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Checklist for Family Leucosiidae Samouelle, 1819 (nut crabs)
and region Continents


Class
 
Malacostraca
Latreille, 1802
Higher Crustaceans, Malacostracans
734
7366
51296
Families
Genera
Species
 
Taxom
 

Subclass
 
Eumalacostraca
Grobben, 1892
716
7312
51144
Families
Genera
Species
 
Taxom
 

Superorder
 
Eucarida
Calman, 1904
268
3165
24238
Families
Genera
Species
 
Taxom
 

Order
 
Decapoda
Latreille, 1802
Crabs, Crayfishes, Decapod, Decapods, Lobsters, Prawns
265
3148
24112
Families
Genera
Species
 
Taxom
 

Suborder
 
Pleocyemata
Burkenroad, 1963
232
2987
23012
Families
Genera
Species
 
Taxom
 

Infraorder
 
Brachyura
Linnaeus, 1758
Crabs, Short-tailed crabs, True crabs
131
1846
11236
Families
Genera
Species
 
Taxom
 

Section
 
Eubrachyura
de Saint Laurent, 1980
106
1646
10355
Families
Genera
Species
 
Taxom
 

Subsection
 
Heterotremata
Guinot, 1977
85
1358
8228
Families
Genera
Species
 
Taxom
 

Superfamily
 
Leucosioidea
Samouelle, 1819
2
90
683
Families
Genera
Species
 
Taxom
 
FamilyLeucosiidae
Samouelle, 1819
Nut crabs, Pebble crabs, Purse crabs
88
669
Genera
Species
 
Taxom
 


Genera for Continents
88 Genera with 669 Species
#ABCDEFGHIKLMNOPRSTU
Acanthilia Galil, 18861SpeciesTaxom
Actaeomorpha Miers, 19354SpeciesTaxom
Afrophila Galil, 18551SpeciesTaxom
Alox C. G. S. Tan..., 199511SpeciesTaxom
Ampliura Morris & Col..., 19911SpeciesTaxom
Ancylodactyla Galil, 19903SpeciesTaxom
Arcania Leach, 184127SpeciesTaxom
Atlantolocia Galil, 18811SpeciesTaxom
Atlantophila Galil, 18811SpeciesTaxom
Atlantotlos Doflein, 19041SpeciesTaxom
Bellidilia Kinahan, 18563SpeciesTaxom
Callidactylus Stimpson, 18711SpeciesTaxom
Cateios C. G. S. Tan..., 18841SpeciesTaxom
Coleusia Galil, 18016SpeciesTaxom
Coralliocryptus Komai & Ng, 20121SpeciesTaxom
Cryptocnemus Stimpson, 192720SpeciesTaxom
Dolos C. G. S. Tan..., 18741SpeciesTaxom
Duncanitrix Schweitzer, ..., 2011
0SpeciesTaxom
Dusa Münster, 18395SpeciesTaxom
Ebalia Leach, 189590SpeciesTaxom
Ebaliopsis Ihle, 19422SpeciesTaxom
Euclosia Galil, 20078SpeciesTaxom
Euclosiana Galil & Ng, 20079SpeciesTaxom
Favus Lanchester, 19001SpeciesTaxom
Galilia Ng & Richer ..., 20142SpeciesTaxom
Gemmacarcinus Müller & Col..., 19911SpeciesTaxom
Gemmellarocarcinus Checchia-Ris..., 19051SpeciesTaxom
Hepatinulus Ristori, 1886
0SpeciesTaxom
Heterolithadia Alcock, 18931SpeciesTaxom
Heteronucia Alcock, 190612SpeciesTaxom
Hiplyra Galil, 18307SpeciesTaxom
Ihleus Ovaere, 19892SpeciesTaxom
Ilia Leach, 18817SpeciesTaxom
Iliacantha Stimpson, 18717SpeciesTaxom
Iphis Leach, 18522SpeciesTaxom
Ixa Leach, 187911SpeciesTaxom
Ixoides MacGilchrist, 19051SpeciesTaxom
Kabutos Komatsu & Ng, 19551SpeciesTaxom
Leucisca MacLeay, 18383SpeciesTaxom
Leucosia Weber, 187583SpeciesTaxom
Leucosides Rathbun, 19112SpeciesTaxom
Leucosilia Bell, 19034SpeciesTaxom
Lissomorpha Ward, 19331SpeciesTaxom
Lithadia Bell, 19738SpeciesTaxom
Lyphira Galil, 20092SpeciesTaxom
Merocryptoides T. Sakai, 20013SpeciesTaxom
Merocryptus A. Milne-Edw..., 18985SpeciesTaxom
Myra Leach, 200131SpeciesTaxom
Myrine Galil, 18753SpeciesTaxom
Myrodes Bell, 18551SpeciesTaxom
Myropsis Stimpson, 18711SpeciesTaxom
Naesa Leach, 18181SpeciesTaxom
Nobiliella Komatsu & Ta..., 20191SpeciesTaxom
Nucia Dana, 187414SpeciesTaxom
Nucilobus Morris & Col..., 19911SpeciesTaxom
Nuciops Serène & Soh, 19181SpeciesTaxom
Nursia Leach, 191819SpeciesTaxom
Nursilia Bell, 18964SpeciesTaxom
Oreophorus Rüppell, 19897SpeciesTaxom
Oreotlos Ihle, 199311SpeciesTaxom
Orientotlos T. Sakai, 19801SpeciesTaxom
Palaeomyra A. Milne-Edw..., 18611SpeciesTaxom
Paranursia Serène & Soh, 18551SpeciesTaxom
Parilia Wood-Mason &..., 19615SpeciesTaxom
Persephona Leach, 189418SpeciesTaxom
Philyra Leach, 189354SpeciesTaxom
Phlyxia Bell, 18804SpeciesTaxom
Praebebalia Rathbun, 197715SpeciesTaxom
Praosia C. G. S. Tan..., 19931SpeciesTaxom
Pseudomyra Capart, 19511SpeciesTaxom
Pseudophilyra Miers, 013SpeciesTaxom
Pterocarcinus Blow, 20031SpeciesTaxom
Pyrhila Galil, 18412SpeciesTaxom
Randallia Stimpson, 198923SpeciesTaxom
Raylilia Galil, 19955SpeciesTaxom
Ryphila Galil, 18933SpeciesTaxom
Seulocia Galil, 185810SpeciesTaxom
Soceulia Galil, 18554SpeciesTaxom
Speleophorus A. Milne-Edw..., 18551SpeciesTaxom
Speloeophorus A. Milne-Edw..., 19356SpeciesTaxom
Tanaoa Galil, 19899SpeciesTaxom
Tlos Adams & White, 18493SpeciesTaxom
Tokoyo Galil, 18962SpeciesTaxom
Toru Galil, 19614SpeciesTaxom
Typilobus Stoliczka, 20016SpeciesTaxom
Uhlias Stimpson, 18712SpeciesTaxom
Urashima Galil, 19611SpeciesTaxom
Urnalana Galil, 185522SpeciesTaxom


