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Checklist for Tribe Scarabaeini Latreille, 1802
and region Ecozones


Class
 		Insecta
Linnaeus, 1758
Insects, True insects		814
32230
411245		Families
Genera
Species		 
Taxom
 

Subclass
 		Pterygota
Lang, 1888
Winged insects		809
32201
411191		Families
Genera
Species		 
Taxom
 

Infraclass
 		Neoptera
van der Wulp, 1890
Modern Wing-folding insects		776
31720
407430		Families
Genera
Species		 
Taxom
 

Superorder
 		Holometabola
Heider, 1889
Holometabolous Insects		508
27640
383593		Families
Genera
Species		 
Taxom
 

Order
 		Coleoptera
Linnaeus, 1758
Beetles, Coleopterans		152
13621
245492		Families
Genera
Species		 
Taxom
 

Suborder
 		Polyphaga
Emery, 1886
Polyphags		138
11892
200777		Families
Genera
Species		 
Taxom
 

Infraorder
 		Scarabaeiformia
Crowson, 1960
18
2639
43663		Families
Genera
Species		 
Taxom
 

Superfamily
 		Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles		18
2639
43663		Families
Genera
Species		 
Taxom
 
FamilyScarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles		2300
39341		Genera
Species		 
Taxom
 
SubfamilyScarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles		226
6709		Genera
Species		 
Taxom
 
TribeScarabaeini
Latreille, 1802
8
226		Genera
Species		 
Taxom
 


Genera for Ecozones
8 Genera with 226 Species
Drepanopodus Janssens, 19012SpeciesTaxom
Kheper Janssens, 198633SpeciesTaxom
Mnematidium Ritsema, 18452SpeciesTaxom
Mnematium MacLeay, 18322SpeciesTaxom
Pachylomera Griffith & P..., 18742SpeciesTaxom
Pachysoma MacLeay, 195317SpeciesTaxom
Scarabaeus Linnaeus, 1901168SpeciesTaxom
Sceliages Westwood, 18447SpeciesTaxom


