Checklist for Family Varunidae H. Milne-Edwards, 1853
and region Continents
Class
| Malacostraca
Latreille, 1802
Higher Crustaceans, Malacostracans | 734
7366
51296 | Families
Genera
Species |
Taxom
|
Subclass
| Eumalacostraca
Grobben, 1892
| 716
7312
51144 | Families
Genera
Species |
Taxom
|
Superorder
| Eucarida
Calman, 1904
| 268
3165
24238 | Families
Genera
Species |
Taxom
|
Order
| Decapoda
Latreille, 1802
Crabs, Crayfishes, Decapod, Decapods, Lobsters, Prawns | 265
3148
24112 | Families
Genera
Species |
Taxom
|
Suborder
| Pleocyemata
Burkenroad, 1963
| 232
2987
23012 | Families
Genera
Species |
Taxom
|
Infraorder
| Brachyura
Linnaeus, 1758
Crabs, Short-tailed crabs, True crabs | 131
1846
11236 | Families
Genera
Species |
Taxom
|
Section
| Eubrachyura
de Saint Laurent, 1980
| 106
1646
10355 | Families
Genera
Species |
Taxom
|
Subsection
| Thoracotremata
Guinot, 1977
| 20
287
2126 | Families
Genera
Species |
Taxom
|
Superfamily
| Grapsoidea
MacLeay, 1838
| 8
126
984 | Families
Genera
Species |
Taxom
| | Family | Varunidae
H. Milne-Edwards, 1853
| 39
225 | Genera
Species |
Taxom
|
Genera for Continents
39 Genera with 225 Species #ABCEGHMNOPSTUV
| Acmaeopleura Stimpson, 1974 | | 5 | Species | Taxom | | Asthenognathus Stimpson, 2004 | | 6 | Species | Taxom | | Austrohelice Sakai, Türka..., 2008 | | 2 | Species | Taxom | | Brachynotus De Haan, 1853 | | 17 | Species | Taxom | | Chasmagnathus De Haan, 0 | | 7 | Species | Taxom | | Cyclograpsus H. Milne-Edw..., 1837 | | 22 | Species | Taxom | | Cyrtograpsus Dana, 1867 | | 5 | Species | Taxom | | Eriocheir De Haan, 1938 | | 11 | Species | Taxom | | Gaetice Gistel, 1939 | | 3 | Species | Taxom | | Globihexapus Schweitzer &..., 2001 | | 1 | Species | Taxom | | Gopkittisak Naruse & Clark, 1976 | | 2 | Species | Taxom | | Helicana K. Sakai & Y..., 1931 | | 3 | Species | Taxom | | Helice De Haan, 1931 | | 18 | Species | Taxom | | Helograpsus Campbell & G..., 1899 | | 1 | Species | Taxom | | Hemigrapsus Dana, 2005 | | 16 | Species | Taxom | | Metaplax H. Milne Edw..., 1950 | | 18 | Species | Taxom | | Miosesarma Karasawa, 1989 | | 1 | Species | Taxom | | Neoeriocheir T. Sakai, 1913 | | 1 | Species | Taxom | | Neohelice Sakai, Türka..., 1851 | | 1 | Species | Taxom | | Noarograpsus N. K. Ng, Ma..., 1991 | | 1 | Species | Taxom | | Orcovita Ng & Tomascik, 1994 | | 8 | Species | Taxom | | Otognathon Ng & Stevcic, 2009 | | 1 | Species | Taxom | | Paragrapsus H. Milne Edw..., 1837 | | 3 | Species | Taxom | | Parahelice K. Sakai, Tü..., 2006 | | 4 | Species | Taxom | | Parapyxidognathus Ward, 1888 | | 1 | Species | Taxom | | Platyeriocheir N. K. Ng, Gu..., 1995 | | 1 | Species | Taxom | | Pseudogaetice Davie & N. K..., 1923 | | 1 | Species | Taxom | | Pseudograpsus H. Milne-Edw..., 1798 | | 11 | Species | Taxom | | Pseudohelice Sakai, Türka..., 1851 | | 5 | Species | Taxom | | Ptychognathus Stimpson, 1939 | | 29 | Species | Taxom | | Pyxidognathus A. Milne-Edw..., 1918 | | 3 | Species | Taxom | | Scutumara Ng & Nakasone, 1972 | | 3 | Species | Taxom | | Sestrostoma Davie & Ng, 1974 | | 3 | Species | Taxom | | Shurebus Verany, 1847 | | 1 | Species | Taxom | | Tetragrapsus Rathbun, 1894 | | 1 | Species | Taxom | | Thalassograpsus Tweedie, 1892 | | 1 | Species | Taxom | | Utica White, 1881 | | 7 | Species | Taxom | | Vanua Rathbun, 1945 | | 1 | Species | Taxom | | Varuna H. Milne Edw..., 1986 | | 5 | Species | Taxom |
Species list for Continents
225 Species and 5 Subspecies | Acmaeopleura balssi (Shen, 1932) | ⇒ Sestrostoma balssi Shen, 1932 | | Acmaeopleura depressum (Sakai, 1965) | ⇒ Sestrostoma depressum Sakai, 1965 | | Acmaeopleura parvula Stimpson, 1858 | | | Acmaeopleura rotunda Rathbun, 1909 | | | Acmaeopleura toriumii (Takeda, 1974) | ⇒ Sestrostoma toriumii Takeda, 1974 | | Asthenognathus atlanticus Monod, 1933 | | | Asthenognathus cornishorum C. Schweitze..., 1999 | | | Asthenognathus gallardoi (Serène & Soh, 1976) | ⇒ Gopkittisak gallardoi Serène & Soh, 1976 | | Asthenognathus hexagonum Rathbun, 1909 | | | Asthenognathus inaequipes Stimpson, 1858 | | | Asthenognathus microspinus ♰ S. Casadio, 2004 | | | Austrohelice crassa Dana, 1851 | Tunneling Mud Crab. Burrowing Mud Crab | | Austrohelice manneringi ♰ R. M. Feldmann, 2008 | | | Brachynotus atlanticus Forest, 1957 | | | Brachynotus brevidigitatus (Yokoya, 1928) | ⇒ Hemigrapsus penicillatus De Haan, 1835 | | Brachynotus corallinus ♰ Beschin, 2007 | | | Brachynotus edwardsii (Hilgendorf, 1882) | ⇒ Hemigrapsus sexdentatus Milne-Edwards, 1837 | | Brachynotus februarius Muller, 1974 | | | Brachynotus foresti Zariquiey Al..., 1968 | | | Brachynotus gemmellari (Rizza, 1839) | ⇒ Brachynotus gemmellaroi Rizza, 1839 | | Brachynotus gemmellaroi Rizza, 1839 | | | Brachynotus harpax (Hilgendorf, 1892) | ⇒ Thalassograpsus harpax Hilgendorf, 1892 | | Brachynotus jouyi (Rathbun, 1894) | ⇒ Tetragrapsus jouyi Rathbun, 1894 | | Brachynotus lucasi (Pesta, 1918) | ⇒ Brachynotus sexdentatus Risso, 1827 | | Brachynotus octodentatus | | | Brachynotus penicillatus De Haan, 1835 | | | Brachynotus sanguineus | | | Brachynotus sexdentatus Risso, 1827 | Mediterranean crab | | Brachynotus sexdentatus gemmellaroi | | | Brachynotus spinosus H. Milne Edw..., 1853 | Little Shore Crab | | Chasmagnathus convexus (De Haan, 1835) | ⇒ Helograpsus haswellianus Whitelegge, 1899 | | Chasmagnathus granulata (Dana, 1851) | ⇒ Neohelice granulata Dana, 1851 | | Chasmagnathus granulatus Dana, 1851 | | | Chasmagnathus hararlliamus | | | Chasmagnathus haswellianus (Whitelegge, 1899) | ⇒ Helograpsus haswellianus Whitelegge, 1899 | | Chasmagnathus laevis (Dana, 1851) | ⇒ Paragrapsus laevis Dana, 1851 | | Chasmagnathus quadridentatus | | | Cyclograpsus audouinii H. Milne Edw..., 1837 | | | Cyclograpsus barbatus MacLeay, 1838 | | | Cyclograpsus cinereus Dana, 1851 | | | Cyclograpsus crenulatus (H. Milne Edw..., 1837) | ⇒ Hemigrapsus crenulatus Milne Edwards, 1837 | | Cyclograpsus escondidensis Rathbun, 1933 | | | Cyclograpsus gaimardii (H. Milne Edw..., 1837) | ⇒ Paragrapsus gaimardii H. Milne Edw..., 1837 | | Cyclograpsus granulatus Dana, 1851 | | | Cyclograpsus granulosus H. Milne Edw..., 1853 | Purple mottled shore crab | | Cyclograpsus henshawi Rathbun, 1902 | Aupaka | | Cyclograpsus insularum Campbell & G..., 1966 | Northern smooth shore crab | | Cyclograpsus integer H. Milne Edw..., 1837 | Globose shore crab | | Cyclograpsus intermedius Ortmann, 1894 | | | Cyclograpsus lavauseii | | | Cyclograpsus lavauxi H. Milne Edw..., 1853 | Smooth shore crab | | Cyclograpsus longipes Stimpson, 1858 | | | Cyclograpsus lucidus Dai, Yang, S..., 1986 | | | Cyclograpsus occidentalis (A. Milne-Edw..., 1878) | ⇒ Cyclograpsus integer H. Milne Edw..., 1837 | | Cyclograpsus parvulus (De Man, 1896) | ⇒ Cyclograpsus integer H. Milne Edw..., 1837 | | Cyclograpsus punctatum | | | Cyclograpsus punctatus H. Milne Edw..., 1837 | | | Cyclograpsus quadridentatus (H. Milne Edw..., 1837) | ⇒ Paragrapsus quadridentatus H. Milne Edw..., 1837 | | Cyclograpsus sexdentatus (H. Milne Edw..., 1837) | ⇒ Hemigrapsus sexdentatus Milne-Edwards, 1837 | | Cyrtograpsus affinis Dana, 1851 | | | Cyrtograpsus altimanus Rathbun, 1914 | | | Cyrtograpsus angulatus Dana, 1851 | | | Cyrtograpsus cirripes Smith, 1800 | | | Cyrtograpsus murchisonii Carruthers, 1867 | | | Eriocheir formosa (T.Y. Chan , ..., 1995) | ⇒ Eriocheir japonica De Haan, 1835 | | Eriocheir hepuensis Dai, 1991 | | | Eriocheir japonica De Haan, 1835 | Japanese mitten crab | | Eriocheir japonicus (De Haan, 1885) | ⇒ Eriocheir japonica De Haan, 1835 | | Eriocheir japoniscus | | | Eriocheir leptognathus (Rathbun, 1913) | ⇒ Neoeriocheir leptognathus Rathbun, 1913 | | Eriocheir ogasawaraensis T. Komai , Y..., 2006 | | | Eriocheir opilio | | | Eriocheir rectus (Stimpson, 1858) | ⇒ Eriocheir japonica De Haan, 1835 | | Eriocheir sinensis Milne-Edwards, 1854 | Big sluice crab | | Eriocheir sinensis acutifrons (Panning, 1938) | ⇒ Eriocheir sinensis Milne-Edwards, 1854 | | Gaetice americanus (Rathbun, 1923) | ⇒ Pseudogaetice americanus Rathbun, 1923 | | Gaetice depressus De Haan, 1833 | | | Gaetice ungulatus Sakai, 1939 | | | Globihexapus paxillus C. Schweitze..., 2001 | | | Gopkittisak angustum Komai, 2011 | | | Gopkittisak gallardoi Serène & Soh, 1976 | | | Helicana doerjesi K. Sakai, Tü..., 2006 | | | Helicana japonica K. Sakai & Y..., 1980 | | | Helicana wuana Rathbun, 1931 | | | Helice crana | | | Helice crassa (Dana, 1851) | ⇒ Austrohelice crassa Dana, 1851 | | Helice evassa | | | Helice formosensis Rathbun, 1931 | | | Helice japonica (K. Sakai & Y..., 1980) | ⇒ Helicana japonica K. Sakai & Y..., 1980 | | Helice latimera Parisi, 1918 | | | Helice latreillei (H. Milne Edw..., 1837) | ⇒ Helice tridens De Haan, 1835 | | Helice leachi (Hess, 1865) | ⇒ Pseudohelice subquadrata Dana, 1851 | | Helice leachii (Hess, 1865) | ⇒ Pseudohelice subquadrata Dana, 1851 | | Helice pingi | | | Helice spinicarpa De Haan, 1835 | | | Helice subquadrata Dana, 1851 | | | Helice tientsinensis Rathbun, 1931 | | | Helice tridens De Haan, 1835 | | | Helice tridens pingi (Rathbun, 1931) | ⇒ Helice latimera Parisi, 1918 | | Helice tridens sheni (Sakai, 1939) | ⇒ Helicana wuana Rathbun, 1931 | | Helice tridens wuana Rathbun, 1931 | | | Helice wuana (Rathbun, 1931) | ⇒ Helicana wuana Rathbun, 1931 | | Helograpsus haswellianus Whitelegge, 1899 | Haswell's Shore Crab | | Hemigrapsus affinis Dana, 1851 | | | Hemigrapsus crassimanus Dana, 1851 | | | Hemigrapsus crenulatus Milne Edwards, 1837 | Hairy-handed crab | | Hemigrapsus edwardsi (Hilgendorf, 1882) | ⇒ Hemigrapsus takanoi Asakura & Wa..., 2005 | | Hemigrapsus estellinensis Creel, 1964 | | | Hemigrapsus lobulatus (Manuel, Gonz..., 1991) | ⇒ Noarograpsus lobulatus Manuel, Gonz..., 1991 | | Hemigrapsus longitarsis Miers, 1879 | | | Hemigrapsus nudus Dana, 1851 | Naked Shore Crab | | Hemigrapsus octodentatus H. Milne Edw..., 1837 | | | Hemigrapsus oregonensis Dana, 1851 | Green shore crab | | Hemigrapsus penicillatus De Haan, 1835 | Common Japanese intertidal crab | | Hemigrapsus sanguineus De Haan, 1835 | Asian shore crab | | Hemigrapsus sexdentatus Milne-Edwards, 1837 | Common Rock Crab | | Hemigrapsus sinensis Rathbun, 1931 | | | Hemigrapsus takanoi Asakura & Wa..., 2005 | Asian Brush Crab | | Hemigrapsus tanakoi (Asakura & Wa..., 2005) | ⇒ Hemigrapsus takanoi Asakura & Wa..., 2005 | | Metaplax crassipes (De Man, 1892) | ⇒ Metaplax elegans De Man, 1888 | | Metaplax crenulata Gerstaecker, 1856 | | | Metaplax crenulatus | | | Metaplax dentipes Heller, 1865 | | | Metaplax distincta H. Milne Edw..., 1852 | | | Metaplax distinguendus | | | Metaplax elegans De Man, 1888 | | | Metaplax gocongensis Davie & Nguyen, 2003 | | | Metaplax hirsutimana (Grant & McCu..., 1906) | ⇒ Australoplax tridentata A. Milne-Edw..., 1873 | | Metaplax hirsutimanus | | | Metaplax hirtipes (Heller, 1865) | ⇒ Hemiplax hirtipes Heller, 1865 | | Metaplax indica H. Milne Edw..., 1852 | | | Metaplax intermedia De Man, 1888 | | | Metaplax longipes Stimpson, 1858 | | | Metaplax occidentalis Pretzmann, 1971 | | | Metaplax sheni Gordon, 1930 | | | Metaplax takahashii Sakai, 1939 | | | Metaplax tredecim Tweedie, 1950 | | | Miosesarma japonicum ♰ Karasawa, 1989 | | | Neoeriocheir leptognathus Rathbun, 1913 | | | Neohelice granulata Dana, 1851 | Cangrejo Granuloso | | Noarograpsus lobulatus Manuel, Gonz..., 1991 | | | Orcovita fictilia Ng, Guinot &..., 1996 | | | Orcovita gracilipes Ng, Guinot &..., 1996 | | | Orcovita hicksi Davie & Ng., 2012 | | | Orcovita mcneiceae Ng & Ng, 2002 | | | Orcovita miruku Naruse & Tamura, 2006 | | | Orcovita mollitia Ng, Guinot &..., 1996 | | | Orcovita orchardorum Davie & Ng., 2012 | | | Orcovita saltatrix Ng & Tomascik, 1994 | | | Otognathon uru N. K. Ng, Ko..., 2009 | | | Paragrapsus gaimardii H. Milne Edw..., 1837 | Red-spotted Shore Crab | | Paragrapsus laevis Dana, 1851 | Smooth shore crab | | Paragrapsus quadridentatus H. Milne Edw..., 1837 | Four-toothed Shore Crab | | Parahelice balssi K. Sakai, TÃ..., 2006 | | | Parahelice daviei K. Sakai, Tü..., 2006 | | | Parahelice pilimana A. Milne-Edw..., 1873 | | | Parahelice pilosa K. Sakai, Tü..., 2006 | | | Parapyxidognathus deianira De Man, 1888 | | | Platyeriocheir formosa Chan, Hung & Yu, 1995 | | | Pseudogaetice americanus Rathbun, 1923 | | | Pseudograpsus albus Stimpson, 1858 | Little White Crab | | Pseudograpsus barbatus (H. Milne Edw..., 1853) | ⇒ Pseudograpsus setosus Fabricius, 1798 | | Pseudograpsus crassus A. Milne-Edw..., 1868 | | | Pseudograpsus elongatus A. Milne-Edw..., 1873 | | | Pseudograpsus erythraeus (Kossman, 1877) | ⇒ Pseudograpsus elongatus A. Milne-Edw..., 1873 | | Pseudograpsus intermedius Chhapgar, 1955 | | | Pseudograpsus laniger (Tesch, 1918) | ⇒ Scutumara laniger Tesch, 1918 | | Pseudograpsus nudus (Stimpson, 1858) | ⇒ Hemigrapsus nudus Dana, 1851 | | Pseudograpsus oregonensis (Dana, 1851) | ⇒ Hemigrapsus oregonensis Dana, 1851 | | Pseudograpsus penicilliger Latreille, 1817 | | | Pseudograpsus setosus Fabricius, 1798 | | | Pseudohelice balssi (K. Sakai, Tü..., 2006) | ