Checklist for Superfamily Scarabaeoidea Latreille, 1802 (Bess Beetles)
and region Maranhão (Northeast Brazil, Brazil)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 158
810
1285 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 158
810
1285 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 150
800
1274 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 90
528
853 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 20
144
174 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 17
134
159 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 4
41
88 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 4
41
88 | Families
Genera
Species |
Taxom
|
List of families for Maranhão
4 Families width 88 Species and 2 Subspecies
Genera for Maranhão
41 Genera with 88 Species #ABCDEGILMNOPST
| Anomiopus Westwood, 1891 | | 2 | Species | Taxom | | Ataenius Harold, 1962 | | 3 | Species | Taxom | | Ateuchus Weber, 1868 | | 1 | Species | Taxom | | Athyreus MacLeay, 1978 | | 1 | Species | Taxom | | Bolax Fischer von ..., 2011 | | 1 | Species | Taxom | | Bothynus Hope, 1968 | | 2 | Species | Taxom | | Canthidium Erichson, 1913 | | 3 | Species | Taxom | | Canthon Hoffmannsegg, 2019 | Tumblebug | 4 | Species | Taxom | | Chlorota Burmeister, 2020 | | 1 | Species | Taxom | | Coelosis Hope, 1779 | | 1 | Species | Taxom | | Coprophanaeus Olsoufieff, 1924 | | 3 | Species | Taxom | | Cryptogenius Westwood, 2023 | | 1 | Species | Taxom | | Cyclocephala Dejean, 1847 | Masked Chafers | 4 | Species | Taxom | | Deltochilum Eschscholtz, 1939 | | 3 | Species | Taxom | | Dendropaemon Perty, 2016 | | 2 | Species | Taxom | | Diabroctis Gistl, 1758 | | 1 | Species | Taxom | | Dichotomius Hope, 2019 | | 9 | Species | Taxom | | Digitonthophagus Balthasar, 1787 | | 1 | Species | Taxom | Dyscinetus Harold, 1869 | Rice Beetles | 0 | Species | Taxom | | Euligyrus Casey, 1968 | | 2 | Species | Taxom | | Eurysternus Dalman, 2009 | | 3 | Species | Taxom | | Gymnetis MacLeay, 1800 | | 6 | Species | Taxom | | Iarupea Martínez, 1973 | | 2 | Species | Taxom | | Isocopris Pereira & Ma..., 1824 | | 2 | Species | Taxom | | Labarrus Mulsant and Rey, 0 | | 1 | Species | Taxom | | Ligyrus Erichson, 1801 | | 1 | Species | Taxom | | Liogenys Guérin-Ménév..., 2019 | | 4 | Species | Taxom | | Megasoma Kirby, 1758 | Elephant beetles | 1 | Species | Taxom | | Minisiderus Arrow, 2005 | | 2 | Species | Taxom | | Neoathyreus Howden & Mar..., 1985 | | 1 | Species | Taxom | | Omorgus Erichson, 1775 | Hastate Hide Beetles | 2 | Species | Taxom | | Ontherus Erichson, 1829 | | 1 | Species | Taxom | | Onthophagus Latreille, 1815 | | 1 | Species | Taxom | | Oxyligyrus Arrow, 1789 | | 3 | Species | Taxom | | Parathyreus Howden & Mar..., 1843 | | 2 | Species | Taxom | | Podischnus Burmeister, 2020 | | 1 | Species | Taxom | | Pseudocanthon Bates, 1846 | | 1 | Species | Taxom | | Spodochlamys Burmeister, 2011 | | 3 | Species | Taxom | | Sylvicanthon Halffter & M..., 2018 | | 3 | Species | Taxom | | Tetraechma Agassiz, 1813 | | 2 | Species | Taxom | | Trichillum Harold, 1880 | | 1 | Species | Taxom |
Species list for Maranhão
88 Species and 2 Subspecies | Anomiopus pereirai Martínez, 1955 | | | Anomiopus simplex Waterhouse, 1891 | | | Ataenius morator Harold, 1869 | | | Ataenius platensis Blanchard, 1846 | | | Ataenius scabrelloides Petrovitz, 1962 | Blennies | | Ateuchus semicribratus Harold, 1868 | | | Athyreus acuticornis Howden, 1978 | | | Bolax genieri Soula, 2011 | | | Bothynus alvarengai Martínez, 1983 | | | Bothynus horridus Endrödi, 1968 | | | Canthidium decoratum Perty, 1830 | | | Canthidium humerale Germar, 1813 | | | Canthidium manni Arrow, 1913 | | | Canthon cinctellus Germar, 1824 | | | Canthon septemmaculatus histrio Lepeletier &..., 1828 | | | Canthon simulans Martínez, 1950 | | | Canthon ziggy Vieira, Vaz-..., 2019 | | | Chlorota cleidecostae Medeiros & G..., 2020 | | | Coelosis bicornis Leske, 1779 | | | Coprophanaeus (Metallophanaeus) pertyi Olsufieff, 1924 | | | Coprophanaeus acrisius MacLeay, 1819 | | | Coprophanaeus cyanescens Olsufieff, 1924 | | | Cryptogenius