Species list for Ecozones
226 Species and 8 Subspecies
Drepanopodus costatus Wiedemann, 1823
Drepanopodus proximus Péringuey, 1901
Kheper aegyptiorum Latreille, 1823
Kheper aegyptiorum aegyptiorum
Kheper aegyptiorum purpurascens Gerstaecker, 1871
Kheper aeratus Gerstaecker, 1871
Kheper asceticus Gillet, 1907
Kheper bonellii MacLeay, 1821
Kheper clericus Boheman, 1857
Kheper cupreus Castelnau, 1840
Kheper cuvieri MacLeay, 1821
Kheper devotus Redtenbacher, 1848
Kheper erichsoni Harold, 1867
Kheper festivus Harold, 1868
Kheper festivus festivus
Kheper festivus nicolasi Moretto, 1998
Kheper festivus simoni Moretto, 1998
Kheper festivus wellsi Moretto, 1998
Kheper intermedius Gillet, 1909
Kheper kalaharicus Davis, Desch..., 2011
Kheper laevistriatus Fairmaire, 1893
Kheper lamarcki MacLeay, 1821
Kheper nigroaeneus Boheman, 1857
Kheper paganus Harold, 1878
Kheper platynotus Bates, 1888
Kheper porosus Bates, 1888
Kheper prodigiosus Erichson, 1843
Kheper pustulosus Gerstaecker, 1871
Kheper rolciki Pokorný & Zí..., 2015
Kheper sanctus Fabricius, 1798
Kheper subaeneus Harold, 1869
Kheper vanderkelleni Lansberge, 1886
Kheper venerabilis Harold, 1871
Kheper vethi Lansberge, 1886
Kheper zurstrasseni Davis, 1986
Mnematidium multidentatum (Klug, 1845)⇒ Scarabaeus multidentatum Klug, 1845
Mnematidium multidentatus Klug, 1845
Mnematium ritchiei (MacLeay, 1821)⇒ Scarabaeus ritchiei MacLeay, 1821
Mnematium silenus Gray, 1832
Pachylomera femoralis Kirby, 1828
Pachylomera opaca Lansberge, 1874
Pachysoma aesculapius Olivier, 1789
Pachysoma bennigseni Felsche, 1907
Pachysoma denticolle Péringuey, 1888
Pachysoma denticollis Péringuey, 1888
Pachysoma endroedyi Harrison, Sc..., 2003
Pachysoma fitzsimonsi Ferreira, 1953
Pachysoma gariepinum Ferreira, 1953
Pachysoma gariepinus Ferreira, 1953
Pachysoma glentoni Harrison, Sc..., 2003
Pachysoma hippocrates MacLeay, 1821
Pachysoma rodriguesi Ferreira, 1953
Pachysoma rotundigena (Felsche, 1907)⇒ Scarabaeus rotundigenus Felsche, 1907
Pachysoma rotundigenus Felsche, 1907
Pachysoma schinzi Fairmaire, 1888
Pachysoma striatum (Castelnau, 1840)⇒ Scarabaeus striatus Castelnau, 1840
Pachysoma striatus Castelnau, 1840
Pachysoma valeflorae Ferreira, 1953
Scarabaeus (Ateuchetus) armeniacus Ménétriés, 1832
Scarabaeus (Ateuchetus) cicatricosus (Lucas, 1846)⇒ Scarabaeus palemo Olivier, 1789
Scarabaeus (Ateuchetus) laticollis Linnaeus, 1767
Scarabaeus (Ateuchetus) puncticollis (Latreille, 1819)⇒ Geotrupes spiniger Marsham, 1802
Scarabaeus (Ateuchetus) semipunctatus Fabricius, 1792
Scarabaeus (Ateuchetus) variolosus (Fabricius, 1787)⇒ Ateuchetus semipunctatus Fabricius, 1792
Scarabaeus (Scarabaeus) babori Balthasar, 1934
Scarabaeus (Scarabaeus) bannuensis Janssens, 1940
Scarabaeus (Scarabaeus) carinatus (Gebler, 1841)⇒ Calcodes carinatus Linnaeus, 1758
Scarabaeus (Scarabaeus) pius Illiger, 1803
Scarabaeus (Scarabaeus) sacer (Linnaeus, 1758)⇒ Scarabaeus pius Illiger, 1803
Scarabaeus (Scarabaeus) typhon (Fischer von ..., 1823)⇒ Megasoma gyas Herbst, 1785
Scarabaeus (Scarabaeus) winkleri Stolfa, 1938
Scarabaeus acuticollis Motschulsky, 1849
Scarabaeus aegyptiacus Stolfa, 1938
Scarabaeus aegyptiorum Latreille, 1827
Scarabaeus affinis (Sturm, 1800)⇒ Onthophagus vacca Linnaeus, 1767
Scarabaeus alienus Péringuey, 1901
Scarabaeus alluaudi (Théry, 1932)⇒ Ateuchetus laticollis Linnaeus, 1767
Scarabaeus ambiguus (Boheman, 1857)⇒ Gnorimus variabilis Linnaeus, 1758
Scarabaeus anderseni Waterhouse, 1890
Scarabaeus andreaei zur Strassen, 1963
Scarabaeus andrewesi Felsche, 1907
Scarabaeus basuto zur Strassen, 1962
Scarabaeus bohemani Harold, 1868
Scarabaeus bornemisszai zur Strassen, 1980
Scarabaeus bornemizzai zur Strassen, 1980
Scarabaeus bottoi Gridelli, 1930
Scarabaeus brahminus Castelnau, 1840
Scarabaeus caffer Boheman, 1857
Scarabaeus canaliculatus Fairmaire, 1888
Scarabaeus cancer Arrow, 1919
Scarabaeus castaneipennis A. Costa, 1853
Scarabaeus clanceyi Ferreira, 1954
Scarabaeus cognatus Péringuey, 1901
Scarabaeus compressicornis (Klug, 1845)⇒ Scarabaeus cristatus Fabricius, 1775
Scarabaeus confluidens Fleischer, 1925
Scarabaeus confusus Müller, 1942
Scarabaeus convexus Hausmann, 1807
Scarabaeus corinthius Fairmaire, 1887
Scarabaeus cornifrons Laporte, 1840
Scarabaeus costatus Wiedemann, 1823
Scarabaeus crenatus DeGeer, 1778
Scarabaeus cristatus Fabricius, 1775
Scarabaeus cuprescens (Gillet, 1907)⇒ Kheper aegyptiorum Latreille, 1823
Scarabaeus damarensis Janssens, 1940
Scarabaeus degeeri MacLeay, 1821
Scarabaeus deludens zur Strassen, 1961
Scarabaeus devotus Redtenbacher, 1848
Scarabaeus digitatus (Motschulsky, 1849)⇒ Scarabaeus pius Illiger, 1803
Scarabaeus dioscoridis Petrovitz, 1962
Scarabaeus dufresnei MacLeay, 1821
Scarabaeus ebenus Klug, 1855
Scarabaeus edentulus Mulsant, 1842
Scarabaeus erichsoni Harold, 1867
Scarabaeus europaeus Motschulsky, 1849
Scarabaeus flavicornis Boheman, 1860
Scarabaeus frankenbergeri (Balthasar, 1934)⇒ Scarabaeus wilsoni Waterhouse, 1890
Scarabaeus fraterculus Kolbe, 1895
Scarabaeus fritschi Harold, 1868
Scarabaeus fulgidus (Olivier, 1789)⇒ Kheper aegyptiorum Latreille, 1823
Scarabaeus funebris Boheman, 1857
Scarabaeus galenus Westwood, 1847
Scarabaeus gangeticus Castelnau, 1840
Scarabaeus ganglbaueri (Reitter, 1892)⇒ Scarabaeus wilsoni Waterhouse, 1890
Scarabaeus geminogalenus Davis & Desc..., 2017
Scarabaeus gilleti Janssens, 1940
Scarabaeus goryi Castelnau, 1840
Scarabaeus gracai Ferreira, 1952
Scarabaeus heqvisti zur Strassen, 1962
Scarabaeus hottentorum Péringuey, 1901
Scarabaeus hypocrita (Castelnau, 1840)⇒ Sericotrupes niger Marsham, 1802
Scarabaeus impius Fabricius, 1801
Scarabaeus indicus Motschulsky, 1864
Scarabaeus indistinctus Seabra, 1907
Scarabaeus inermis (Mulsant, 1842)⇒ Lucanus cervus Linnaeus, 1758
Scarabaeus infirmus (Fischer von ..., 1823)⇒ Scarabaeus pius Illiger, 1803
Scarabaeus inoportunus Ferreira, 1953
Scarabaeus inquisitus Péringuey, 1908
Scarabaeus interstitialis Boheman, 1857
Scarabaeus intricatus Fabricius, 1801
Scarabaeus irakensis Stolfa, 1938
Scarabaeus isidis Castelnau, 1840
Scarabaeus jalof Castelnau, 1840
Scarabaeus janssensi Balthasar, 1940
Scarabaeus janssensianus (Auber, 1952)⇒ Escarabaeus bannuensis Janssens, 1940
Scarabaeus karae Davis & Desc..., 2016
Scarabaeus karlwerneri Nicolas & Mo..., 2002
Scarabaeus karrooensis zur Strassen, 1961
Scarabaeus knobeli Ferreira, 1958
Scarabaeus kochi Ferreira, 1952
Scarabaeus krutai Maly & Montr..., 2011
Scarabaeus kwiluensis Janssens, 1940
Scarabaeus laevicollis (Mulsant, 1842)⇒ Ateuchetus laticollis Linnaeus, 1767
Scarabaeus laevifrons Fairmaire, 1884
Scarabaeus laevigatus (Kolbe, 1887)⇒ Scarabaeus goryi Castelnau, 1840
Scarabaeus licitus Ferreira, 1954
Scarabaeus lucidulus Boheman, 1860
Scarabaeus minor (Ladeiro, 1950)⇒ Ateuchetus laticollis Linnaeus, 1767
Scarabaeus minutus (Fabricius, 1792)⇒ Sisyphus longipes Olivier, 1789
Scarabaeus monachus (Faldermann, 1835)⇒ Scarabaeus pius Illiger, 1803
Scarabaeus montreuili meridionalis Moretto, 2023
Scarabaeus montreuili montreuili
Scarabaeus multidentatum Klug, 1845