Species list for Continents
669 Species and 10 Subspecies
Acanthilia intermedia Miers, 1886Granulose purse crab
Actaeomorpha alvae Boone, 1934
Actaeomorpha erosa Miers, 1877Pointtip twinpod
Actaeomorpha morum (Alcock, 1896)⇒ Drachiella morum Alcock, 1896
Actaeomorpha punctata Edmondson, 1935
Afrophila punctata Bell, 1855
Alox antheos C. G. S. Tan..., 1995
Alox glene C. G. S. Tan..., 1995
Alox latusoides Sakai, 1937
Alox liklik N.g.Galil, 2015
Alox naispela N.g.Galil, 2015Naispela Crab
Alox ornatum Ihle, 1918
Alox patella Alcock, 1896
Alox rugosum Stimpson, 1858
Alox somphos C. G. S. Tan..., 1995
Alox tormos Galil & Ng, 2009
Alox zalion C. G. S. Tan..., 1995
Ampliura simplex Morris & Col..., 1991
Ancylodactyla elata Zarenkov, 1994
Ancylodactyla elongata Zarenkov, 1969
Ancylodactyla nana Zarenkov, 1990
Arcania (Iphis) quinquespinosa (Alcock & And..., 1894)⇒ Arcania gracilis Henderson, 1893
Arcania (Iphis) septemspinosa Fabricius, 1787
Arcania brevifrons Chen, 1989
Arcania cornigera Naruse, 2014
Arcania cornuta MacGilchrist, 1905
Arcania echinata Galil, 2001
Arcania elongata Yokoya, 1933
Arcania erinacea Fabricius, 1787
Arcania erinaceus
Arcania foliolata Galil, 2001
Arcania fungilifera Galil, 2001
Arcania globata Stimpson, 1858
Arcania gracilipes (Bell, 1855)⇒ Raylilia gracilipes Bell, 1855
Arcania gracilis Henderson, 1893
Arcania granulosa (Miers, 1877)⇒ Arcania undecimspinosa De Haan, 1841
Arcania heptacantha De Haan, 1861
Arcania muricata Galil, 2001
Arcania novemspinosa Lichtenstein, 1816
Arcania novemspinosa aspera Miers, 1880
Arcania pulcherrima (Haswell, 1879)⇒ Ixa pulcherrima Haswell, 1879
Arcania sagamiensis Sakai, 1969
Arcania siamensis (Rathbun, 1909)⇒ Arcania septemspinosa Fabricius, 1787
Arcania spinixa (Zarenkov, 1994)⇒ Arcania cornuta MacGilchrist, 1905
Arcania spinosa
Arcania tropicalis Naruse, 2014
Arcania tuberculata Bell, 1855
Arcania undecimspinosa De Haan, 1841Three-spined Box Crab
Atlantolocia laevidorsalis Miers, 1881
Atlantophila cristata Miers, 1881
Atlantotlos rhombifer Doflein, 1904
Bellidilia cheesmani Filhol, 1886
Bellidilia laevis Bell, 1855Smooth Pebble Crab
Bellidilia undecimspinosa Kinahan, 1856Large Pebble Crab
Callidactylus asper Stimpson, 1871Spurfinger purse crab
Cateios frontalis Miers, 1884
Coleusia biannulata Tyndale-Bisc..., 1962
Coleusia janani Giraldes, Al..., 2017
Coleusia magna Tyndale-Bisc..., 1962
Coleusia rangita Galil, 2006
Coleusia signata Paul´son, 1875Pebble Crab
Coleusia urania Herbst, 1801
Coralliocryptus caementa Komai & Ng, 2012
Cryptocnemus aberrans Balss, 1938
Cryptocnemus calmani Ihle, 1915
Cryptocnemus crenulatus Grant & McCu..., 1906
Cryptocnemus grandidieri A. Milne-Edw..., 1865
Cryptocnemus haddoni Calman, 1900
Cryptocnemus hemispheroides Campbell, 1971
Cryptocnemus holdsworthi Miers, 1877
Cryptocnemus kamekii Sakai, 1961
Cryptocnemus macrognathus Ihle, 1918
Cryptocnemus marginatus Sakai, 1983
Cryptocnemus mortenseni Rathbun, 1909
Cryptocnemus obolus Ortmann, 1892
Cryptocnemus pentagonus Stimpson, 1858
Cryptocnemus planus Ward, 1933
Cryptocnemus siamensis Serène & Soh, 1976
Cryptocnemus stimpsoni Ihle, 1915
Cryptocnemus trapezoides Ihle, 1915
Cryptocnemus trigonus Komatsu & Ta..., 2000
Cryptocnemus tuberosus Klunzinger, 1906
Cryptocnemus vincentianus Hale, 1927Tortoise-crab
Dolos petraeus A. Milne-Edw..., 1874
Dusa araneae ♰ Schweigert, 2002
Dusa bronni Oppel, 1862
Dusa denticulata ♰ Munster, 1839
Dusa longipes ♰ Pinna, 1974
Dusa monocera ♰ Munster, 1839
Ebalia (Phlyxia) crassipes Bell, 1885
Ebalia (Phlyxia) deutipons
Ebalia (Phlyxia) intermedia Miers, 1886Smooth Nut Crab
Ebalia (Phlyxia) orbicularis
Ebalia (Phlyxia) quadridentata Gray, 1831
Ebalia quadridentata spinifera Miers, 1886
Ebalia undecispinosa orbicularis
Ebalia abdominalis Nobili, 1906
Ebalia affinis Miers, 1881
Ebalia agglomus Barnard, 1955
Ebalia algirica (Lucas, 1846)⇒ Ebalia edwardsii Costa, 1838
Ebalia ambigua (Bouvier, 1940)⇒ Ebalia edwardsii Costa, 1838
Ebalia americana (Rathbun, 1894)⇒ Randallia americana Rathbun, 1894
Ebalia barnardi Stebbing, 1914
Ebalia brevimana Campbell, 1971
Ebalia bryerii (Leach, 1817)⇒ Ebalia tumefacta Montagu, 1808
Ebalia cariosa Stimpson, 1860Sculptured clutch crab
Ebalia cheesmani (Filhol, 1886)⇒ Bellidilia cheesmani Filhol, 1886
Ebalia clarionensis Rathbun, 1935
Ebalia cranchi
Ebalia cranchii Leach, 1817Cranch´s nut crab
Ebalia cristata Rathbun, 1898
Ebalia cryptocnemoides Takeda & Miyake, 1972
Ebalia dentifrons Miers, 1886
Ebalia deshayesi Lucas, 1846
Ebalia dimorphoides Sakai, 1963
Ebalia edwardsi (A. Milne-Edw..., 1900)⇒ Ebalia deshayesi Lucas, 1846
Ebalia edwardsii Costa, 1838Edward's Nut Crab
Ebalia erosa (A. Milne Edw..., 1873)⇒ Ebaliopsis erosa A. Milne-Edw..., 1873
Ebalia fasciata (Ihle, 1918)⇒ Praebebalia fasciata Ihle, 1918
Ebalia glans Alcock, 1896
Ebalia globulosa Müller, 1976
Ebalia glomus Stebbing, 1921
Ebalia gotoensis (Rathbun, 1932)⇒ Ebalia longimana Ortmann, 1892
Ebalia granulata Rüppell, 1830
Ebalia granulosa H. Milne Edw..., 1837
Ebalia hancocki Rathbun, 1933Oxystomatous crabs
Ebalia hayamaensis Sakai, 1963
Ebalia heterochalaza Kemp, 1918
Ebalia humilis Takeda, 1977
Ebalia hungarica ♰ P. Muller, 1974
Ebalia japonica (Rathbun, 1932)⇒ Ebalia tuberculosa Milne-Edwards, 1873
Ebalia jordani Rathbun, 1906
Ebalia kakegawaensis ♰ Karasawa, 1993
Ebalia lacertosa Nobili, 1906
Ebalia laevis Bell, 1885
Ebalia lambriformis Bell, 1885
Ebalia longimana Ortmann, 1892
Ebalia longispinosa Ihle, 1918
Ebalia magdalenensis Rathbun, 1933
Ebalia maldivensis Borradaile, 1903
Ebalia malefactrix (Kemp, 1915)⇒ Philyra malefactrix Kemp, 1915
Ebalia miliaris (A. Milne-Edw..., 1873)⇒ Nucia miliaris A. Milne-Edw..., 1873
Ebalia nana Ihle, 1918
Ebalia nobilii Balss, 1916
Ebalia nudipes Sakai, 1963
Ebalia nux A. Milne-Edw..., 1883
Ebalia orientalis Kossmann, 1877
Ebalia pennantii (Leach, 1817)⇒ Ebalia tuberosa Pennant, 1777
Ebalia philippinensis Chen, 1989
Ebalia pondoensis Barnard, 1955
Ebalia postulans Stebbing, 1910
Ebalia pulchella (A. Milne-Edw..., 1873)⇒ Nucia pulchella A. Milne-Edw..., 1873
Ebalia punctulata Sakai, 1983
Ebalia quadrata A. Milne-Edw..., 1873
Ebalia ramsayi Haswell, 1880
Ebalia rhomboidalis Miers, 1879
Ebalia rotundata A. Milne-Edw..., 1880
Ebalia sakaii Takeda & Miyake, 1972
Ebalia salamensis (Doflein, 1904)⇒ Ebalia tuberculosa Milne-Edwards, 1873
Ebalia scabra (Dai, Yang, S..., 1984)⇒ Philyra scabra Dai, Yang, S..., 1984
Ebalia scabriuscula Ortmann, 1892
Ebalia scandens Stebbing, 1910
Ebalia serenei Chen, 1989
Ebalia spinifera Miers, 1886
Ebalia spinosa ♰ A. Milne-Edw..., 1873
Ebalia stellaris Naruse & Ng, 2006
Ebalia stimpsoni A. Milne-Edw..., 1880Thinarm clutch crab
Ebalia stimpsonii (A. Milne-Edw..., 1880)⇒ Ebalia stimpsoni A. Milne-Edw..., 1880
Ebalia takedai Komatsu, Man..., 2012
Ebalia tosaensis Sakai, 1963
Ebalia tuberculata ♰ (Miers, 1881)⇒ Ebalia yokoyai Sakai, 1965
Ebalia tuberculosa Milne-Edwards, 1873Nut-crab
Ebalia tuberosa Pennant, 1777Pennant´s nut crab
Ebalia tumefacta Montagu, 1808Bryer's Nut Crab
Ebalia undecimspinosa
Ebalia webberi Komatsu & Ta..., 2007
Ebalia woodmasoni Alcock, 1896
Ebalia yokoyai Sakai, 1965
Ebalia ypsilon Ortmann, 1895
Ebaliopsis erosa A. Milne-Edw..., 1873
Ebaliopsis viadieri Ward, 1942
Euclosia concinna (Galil, 2003)⇒ Euclosiana concinna Galil, 2003
Euclosia crosnieri (Chen, 1989)⇒ Euclosiana crosnieri Chen, 1989
Euclosia exquisita (Galil, 2003)⇒ Euclosiana exquisita Galil, 2003
Euclosia obtusifrons (De Haan, 1841)⇒ Euclosiana obtusifrons De Haan, 1841
Euclosia scitula (Galil, 2003)⇒ Euclosiana scitula Galil, 2003
Euclosia tornatilia (Galil, 2003)⇒ Euclosiana tornatilia Galil, 2003
Euclosia unidentata (De Haan, 1841)⇒ Euclosiana unidentata De Haan, 1841
Euclosia vella (Galil, 2007)⇒ Euclosiana vella Galil, 2007
Euclosiana concinna Galil, 2003
Euclosiana crosnieri Chen, 1989
Euclosiana exquisita Galil, 2003
Euclosiana guinotae Galil & Ng, 2010
Euclosiana obtusifrons De Haan, 1841
Euclosiana scitula Galil, 2003
Euclosiana tornatilia Galil, 2003
Euclosiana unidentata De Haan, 1841One Teeth Peeble Crab
Euclosiana vella Galil, 2007
Favus granulatus Lanchester, 1900
Galilia narusei Ng & Richer ..., 2007
