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Checklist for Superfamily Staphylinoidea Latreille, 1802 (Carrion Beetles)
and region Quaternary (Cenozoic)


Class
 		Insecta
Linnaeus, 1758
Insects, True insects		193
796
1475		Families
Genera
Species		 
Taxom
 

Subclass
 		Pterygota
Lang, 1888
Winged insects		192
795
1474		Families
Genera
Species		 
Taxom
 

Infraclass
 		Neoptera
van der Wulp, 1890
Modern Wing-folding insects		188
791
1469		Families
Genera
Species		 
Taxom
 

Superorder
 		Holometabola
Heider, 1889
Holometabolous Insects		141
711
1383		Families
Genera
Species		 
Taxom
 

Order
 		Coleoptera
Linnaeus, 1758
Beetles, Coleopterans		64
511
1161		Families
Genera
Species		 
Taxom
 

Suborder
 		Polyphaga
Emery, 1886
Polyphags		57
425
740		Families
Genera
Species		 
Taxom
 

Infraorder
 		Staphyliniformia
Lameere, 1900
10
120
251		Families
Genera
Species		 
Taxom
 

Superfamily
 		Staphylinoidea
Latreille, 1802
Carrion Beetles, Fungus Beetles, Rove Beetles		7
97
188		Families
Genera
Species		 
Taxom
 


List of families for Quaternary
7 Families width 188 Species and 2 Subspecies
Staphylinidae Lameere, 1900Rove beetles146SpeciesTaxom
Silphidae Latreille, 1807Burying And Carrion Beetles24SpeciesTaxom
Leiodidae Fleming, 1821Round fungus beetles6SpeciesTaxom
Scydmaenidae Leach, 1815Ant-like stone beetles1SpeciesTaxom
Hydraenidae (Mulsant, 1844)Cascade Beetles10SpeciesTaxom
Ptiliidae Heer, 1843Feather-wing beetles0SpeciesTaxom
Agyrtidae Thomson, 1859Primitive carrion beetles1SpeciesTaxom


Genera for Quaternary
97 Genera with 188 Species
#ABCDEFGHIKLMNOPQRSTUX
Abdiunguis Park and Wagner, 1962
0SpeciesTaxom
Acidota Stephens, 18382SpeciesTaxom
Aclypea Reitter, 18947SpeciesTaxom
Acrulia Thomson, 18531SpeciesTaxom
Actium Casey, 1886
0SpeciesTaxom
Agaricophagus Schmidt, 18411SpeciesTaxom
Agathidium Panzer, 18072SpeciesTaxom
Aleochara Gravenhorst, 1802
0SpeciesTaxom
Anisotoma Panzer, 17921SpeciesTaxom
Anotylus Thomson, 18777SpeciesTaxom
Anthobium Leach, 1819
0SpeciesTaxom
Aploderus Stephens, 18021SpeciesTaxom
Arpedium Erichson, 18953SpeciesTaxom
Articerus Dalman, 18261SpeciesTaxom
Artochia Casey, 18931SpeciesTaxom
Atheta Thomson, 1858
Vexing Rove Beetles0SpeciesTaxom
Atrecus Jacquelin du..., 18371SpeciesTaxom
Bledius Samouelle, 18636SpeciesTaxom
Blitophaga Reitter, 02SpeciesTaxom
Bolitobius Samouelle, 1819
0SpeciesTaxom
Boreaphilus Sahlberg, 18321SpeciesTaxom
Carpelimus Kirby, 1819
0SpeciesTaxom
Cholevinus Reitter, 19231SpeciesTaxom
Coryphium Stephens, 18341SpeciesTaxom
Cryptobium Mannerheim, 19002SpeciesTaxom
Deleaster Erichson, 18571SpeciesTaxom
Dianous Samouelle, 18741SpeciesTaxom
Dinothenarus C.G. Thomson, 17741SpeciesTaxom
Elacatophora Schaufuss, 20041SpeciesTaxom
Euaesthetus Gravenhorst, 18441SpeciesTaxom
Eucnecosum Reitter, 18633SpeciesTaxom
Eusilpha Semenov, 18621SpeciesTaxom
Eusphalerum Kraatz, 1857
0SpeciesTaxom
Eusteniamorpha Cameron, 20001SpeciesTaxom
Foveoscapha Park and Wagner, 1962
0SpeciesTaxom
Geodromicus Redtenbacher, 18952SpeciesTaxom
Gymnusa Gravenhorst, 18942SpeciesTaxom
Gyrohypnus Samouelle, 17891SpeciesTaxom
Hapalaraea C.G. Thomson, 1858
0SpeciesTaxom
Heterosilpha Portevin, 18231SpeciesTaxom
Holoboreaphilus Campbell, 18782SpeciesTaxom
Hydnobius Schmidt, 18071SpeciesTaxom
Hydraena Kugelann, 19204SpeciesTaxom
Ischnosoma Stephens, 18771SpeciesTaxom
Kalissus LeConte, 18741SpeciesTaxom
Lathrobium Gravenhorst, 18906SpeciesTaxom
Leptotyphlus Fauvel, 19841SpeciesTaxom
Lesteva Latreille, 18631SpeciesTaxom
Limnebius Leach, 17941SpeciesTaxom
Lomechusoides Tottenham, 17751SpeciesTaxom
Lordithon C.G. Thomson, 18631SpeciesTaxom
Megarthrus Curtis, 1829
0SpeciesTaxom
Micralymma Westwood, 18571SpeciesTaxom
Microedus LeConte, 18741SpeciesTaxom
Micropeplus Latreille, 18028SpeciesTaxom
Mycetoporus Mannerheim, 19571SpeciesTaxom
Neohypnus Coiffait & Saiz, 1964
0SpeciesTaxom
Nicrophorus Fabricius, 1843Burying Beetle3SpeciesTaxom
Nitidotachinus Campbell, 18772SpeciesTaxom
Ochthebius Leach, 18535SpeciesTaxom
Ochthephilum Stephens, 19002SpeciesTaxom
Ocypus Leach, 18022SpeciesTaxom
Olophrum Erichson, 181712SpeciesTaxom
Omalium Gravenhorst, 1802
0SpeciesTaxom
Orobanus LeConte, 1878
0SpeciesTaxom
Osorius Latreille, 18371SpeciesTaxom
Oxyporus Bourdot & Ga..., 18901SpeciesTaxom
Oxytelus Gravenhorst, 18774SpeciesTaxom
Paederus Fabricius, 18021SpeciesTaxom
Pelecomalium Casey, 18431SpeciesTaxom
Philonthus Stephens, 19003SpeciesTaxom
Phosphuga Leach, 17581SpeciesTaxom
Phyllodrepa Thomson, 1859
0SpeciesTaxom
Platystethus Mannerheim, 18405SpeciesTaxom
Proscapheus1SpeciesTaxom
Pselaphaulax Reitter, 17921SpeciesTaxom
Pselaphus Herbst, 17921SpeciesTaxom
Pselaptrichus Brendel, 1889
0SpeciesTaxom
Pseudopsis Newman, 18341SpeciesTaxom
Ptenidium Erichson, 1845
0SpeciesTaxom
Pteroloma Gyllenhal, 18101SpeciesTaxom
Pycnoglypta C.G. Thomson, 1813winter rove beetle3SpeciesTaxom
Quedius Stephens, 19003SpeciesTaxom
Reichenbachia Leach, 1825
0SpeciesTaxom
Silpha Linnaeus, 17983SpeciesTaxom
Sonoma Casey, 19621SpeciesTaxom
Staphylinus Linnaeus, 17742SpeciesTaxom
Stenus Latreille, 188415SpeciesTaxom
Stilicus Berthold, 18391SpeciesTaxom
Syntomium Curtiss, 19571SpeciesTaxom
Tachinus Gravenhorst, 195713SpeciesTaxom
Tachyporus Gravenhorst, 19363SpeciesTaxom
Temnodera Hope, 18371SpeciesTaxom
Tetrascapha Schuster and..., 1957
0SpeciesTaxom
Thanatophilus Leach, 18535SpeciesTaxom
Unamis Casey, 1893
0SpeciesTaxom
Xantholinus Dejean, 19262SpeciesTaxom


Species list for Quaternary
188 Species and 2 Subspecies
Acidota crenata Fabricius, 1792Staph Beetle
Acidota quadrata Zetterstedt, 1838Staph beetle
Aclypea bituberosa LeConte, 1859Anchor worm
Aclypea calva Reitter, 1890
Aclypea jacutica Rjabukhin, 1990
Aclypea opaca Linnaeus, 1758Beet Carrion Beetle
Aclypea reitteri Portevin, 1926
Aclypea sericea Zoubkoff, 1833
Aclypea vetusta A.M.�omnick..., 1894
Acrulia tumidula Maklin, 1853
Agaricophagus cephalotes Schmidt, 1841
Agathidium (Agathidium) laevigatum Erichson, 1845
Agathidium (Neoceble) marginatum Sturm, 1807
Anisotoma castanea Herbst, 1792
Anotylus (Oxytelops) inustus Gravenhorst, 1806
Anotylus (Oxytelops) nitidulus Gravenhorst, 1802
Anotylus (Oxytelops) rugosus Fabricius, 1775Rove beetle
Anotylus (Oxytelops) sculpturatus Gravenhorst, 1806
Anotylus (Oxytelops) tetracarinatus Block, 1799Rove beetle
Anotylus (Oxytelops) tetratoma Czwalina, 1871
Anotylus gibbulus Eppelsheim, 1877
Aploderus caelatus Gravenhorst, 1802
Arpedium brachypterum Gravenhorst, 1802Staph beetle
Arpedium cribratum Fauvel, 1878
Arpedium stillicidii ♰ S. H. Scudder, 1895
Articerus armatus ♰ J. W. Dalman, 1826
Artochia productifrons Casey, 1893
Atrecus macrocephalus Nordmann, 1837
Bledius (Astycops) talpa Gyllenhal, 1810
Bledius (Hesperophilus) arcticus J. Sahlberg, 1890
Bledius (Hesperophilus) littoralis Heer, 1839
Bledius glaciatus ♰ S. H. Scudder, 1890Uruguay waterprimrose
Bledius litoralis Heer, 1839
Bledius suturalis LeConte, 1863
Blitophaga reitteri Portevin, 1926
Blitophaga vetusta
Boreaphilus henningianus Sahlberg, 1832Staph beetle
Cholevinus sibiricus Jeannel, 1923
Coryphium angusticolle Stephens, 1834
Cryptobium cinctum ♰ S. H. Scudder, 1900
Cryptobium detectum Scudder, 1900
Deleaster dryophilus Gistel, 1857
Dianous nitidulus (LeConte, 1874)⇒ Dianous dubiosus Puthz, 2000
Dinothenarus (Dinothenarus) pubescens De Geer, 1774
Elacatophora apion Jaloszynski, 2004
Euaesthetus laeviusculus Mannerheim, 1844Staph beetle
Eucnecosum brachypterum Gravenhorst, 1802
Eucnecosum brunnescens Sahlberg, 1871
Eucnecosum tenue LeConte, 1863Staph beetle
Eusilpha japonica Motschulsky, 1862
Eusteniamorpha aceriabdomena Lin, 2000
Geodromicus (Geodromicus) plagiatus (Fabricius, 1798)⇒ Geodromicus nigrita P. W. J. Mül..., 1821
Geodromicus stiricidii ♰ S. H. Scudder, 1895
Gymnusa absens ♰ S. H. Scudder, 1900
Gymnusa antiqua ♰ A. M. Lomnicki, 1894
Gyrohypnus punctulatus (Paykull, 1789)⇒ Gyrohypnus liebei Scheerpeltz, 1926
Heterosilpha ramosa Say, 1823Garden Carrion Beetle
Holoboreaphilus nordenskioeldi Maklin, 1878Staph beetle
Holoboreaphilus nordenskioldi Mäklin, 1878
Hydnobius punctatus Sturm, 1807
Hydraena (Hydraena) riparia Kugelann, 1794Southern threecornerjack
Hydraena circulata Perkins, 1980
Hydraena coopei Angus, 1997
Hydraena reidiana ♰ Lesne, 1920
Ischnosoma pictum Horn, 1877
Kalissus nitidus LeConte, 1874
Lathrobium antiquatum ♰ S. H. Scudder, 1900
Lathrobium debilitatum ♰ S. H. Scudder, 1900
Lathrobium exesum ♰ S. H. Scudder, 1900
Lathrobium frustum ♰ S. H. Scudder, 1900
Lathrobium inhibitum ♰ S. H. Scudder, 1900
Lathrobium interglaciale ♰ S. H. Scudder, 1890
Leptotyphlus omessae mutilus Orousset, 1984
Lesteva pallipes LeConte, 1863
Limnebius truncatellus Thunberg, 1794
Lomechusoides strumosa Fabricius, 1775
Lordithon longiceps LeConte, 1863
Micralymma brevilingue Schiødte, 1857
Microedus austinianus LeConte, 1874
Micropeplus caelatus Erichson, 1839
Micropeplus cribratus LeConte, 1863
Micropeplus fulvus Erichson, 1840
Micropeplus laticollis Mäklin, 1853
Micropeplus nelsoni Campbell, 1989
Micropeplus porcatus Paykull, 1789
Micropeplus punctatus (LeConte, 1877)⇒ Micropeplus taiwanensis Compbell, 1992
Micropeplus staphylinoides Marsham, 1802
Mycetoporus punctatissimus Hatch, 1957
Nicrophorus guttula Motschulsky, 1845Yellow-bellied Burying Beetle
Nicrophorus marginatus Fabricius, 1801Margined Burying Beetle
Nicrophorus nigrita Mannerheim, 1843Arizona cottonwood
Nitidotachinus lanei Hatch, 1957
Nitidotachinus tachyporoides Horn, 1877
Ochthebius (Asiobates) bicolon (Germar, 1824)⇒ Ochthebius dilatatus Stephens, 1829
Ochthebius (Asiobates) dilatatus Stephens, 1829Jumping-bean sapium
Ochthebius (Asiobates) discretus LeConte, 1878
Ochthebius (Asiobates) minimus (Fabricius, 1792)⇒ Protochthebius jagthanae Champion, 1921
Ochthebius holmbergi (Mannerheim, 1853)⇒ Ochthebius marinus Paykull, 1798
Ochthephilum cinctum Scudder, 1900
Ochthephilum detectum Scudder, 1900
Ocypus (Ocypus) olens Müller, 1764Cocktail
Ocypus (Pseudocypus) fuscatus Gravenhorst, 1802
Olophrum arcanum ♰ S. H. Scudder, 1900
Olophrum assimile Paykull, 1800
Olophrum boreale Paykull, 1792Staph beetle
Olophrum celatum ♰ S. H. Scudder, 1900
Olophrum consimile Gyllenhal, 1810Staph beetle
Olophrum dejectum ♰ S. H. Scudder, 1900
Olophrum fuscum Gravenhorst, 1806
Olophrum henryi ♰ Herman, 2001
Olophrum interglaciale ♰ E. Mjöberg, 1904
Olophrum latum Maklin, 1853Staph beetle
Olophrum obtectum Erichson, 1840
Olophrum rotundicolle Sahlberg, 1817Staph beetle
Osorius brunnicornis ♰ Hope, 1837
Oxyporus stiriacus ♰ S. H. Scudder, 1890
Oxytelus (Oxytelus) fulvipes Erichson, 1839
Oxytelus (Tanycraerus) laqueatus Marsham, 1802Rove beetle
Oxytelus gibbulus Eppelsheim, 1877
Oxytelus niger Le Conte, 1877
Paederus (Paederus) riparius Gravenhorst, 1802
Pelecomalium testaceum Mannerheim, 1843
Philonthus (Philonthus) linki Solsky, 1867
Philonthus (Philonthus) rotundicollis Ménétriés, 1832
Philonthus (Trionthus) claudus ♰ S. H. Scudder, 1900
Phosphuga atrata Linnaeus, 1758Black Carrion Beetle
Platystethus (Craetopycrus) cornutus Gravenhorst, 1802
Platystethus (Craetopycrus) nitens C. R. Sahlberg, 1832
Platystethus (Craetopycrus) nodifrons Mannerheim, 1830
Platystethus (Platystethus) arenarius Fourcroy, 1785
Platystethus americanus Erichson, 1840Big spore andreaeobryum moss
Proscapheus zhangla Háva, J.Schn..., 1999
Pselaphaulax dresdensis Herbst, 1792
Pselaphus heisei Herbst, 1792
Pseudopsis sulcata Newman, 1834Staph beetle
Pteroloma forsstromii Gyllenhal, 1810
Pycnoglypta aptera Campbell, 1983
Pycnoglypta campbelli Gusarov, 1995
Pycnoglypta lurida Gyllenhal, 1813Staph beetle
Quedius (Microsaurus) uteanus Casey, 1915
Quedius (Raphirus) boops Gravenhorst, 1802
Quedius (Raphirus) deperditus ♰ S. H. Scudder, 1900
Silpha obscura Linnaeus, 1758Gray bushmallow
Silpha perforata Gebler, 1832
Silpha tristis Illiger, 1798Dwarf sand verbena
Sonoma margemina Park & Wagner, 1962
Staphylinus olens (Müller, 1764)⇒ Ocypus olens olens O. Müller, 1764
Staphylinus pubescens (De Geer, 1774)⇒ Dinothenarus pubescens pubescens De Geer, 1774
Stenus (Hemistenus) pubescens Stephens, 1833Staph beetle
Stenus (Nestus) assequens Rey, 1884
Stenus (Nestus) canaliculatus Gyllenhal, 1827Amaumau fern
Stenus (Nestus) carbonarius Gyllenhal, 1827
Stenus (Nestus) mammops Casey, 1884
Stenus (Nestus) melanarius Stephens, 1833
Stenus (Nestus) sordidus Puthz, 1988
Stenus (Stenus) biguttatus Linnaeus, 1758Cornish mallow
Stenus (Stenus) comma LeConte, 1863
Stenus (Stenus) juno Paykull, 1789Richard's macromitrium moss
Stenus (Tesnus) brunnipes Stephens, 1833
Stenus (Tesnus) hyperboreus (Sahlberg, 1876)⇒ Stenus hyperboreus Sahlberg, 1876
Stenus (Tesnus) vinnulus Casey, 1884
Stenus pinguis Casey, 1884
Stenus scrupeus Casey, 1884
Stilicus similis (Erichson, 1839)⇒ Rugilus similis Erichson, 1839
Syntomium malkini Hatch, 1957
Tachinus (Drymoporus) elongatus Gyllenhal, 1810Staph beetle
Tachinus (Tachinus) angustatus Horn, 1877
Tachinus (Tachinus) crotchii Horn, 1877
Tachinus (Tachinus) fimbriatus Gravenhorst, 1802
Tachinus (Tachinus) frigidus Erichson, 1839
Tachinus (Tachinus) instabilis Mäklin, 1853Staph beetle
Tachinus (Tachinus) luridus Erichson, 1840
Tachinus (Tachinus) mimus Horn, 1877
Tachinus (Tachinus) schwarzi Horn, 1877
Tachinus apertus
Tachinus arcticus Motschulsky, 1860
Tachinus tachyporoides Horn, 1877
Tachinus thruppi (Hatch, 1957)⇒ Tachinus frigidus Erichson, 1839
Tachyporus borealis Campbell, 1979
Tachyporus nimbicola Campbell, 1979
Tachyporus rulomus Blackwelder, 1936
Temnodera testacea ♰ Hope, 1837
Thanatophilus baicalicus (Motschulsky, 1860)⇒ Thanatophilus trituberculatus Kirby, 1837
Thanatophilus coloradensis Wickham, 1902
Thanatophilus dispar Herbst, 1793
Thanatophilus lapponicus Herbst, 1793Northern Carrion Beetle
Thanatophilus sagax Mannerheim, 1853
Xantholinus (Xantholinus) linearis (Olivier, 1795)⇒ Neohypnus lineatus Wendeler, 1924
Xantholinus linearis anglicus Lesne, 1926

Subnodes of Staphylinoidea with occurrences in Quaternary
FamilyAgyrtidae
Thomson, 1859
Primitive carrion beetles		1
1		Genus
Species
Taxom

FamilyHydraenidae
Mulsant, 1844
Cascade Beetles, Hydraenid beetles, minute moss beetle, Minute moss beetles		3
10		Genera
Species
Taxom

FamilyLeiodidae
Fleming, 1821
Round fungus beetles		5
6		Genera
Species
Taxom

FamilyPtiliidae
Heer, 1843
Feather-wing beetles, Feather-winged beetles, Featherwing beetles, Featherwinged Beetles, Horseshoe crab beetles		1
0		Genus
Species
Taxom

FamilyScydmaenidae
Leach, 1815
Ant-like stone beetles, Scydmaenines		1
1		Genus
Species
Taxom

FamilySilphidae
Latreille, 1807
Burying And Carrion Beetles, Burying beetles, Carrion Beetle, Carrion beetles, Large carrion beetles, Sexton beetles		9
24		Genera
Species
Taxom

FamilyStaphylinidae
Lameere, 1900
Rove beetles, Short-winged mold beetles, Staphs		77
146		Genera
Species
Taxom



Links, sources and further information
Staphylinoidea at artfakta.se
Staphylinoidea at dyntaxa.se
Staphylinoidea at itis.gov