westwoodi Costa-Silva ..., 2023 | | | Cyclocephala atricapilla Mannerheim, 1829 | | | Cyclocephala colasi Endrödi, 1964 | | | Cyclocephala latericia Höhne, 1923 | | | Cyclocephala testacea Burmeister, 1847 | Twister | | Deltochilum carinatum Westwood, 1837 | | | Deltochilum orbiculare Lansberge, 1874 | | | Deltochilum pseudoicarus Balthasar, 1939 | | | Dendropaemon (Titthopaemon) denticollis Felsche, 1909 | | | Dendropaemon carinifer Génier & Arn..., 2016 | | | Diabroctis mimas Linnaeus, 1758 | | | Dichotomius (Homocanthonides) smaragdinus Perty, 1830 | | | Dichotomius (Luederwaldtinia) geminatus Arrow, 1913 | | | Dichotomius (Luederwaldtinia) nisus Olivier, 1789 | | | Dichotomius (Luederwaldtinia) semisquamosus Curtis, 1845 | | | Dichotomius (Luederwaldtinia) sexdentatus Luederwaldt, 1925 | | | Dichotomius bos Blanchard, 1846 | | | Dichotomius carinatus Luederwaldt, 1925 | | | Dichotomius nimuendajui Luederwaldt, 1925 | | | Dichotomius rondoniaensis Nunes & Vaz-..., 2019 | | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Euligyrus ebenus Degeer, 1774 | | | Euligyrus similis Endrödi, 1968 | | | Eurysternus caribaeus Herbst, 1789 | | | Eurysternus fallaciosus Génier, 2009 | | | Eurysternus nigrovirens Génier, 2009 | | | Gymnetis aurantivittae Ratcliffe, 2018 | | | Gymnetis buckleyi (Janson, 1875) | ⇒ Gymnetis hieroglyphica buckleyi Janson, 1879 | | Gymnetis flava (Burmeister, 1842) | ⇒ Gymnetis xanthospila Schaum, 1844 | | Gymnetis holosericea (Voet, 1779) | ⇒ Gymnetis xanthospila Schaum, 1844 | | Gymnetis margineguttata Gory & Perch..., 1833 | | | Gymnetis rufilatris Illiger, 1800 | | | Iarupea luisae Stebnicka, 2007 | | | Iarupea serratipennis Petrovitz, 1973 | | | Isocopris hypocrita Lucas, 1859 | | | Isocopris inhiatus Germar, 1824 | | | Labarrus lividus lividus | | | Ligyrus cuniculus (Fabricius, 1801) | ⇒ Tomarus cuniculus Fabricius, 1801 | | Liogenys bidentata Burmeister, 1855 | | | Liogenys cavifrons Cherman, 2017 | | | Liogenys pilosipennis Moser, 1918 | | | Liogenys variabilis Cherman, 2019 | | | Megasoma actaeon Linnaeus, 1758 | Actaeon beetle | | Minisiderus elyanae Dechambre, 2009 | | | Minisiderus mielkeorum Grossi & Grossi, 2005 | | | Neoathyreus (Neoathyreus) brazilensis Howden, 1985 | | | Omorgus (Haroldomorgus) batesi Harold, 1872 | | | Omorgus (Omorgus) suberosus Fabricius, 1775 | Hide beetle | | Ontherus appendiculatus Mannerheim, 1829 | | | Onthophagus hircus Billberg, 1815 | | | Oxyligyrus acutus Endrödi, 1966 | | | Oxyligyrus limeirai Pinto-Júnior..., 2019 | | | Oxyligyrus zoilus Olivier, 1789 | | | Parathyreus fissicollis Arrow, 1913 | | | Parathyreus trituberculatus Klug, 1843 | | | Podischnus limeirai Duarte & Grossi, 2020 | | | Pseudocanthon xanthurus Blanchard, 1846 | | | Spodochlamys caesarea Burmeister, 1855 | | | Spodochlamys iheringi Ohaus, 1905 | | | Spodochlamys marahuaca Jameson & Ra..., 2011 | | | Sylvicanthon enkerlini Cupello & Va..., 2018 | | | Sylvicanthon proseni Martínez, 1949 | | | Sylvicanthon seag Cupello & Va..., 2018 | | | Tetraechma chlorophana Mannerheim, 1828 | | | Tetraechma liturata Germar, 1813 | | | Trichillum externepunctatum Preudhomme d..., 1880 | |
Subnodes of Scarabaeoidea with occurrences in Maranhão | Family | Geotrupidae
Latreille, 1802
Dor beetles, Dung beetles, Earth-Boring Dung Beetle, Earth-boring dung beetles, Earth-boring scarab beetles | 3
4 | Genera
Species |
Taxom
| | Family | Hybosoridae
Erichson, 1847
Pill scarab beetles, scavenger scarab beetle, Scavenger scarab beetles, Scavenging scarab beetles | 1
1 | Genus
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Monkey beetles, Rain beetles, Scarab Beetle, Scarab beetles, Scarabs, Tumble bugs | 36
81 | Genera
Species |
Taxom
| | Family | Trogidae
MacLeay, 1819
Gnawer beetles, Hide, hide beetle, Hide beetles, Keratin Beetles, Skin and hide beetles, Skin beetles | 1
2 | Genus
Species |
Taxom
|
Links, sources and further information Scarabaeoidea at scarabeidi.it
Scarabaeoidea at artfakta.se
Scarabaeoidea at dyntaxa.se
Scarabaeoidea at itis.gov
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