Scarabaeus natalensis zur Strassen, 1962
Scarabaeus nepos (Fairmaire, 1884)⇒ Scarabaeus goryi Castelnau, 1840
Scarabaeus nigricans (Gillet, 1907)⇒ Kheper aegyptiorum Latreille, 1823
Scarabaeus nigrociliatus (Motschulsky, 1849)⇒ Ateuchetus armeniacus Ménétriés..., 1832
Scarabaeus nitidicollis Lansberge, 1882
Scarabaeus nudifrons Fischer von ..., 1823
Scarabaeus obsoletepunctatus Balthasar, 1940
Scarabaeus opacipennis Fairmaire, 1884
Scarabaeus osiridis Moretto, 2023
Scarabaeus pabulator Péringuey, 1908
Scarabaeus palemo Olivier, 1789
Scarabaeus parumpunctatus Klug, 1845
Scarabaeus parvulus Boheman, 1860
Scarabaeus peregrinus Kolbe, 1886
Scarabaeus piliventris zur Strassen, 1962
Scarabaeus planifrons Fairmaire, 1884
Scarabaeus platychilus Fischer von ..., 1823
Scarabaeus plausibilis Péringuey, 1892
Scarabaeus poggei Waterhouse, 1890
Scarabaeus proboscideus Guérin-Ménev..., 1844
Scarabaeus profanus Fabricius, 1792
Scarabaeus proximus (Péringuey, 1901)⇒ Drepanopodus proximus Péringuey, 1901
Scarabaeus pseudogangeticus Moretto, 2023
Scarabaeus punctulatus (Mulsant, 1842)⇒ Nialus varians Duftschmid, 1805
Scarabaeus retusus (Fabricius, 1781)⇒ Temnorrhynchus retusus Fabricius, 1781
Scarabaeus ritchiei MacLeay, 1821
Scarabaeus rixosus Péringuey, 1901
Scarabaeus rotundipenne Holdhaus, 1919
Scarabaeus rubripennis Boheman, 1860
Scarabaeus rufipes (Seabra, 1907)⇒ Acrossus rufipes Linnaeus, 1758
Scarabaeus rugosus (Hausmann, 1807)⇒ Onthophagus taurus Schreber, 1759
Scarabaeus rusticus Boheman, 1857
Scarabaeus sanctus Fabricius, 1798
Scarabaeus sanguinolentus (Gmelin, 1790)⇒ Gametis versicolor Fabricius, 1775
Scarabaeus scholtzi Mostert & Holm, 1982
Scarabaeus schulzeae Ferreira, 1969
Scarabaeus semilunatoides (Baguena, 1955)⇒ Ateuchetus cicatricosus Lucas, 1846
Scarabaeus semilunatus (Xambeu, 1893)⇒ Ateuchetus laticollis Linnaeus, 1767
Scarabaeus senaariensis Laporte, 1840
Scarabaeus sericeus (Motschulsky, 1849)⇒ Ateuchetus puncticollis Latreille, 1819
Scarabaeus serratus (Fourcroy, 1785)⇒ Scarabaeus laticollis Linnaeus, 1767
Scarabaeus silenus (Gray, 1832)⇒ Phyllognathus excavatus Forster, 1771
Scarabaeus spencei MacLeay, 1821
Scarabaeus spretus zur Strassen, 1962
Scarabaeus stigmaticus (Fairmaire, 1887)⇒ Kheper aegyptiorum Latreille, 1823
Scarabaeus striatopunctatus Leoni, 1911
Scarabaeus striolatus (Reitter, 1893)⇒ Ateuchetus laticollis Linnaeus, 1767
Scarabaeus subinermis (Mulsant, 1842)⇒ Scarabaeus semipunctatus Fabricius, 1792
Scarabaeus substriatus (Mulsant, 1842)⇒ Scarabaeus semipunctatus Fabricius, 1792
Scarabaeus subsulcatus (Mulsant, 1842)⇒ Scarabaeus pius Illiger, 1803
Scarabaeus sulcipennis Quedenfeldt, 1888
Scarabaeus suri Hausmann, 1807
Scarabaeus tonckeri Boucomont, 1923
Scarabaeus transcaspicus Stolfa, 1938
Scarabaeus vansoni Ferreira, 1958
Scarabaeus vethi Lansberge, 1886
Scarabaeus viator Péringuey, 1901
Scarabaeus vicinus (Janssens, 1940)⇒ Lucanus cervus Linnaeus, 1758
Scarabaeus villiersi Baraud, 1973
Scarabaeus westwoodi (Harold, 1869)⇒ Scarabaeus galenus Westwood, 1847
Scarabaeus wilsoni Waterhouse, 1890
Scarabaeus xavieri Ferreira, 1968
Scarabaeus zagrosensis Montreuil, 2006
Scarabaeus zambesianus Péringuey, 1901
Sceliages adamastor Lepeletier &..., 1828
Sceliages augias Gillet, 1908
Sceliages brittoni zur Strassen, 1965
Sceliages difficilis zur Strassen, 1965
Sceliages gagates Shipp, 1895
Sceliages granulatus Forgie & Gre..., 2002
Sceliages hippias Westwood, 1844


Links, sources and further information
Scarabaeini at artfakta.se