 | | | Euonthophagus crocatus clementi Baraud, 1960 | | | Euonthophagus crocatus crocatus | | | Euonthophagus difformis Mulsant, 1842 | | | Euonthophagus dorbignyi Olsoufieff, 1900 | | | Euonthophagus dorbignyi neutto Olsoufieff, 1900 | | | Euonthophagus dubius Mulsant, 1842 | | | Euonthophagus felschei Reitter, 1893 | | | Euonthophagus ganglbaueri Reitter, 1891 | | | Euonthophagus gibbosus Scriba, 1790 | | | Euonthophagus hannas (J. R. Sahlberg, 1908) | ⇒ Euonthophagus tissoni Reitter, 1906 | | Euonthophagus hazariensis Kabakov, 1977 | | | Euonthophagus hybneri Fabricius, 1792 | | | Euonthophagus imitator Reitter, 1891 | | | Euonthophagus koshantschikoffi Reitter, 1891 | | | Euonthophagus laghmanicus Kabakov, 1977 | | | Euonthophagus loeffleri Petrovitz, 1965 | | | Euonthophagus maindroni d´Orbigny, 1897 | | | Euonthophagus markaziensis (Petrovitz, 1980) | ⇒ Onthophagus aeneobrunneus Kabakov, 1977 | | Euonthophagus mostafatsairi Palestrini, ..., 1979 | | | Euonthophagus mostafatsairii Palestrini, ..., 1979 | | | Euonthophagus ochripennis d´Orbigny, 1898 | | | Euonthophagus orcas Ménétriés, 1838 | | | Euonthophagus pentaceros Olsoufieff, 1900 | | | Euonthophagus peropacus Kirschenblatt, 1935 | | | Euonthophagus pertinax Balthasar, 1959 | | | Euonthophagus rapillyi (Baraud, 1984) | ⇒ Onthophagus conterminus Petrovitz, 1971 | | Euonthophagus rechingeriorum (Mandl, 1976) | ⇒ Euonthophagus amyntas auchenia Redtenbacher, 1850 | | Euonthophagus schnabeli Splichal, 1910 | | | Euonthophagus semirufus d´Orbigny, 1898 | | | Euonthophagus subviolaceus Ménétriés, 1832 | | | Euonthophagus sulcicollis Reitter, 1892 | | | Euonthophagus sycophanta Mulsant, 1842 | | | Euonthophagus tages A. G. Olivier, 1789 | | | Euonthophagus tissoni Reitter, 1906 | | | Euonthophagus umbrinus Mulsant, 1842 | | | Euonthophagus unituberculatus Mulsant, 1842 | | | Euonthophagus verberatus Ziani, 2006 | | | Euonthophagus weisei Reitter, 1891 | | | Euonthophagus yama Ziani, 2006 | | | Euonthophagus yemenicus Balthasar, 1946 | | | Haroldius heimi Wasmann, 1918 | | | Haroldius hwangi Masumoto, Le..., 2005 | | | Haroldius jechaer Král & Hájek, 2012 | | | Haroldius lassallei Cambefort, 1986 | | | Haroldius nigrita Wassmann, 1918 | | | Haroldius perroti Paulian, 1939 | | | Haroldius stevensi Arrow, 1931 | | | Haroldius turnai Kral, 2003 | | | Haroldius uenoi Masumoto & Yin, 1993 | | | Onthophagus (Afrostrandius) granulipennis (Lansberge, 1886) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus (Altonthophagus) cervenkai Kabakov, 2008 | | | Onthophagus (Altonthophagus) inelegans Balthasar, 1935 | | | Onthophagus (Altonthophagus) kozlovi Kabakov, 1990 | | | Onthophagus (Altonthophagus) kukunorensis Kabakov, 1990 | | | Onthophagus (Altonthophagus) marmotae Kabakov, 1990 | | | Onthophagus (Altonthophagus) sibiricus Harold, 1877 | | | Onthophagus (Altonthophagus) turpidoides Kabakov, 2008 | | | Onthophagus (Amphionthophagus) falzonii Goidanich, 1926 | | | Onthophagus (Amphionthophagus) melitaeus Fabricius, 1798 | | | Onthophagus (Colobonthophagus) aenescens Wiedemann, 1823 | | | Onthophagus (Colobonthophagus) arunensis Scheuern, 1995 | | | Onthophagus (Colobonthophagus) dama Fabricius, 1798 | | | Onthophagus (Colobonthophagus) occipitalis Lansberge, 1885 | | | Onthophagus (Colobonthophagus) paliceps Arrow, 1931 | | | Onthophagus (Colobonthophagus) piceorufulus Kabakov, 1994 | | | Onthophagus (Colobonthophagus) quadridentatus (Fabricius, 1798) | ⇒ Onthophagus fissicornis Stevens, 1809 | | Onthophagus (Colobonthophagus) tragus Fabricius, 1792 | | | Onthophagus (Euonthophagus) aeneobrunneus Kabakov, 1977 | | | Onthophagus amyntas auchenia Redtenbacher, 1850 | | | Onthophagus (Euonthophagus) conterminus Petrovitz, 1971 | | | Onthophagus (Furconthophagus) amicus Gillet, 1925 | | | Onthophagus (Furconthophagus) boucomonti Paulian, 1931 | | | Onthophagus (Furconthophagus) furcatus (Fabricius, 1781) | ⇒ Onthophagus furcaticeps Masters, 1886 | | Onthophagus (Furconthophagus) lilliputanus Lansberge, 1883 | | | Onthophagus (Furconthophagus) schoolmeestersi Ochi & Kon, 2007 | | | Onthophagus (Gibbonthophagus) apicetinctus (d´Orbigny, 1900) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus (Gibbonthophagus) atripennis Waterhouse, 1875 | | | Onthophagus (Gibbonthophagus) chineicus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) duporti Boucomont, 1914 | | | Onthophagus (Gibbonthophagus) euryceros Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) gracilipes Boucomont, 1914 | | | Onthophagus (Gibbonthophagus) kentingensis Nomura, 1973 | | | Onthophagus (Gibbonthophagus) lenis Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) limbatus Herbst, 1789 | | | Onthophagus (Gibbonthophagus) luridipennis Boheman, 1858 | | | Onthophagus (Gibbonthophagus) privus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) taurinus White, 1844 | | | Onthophagus (Gibbonthophagus) viduus (Harold, 1875) | ⇒ Onthophagus orbus Boucomont, 1924 | | Onthophagus (Indachorius) arai Masumoto, 1989 | | | Onthophagus (Indachorius) baykanus Kabakov, 1994 | | | Onthophagus (Indachorius) bonengus Kabakov, 1994 | | | Onthophagus (Indachorius) chinensis Balthasar, 1952 | | | Onthophagus (Indachorius) koshunensis Balthasar, 1941 | | | Onthophagus (Indachorius) octonaevus Kabakov, 1994 | | | Onthophagus (Indachorius) platypus Zhang, 1997 | | | Onthophagus (Indachorius) subsapaensis Kabakov, 1994 | | | Onthophagus (Indachorius) suginoi Ochi, 1984 | | | Onthophagus (Indachorius) tiamicus Kabakov, 1994 | | | Onthophagus (Macronthophagus) cludtsi Ochi, 2003 | | | Onthophagus (Macronthophagus) diabolicus Harold, 1877 | | | Onthophagus (Macronthophagus) manipurensis Arrow, 1907 | | | Onthophagus (Macronthophagus) rubricollis Hope, 1831 | | | Onthophagus (Matashia) anguliceps (Boucomont, 1914) | ⇒ Onthophagus avocetta Arrow, 1933 | | Onthophagus (Matashia) yubarinus (Matsumura, 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus (Micronthophagus) falsivigilans Masumoto, 1995 | | | Onthophagus (Micronthophagus) konoi Matsumura, 1938 | | | Onthophagus (Micronthophagus) quaestus Sharp, 1875 | | | Onthophagus (Micronthophagus) vigilans Boucomont, 1921 | | | Onthophagus (Onthophagiellus) kondaoensis Kabakov, 1994 | | | Onthophagus (Onthophagiellus) popovi Kabakov, 1983 | | | Onthophagus (Onthophagiellus) taiyaruensis Masumoto, 1977 | | | Onthophagus (Onthophagiellus) unguiculatus Kabakov, 1994 | | | Onthophagus (Onthophagus) abacus Boucomont, 1921 | | | Onthophagus (Onthophagus) acuticollis (Gillet, 1927) | ⇒ Onthophagus amphioxus Arrow, 1933 | | Onthophagus (Onthophagus) acuticornis Endrödi, 1973 | | | Onthophagus (Onthophagus) alternans (Raffray, 1877) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus (Onthophagus) amirus Kabakov, 1982 | | | Onthophagus (Onthophagus) anguicorius Boucomont, 1914 | | | Onthophagus (Onthophagus) arnoldii Kabakov, 1982 | | | Onthophagus (Onthophagus) barbieri Paulian, 1978 | | | Onthophagus (Onthophagus) dapcauensis Boucomont, 1921 | | | Onthophagus (Onthophagus) dubernardi Boucomont, 1914 | | | Onthophagus (Onthophagus) finschi Harold, 1877 | | | Onthophagus (Onthophagus) fissicornis Stevens, 1809 | | | Onthophagus (Onthophagus) formosanus Gillet, 1924 | | | Onthophagus (Onthophagus) gibbulus Pallas, 1781 | | | Onthophagus (Onthophagus) gorochovi Kabakov, 1998 | | | Onthophagus (Onthophagus) hajimei Masumoto, 1984 | | | Onthophagus (Onthophagus) hastifer Lansberge, 1885 | | | Onthophagus (Onthophagus) hayashii Masumoto, 1991 | | | Onthophagus (Onthophagus) hecate Panzer, 1794 | Scooped Scarab | | Onthophagus (Onthophagus) illyricus Scopoli, 1763 | | | Onthophagus (Onthophagus) inermifrons (d´Orbigny, 1902) | ⇒ Onthophagus margaritifer d´Orbigny, 1898 | | Onthophagus (Onthophagus) ivae Balthasar, 1963 | | | Onthophagus (Onthophagus) kaszabi (Balthasar, 1963) | ⇒ Onthophagus vitulus Péringuey, 1901 | | Onthophagus (Onthophagus) kinhthaicus Kabakov, 1998 | | | Onthophagus (Onthophagus) koryoensis Kim, 1985 | | | Onthophagus (Onthophagus) kryzhanovskii (Kabakov, 1982) | ⇒ Onthophagus zuvandi Qaryagdy, 1939 | | Onthophagus (Onthophagus) kuatunensis Balthasar, 1942 | | | Onthophagus (Onthophagus) lutosopictus Fairmaire, 1897 | | | Onthophagus (Onthophagus) margaritifer d´Orbigny, 1898 | | | Onthophagus (Onthophagus) masumotoi Ochi, 1985 | | | Onthophagus (Onthophagus) mendicus Gillet, 1924 | | | Onthophagus (Onthophagus) mulleri van Lansberge, 1883 | | | Onthophagus (Onthophagus) nasonis Kabakov, 1998 | | | Onthophagus (Onthophagus) nuchicornis Linnaeus, 1758 | | | Onthophagus (Onthophagus) nurestanicus Kabakov, 1982 | | | Onthophagus (Onthophagus) orientalis (Harold, 1868) | ⇒ Onthophagus fissinasus Fairmaire, 1895 | | Onthophagus (Onthophagus) paraprotzenkoi (Tesar, 1967) | ⇒ Onthophagus finschi Harold, 1877 | | Onthophagus (Onthophagus) penicillatus (Harold, 1879) | ⇒ Onthophagus olsoufieffi Boucomont, 1924 | | Onthophagus (Onthophagus) poephagus Kabakov, 1979 | | | Onthophagus (Onthophagus) proletarius Harold, 1875 | | | Onthophagus (Onthophagus) protzenkoi (Tesar, 1967) | ⇒ Onthophagus finschi Harold, 1877 | | Onthophagus (Onthophagus) pseudoarmatus Balthasar, 1944 | | | Onthophagus (Onthophagus) punctator Reitter, 1893 | | | Onthophagus (Onthophagus) rechingeri (Petrovitz, 1980) | ⇒ Onthophagus zuvandi Qaryagdy, 1939 | | Onthophagus (Onthophagus) roubali Balthasar, 1935 | | | Onthophagus (Onthophagus) rufimanus Kabakov, 1982 | | | Onthophagus (Onthophagus) rugulosus Harold, 1886 | | | Onthophagus (Onthophagus) sakainoi Masumoto, 1991 | | | Onthophagus (Onthophagus) sauteri Gillet, 1924 | | | Onthophagus (Onthophagus) setchan Masumoto, 1984 | | | Onthophagus (Onthophagus) sobrius Balthasar, 1959 | | | Onthophagus (Onthophagus) striatulus Palisot de B..., 1809 | Surf blenny | | Onthophagus (Onthophagus) subaeneus (Palisot de B..., 1811) | ⇒ Onthophagus striatulus Palisot de B..., 1809 | | Onthophagus (Onthophagus) tatsienluensis Balthasar, 1942 | | | Onthophagus (Onthophagus) taurus Schreber, 1759 | Bull Headed Dung Beetle | | Onthophagus (Onthophagus) terminatus Eschscholtz, 1822 | | | Onthophagus (Onthophagus) tritinctus Boucomont, 1914 | | | Onthophagus (Onthophagus) trituber Wiedemann, 1823 | | | Onthophagus trituber jacobsoni Kabakov, 2006 | | | Onthophagus (Onthophagus) truncaticornis (Schaller, 1783) | ⇒ Onthophagus quadrinodosus Fahraeus, 1857 | | Onthophagus (Onthophagus) vitulus Péringuey, 1901 | | | Onthophagus (Onthophagus) yangi Masumoto, Ts..., 2006 | | | Onthophagus (Palaeonthophagus) afghanus Petrovitz, 1961 | | | Onthophagus (Palaeonthophagus) albarracinus Baraud, 1979 | | | Onthophagus (Palaeonthophagus) angorensis Petrovitz, 1963 | | | Onthophagus (Palaeonthophagus) baraudi Nicolas, 1964 | | | Onthophagus (Palaeonthophagus) bytinskii Balthasar, 1960 | | | Onthophagus (Palaeonthophagus) clitellifer Reitter, 1894 | | | Onthophagus (Palaeonthophagus) coenobita Herbst, 1783 | | | Onthophagus (Palaeonthophagus) conspersus Reitter, 1893 | | | Onthophagus (Palaeonthophagus) cruciatus (Ménétriés, 1832) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus (Palaeonthophagus) dacatrai Pittino, 2004 | | | Onthophagus (Palaeonthophagus) dellacasai Pittino & Ma..., 1981 | | | Onthophagus (Palaeonthophagus) eulaminicornis Pittino, 2006 | | | Onthophagus (Palaeonthophagus) excisus Reiche & Saulcy, 1856 | | | Onthophagus (Palaeonthophagus) excubitor Ziani & Gudenzi, 2006 | | | Onthophagus (Palaeonthophagus) finschi Harold, 1877 | | | Onthophagus (Palaeonthophagus) fissicornis Steven, 1809 | | | Onthophagus (Palaeonthophagus) fortigibber Reitter, 1909 | | | Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790) | ⇒ Onthophagus similis Scriba, 1790 | | Onthophagus (Palaeonthophagus) furciceps Marseul, 1869 | | | Onthophagus (Palaeonthophagus) gibbulus Pallas, 1781 | | | Onthophagus gibbulus gibbulus | | | Onthophagus gibbulus rostrifer Reitter, 1892 | | | Onthophagus (Palaeonthophagus) gorodinskii Kabakov, 2008 | | | Onthophagus (Palaeonthophagus) grossepunctatus Reitter, 1905 | | | Onthophagus (Palaeonthophagus) haroldi Ballion, 1870 | | | Onthophagus (Palaeonthophagus) hissariensis Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) isikdagensis Pittino, 2004 | | | Onthophagus (Palaeonthophagus) joannae Goljan, 1953 | | | Onthophagus (Palaeonthophagus) kindermanni Harold, 1877 | | | Onthophagus (Palaeonthophagus) kolenatii Reitter, 1893 | | | Onthophagus (Palaeonthophagus) laticornis Gebler, 1823 | | | Onthophagus (Palaeonthophagus) latigena d´Orbigny, 1897 | | | Onthophagus (Palaeonthophagus) lemur Fabricius, 1781 | | | Onthophagus (Palaeonthophagus) leucostigma Steven, 1811 | | | Onthophagus (Palaeonthophagus) lucidus Illiger, 1800 | | | Onthophagus (Palaeonthophagus) macedonicus Miksic, 1959 | | | Onthophagus (Palaeonthophagus) marginalis Gebler, 1817 | | | Onthophagus marginalis marginalis | | | Onthophagus marginalis przewalski Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) massai Baraud, 1975 | | | Onthophagus (Palaeonthophagus) medius (Kugelann, 1792) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus (Palaeonthophagus) medvedevi Kabakov, 1982 | | | Onthophagus (Palaeonthophagus) merdarius Chevrolat, 1865 | | | Onthophagus (Palaeonthophagus) muelleri (Lansberge, 1883) | ⇒ Onthophagus statzi Krell, 1990 | | Onthophagus (Palaeonthophagus) nikolajevi Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) novaki Boucomont, 1927 | | | Onthophagus (Palaeonthophagus) nuchicornis Linnaeus, 1758 | Mottled dung beetle | | Onthophagus (Palaeonthophagus) olsoufieffi Boucomont, 1924 | | | Onthophagus (Palaeonthophagus) opacicollis Reitter, 1893 | | | Onthophagus (Palaeonthophagus) ovatus Linnaeus, 1767 | | | Onthophagus (Palaeonthophagus) panici Petrovitz, 1964 | | | Onthophagus (Palaeonthophagus) parmatus Reitter, 1892 | | | Onthophagus (Palaeonthophagus) petrovitzi (Frey, 1975) | ⇒ Onthophagus petrovitzianus Pittino, 2006 | | Onthophagus (Palaeonthophagus) petrovitzianus Pittino, 2006 | | | Onthophagus (Palaeonthophagus) ponticus Harold, 1883 | | | Onthophagus (Palaeonthophagus) pseudocaccobius Reitter, 1889 | | | Onthophagus (Palaeonthophagus) psychopompus Ziani & Ghar..., 2010 | | | Onthophagus (Palaeonthophagus) pygargus Motschulsky, 1845 | | | Onthophagus (Palaeonthophagus) quadrinodus Reitter, 1896 | | | Onthophagus (Palaeonthophagus) rachelis Martín Piera, 1985 | | | Onthophagus (Palaeonthophagus) rakovici Pittino, 2004 | | | Onthophagus (Palaeonthophagus) rodentium Pittino, 2004 | | | Onthophagus (Palaeonthophagus) ruficapillus Brullé, 1832 | | | Onthophagus ruficapillus guilanensis (Pittino, 1982) | ⇒ Onthophagus ruficapillus Brullé, 1832 | | Onthophagus ruficapillus ruficapillus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) samai Ziani, 2011 | | | Onthophagus (Palaeonthophagus) scabriusculus Harold, 1873 | | | Onthophagus (Palaeonthophagus) semicornis Panzer, 1798 | | | Onthophagus (Palaeonthophagus) sericatus Reitter, 1893 | | | Onthophagus (Palaeonthophagus) shapovalovi Gusakov, 2012 | | | Onthophagus (Palaeonthophagus) similis (Scriba, 1790) | ⇒ Onthophagus fracticornis Preyssler, 1790 | | Onthophagus (Palaeonthophagus) speculifer Solsky, 1876 | | | Onthophagus (Palaeonthophagus) strabo Reitter, 1892 | | | Onthophagus (Palaeonthophagus) stylocerus Graells, 1851 | | | Onthophagus (Palaeonthophagus) suturellus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) tesquorum Semenov & Me..., 1927 | | | Onthophagus (Palaeonthophagus) trigibber Reitter, 1893 | | | Onthophagus (Palaeonthophagus) trispinus Reitter, 1892 | | | Onthophagus (Palaeonthophagus) truchmenus Kolenati, 1846 | | | Onthophagus truchmenus iranicus Kabakov, 2006 | | | Onthophagus truchmenus truchmenus | | | Onthophagus truchmenus turkmeniacus Kabakov, 2006 | | | Onthophagus truchmenus xerxes Petrovitz, 1965 | | | Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767 | | | Onthophagus (Palaeonthophagus) verticicornis (Laicharting, 1781) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus (Palaeonthophagus) viridis Ménétriés, 1832 | | | Onthophagus (Palaeonthophagus) vitulus (Fabricius, 1776) | ⇒ Onthophagus immundus Boheman, 1858 | | Onthophagus (Palaeonthophagus) vlasovi Medvedev, 1958 | | | Onthophagus (Palaeonthophagus) zuninoi Martín Piera, 1985 | | | Onthophagus (Paleonthophagus) andalusicus Waltl, 1835 | | | Onthophagus andalusicus andalusicus | | | Onthophagus andalusicus persianus Olsoufieff, 1900 | | | Onthophagus (Paleonthophagus) fracticornis Preyssler, 1790 | | | Onthophagus (Paleonthophagus) nuchicornis Linnaeus, 1758 | | | Onthophagus (Paleonthophagus) similis Scriba, 1790 | | | Onthophagus (Paleonthophagus) vacca Linnaeus, 1767 | | | Onthophagus (Paraphanaeomorphus) argyropygus Gillet, 1927 | | | Onthophagus (Paraphanaeomorphus) carinensis Boucomont, 1914 | | | Onthophagus (Paraphanaeomorphus) janushevi Kabakov, 1983 | | | Onthophagus (Paraphanaeomorphus) napolovi Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) pseudojavanus Paulian, 1931 | | | Onthophagus (Paraphanaeomorphus) quadricolor Kabakov, 1983 | | | Onthophagus (Paraphanaeomorphus) semipiceus Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) strnadi Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) vaulogeri Boucomont, 1923 | | | Onthophagus (Parascatonomus) carnarius Nomura, 1976 | | | Onthophagus (Parascatonomus) discedens Sharp, 1875 | | | Onthophagus (Parascatonomus) laotianus Boucomont, 1919 | | | Onthophagus (Parascatonomus) miyakei Ochi & Araya, 1992 | | | Onthophagus (Parascatonomus) miyakoinsularis Ochi, Y. Miy..., 1999 | | | Onthophagus (Parascatonomus) murasakianus Nomura, 1976 | | | Onthophagus (Parascatonomus) panfilovi Kabakov, 1983 | | | Onthophagus (Parascatonomus) politus Fabricius, 1798 | | | Onthophagus (Parascatonomus) tricornis (Wiedemann, 1823) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus (Parentius) atricapillus d´Orbigny, 1908 | | | Onthophagus (Parentius) emarginatus Mulsant, 1842 | | | Onthophagus (Parentius) hispanicus Baraud, 1963 | | | Onthophagus (Parentius) kabakovi Martin-Piera, 1985 | | | Onthophagus (Parentius) punctatus Illiger, 1803 | | | Onthophagus (Phalops) tricuspis (Semenov, 1900) | ⇒ Onthophagus fortigibber Reitter, 1909 | | Onthophagus (Phanaeomorphus) ater Waterhouse, 1875 | | | Onthophagus (Phanaeomorphus) cernyi (Balthasar, 1935) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus (Phanaeomorphus) lenzi Harold, 1875 | | | Onthophagus potanini hiurai Ochi, 1984 | | | Onthophagus (Proagoderus) quadriarmatus Fairmaire, 1892 | | | Onthophagus (Relictonthophagus) emarginatus (Mulsant, 1842) | ⇒ Onthophagus rubrimaculatus MacLeay, 1864 | | Onthophagus (Relictonthophagus) nigellus (Illiger, 1803) | ⇒ Caccobius histeroides Ménétriés, 1832 | | Onthophagus (Relictonthophagus) punctatus (Illiger, 1803) | ⇒ Onthophagus emarginatus Mulsant, 1842 | | Onthophagus (Serrophorus) avocettoides Kabakov, 1983 | | | Onthophagus (Serrophorus) laevis (Harold, 1880) | ⇒ Onthophagus laevissimus d´Orbigny, 1905 | | Onthophagus laevis asiaticus Boucomont, 1919 | | | Onthophagus laevis laevis | | | Onthophagus laevis lampromelas Fairmaire, 1891 | | | Onthophagus laevis stevensi Arrow, 1931 | | | Onthophagus (Serrophorus) oblongus Zhang, 1997 | | | Onthophagus (Serrophorus) rectecornutus Lansberge, 1883 | | | Onthophagus (Serrophorus) sagittarius Fabricius, 1775 | Sagittarius dung beetle | | Onthophagus (Strandius) changshouensis Zhang, 1997 | | | Onthophagus (Strandius) hiurai (Ochi, 1984) | ⇒ Onthophagus potanini hiurai Ochi, 1984 | | Onthophagus (Strandius) japonicus Harold, 1875 | | | Onthophagus (Strandius) kuraruanus (Matsumura, 1938) | ⇒ Onthophagus lenzi Harold, 1875 | | Onthophagus (Strandius) obliquus Olivier, 1789 | | | Onthophagus (Strandius) potanini R. C. Brusca..., 1990 | | | Onthophagus potanini hiurai Ochi, 1984 | | | Onthophagus potanini potanini | | | Onthophagus (Strandius) subansiriensis Biswas, 1979 | | | Onthophagus (Strandius) taiwanus Nomura, 1973 | | | Onthophagus (Strandius) yakushimanus Nakane, 1983 | | | Onthophagus (Trichonthophagus) dynastoides Arrow, 1931 | | | Onthophagus (Trichonthophagus) hirtus Illiger, 1803 | | | Onthophagus (Trichonthophagus) maki Illiger, 1803 | | | Onthophagus (Trichonthophagus) tarandus Fabricius, 1792 | | | Onthophagus abreui Arrow, 1931 | | | Onthophagus acornis Geoffroy, 1785 | | | Onthophagus acrisius Balthasar, 1946 | | | Onthophagus actaeon Balthasar, 1946 | | | Onthophagus acutus d´Orbigny, 1908 | | | Onthophagus aeneopiceus d´Orbigny, 1902 | | | Onthophagus aeneus A. G. Olivier, 1789 | | | Onthophagus aequiperus Matsumura & ..., 1937 | | | Onthophagus aerarius Reitter, 1892 | | | Onthophagus aeruginosus Roth, 1851 | | | Onthophagus affinis Gillet, 1930 | | | Onthophagus agaricophilus Arrow, 1931 | | | Onthophagus agilis (Matsumura, 1938) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus akinini Koenig, 1889 | | | Onthophagus akirai (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus albinae (Reitter, 1898) | ⇒ Onthophagus formaneki Reitter, 1897 | | Onthophagus aleppensis Redtenbacher, 1843 | | | Onthophagus aleppensis aleppensis | | | Onthophagus aleppensis hyrcanicus Kabakov, 1982 | | | Onthophagus alpinus (Kolenati, 1846) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus alternatus (d´Orbigny, 1902) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus amamiensis Nomura, 1965 | | | Onthophagus amamiensis amamiensis | | | Onthophagus amamiensis matsumurai Ochi, Kawaha..., 2019 | | | Onthophagus amenthus Matsumura, 1937 | | | Onthophagus amphioxus Arrow, 1933 | | | Onthophagus amplexus Sharp, 1875 | | | Onthophagus amycoides Kabakov, 2014 | | | Onthophagus analis (Lucas, 1846) | ⇒ Onthophagus sellatus Klug, 1845 | | Onthophagus anatolicus Petrovitz, 1962 | | | Onthophagus andreji Prokofiev, 2014 | | | Onthophagus angulatus (Redtenbacher, 1848) | ⇒ Onthophagus gagates Hope, 1831 | | Onthophagus angulicornis (Mulsant, 1842) | ⇒ Onthophagus semicornis Panzer, 1798 | | Onthophagus anonymus Delabie, 1956 | | | Onthophagus antilope (Motschulsky, 1845) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus aokii Nomura, 1976 | | | Onthophagus araxicola (Reitter, 1893) | ⇒ Onthophagus truchmenus Kolenati, 1846 | | Onthophagus arboreus Arrow, 1931 | | | Onthophagus arcifer d´Orbigny, 1904 | | | Onthophagus armatus Blanchard, 1853 | | | Onthophagus arunachalensis Biswas & Cha..., 1985 | | | Onthophagus asiaticus Endrödi, 1973 | | | Onthophagus asper MacLeay, 1864 | | | Onthophagus asperrimus d´Orbigny, 1902 | | | Onthophagus assimilis Hoppe, 1795 | | | Onthophagus athenae (Goidanich, 1925) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus atropolitus d´Orbigny, 1902 | | | Onthophagus australis Guérin-Ménev..., 1830 | Southern dung beetle | | Onthophagus austriacus (Panzer, 1793) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus aviculus Arrow, 1931 | | | Onthophagus avocetta Arrow, 1933 | | | Onthophagus balawaicus Scheuern, 1995 | | | Onthophagus balthasari Vsetecka, 1939 | | | Onthophagus basalis (Mulsant, 1842) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus basicruentatus Goidanich, 1926 | | | Onthophagus basipustulatus Heyden, 1879 | | | Onthophagus beesoni Arrow, 1931 | | | Onthophagus bengalensis Harold, 1886 | | | Onthophagus bengali Gordon & Opp..., 1977 | | | Onthophagus biarcuatus (Lansberge, 1882) | ⇒ Onthophagus sticticus Harold, 1867 | | Onthophagus bicolor (Raffray, 1877) | ⇒ Onthophagus flavolimbatus Klug, 1855 | | Onthophagus bicornutus Mulsant, 1842 | | | Onthophagus bicristatus d´Orbigny, 1905 | | | Onthophagus bidentarus Mulsant, 1842 | | | Onthophagus biexcavatus (d´Orbigny, 1898) | ⇒ Onthophagus tarandus Fabricius, 1792 | | Onthophagus bifasciatus Fabricius, 1781 | | | Onthophagus bifenestratus (Reitter, 1898) | ⇒ Onthophagus formaneki Reitter, 1897 | | Onthophagus bifossus (d´Orbigny, 1902) | ⇒ Onthophagus spinifex Fabricius, 1781 | | Onthophagus birmanicus (Harold, 1879) | ⇒ Onthophagus bifasciatus Fabricius, 1781 | | Onthophagus bison Boucomont, 1919 | | | Onthophagus bituberculatus (Olivier, 1789) | ⇒ Onthophagus acutus d´Orbigny, 1908 | | Onthophagus bivertex Heyden, 1887 | | | Onthophagus blanchardi Harold, 1869 | | | Onthophagus bonsae Zunino, 1976 | | | Onthophagus bos (Villa, 1833) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus boucomontianus Balthasar, 1935 | | | Onthophagus bourgognei Paulian, 1945 | | | Onthophagus bovillus (Mulsant, 1842) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus brevicornis Fahraeus, 1857 | | | Onthophagus brevipes Herbst, 1783 | | | Onthophagus brisouti (d´Orbigny, 1897) | ⇒ Onthophagus illyricus Scopoli, 1763 | | Onthophagus brittoni Paulian, 1948 | | | Onthophagus brutus Arrow, 1931 | | | Onthophagus bufo (Arrow, 1907) | ⇒ Onthophagus semicinctus d´Orbigny, 1897 | | Onthophagus cameloides d´Orbigny, 1900 | | | Onthophagus camelus Fabricius, 1787 | | | Onthophagus campestris Masumoto, Ki..., 2018 | | | Onthophagus capella (Kirby, 1818) | ⇒ Onthophagus ramosellus Bates, 1891 | | Onthophagus capilossus d´Orbigny, 1905 | | | Onthophagus capitosus Harold, 1867 | | | Onthophagus capra Fabricius, 1787 | | | Onthophagus capreolus (Mulsant, 1842) | ⇒ Onthophagus finschi Harold, 1877 | | Onthophagus carbunculus (Boucomont, 1921) | ⇒ Onthophagus lilliputanus Lansberge, 1883 | | Onthophagus carpanetoi Pittino, 1982 | | | Onthophagus castanonotus Seabra, 1907 | | | Onthophagus centricornis Fabricius, 1798 | | | Onthophagus centromaculatus Redtenbacher, 1843 | | | Onthophagus cervus (Fabricius, 1798) | ⇒ Onthophagus falsus Gillet, 1925 | | Onthophagus chibanus Matsumura & ..., 1937 | | | Onthophagus chosensis Matsumura, 1937 | | | Onthophagus chrysurus Arrow, 1931 | | | Onthophagus chuzenjianus (Matsumura & ..., 1937) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus cicatricosus (Balthasar, 1935) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus cinctus (Dallatorre, 1879) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus circulator Reitter, 1891 | | | Onthophagus circulifer Arrow, 1931 | | | Onthophagus circumcinctus (Escalera, 1914) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus circumscriptus (Faldermann, 1835) | ⇒ Onthophagus marginalis Gebler, 1817 | | Onthophagus citellorum (Petrovitz, 1967) | ⇒ Onthophagus vitulus Péringuey, 1901 | | Onthophagus clermonti Paulian, 1931 | | | Onthophagus cognatus Boucomont, 1921 | | | Onthophagus collaris (Kabakov, 1983) | ⇒ Proagoderus loricatus Klug, 1855 | | Onthophagus commottoides Kabakov, 1998 | | | Onthophagus compactus Arrow, 1933 | | | Onthophagus concolor Sharp, 1878 | | | Onthophagus concursator (Balthasar, 1960) | ⇒ Onthophagus sutleinensis Splichal, 1910 | | Onthophagus confluens (d´Orbigny, 1902) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus confucius (Goidanich, 1926) | ⇒ Onthophagus solivagus Harold, 1886 | | Onthophagus conjugata (Seabra, 1907) | ⇒ Onthophagus hirtus Illiger, 1803 | | Onthophagus conspurcatus Geoffroy, 1785 | | | Onthophagus convexicollis Boheman, 1858 | | | Onthophagus coomani Paulian, 1931 | | | Onthophagus coracinoides Kabakov, 1983 | | | Onthophagus coracinus Boucomont, 1914 | | | Onthophagus cornicollis d´Orbigny, 1902 | | | Onthophagus corniger Geoffroy, 1785 | | | Onthophagus corvus Fabricius, 1798 | | | Onthophagus crassicollis Boucomont, 1913 | | | Onthophagus crassus ♰ (Heer, 1862) | ⇒ Parascatonomus tricornis Wiedemann, 1823 | | Onthophagus cruoreus Schrank, 1798 | | | Onthophagus cultrifer (Harold, 1886) | ⇒ Onthophagus falcifer Harold, 1880 | | Onthophagus cupreiceps Arrow, 1907 | | | Onthophagus cupreolus (Reitter, 1894) | ⇒ Onthophagus punctatus Illiger, 1803 | | Onthophagus curvicinctus (Mulsant, 1842) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus curvispina Reitter, 1893 | | | Onthophagus cuspidiusculus (Mulsant, 1842) | ⇒ Onthophagus coenobita Herbst, 1783 | | Onthophagus cutai (Roubal, 1933) | ⇒ Onthophagus ruficapillus Brullé, 1832 | | Onthophagus cyaneus Kabakov, 2006 | | | Onthophagus decempunctatus Schaller, 1783 | | | Onthophagus decipiens (Mulsant, 1842) | ⇒ Onthophagus semicornis Panzer, 1798 | | Onthophagus deflexicollis Lansberge, 1883 | | | Onthophagus degener Mulsant, 1842 | | | Onthophagus deletus (Reitter, 1893) | ⇒ Onthophagus sibiricus Harold, 1877 | | Onthophagus delkeskampi (Frey, 1954) | ⇒ Onthophagus mirandus Arrow, 1931 | | Onthophagus depolii (Goidanich, 1925) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus difficilis (Le Guillou, 1844) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus digitatus Arrow, 1931 | | | Onthophagus dillwynii (Stephens, 1830) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus discoideus A. G. Olivier, 1789 | | | Onthophagus dissentaneus Balthasar, 1964 | | | Onthophagus distinctus (Fuente, 1910) | ⇒ Onthophagus latigena d´Orbigny, 1897 | | Onthophagus distinguendus Mulsant, 1842 | | | Onthophagus diversicornis (Kirschenblatt, 1935) | ⇒ Onthophagus aerarius Reitter, 1892 | | Onthophagus dorsofasciatus Fairmaire, 1893 | | | Onthophagus dorsosignatus d´Orbigny, 1898 | | | Onthophagus egenus (Harold, 1877) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus egurianus Matsumura, 1938 | | | Onthophagus ensifer Boucomont, 1914 | | | Onthophagus erectus Wiedemann, 1819 | | | Onthophagus euthymi (Mulsant & Go..., 1875) | ⇒ Onthophagus sellatus Klug, 1845 | | Onthophagus expansicollis (Fairmaire, 1891) | ⇒ Onthophagus rugulosus Harold, 1886 | | Onthophagus expansicornis Bates, 1891 | | | Onthophagus exquisitus Arrow, 1931 | | | Onthophagus fabricii Waterhouse, 1894 | | | Onthophagus falcifer Harold, 1880 | | | Onthophagus falsus Gillet, 1925 | | | Onthophagus fanatus Laporte, 1840 | | | Onthophagus fasciatus (Boucomont, 1914) | ⇒ Onthophagus fasciolatus Boucomont, 1924 | | Onthophagus fasciolatus Boucomont, 1924 | | | Onthophagus femineus (Mulsant, 1842) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus filicornis Harold, 1873 | | | Onthophagus fissinasus Fairmaire, 1895 | | | Onthophagus flagrans Reitter, 1893 | | | Onthophagus flavescens Seabra, 1907 | | | Onthophagus flavofasciatus (d´Orbigny, 1900) | ⇒ Onthophagus dorsosignatus d´Orbigny, 1898 | | Onthophagus flavolimbatus Klug, 1855 | | | Onthophagus flavonotatus d´Orbigny, 1900 | | | Onthophagus fodiens Waterhouse, 1875 | | | Onthophagus formaneki Reitter, 1897 | | | Onthophagus fossatus Castelnau, 1840 | | | Onthophagus fossor Arrow, 1931 | | | Onthophagus foveolatus (Harold, 1877) | ⇒ Parascatonomus rudis Sharp, 1875 | | Onthophagus frontalis Raffray, 1877 | | | Onthophagus frugivorus Arrow, 1931 | | | Onthophagus fucatrus (Mulsant, 1842) | ⇒ Onthophagus ovatus Linnaeus, 1767 | | Onthophagus fukiensis Balthasar, 1942 | | | Onthophagus fulgens Brahm, 1790 | | | Onthophagus fulvicornis (Motschulsky, 1853) | ⇒ Onthophagus tragus Fabricius, 1792 | | Onthophagus fulvipes (Faldermann, 1835) | ⇒ Onthophagus coenobita Herbst, 1783 | | Onthophagus funebris Boucomont, 1919 | | | Onthophagus furcaticeps Masters, 1886 | | | Onthophagus furcatoides (Lansberge, 1886) | ⇒ Onthophagus neofurcatus Goidanich, 1926 | | Onthophagus furcicollis (Arrow, 1931) | ⇒ Onthophagus hecate Panzer, 1794 | | Onthophagus furcicornis (Reitter, 1893) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus furcillifer Bates, 1891 | | | Onthophagus fuscipennis (Mulsant, 1842) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus fuscopunctatus (Fabricius, 1798) | ⇒ Onthophagus luridipennis Boheman, 1858 | | Onthophagus fuscopunctulatus (Boucomont, 1914) | ⇒ Onthophagus luridipennis Boheman, 1858 | | Onthophagus futsukaichianus Matsumura, 1937 | | | Onthophagus gagates Hope, 1831 | | | Onthophagus gagatoides Kabakov, 2006 | | | Onthophagus gangeticus Gillet, 1925 | | | Onthophagus germanus Gillet, 1927 | | | Onthophagus germari (Depoli, 1917) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus ginyunensis Vsetecka, 1942 | | | Onthophagus glabra (Seabra, 1907) | ⇒ Onthophagus maki Illiger, 1803 | | Onthophagus glandicolor (Mulsant, 1842) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus glareanus (Gistel, 1857) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus glasunovi Koshantschikov, 1894 | | | Onthophagus glasunowi D. Koshantsc..., 1894 | | | Onthophagus gorhami (Shipp, 1895) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus gracilicornis (Raffray, 1877) | ⇒ Onthophagus filicornis Harold, 1873 | | Onthophagus granulifer (Harold, 1886) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus granulosus (Koshantschikov, 1894) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus gratus Arrow, 1931 | | | Onthophagus griseosetosus Arrow, 1931 | | | Onthophagus guilanensis (Pittino, 1982) | ⇒ Onthophagus ruficapillus Brullé, 1832 | | Onthophagus hamaticeps Arrow, 1931 | | | Onthophagus herbsti Brahm, 1790 | | | Onthophagus hermonensis Baraud, 1982 | | | Onthophagus heydeni Harold, 1875 | | | Onthophagus heyrovskyi Vsetecka, 1943 | | | Onthophagus hikosanus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus hindu Arrow, 1931 | | | Onthophagus hingstoni Arrow, 1931 | | | Onthophagus hirtulus (Reitter, 1893) | ⇒ Onthophagus hirtus Illiger, 1803 | | Onthophagus horni Balthasar, 1935 | | | Onthophagus hrubanti (Balthasar, 1955) | ⇒ Onthophagus ramosellus Bates, 1891 | | Onthophagus humeralis (Miksic, 1950) | ⇒ Onthophagus vitulus Péringuey, 1901 | | Onthophagus hvangeus Stebnicka, 1973 | | | Onthophagus hypertropis (Olsoufieff, 1900) | ⇒ Onthophagus tragus Fabricius, 1792 | | Onthophagus hyrcanius Kabakov, 1982 | | | Onthophagus hystrix Boucomont, 1914 | | | Onthophagus hyuganus Matsumura & ..., 1937 | | | Onthophagus ibex (Fabricius, 1792) | ⇒ Onthophagus tragus Fabricius, 1792 | | Onthophagus ibonus (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus ibrydus O. G. Costa, 1828 | | | Onthophagus immaculatus (Mulsant, 1842) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus immundus Boheman, 1858 | | | Onthophagus impuncticollis (Frey, 1950) | ⇒ Onthophagus atropolitus d´Orbigny, 1902 | | Onthophagus incertus (d´Orbigny, 1897) | ⇒ Onthophagus asperrimus d´Orbigny, 1902 | | Onthophagus incollaris Kabakov, 2008 | | | Onthophagus indistinctus (Mulsant, 1842) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus indosinicus Boucomont, 1921 | | | Onthophagus infucatus Harold, 1877 | | | Onthophagus infuscata (Seabra, 1907) | ⇒ Onthophagus hirtus Illiger, 1803 | | Onthophagus infuscatus Klug, 1845 | | | Onthophagus inquinatus (Goidanich, 1925) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus intercepta (Seabra, 1907) | ⇒ Onthophagus maki Illiger, 1803 | | Onthophagus intermedius (Mulsant, 1842) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus irroratus Faldermann, 1835 | | | Onthophagus italicus (Goidanich, 1925) | ⇒ Onthophagus andalusicus Waltl, 1835 | | Onthophagus itoi Nomura, 1976 | | | Onthophagus javanensis Balthasar, 1963 | | | Onthophagus javanus (Lansberge, 1883) | ⇒ Onthophagus javanensis Balthasar, 1963 | | Onthophagus jeannelianus Paulian, 1945 | | | Onthophagus jeannelianus shokhini Kabakov, 2014 | | | Onthophagus jedensis Matsumura, 1937 | | | Onthophagus jiupengensis Masumoto, La..., 2017 | | | Onthophagus jononis Matsumura, 1936 | | | Onthophagus jucundus (Arrow, 1931) | ⇒ Onthophagus cognatus Boucomont, 1921 | | Onthophagus jugicola d´Orbigny, 1902 | | | Onthophagus kabulicus (Balthasar, 1971) | ⇒ Onthophagus conspersus Reitter, 1893 | | Onthophagus kandai (Matsumura, 1937) | ⇒ Onthophagus olsoufieffi Boucomont, 1924 | | Onthophagus kashmirensis Balthasar, 1966 | | | Onthophagus kawarinus (Matsumura & ..., 1937) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus kirgisicus (Protzenko, 1963) | ⇒ Onthophagus silus Balthasar, 1960 | | Onthophagus klapperichi Balthasar, 1952 | | | Onthophagus kleinei Balthasar, 1935 | | | Onthophagus kogatanus (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus koma Matsumura, 1937 | | | Onthophagus komabellus Matsumura & ..., 1937 | | | Onthophagus komareki Balthasar, 1935 | | | Onthophagus kontumicus Kabakov, 1983 | | | Onthophagus kozunonis (Matsumura, 1937) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus krali (Balthasar, 1963) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus kulti Balthasar, 1952 | | | Onthophagus kuluensis Bates, 1891 | | | Onthophagus kumaonensis Arrow, 1931 | | | Onthophagus laevicollis (Lansberge, 1883) | ⇒ Parascatonomus laevicollis Lansberge, 1883 | | Onthophagus laevigatus Fabricius, 1798 | | | Onthophagus laevissimus d´Orbigny, 1905 | | | Onthophagus lahorensis Kabakov, 2008 | | | Onthophagus lamellatus (Boucomont, 1924) | ⇒ Onthophagus lunatus Harold, 1868 | | Onthophagus laminatus (MacLeay, 1864) | ⇒ Onthophagus lunatus Harold, 1868 | | Onthophagus laminiger Mulsant, 1842 | | | Onthophagus lampromelas (Fairmaire, 1891) | ⇒ Onthophagus laevis lampromelas Fairmaire, 1891 | | Onthophagus lamyi Paulian, 1945 | | | Onthophagus lapillus Arrow, 1931 | | | Onthophagus latipennis d´Orbigny, 1897 | | | Onthophagus latiplaga (Reitter, 1897) | ⇒ Onthophagus truchmenus Kolenati, 1846 | | Onthophagus lemuroides d´Orbigny, 1898 | | | Onthophagus lenzii Harold, 1874 | | | Onthophagus leucomelas (Solsky, 1876) | ⇒ Onthophagus leucostigma Steven, 1811 | | Onthophagus lineatus (Mulsant, 1842) | ⇒ Onthophagus transcaspicus Koenig, 1888 | | Onthophagus lineolatus (Mulsant, 1842) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus liothorax Koshantschikov, 1894 | | | Onthophagus liui Masumoto, Oc..., 2014 | | | Onthophagus loricatus Klug, 1855 | | | Onthophagus lunatus Harold, 1868 | | | Onthophagus luridus (Paulian, 1933) | ⇒ Onthophagus rectecornutus Lansberge, 1883 | | Onthophagus lusitanica (Seabra, 1907) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus luteipennis Wiedemann, 1823 | | | Onthophagus luzonicus (Lansberge, 1883) | ⇒ Onthophagus armatus Blanchard, 1853 | | Onthophagus magnini Paulian, 1933 | | | Onthophagus mandarinus (Harold, 1859) | ⇒ Onthophagus tragus Fabricius, 1792 | | Onthophagus marani (Vsetecka, 1942) | ⇒ Parascatonomus acuticollis Gillet, 1927 | | Onthophagus marginatus (Castelnau, 1840) | ⇒ Onthophagus andalusicus Waltl, 1835 | | Onthophagus marginithorax (Vsetecka, 1939) | ⇒ Onthophagus lenzi Harold, 1875 | | Onthophagus marmoratus (Ménétriés, 1832) | ⇒ Onthophagus marginalis Gebler, 1817 | | Onthophagus marxi (Splichal, 1910) | ⇒ Onthophagus sutleinensis Splichal, 1910 | | Onthophagus matsuokai (Matsumura & ..., 1937) | ⇒ Onthophagus sauteri Gillet, 1924 | | Onthophagus mauritii Boucomont, 1919 | | | Onthophagus maurus (Lucas, 1846) | ⇒ Onthophagus andalusicus Waltl, 1835 | | Onthophagus mediofasciatus (d´Orbigny, 1900) | ⇒ Onthophagus nitidulus Klug, 1845 | | Onthophagus melanocephalus Klug, 1845 | | | Onthophagus mendax (Mulsant, 1842) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus menetriesii (Faldermann, 1835) | ⇒ Onthophagus illyricus Scopoli, 1763 | | Onthophagus micellus Matsumura, 1937 | | | Onthophagus midorianus (Matsumura, 1938) | ⇒ Parascatonomus anguicorius Boucomont, 1914 | | Onthophagus millingeni d´Orbigny, 1900 | | | Onthophagus minoi Matsumura & ..., 1937 | | | Onthophagus minokuchianus (Matsumura, 1934) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus minutus (Hausmann, 1807) | ⇒ Onthophagus centricornis Fabricius, 1798 | | Onthophagus mirandus Arrow, 1931 | | | Onthophagus missor (Balthasar, 1939) | ⇒ Onthophagus carinensis Boucomont, 1914 | | Onthophagus misujianus Matsumura & ..., 1937 | | | Onthophagus miyabei (Matsumura, 1937) | ⇒ Onthophagus punctator Reitter, 1893 | | Onthophagus miyazakianus Matsumura & ..., 1937 | | | Onthophagus mniszekii Hochhuth, 1873 | | | Onthophagus moerens Walker, 1858 | | | Onthophagus mopsus Fabricius, 1792 | | | Onthophagus moreleti Baraud, 1980 | | | Onthophagus morio (Brullé, 1832) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus mushana Matsumura, 1938 | | | Onthophagus mushensis Matsumura, 1938 | | | Onthophagus mutabilis (Mulsant, 1842) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus nagasawai Matsumura, 1938 | | | Onthophagus nakatomii (Matsumura, 1937) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus nanus (Harold, 1878) | ⇒ Onthophagus waterhousei Gillet, 1927 | | Onthophagus narashinensis Matsumura & ..., 1936 | | | Onthophagus nasalis Arrow, 1931 | | | Onthophagus nasutus Guérin-Ménev..., 1855 | | | Onthophagus naviculifer Boucomont, 1919 | | | Onthophagus nebulosus Reiche, 1864 | | | Onthophagus necessarius Reitter, 1893 | | | Onthophagus necrophagus Arrow, 1931 | | | Onthophagus nefarius Balthasar, 1941 | | | Onthophagus neofurcatus Goidanich, 1926 | | | Onthophagus neptunulus (Lansberge, 1883) | ⇒ Onthophagus tragus Fabricius, 1792 | | Onthophagus nigrimargo (Goidanich, 1926) | ⇒ Onthophagus marginalis Gebler, 1817 | | Onthophagus nigrovirescens (Mulsant, 1842) | ⇒ Euonthophagus amyntas Olivier, 1789 | | Onthophagus nitidicollis (Brullé, 1832) | ⇒ Onthophagus lucidus Illiger, 1800 | | Onthophagus nitidulus Klug, 1845 | | | Onthophagus nitidus Waterhouse, 1875 | | | Onthophagus noctivagus (Harold, 1877) | ⇒ Onthophagus reticulatus d´Orbigny, 1902 | | Onthophagus nocturnus (Reitter, 1893) | ⇒ Onthophagus sibiricus Harold, 1877 | | Onthophagus novemmaculatus Gmelin, 1788 | | | Onthophagus nuchidens Fabricius, 1798 | | | Onthophagus numidicus d´Orbigny, 1908 | | | Onthophagus nutans (Roth, 1851) | ⇒ Onthophagus schaufussi Harold, 1867 | | Onthophagus oberthueri d´Orbigny, 1898 | | | Onthophagus oberthuri d´Orbigny, 1898 | | | Onthophagus obesus (Fahraeus, 1857) | ⇒ Onthophagus vinctus Erichson, 1843 | | Onthophagus obliviosus Balthasar, 1935 | | | Onthophagus obscuratus (Heyden, 1890) | ⇒ Onthophagus maki Illiger, 1803 | | Onthophagus obsoletus (Fairmaire, 1875) | ⇒ Onthophagus transcaspicus Koenig, 1888 | | Onthophagus obtusicornis (Fahraeus, 1857) | ⇒ Onthophagus marginalis przewalski Kabakov, 2006 | | Onthophagus obtusus Wiedemann, 1819 | | | Onthophagus ocellatopunctatus Waterhouse, 1875 | | | Onthophagus ochreatus d´Orbigny, 1897 | | | Onthophagus oculatus Arrow, 1931 | | | Onthophagus ohbayashii Nomura, 1939 | | | Onthophagus oishii Matsumura & ..., 1937 | | | Onthophagus okinawanus Nomura, 1976 | | | Onthophagus okushirianus Matsumura, 1937 | | | Onthophagus oniellus (Matsumura, 1937) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus orbus Boucomont, 1924 | | | Onthophagus oryx Fabricius, 1792 | | | Onthophagus osellai Pittino, 1982 | | | Onthophagus oshimanus Nakane, 1960 | | | Onthophagus pacificus Lansberge, 1885 | | | Onthophagus pactolus Fabricius, 1787 | | | Onthophagus parvulus (Fabricius, 1798) | ⇒ Onthophagus frontalis Raffray, 1877 | | Onthophagus pauperatus Mulsant, 1842 | | | Onthophagus peguanus (Boucomont, 1914) | ⇒ Onthophagus pacificus Lansberge, 1885 | | Onthophagus persianus (Olsoufieff, 1900) | ⇒ Onthophagus andalusicus persianus Olsoufieff, 1900 | | Onthophagus phanaeiformis Boucomont, 1914 | | | Onthophagus pictus (Reitter, 1893) | ⇒ Oniticellus tessellatus Harold, 1879 | | Onthophagus piffli Petrovitz, 1961 | | | Onthophagus piliger (Mulsant, 1842) | ⇒ Onthophagus illyricus Scopoli, 1763 | | Onthophagus planicornis Herbst, 1789 | | | Onthophagus planifrons Frey, 1962 | | | Onthophagus pljushtchi Ivanova, 2012 | | | Onthophagus plurimacula (Depoli, 1924) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus poeophagus Kabakov, 1979 | | | Onthophagus porcus Arrow, 1931 | | | Onthophagus procurvus Balthasar, 1935 | | | Onthophagus productus Arrow, 1907 | | | Onthophagus prolixus (Walker, 1858) | ⇒ Onthophagus unifasciatus Schaller, 1783 | | Onthophagus propinquus (MacLeay, 1888) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus pseudobrutus Kabakov, 1983 | | | Onthophagus pseudoconvexicollis Bai & Yang, 2016 | | | Onthophagus pseudocoracinus Kabakov, 1983 | | | Onthophagus pseudojaponicus Balthasar, 1941 | | | Onthophagus pseudojaponicus fukiensis Balthasar, 1941 | | | Onthophagus pseudojaponicus pseudojaponicus | | | Onthophagus puetalis Matsumura, 1938 | | | Onthophagus pumilus Kabakov, 2014 | | | Onthophagus punctatominiatus (Goidanich, 1925) | ⇒ Onthophagus lucidus Illiger, 1800 | | Onthophagus pupillatus Kolbe, 1886 | | | Onthophagus purpurascens Boucomont, 1914 | | | Onthophagus putealis (Matsumura, 1938) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus pygialis (Thomson, 1858) | ⇒ Onthophagus sellatus Klug, 1845 | | Onthophagus quadricornis Fabricius, 1801 | | | Onthophagus quadrinodosus Fahraeus, 1857 | | | Onthophagus quadrituberculatus Laicharting, 1781 | | | Onthophagus quadrum Kugelann, 1792 | | | Onthophagus ramosellus Bates, 1891 | | | Onthophagus ramosus Wiedemann, 1823 | | | Onthophagus rana Arrow, 1931 | | | Onthophagus recticornis Leske, 1785 | | | Onthophagus rectinicornis (Fairmaire, 1878) | ⇒ Parascatonomus tricornis Wiedemann, 1823 | | Onthophagus reflexicornis (Redtenbacher, 1867) | ⇒ Onthophagus spinifex Fabricius, 1781 | | Onthophagus reichardti (Medvedev, 1928) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus remotus Harold, 1879 | | | Onthophagus reticulatus d´Orbigny, 1902 | | | Onthophagus rhinoceros (Melsheimer, 1845) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus roessneri Ziani, 2016 | | | Onthophagus rostrifer (Reitter, 1893) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus royi Biswas & Cha..., 1985 | | | Onthophagus rubellus d´Orbigny, 1908 | | | Onthophagus rubrescens (Blanchard, 1843) | ⇒ Onthophagus stylocerus Graells, 1851 | | Onthophagus rubrimaculatus MacLeay, 1864 | | | Onthophagus rubripennis (Arrow, 1907) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus rubripes (Mulsant, 1842) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus rubromaculatus Kolbe, 1886 | | | Onthophagus rubrotinctus (d´Orbigny, 1900) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus rudis Sharp, 1875 | | | Onthophagus rufipes (Ménétriés, 1832) | ⇒ Onthophagus illyricus Scopoli, 1763 | | Onthophagus rugicollis Harold, 1880 | | | Onthophagus rugosus (Kirby, 1818) | ⇒ Onthophagus tenebrosus Harold, 1871 | | Onthophagus rugulipennis Fairmaire, 1887 | | | Onthophagus rutilipennis Reitter, 1893 | | | Onthophagus sacharovskii Olsoufieff, 1918 | | | Onthophagus saharae (Bedel, 1911) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus saiga (Ballion, 1870) | ⇒ Onthophagus pygargus Motschulsky, 1845 | | Onthophagus saigonensis Boucomont, 1924 | | | Onthophagus sakishimanus Nomura, 1976 | | | Onthophagus saudiensis Frey, 1962 | | | Onthophagus scaber Roth, 1851 | | | Onthophagus schaefernai Balthasar, 1935 | | | Onthophagus schatzmayri (Pierotti, 1959) | ⇒ Onthophagus opacicollis Reitter, 1893 | | Onthophagus schaufussi Harold, 1867 | | | Onthophagus schawalleri Scheuern, 1996 | | | Onthophagus secundarius Roth, 1851 | | | Onthophagus sellatus Klug, 1845 | | | Onthophagus semicinctus d´Orbigny, 1897 | | | Onthophagus semiflavus (Boheman, 1860) | ⇒ Onthophagus flavolimbatus Klug, 1855 | | Onthophagus senescens Péringuey, 1908 | | | Onthophagus senex Boucomont, 1914 | | | Onthophagus seniculus (Fabricius, 1781) | ⇒ Onthophagus senescens Péringuey, 1908 | | Onthophagus shaykh Montanaro & ..., 2022 | | | Onthophagus shibatai Nakane, 1960 | | | Onthophagus shigeoi Matsumura, 1937 | | | Onthophagus shinanensis (Matsumura, 1934) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus shirakii Nakane, 1960 | | | Onthophagus shoyozanus (Matsumura, 1937) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus shurianus Matsumura & ..., 1937 | | | Onthophagus sikkimensis Gillet, 1925 | | | Onthophagus silus Balthasar, 1960 | | | Onthophagus simius Reitter, 1893 | | | Onthophagus simplicifrons Reitter, 1893 | | | Onthophagus sinicus Zhang & Wang, 1997 | | | Onthophagus smeei (Harold, 1875) | ⇒ Proagoderus amplexus Sharp, 1875 | | Onthophagus smetanai Balthasar, 1952 | | | Onthophagus sobosanus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus solidus (Gillet, 1927) | ⇒ Onthophagus compactus Arrow, 1933 | | Onthophagus solivagus Harold, 1886 | | | Onthophagus sonani Miwa, 1930 | | | Onthophagus songsokensis Biswas & Cha..., 1985 | | | Onthophagus sparsulus Reitter, 1893 | | | Onthophagus spinifer A. G. Olivier, 1789 | | | Onthophagus spinifex Fabricius, 1781 | | | Onthophagus spitiensis (Gillet, 1925) | ⇒ Onthophagus troglodyta Wiedemann, 1823 | | Onthophagus spurius Matsumura, 1937 | | | Onthophagus statzi ♰ Krell, 1990 | | | Onthophagus sternalis Arrow, 1931 | | | Onthophagus sternax Balthasar, 1959 | | | Onthophagus stevensi Arrow, 1931 | | | Onthophagus sticticus Harold, 1867 | | | Onthophagus strandi Balthasar, 1935 | | | Onthophagus strigatus (Mulsant, 1842) | ⇒ Onthophagus maki Illiger, 1803 | | Onthophagus subaenescens (Goidanich, 1926) | ⇒ Onthophagus ovatus Linnaeus, 1767 | | Onthophagus subconvexus Seabra, 1907 | | | Onthophagus subcribratus Arrow, 1907 | | | Onthophagus sublaminatus Mulsant, 1842 | | | Onthophagus sublineatus (d´Orbigny, 1900) | ⇒ Onthophagus turpidus Reitter, 1887 | | Onthophagus sublineolatus (Mulsant, 1842) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus submarginalis (Sahlberg, 1926) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus subprominulus (Mulsant, 1842) | ⇒ Onthophagus coenobita Herbst, 1783 | | Onthophagus subrecticornis Mulsant, 1842 | | | Onthophagus suermelii Petrovitz, 1963 | | | Onthophagus sugillatus Klug, 1855 | | | Onthophagus sulci Balthasar, 1935 | | | Onthophagus sunanicus (Stebnicka, 1973) | ⇒ Onthophagus punctator Reitter, 1893 | | Onthophagus susterai Balthasar, 1952 | | | Onthophagus sutleinensis Splichal, 1910 | | | Onthophagus suturalis (Péringuey, 1888) | ⇒ Onthophagus trituber Wiedemann, 1823 | | Onthophagus suturatus (Germar, 1824) | ⇒ Onthophagus tarandus Fabricius, 1792 | | Onthophagus sycophanta (Fairmaire, 1887) | ⇒ Euonthophagus gibbosus Scriba, 1790 | | Onthophagus tabidus Balthasar, 1935 | | | Onthophagus taiheizanus Matsumura, 1937 | | | Onthophagus takabayashii Matsumura, 1937 | | | Onthophagus taschenbergi Gillet, 1925 | | | Onthophagus tenebrosus Harold, 1871 | | | Onthophagus tenuicornis (Klug, 1855) | ⇒ Onthophagus aeruginosus Roth, 1851 | | Onthophagus tessulatus Harold, 1871 | | | Onthophagus tetraspilus (Fairmaire, 1887) | ⇒ Onthophagus nitidulus Klug, 1845 | | Onthophagus tibetanus Arrow, 1907 | | | Onthophagus tirapensis Biswas & Cha..., 1985 | | | Onthophagus tonsus d´Orbigny, 1902 | | | Onthophagus tragoides Boucomont, 1914 | | | Onthophagus transbaicalicus (Csiki, 1901) | ⇒ Onthophagus clitellifer Reitter, 1894 | | Onthophagus transcaspicus Koenig, 1888 | Gazella scarab | | Onthophagus trapezicornis d´Orbigny, 1902 | | | Onthophagus triceratops Arrow, 1913 | | | Onthophagus tricolor Boucomont, 1914 | | | Onthophagus tricuspidus Mulsant, 1842 | | | Onthophagus trihastatus (Reitter, 1909) | ⇒ Onthophagus oberthuri d´Orbigny, 1898 | | Onthophagus trinominatus Goidanich, 1926 | | | Onthophagus tripolitanus Heyden, 1890 | | | Onthophagus trituberculatus Schrank, 1798 | | | Onthophagus trochiscobius (Kolenati, 1846) | ⇒ Onthophagus suturellus Brullé, 1832 | | Onthophagus troglodyta Wiedemann, 1823 | | | Onthophagus tsunoellus Matsumura, 1938 | | | Onthophagus turfanicus Balthasar, 1971 | | | Onthophagus turmalis (Gillet, 1924) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus turpidus Reitter, 1887 | | | Onthophagus uedanus (Matsumura, 1937) | ⇒ Onthophagus olsoufieffi Boucomont, 1924 | | Onthophagus umbilicopunctatus (Medvedev, 1959) | ⇒ Onthophagus trispinus Reitter, 1892 | | Onthophagus umenoi (Matsumura, 1938) | ⇒ Onthophagus trituber Wiedemann, 1823 | | Onthophagus umeonoi Matsumura, 1938 | | | Onthophagus unifasciatus Schaller, 1783 | | | Onthophagus uniformis Heyden, 1886 | | | Onthophagus unxovi (Bojachew, 1930) | ⇒ Onthophagus fortigibber Reitter, 1909 | | Onthophagus urellus Boucomont, 1919 | | | Onthophagus urus (Harold, 1867) | ⇒ Onthophagus urusheeri Krell, 2000 | | Onthophagus urusheeri ♰ Krell, 2000 | | | Onthophagus ushiodai (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus variabilis (Mulsant, 1842) | ⇒ Onthophagus maki Illiger, 1803 | | Onthophagus variegatus (Fabricius, 1798) | ⇒ Onthophagus filicornis Harold, 1873 | | Onthophagus venustulus Erichson, 1843 | | | Onthophagus vespillo Boucomont, 1919 | | | Onthophagus vicinus (Harold, 1886) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus vinctus Erichson, 1843 | | | Onthophagus virescens Harold, 1867 | | | Onthophagus viridilimbatus (Olsoufieff, 1907) | ⇒ Onthophagus haroldi Ballion, 1870 | | Onthophagus viriditinctus Reitter, 1893 | | | Onthophagus vulneratus (Mulsant, 1842) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus vulpinus Arrow, 1931 | | | Onthophagus waterhousei Gillet, 1927 | | | Onthophagus xerxes Petrovitz, 1965 | | | Onthophagus xiphias (Solis & Kohl..., 2003) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus yaeyamanus Nomura, 1964 | | | Onthophagus yakuinsulanus Nakane, 1984 | | | Onthophagus yanoi Matsumura, 1938 | | | Onthophagus yaoi Masumoto, Oc..., 2014 | | | Onthophagus yohenai (Matsumura, 1937) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus yugianus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus yumigatanus Matsumura, 1937 | | | Onthophagus yunnanus Boucomont, 1912 | | | Onthophagus zagrosicus Kabakov, 2006 | | | Onthophagus zavreli Balthasar, 1935 | | | Onthophagus zimmermanni Balthasar, 1959 | | | Onthophagus zinovskyi Qaryagdy, 1939 | | | Onthophagus zolotarevi (Olsoufieff, 1907) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus zubaci (Balthasar, 1932) | ⇒ Onthophagus dama Fabricius, 1798 | | Onthophagus zuvandi Qaryagdy, 1939 | | | Phalops candezei Lansberge, 1883 | | | Phalops chalybeus Klug, 1845 | | | Phalops cyanescens d´Orbigny, 1897 | | | Phalops divisus Wiedemann, 1823 | | | Phalops iphis Olivier, 1789 | | | Phalops olivaceus Lansberge, 1883 | | | Proagoderus amplexus Sharp, 1875 | | | Proagoderus loricatus Klug, 1855 | | | Proagoderus pactolus Fabricius, 1787 | | | Proagoderus quadriarmatus Fairmaire, 1892 | | | Proagoderus worsissa Roth, 1851 | | | Proagoderus yunnanus Boucomont, 1912 | |
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