Galilia petricola Komai & Tsuc..., 2014
Gemmacarcinus fossatus ♰ Muller, 1991
Gemmellarocarcinus loerentheyi Checchia-Ris..., 1905
Heterolithadia fallax Henderson, 1893
Heteronucia angulata Barnard, 1947
Heteronucia fujitai Komatsu & Ta..., 2009Leucosiid Crab
Heteronucia globata Sakai, 1963
Heteronucia granulata Komatsu & Ta..., 2005
Heteronucia ingens (Rathbun, 1911)⇒ Nucia ingens Rathbun, 1911
Heteronucia laminata Doflein, 1904
Heteronucia mesanensis Rathbun, 1909
Heteronucia perlata Sakai, 1963Pearl Granulate Purse Crab
Heteronucia spinifera Edmondson, 1951Leucosiid Crab
Heteronucia toyoshioae Komatsu & Ta..., 2005
Heteronucia tuberculata Chen & Türkay, 2001
Heteronucia venusta Nobili, 1906Bullet Crab
Hiplyra elegans Gravier, 1920
Hiplyra longimana A. Milne Edw..., 1874
Hiplyra platycheir De Haan, 1841
Hiplyra ramil Naderloo & Apel, 2012
Hiplyra sagitta Galil, 2009
Hiplyra variegata Rüppell, 1830
Hiplyra varietgata Rüppell, 1830
Ihleus lanatus Alcock, 1896
Ihleus villosus Chen, 1989
Ilia laevidorsalis
Ilia mariannae (Herklots, 1852)⇒ Pariphiculus mariannae Herklots, 1852
Ilia nucleus Linnaeus, 1758Nut-crab
Ilia pliocaenica Ristori, 1891
Ilia polita
Ilia rugulosa (Risso, 1827)⇒ Ilia nucleus Linnaeus, 1758
Ilia spinosa Miers, 1881
Iliacantha hancocki Rathbun, 1935
Iliacantha intermedia (Miers, 1886)⇒ Acanthilia intermedia Miers, 1886
Iliacantha liodactylus Rathbun, 1898
Iliacantha panamanica Todd & Collins, 2005
Iliacantha schmitti Rathbun, 1935
Iliacantha sparsa Stimpson, 1871Shouldered purse crab
Iliacantha subglobosa Stimpson, 1871Longfinger purse crab
Iphis heptacantha (Herklots, 1861)⇒ Arcania heptacantha De Haan, 1861
Iphis longipes (Dana, 1852)⇒ Arcania septemspinosa Fabricius, 1787
Ixa acuta Tyndale-Bisc..., 1962
Ixa canaliculata (Leach, 1817)⇒ Ixa cylindrus Fabricius, 1777
Ixa cylindra Fabricius, 1777
Ixa cylindricus (Fabricius, 1777)⇒ Ixa cylindrus Fabricius, 1777
Ixa cylindrus Fabricius, 1777Purse Crab
Ixa cylindrus megaspis Adams & White, 1849
Ixa edwardsii Lucas, 1858
Ixa inermis Leach, 1817
Ixa megaspis Adams & White, 1849
Ixa monodi Holthuis & G..., 1956
Ixa pulcherrima Haswell, 1879
Ixoides cornutus (MacGilchrist, 1905)⇒ Arcania cornuta MacGilchrist, 1905
Kabutos durandi Serène, 1955
Leucisca halimedophila Galil & Inno..., 2019
Leucisca levigena George & M. ..., 1976
Leucisca squalina MacLeay, 1838
Leucosia affinis Bell, 1855
Leucosia anatum Herbst, 1783Painted Pebble Crab
Leucosia australiensis (Miers, 1880)⇒ Leucosia anatum Herbst, 1783
Leucosia biannulata (Tyndale-Bisc..., 1962)⇒ Coleusia biannulata Tyndale-Bisc..., 1962
Leucosia bikiniensis (Sakai, 1983)⇒ Urnalana elata A. Milne-Edw..., 1873
Leucosia brumea
Leucosia brunnea (Miers, 1877)⇒ Soceulia brunnea Miers, 1877
Leucosia calcarata Collins & al., 2003
Leucosia cheverti Haswell, 1879
Leucosia corallicola Alcock, 1896
Leucosia craniolaris Linnaeus, 1758
Leucosia craniolaris laevimana Miers, 1884
Leucosia crosnieri (Chen, 1989)⇒ Euclosiana crosnieri Chen, 1989
Leucosia cumingi
Leucosia cumingii (Bell, 1855)⇒ Urnalana cumingii Bell, 1855
Leucosia elata (A. Milne-Edw..., 1873)⇒ Urnalana elata A. Milne-Edw..., 1873
Leucosia elatoides (Bouvier, 1915)⇒ Urnalana elatoides Bouvier, 1915
Leucosia foresti (Chen, 1989)⇒ Urnalana foresti Chen, 1989
Leucosia formosensis Sakai, 1937
Leucosia fugax (Fabricius, 1798)⇒ Myra fugax Fabricius, 1798
Leucosia grandis (Chen & Türkay, 2002)⇒ Coleusia urania Herbst, 1801
Leucosia haematostica
Leucosia haematosticta (Adams & White, 1849)⇒ Urnalana haematosticta Adams & White, 1849
Leucosia haswelli Miers, 1886
Leucosia hilaris (Nobili, 1905)⇒ Urnalana hilaris Nobili, 1905
Leucosia hilarus
Leucosia insularis (Takeda & Kurata, 1976)⇒ Urnalana insularis Takeda & Kurata, 1976
Leucosia jecusculum Rathbun, 1911
Leucosia jurinei Saussure, 1835
Leucosia laevimana Miers, 1884
Leucosia leslii Haswell, 1879
Leucosia longibrachia Shen & Chen, 1978
Leucosia longiferus
Leucosia longifrons (De Haan, 1841)⇒ Leucosia anatum Herbst, 1783
Leucosia longifrons pulcherrima
Leucosia longimaculata Chen & Fang, 1991
Leucosia maculata (Stimpson, 1858)⇒ Seulocia rhomboidalis De Haan, 1841
Leucosia magna (Tyndale-Bisc..., 1962)⇒ Coleusia magna Tyndale-Bisc..., 1962
Leucosia major (Chen & Ng, 2003)⇒ Soceulia major Chen & Ng, 2003
Leucosia margaritacea Bell, 1855
Leucosia margaritata (A. Milne-Edw..., 1873)⇒ Urnalana margaritata A. Milne-Edw..., 1873
Leucosia marmorea (White, 1847)⇒ Soceulia marmorea Bell, 1855
Leucosia mimasensis (Sakai, 1969)⇒ Euclosiana obtusifrons De Haan, 1841
Leucosia moresbiensis Haswell, 1880
Leucosia neocaledonica (A. Milne-Edw..., 1873)⇒ Leucosia anatum Herbst, 1783
Leucosia obscura (Bell, 1855)⇒ Leucosia craniolaris Linnaeus, 1758
Leucosia obtusifrons (De Haan, 1841)⇒ Euclosiana obtusifrons De Haan, 1841
Leucosia obtusifrons unidentata
Leucosia ocellata Bell, 1855
Leucosia ornata (Miers, 1877)⇒ Leucosia anatum Herbst, 1783
Leucosia pallida (Bell, 1855)⇒ Leucosia craniolaris Linnaeus, 1758
Leucosia pallida obscura
Leucosia parvimana (Stimpson, 1858)⇒ Leucosia craniolaris Linnaeus, 1758
Leucosia perlata (De Haan, 1841)⇒ Leucosia craniolaris Linnaeus, 1758
Leucosia perryi (Miers, 1876)⇒ Pseudophilyra perryi Miers, 1876
Leucosia persica ♰ F. J. Vega, 2010
Leucosia polita (Hess, 1865)⇒ Leucosia anatum Herbst, 1783
Leucosia prevostiana Desmarest, 1817
Leucosia pseudomargaritata (Chen, 1987)⇒ Urnalana pulchella Bell, 1855
Leucosia pubescens (Miers, 1877)⇒ Seulocia pubescens Miers, 1877
Leucosia pulchella (Bell, 1855)⇒ Urnalana pulchella Bell, 1855
Leucosia pulcherrima (Miers, 1877)⇒ Leucosia anatum Herbst, 1783
Leucosia punctata Bell, 1855
Leucosia purarensis (Ovaere, 1987)⇒ Urnalana purarensis Ovaere, 1987
Leucosia reticulata Miers, 1877
Leucosia rhomboidalis (De Haan, 1850)⇒ Seulocia rhomboidalis De Haan, 1841
Leucosia rubripalma Galil, 2003Pebble Crab
Leucosia sagamiensis (Sakai, 1961)⇒ Urnalana elata A. Milne-Edw..., 1873
Leucosia serenei Morris & Col..., 1991
Leucosia signata (Paulson, 1875)⇒ Coleusia signata Paul´son, 1875
Leucosia sima Alcock, 1896
Leucosia sinica (Shen & Chen, 1978)⇒ Seulocia vittata Stimpson, 1858
Leucosia splendida (Haswell, 1879)⇒ Leucosia anatum Herbst, 1783
Leucosia takamii ♰ Karasawa, 1993
Leucosia tetraodon Bouvier, 1914
Leucosia tricarinata ♰ Martin, 1880
Leucosia tutongensis Morris & Col..., 1991
Leucosia unidentata (De Haan, 1841)⇒ Euclosiana unidentata De Haan, 1841
Leucosia urania Herbst, 1801
Leucosia vittata (Stimpson, 1858)⇒ Seulocia vittata Stimpson, 1858
Leucosia whitei (Bell, 1855)⇒ Urnalana whitei Bell, 1855
Leucosia whitemeei
Leucosia whitmeei Miers, 1875
Leucosides angulata (Rathbun, 1911)⇒ Urnalana angulata Rathbun, 1911
Leucosides jecusculum (Rathbun, 1911)⇒ Leucosia jecusculum Rathbun, 1911
Leucosilia bananensis ♰ M. J. Rathbun, 1918
Leucosilia jurinei
Leucosilia jurinii Saussure, 1853
Leucosilia maldivensis Borradaile, 1903
Lissomorpha haswelli Ward, 1933
Lithadia brasiliensis von Martens, 1872
Lithadia cadaverosa Stimpson, 1871Carinate clutch crab
Lithadia cariosa (Stimpson, 1860)⇒ Ebalia cariosa Stimpson, 1860
Lithadia cumingii Bell, 1855
Lithadia granulosa A. Milne-Edw..., 1880
Lithadia lacunosa (Kingsley, 1879)⇒ Ebalia cariosa Stimpson, 1860
Lithadia obliqua Coêlho, 1973
Lithadia vertiginosa Coêlho, 1973
Lyphira heterograna Ortmann, 1892
Lyphira perplexa Galil, 2009
Merocryptoides frontalis Sakai, 1963
Merocryptoides ohtsukai Komatsu & Ta..., 2001
Merocryptoides peteri Komatsu & Ta..., 2001
Merocryptus boletifer A. Milne-Edw..., 1894
Merocryptus boletisculpta Zarenkov, 1994
Merocryptus durandi Serène, 1955
Merocryptus lambriformis A. Milne-Edw..., 1873
Merocryptus obsoletus A. Milne-Edw..., 1898
Myra affinis Bell, 1855
Myra australis Haswell, 1880
Myra biconica Ihle, 1918
Myra brevimana Alcock, 1896
Myra carinata (Bell, 1855)⇒ Myra fugax Fabricius, 1798
Myra celeris Galil, 2001
Myra coalita (Hilgendorf, 1878)⇒ Myra subgranulata Kossmann, 1877
Myra currax Galil, 2001
Myra curtimana Galil, 2001
Myra darnleyensis (Haswell, 1879)⇒ Myrine kessleri Paulson, 1875
Myra digitata Galil, 2004
Myra elegans Bell, 1855
Myra eudactyla (Bell, 1855)⇒ Myra eudactylus Bell, 1855
Myra eudactylus Bell, 1855
Myra fugax Fabricius, 1798Fleeting Purse Crab
Myra grandis Zarenkov, 1990Pebble Crab
Myra hainanica Chen & Türkay, 2001
Myra intermedia Borradaile, 1902
Myra kessleri Paulson, 1875
Myra longimerus (Chen & Türk..., 2001)⇒ Myra fugax Fabricius, 1798
Myra mammilaris Bell, 1855
Myra mammillaris Bell, 1855
Myra pentacantha (Alcock, 1896)⇒ Myra fugax Fabricius, 1798
Myra pernix Galil, 2001
Myra pugax
Myra subcarinata Morris & Col..., 1991
Myra subgranulata Kossmann, 1877
Myra subovata (Rathbun, 1894)⇒ Persephona subovata Rathbun, 1894
Myra townsendi (Rathbun, 1894)⇒ Persephona townsendi Rathbun, 1894
Myra trispinosa Morris & Col..., 1991
Myra tumidospina Galil, 2001
Myrine acutidens Ihle, 1918
Myrine kessleri Paulson, 1875
Myrine kesslerii Paulson, 1875
Myrodes eudactylus (Bell, 1855)⇒ Myra eudactylus Bell, 1855
Myropsis quinquespinosa Stimpson, 1871Fivespine purse crab
Naesa caudata (Say, 1818)⇒ Paracerceis caudata Say, 1818
Nobiliella margaritata Galil & Inno..., 2019
Nucia borneoensis Morris & Col..., 1991
Nucia bouvieri Ihle, 1918
Nucia calculoides Morris & Col..., 1991
Nucia coxi Morris & Col..., 1991
Nucia gelida (Rathbun, 1907)⇒ Heteronucia venusta Nobili, 1906
Nucia ingens Rathbun, 1911
Nucia miliaris A. Milne-Edw..., 1873
Nucia modestus (Ihle, 1918)⇒ Nuciops modestus Ihle, 1918
Nucia perlata (Sakai, 1963)⇒ Heteronucia perlata Sakai, 1963
Nucia platyspinosa Collins & al., 2003
Nucia pulchella A. Milne-Edw..., 1873
Nucia rosea Nobili, 1906
Nucia speciosa Dana, 1852Leucosiid Crab
Nucia tuberculosa A. Milne-Edw..., 1874
Nucilobus symmetricus Morris & Col..., 1991
Nuciops modestus Ihle, 1918
Nursia abbreviata (Bell, 1855)⇒ Paranursia abbreviata Bell, 1855
Nursia alata Komatsu & Ta..., 1999
Nursia elegans Ihle, 1918
Nursia guinotae Komatsu & Ta..., 2001
Nursia hardwicki
Nursia hardwickii (Leach, 1817)⇒ Nursia lar Fabricius, 1793
Nursia japonica Sakai, 1935
Nursia lar Fabricius, 1793
Nursia mimetica Nobili, 1906
Nursia minor Miers, 1879
Nursia nasata
Nursia nasuta Alcock, 1896
Nursia phylloides Ihle, 1918
Nursia plicata Herbst, 1803
Nursia rhomboidalis Miers, 1879
Nursia sinica (Shen, 1937)⇒ Nursia minor Miers, 1879
Nursia sinuata Miers, 1877
Nursia tuberculata (Rathbun, 1894)⇒ Ebalia cristata Rathbun, 1898
Nursia weberi Ihle, 1918
Nursilia dentata Bell, 1855
Nursilia hardwickii
Nursilia sinica Chen, 1982
Nursilia tonsor Alcock, 1896
Oreophorus crosnieri C. G. S. Tan..., 1995
Oreophorus frontalis (Miers, 1884)⇒ Cateios frontalis Miers, 1884
Oreophorus horridus Rüppell, 1830
Oreophorus ornatus (Ihle, 1918)⇒ Alox ornatum Ihle, 1918
Oreophorus reticulatus Adams & White, 1849
Oreophorus rugosus (Stimpson, 1858)⇒ Alox rugosum Stimpson, 1858
Oreophorus speciosus Chen, 1989
Oreotlos angulatus Rathbun, 1906
Oreotlos bertrandi C. G. S. Tan..., 1995
Oreotlos encymus C. G. S. Tan..., 1993
Oreotlos etor C. G. S. Tan..., 1993
Oreotlos havelocki Laurie, 1906
Oreotlos heuretos C. G. S. Tan..., 1995
Oreotlos lagarodes C. G. S. Tan..., 1995
Oreotlos latus Borradaile, 1903
Oreotlos pala C. G. S. Tan..., 1995
Oreotlos pax C. G. S. Tan..., 1995
Oreotlos potanus C. G. S. Tan..., 1993
Orientotlos iishibai Sakai, 1980
Palaeomyra bispinosa ♰ A. Milne-Edw..., 1861
Paranursia abbreviata Bell, 1855
Parilia alcocki Wood-Mason &..., 1891
Parilia major Sakai, 1961
Parilia ovata Chen, 1984
Parilia pattersoni Ng, Devi & K..., 2018
Parilia tuberculata Sakai, 1961
Persephona aquilonaris Rathbun, 1933
Persephona crinita Rathbun, 1931Pink purse crab
Persephona edwardsii Bell, 1855
Persephona finneganae Rathbun, 1931
Persephona fugax
Persephona guaia (Bell, 1855)⇒ Persephona punctata Linnaeus, 1758
Persephona lichtensteini
Persephona lichtensteinii Leach, 1817
Persephona lichtenstenii
Persephona mediterranea Herbst, 1794Mottled purse crab
Persephona niemeyeri Blow, 2003
Persephona orbicularis Bell, 1855
Persephona punctata Linnaeus, 1758Purse Crab
Persephona punctata aquilonaris Rathbun, 1933
Persephona punctata punctata 1, 0
Persephona rodesae Blow, 2003
Persephona subovata Rathbun, 1894
Persephona townsendi Rathbun, 1894
Philyra acutidens (Chen, 1987)⇒ Lyphira heterograna Ortmann, 1892
Philyra adamsi
Philyra adamsii Bell, 1855
Philyra alcocki Kemp, 1915
Philyra angularis Rathbun, 1924
Philyra carinata (Bell, 1855)⇒ Pyrhila carinata Bell, 1855
Philyra collinsi Karasawa, 2010
Philyra corallicola Alcock, 1896
Philyra cranium Desmarest, 1822
Philyra cristata (Miers, 1881)⇒ Atlantophila cristata Miers, 1881
Philyra fuliginosa Targioni-Toz..., 1877
Philyra globosa
Philyra globulosa (H. Milne Edw..., 1837)⇒ Philyra globus Fabricius, 1775
Philyra globus Fabricius, 1775
Philyra granigera Nobili, 1905
Philyra granulosa Ihle, 1918
Philyra hayasakai ♰ Karasawa, 1992
Philyra heterograna (Ortmann, 1892)⇒ Lyphira heterograna Ortmann, 1892
Philyra hormozganensis ♰ F. J. Vega, 2010
Philyra iriomotensis Sakai, 1983
Philyra kanekoi Sakai, 1934
Philyra laevidorsalis (Miers, 1881)⇒ Atlantolocia laevidorsalis Miers, 1881
Philyra laevis (Bell, 1855)⇒ Bellidilia laevis Bell, 1855
Philyra laminata (Doflein, 1904)⇒ Heteronucia laminata Doflein, 1904
Philyra macrophthalma Bell, 1855
Philyra malefactrix Kemp, 1915
Philyra marginata A. Milne-Edw..., 1873
Philyra minuta (Chen & Türkay, 2001)⇒ Philyra malefactrix Kemp, 1915
Philyra misagoana Sakai, 1937
Philyra nishihirai Takeda & Nak..., 1991
Philyra olivacea Rathbun, 1909
Philyra orbicularis Bell, 1855
Philyra pisum (De Haan, 1841)⇒ Pyrhila pisum De Haan, 1841
Philyra plana ♰ Karasawa, 1989
Philyra platycheir (De Haan, 1841)⇒ Hiplyra platycheir De Haan, 1841
Philyra platycheira
Philyra platychira De Haan, 1841
Philyra polita (Henderson, 1893)⇒ Philyra globus Fabricius, 1775
Philyra porcellanea Herbst, 1783
Philyra punctata (Bell, 1855)⇒ Afrophila punctata Bell, 1855
Philyra rectangularis Miers, 1884
Philyra rudis Miers, 1884
Philyra sagittifera Alcock, 1896
Philyra scabra Dai, Yang, S..., 1984
Philyra scabriuscula Fabricius, 1798
Philyra sexangula Alcock, 1896
Philyra syndactyla Ortmann, 1892
Philyra taekoae Takeda, 1972
Philyra tanakai ♰ Karasawa, 1993
Philyra tridentata ♰ Karasawa, 1993
Philyra trusanensis Collins & al., 2003
Philyra tuberculosa (Stimpson, 1858)⇒ Pyrhila carinata Bell, 1855
Philyra variegata (Rüppell, 1830)⇒ Hiplyra variegata Rüppell, 1830
Philyra verrucosa (Henderson, 1893)⇒ Ryphila verrucosa Henderson, 1893
Phlyxia crassipes (Bell, 1885)⇒ Ebalia crassipes Bell, 1885
Phlyxia erosa (A. Milne-Edw..., 1873)⇒ Ebaliopsis erosa A. Milne-Edw..., 1873
Phlyxia lambriformis (Bell, 1885)⇒ Ebalia lambriformis Bell, 1885
Phlyxia ramsayi (Haswell, 1880)⇒ Ebalia ramsayi Haswell, 1880
Praebebalia dondonae Chen, 1989
Praebebalia extensiva Rathbun, 1911
Praebebalia fasciata Ihle, 1918
Praebebalia fujianensis Chen & Fang, 2000
Praebebalia fungifera Galil, 2015
Praebebalia kumanoensis Sakai, 1983
Praebebalia longidactyla Yokoya, 1933
Praebebalia madagascariensis Galil, 2001
Praebebalia magna Galil, 2001
Praebebalia mosakiana Sakai, 1965
Praebebalia pisiformis Ihle, 1918
Praebebalia semblatae (Chen, 1989)⇒ Ancylodactyla elongata Zarenkov, 1969
Praebebalia septemspinosa Sakai, 1983
Praebebalia sikokuensis Yokoya, 1933
Praebebalia taeniata Takeda, 1977
Praosia punctata C. G. S. Tan..., 1993
Pseudomyra mbizi Capart, 1951
Pseudophilyra burmensis Sakai, 1983
Pseudophilyra deficiens Ihle, 1918
Pseudophilyra intermedia Ihle, 1918
Pseudophilyra melita De Man, 1888
Pseudophilyra parilis Komai, Shime..., 2019
Pseudophilyra perryi Miers, 1876
Pseudophilyra polita Miers, 1884
Pseudophilyra punctulata Chen & Nq, 2003
Pseudophilyra pusilla Henderson, 1893
Pseudophilyra tenuipes Ihle, 1918
Pseudophilyra tridentata Miers, 1879
Pseudophilyra woodmasoni Alcock, 1896
Pseudophilyra woodmasonia
Pterocarcinus baileyi Blow, 2003
Pyrhila carinata Bell, 1855
Pyrhila pisum De Haan, 1841
Randallia agaricias Rathbun, 1898
Randallia americana Rathbun, 1894
Randallia angelica Garth, 1940
Randallia bulligera Rathbun, 1898
Randallia distincta M. J. Rathbun, 1893
Randallia eburnea (Alcock, 1896)⇒ Tokoyo eburnea Alcock, 1896
Randallia gilberti Rathbun, 1906
Randallia glana
Randallia glans (Alcock, 1896)⇒ Ebalia glans Alcock, 1896
Randallia granulata Miers, 1886
Randallia granuloides (Sakai, 1961)⇒ Toru granuloides Sakai, 1961
Randallia laevis Borradaile, 1916
Randallia minuta Rathbun, 1935
Randallia ornata Randall, 1840Globose sand crab
Randallia pila (C. G. S. Tan, 1996)⇒ Toru pilus C. G. S. Tan, 1996
Randallia pustulilabris (Alcock, 1896)⇒ Tanaoa pustulilabris Alcock, 1896
Randallia pustuloides (Sakai, 1961)⇒ Urashima pustuloides Sakai, 1961
Randallia pustulosa (Wood Mason &..., 1891)⇒ Tanaoa pustulosus Wood-Mason &..., 1891
Randallia saitoensis ♰ Karasawa, 1993
Randallia serenei (Richer de Fo..., 1983)⇒ Tanaoa serenei Richer de Fo..., 1983
Randallia speciosus (Chen, 1989)⇒ Tanaoa speciosus Chen, 1989
Randallia trituberculatus (Sakai, 1961)⇒ Toru trituberculatus Sakai, 1961
Randallia villosus (Chen, 1989)⇒ Ihleus villosus Chen, 1989
Raylilia coniculifera Galil, 2001
Raylilia gracilipes Bell, 1855
Raylilia intermedia Komatsu, Man..., 2005
Raylilia mirabilis Zarenkov, 1969
Raylilia uenoi Takeda, 1995
Ryphila bertrandi N.g.Galil, 2015
Ryphila cancellus Herbst, 1783
Ryphila verrucosa Henderson, 1893
Seulocia anahita Galil, 2005
Seulocia cristata Galil, 2005
Seulocia gagulae N.g.Galil, 2015
Seulocia laevimana Miers, 1884
Seulocia latirostrata Shen & Chen, 1978
Seulocia pubescens Miers, 1877Olive Purse Crab
Seulocia pulchra Shen & Chen, 1978
Seulocia rhomboidalis De Haan, 1841
Seulocia truncata Alcock, 1896
Seulocia vittata Stimpson, 1858
Soceulia alainia Galil, 2006
Soceulia brunnea Miers, 1877
Soceulia major Chen & Ng, 2003
Soceulia marmorea Bell, 1855
Speleophorus nodosus Bell, 1855
Speloeophorus digueti Bouvier, 1898
Speloeophorus elevatus Rathbun, 1898Bighole clutch crab
Speloeophorus microspeos Telford, 1980
Speloeophorus nodosus Bell, 1855Twohole clutch crab
Speloeophorus pontifer Stimpson, 1871Smallhole clutch crab
Speloeophorus schmitti Glassell, 1935
Tanaoa distinctus Rathbun, 1894
Tanaoa granulosa Alcock & And..., 1894
Tanaoa kuka N.g.Galil, 2015
Tanaoa nanus Galil, 2003
Tanaoa pustulilabris Alcock, 1896
Tanaoa pustulosus Wood-Mason &..., 1891
Tanaoa retpela N.g.Galil, 2015
Tanaoa serenei Richer de Fo..., 1983
Tanaoa speciosus Chen, 1989
Tlos angulatus (Rathbun, 1906)⇒ Oreotlos angulatus Rathbun, 1906
Tlos latus (Borradaile, 1903)⇒ Oreotlos latus Borradaile, 1903
Tlos muriger Adams & White, 1849
Tokoyo cirrata Galil, 2003
Tokoyo eburnea Alcock, 1896
Toru granuloides Sakai, 1961
Toru pilus C. G. S. Tan, 1996
Toru septimus Galil, 2003
Toru trituberculatus Sakai, 1961
Typilobus belli Quayle & Col..., 1981
Typilobus granulosus ♰ Stoliczka, 1871
Typilobus marginatus Morris & Col..., 1991
Typilobus obscurus Quayle & Col..., 1981
Typilobus sadeki Withers, 1925
Typilobus unispinatus ♰ Martins-Neto, 2001
Uhlias ellipticus Stimpson, 1871
Uhlias limbatus Stimpson, 1871Eroded clutch crab
Urashima pustuloides Sakai, 1961
Urnalana angulata Rathbun, 1911
Urnalana cheverti Haswell, 1880
Urnalana chevreti Haswell, 1880
Urnalana chevretii Haswell, 1880
Urnalana cumingii Bell, 1855
Urnalana elata A. Milne-Edw..., 1873Nut Crab
Urnalana elatoides Bouvier, 1915
Urnalana elatula Galil, 2005
Urnalana erythema Lin, Lai & G..., 2014Pebble Crab
Urnalana flammea N.g.Galil, 2015
Urnalana foresti Chen, 1989
Urnalana granulimera Galil, 2005
Urnalana haematosticta Adams & White, 1849
Urnalana hilaris Nobili, 1905
Urnalana hirsuta Lin, Lai & G..., 2014
Urnalana insularis Takeda & Kurata, 1976
Urnalana margaritata A. Milne-Edw..., 1873
Urnalana parahaematostica Galil, 2005
Urnalana pulchella Bell, 1855
Urnalana purarensis Ovaere, 1987Bulbous Purse Crab
Urnalana thysanotus George & M. ..., 1976
Urnalana whitei Bell, 1855

Subnodes of Leucosiidae with occurrences in Continents
SubfamilyCryptocneminae
Stimpson, 1907
3
24
Genera
Species

Taxom

SubfamilyEbaliinae
Stimpson, 1871
63
480
Genera
Species

Taxom

SubfamilyLeucosiinae
Samouelle, 1819
8
140
Genera
Species

Taxom



Links, sources and further information
Leucosiidae at artfakta.se
Leucosiidae at biodiversity.org.au
Leucosiidae at dyntaxa.se
Leucosiidae at gbif.org
Leucosiidae at irmng.org
Leucosiidae at itis.gov
Leucosiidae at marinespecies.org