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Checklist for Superfamily Scarabaeoidea Latreille, 1802 (Bess Beetles)
and region Neogene (Cenozoic)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
379
1807
3318
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
375
1797
3306
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
355
1717
3172
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
224
1264
2441
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
70
585
1018
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
63
513
865
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
7
51
93
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
7
51
93
Families
Genera
Species
 
Taxom
 


List of families for Neogene
7 Families width 93 Species
Scarabaeidae Latreille, 1802Dung beetles75SpeciesTaxom
Hybosoridae Erichson, 1847Pill scarab beetles6SpeciesTaxom
Lucanidae Latreille, 1804Stag Beetle4SpeciesTaxom
Geotrupidae Latreille, 1802Dor beetles4SpeciesTaxom
Glaphyridae MacLeay, 1819Bumble bee scarab beetle1SpeciesTaxom
Ochodaeidae Mulsant & Rey, 1870Sand-loving scarab beetle1SpeciesTaxom
Passalidae Leach, 1815Bess beetle1SpeciesTaxom


Genera for Neogene
50 Genera with 93 Species
#ACEGHLMOPRSTVZ
Adoretus Laporte, 19942SpeciesTaxom
Anachalcos Hope, 19761SpeciesTaxom
Anaides Westwood, 20061SpeciesTaxom
Anomala Samouelle, 1994Shining Leaf Chafer13SpeciesTaxom
Aphodius Hellwig, 18475SpeciesTaxom
Calcitoryctes Krell, 20111SpeciesTaxom
Canthochilum Chapin, 20162SpeciesTaxom
Ceratocanthus White, 20141SpeciesTaxom
Ceruchus MacLeay, 19111SpeciesTaxom
Cheirotonus Hope, 19891SpeciesTaxom
Copris Geoffroy, 1862Mexican dung beetle4SpeciesTaxom
Coprologus Heer, 18471SpeciesTaxom
Elytridium Heer, 1870
0SpeciesTaxom
Genonota Zhang, 19892SpeciesTaxom
Geotrupes Latreille, 1983Dor Beetle3SpeciesTaxom
Glaphyrus Latreille, 18621SpeciesTaxom
Gnorimus Le Peletier ..., 1862Chafer1SpeciesTaxom
Gymnopleurus Illiger, 18473SpeciesTaxom
Heliocopris Hope, 19681SpeciesTaxom
Holotrichia Hope, 19892SpeciesTaxom
Hybosorus MacLeay, 18621SpeciesTaxom
Lepitrix Lepeletier &..., 18621SpeciesTaxom
Lucanus Scopoli, 19131SpeciesTaxom
Macronota Hoffmansegg, 19891SpeciesTaxom
Magniscarabaeus Hong, 19881SpeciesTaxom
Melolontha Fabricius, 1878Billy witch1SpeciesTaxom
Melolonthites Heer, 18478SpeciesTaxom
Metacatharsius Paulian, 19761SpeciesTaxom
Miocenidorcus Riou, 19991SpeciesTaxom
Mioochodaeus Nikolajev, 18491SpeciesTaxom
Mioserica Zhang, 19891SpeciesTaxom
Oniticellus Serville, 18621SpeciesTaxom
Onthophagus Latreille, 20007SpeciesTaxom
Orrhodomala Zhang, 19891SpeciesTaxom
Oryctes Hellwig, 1758Coconut rhinoceros beetles1SpeciesTaxom
Paleotrichius Poinar, 20111SpeciesTaxom
Petraeianus Zhang, 19891SpeciesTaxom
Petulantis Zhang & al., 19941SpeciesTaxom
Phalops Erichson, 1847
0SpeciesTaxom
Procoilodes Ocampo, 20021SpeciesTaxom
Rhizotrogus Lepeletier &..., 18471SpeciesTaxom
Scarabaeus Linnaeus, 18621SpeciesTaxom
Scelocopris Zhang, 19891SpeciesTaxom
Serica MacLeay, 18621SpeciesTaxom
Serrulus Hong, 19831SpeciesTaxom
Syndesus MacLeay, 20091SpeciesTaxom
Trichius Fabricius, 1862Bee beetles5SpeciesTaxom
Tyrannasorus Ratcliffe & ..., 20011SpeciesTaxom
Valgus Scriba, 18621SpeciesTaxom
Zhangsunia Krell, 19941SpeciesTaxom


Species list for Neogene
93 Species
Adoretus recticlypeus J. F. Zhang, 1989
Adoretus rhinus J. Zhang, 1994
Anachalcos mfwangani R. Paulian, 1976
Anaides howdeni Ocampo, 2006
Anomala amblobelia J. F. Zhang, 1989
Anomala brachytarsia J. Zhang, 1994
Anomala endoxa J. Zhang, 1994
Anomala eversa J. F. Zhang, 1989
Anomala fugax O. Heer, 1862
Anomala furva Y. C. Hong, 1983
Anomala lochmocola J. F. Zhang, 1989
Anomala orcina J. Zhang, 1994
Anomala palaeobrunnea Hong, 1985
Anomala punctulata J. F. Zhang, 1989
Anomala scia J. F. Zhang, 1989
Anomala synemosyna J. F. Zhang, 1989
Anomala ursa J. Zhang, 1994
Aphodius (Acrossus) rufipes (Linnaeus, 1758)⇒ Acrossus rufipes Linnaeus, 1758
Aphodius anteactus Krell, 2000
Aphodius bosniaskii A. Handlirsch, 1907
Aphodius brevipennis (O. Heer, 1862)⇒ Aphodius foetidus Herbst, 1783
Aphodius meyeri Heer, 1847
Calcitoryctes magnificus F. T. Krell, 2011
Canthochilum alleni Tarasov & Va..., 2016
Canthochilum philipsivieorum Tarasov & Va..., 2016
Ceratocanthus emarginatus Poinar, 2014
Ceruchus fuchsii Wickham, 1911
Cheirotonus otai K. Ueda, 1989
Copris (Copris) kartlinus O. N. Kabakov, 1988
Copris druidum O. Heer, 1862
Copris leakeyorum R. Paulian, 1976
Copris subterranea O. Heer, 1862
Coprologus gracilis Heer, 1847
Genonota mochthera J. F. Zhang, 1989
Genonota scola J. F. Zhang, 1989
Geotrupes germari O. Heer, 1862
Geotrupes jiaoyanshanense Y. C. Hong, 1983
Geotrupes jiaoyanshanensis Hong, 1983
Glaphyrus antiquus O. Heer, 1862
Gnorimus aedilis Heer, 1862
Gymnopleurus deperditus O. Heer, 1862
Gymnopleurus rotundatus O. Heer, 1862
Gymnopleurus sisyphus O. Heer, 1847
Heliocopris antiquus I. Fujiyama, 1968
Holotrichia cressona J. Zhang, 1994
Holotrichia spatha J. F. Zhang, 1989
Hybosorus lividus O. Heer, 1862
Lepitrix germanica O. Heer, 1862
Lucanus (Lucanus) fossilis Wickham, 1913
Macronota shandongiana J. F. Zhang, 1989
Magniscarabaeus furvus Hong, 1988
Melolontha solitaria O. Novák, 1878
Melolonthites aciculata O. Heer, 1847
Melolonthites deperdita Heer, 1847
Melolonthites interemtus Cockerell, 1926
Melolonthites kollari Heer, 1847
Melolonthites laterosinuatus Piton, 1935
Melolonthites lavateri Heer, 1847
Melolonthites obsoleta Heer, 1847
Melolonthites parschlugiana Heer, 1847
Metacatharsius rusingae R. Paulian, 1976
Miocenidorcus andancensis B. Riou, 1999
Mioochodaeus proaevus Germar, 1849
Mioserica margelis J. F. Zhang, 1989
Oniticellus amplicollis (O. Heer, 1862)⇒ Helictopleurus neoamplicollis Krell, 2000
Onthophagus bisontinus Heer, 1862
Onthophagus crassus (Heer, 1862)⇒ Parascatonomus tricornis Wiedemann, 1823
Onthophagus ovatulus Heer, 1847
Onthophagus prodromus Heer, 1862
Onthophagus spitsbergeniensis Krell, 2010
Onthophagus urus (Harold, 1867)⇒ Onthophagus urusheeri Krell, 2000
Onthophagus urusheeri Krell, 2000
Orrhodomala protista J. F. Zhang, 1989
Oryctes (Oryctes) nasicornis Linnaeus, 1758European rhinoceros beetle
Paleotrichius dominicanus G. Poinar, 2011
Petraeianus ruderatus J. F. Zhang, 1989
Petulantis yimengensis J. Zhang, 1994
Procoilodes adrastus Ocampo, 2002
Rhizotrogus longimanus Heer, 1847
Scarabaeus proserpinae O. Heer, 1862
Scelocopris enertheus J. F. Zhang, 1989
Serica minutula (O. Heer, 1862)⇒ Gynaecoserica pellecta Brenske, 1896
Serrulus sinicus Hong, 1983
Syndesus ambericus Woodruff, 2009
Trichius aedilis O. Heer, 1862
Trichius amoenus Heer, 1847
Trichius lugubris O. Heer, 1862
Trichius rotundatus O. Heer, 1862
Trichius unifasciatus O. Heer, 1862
Tyrannasorus rex Ratcliffe & ..., 2001
Valgus oeningensis O. Heer, 1862
Zhangsunia extumida J. Zhang, 1994

Subnodes of Scarabaeoidea with occurrences in Neogene
FamilyGeotrupidae
Latreille, 1802
Dor beetles, Dung beetles, Earth-Boring Dung Beetle, Earth-boring dung beetles, Earth-boring scarab beetles
2
4
Genera
Species

Taxom

FamilyGlaphyridae
MacLeay, 1819
bumble bee scarab beetle, Bumble bee scarab beetles, The bumble bee scarab beetles
1
1
Genus
Species

Taxom

FamilyHybosoridae
Erichson, 1847
Pill scarab beetles, scavenger scarab beetle, Scavenger scarab beetles, Scavenging scarab beetles
6
6
Genera
Species

Taxom

FamilyLucanidae
Latreille, 1804
Stag Beetle, Stag beetles
4
4
Genera
Species

Taxom

FamilyOchodaeidae
Mulsant & Rey, 1870
sand-loving scarab beetle, Sand-loving scarab beetles
1
1
Genus
Species

Taxom

FamilyPassalidae
Leach, 1815
bess beetle, Bess beetles, Betsy beetles, Passalid beetles, Patent leather beetles, Peg
1
1
Genus
Species

Taxom

FamilyScarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Monkey beetles, Rain beetles, Scarab Beetle, Scarab beetles, Scarabs, Tumble bugs
35
75
Genera
Species

Taxom



Links, sources and further information
Scarabaeoidea at scarabeidi.it
Scarabaeoidea at artfakta.se
Scarabaeoidea at dyntaxa.se
Scarabaeoidea at itis.gov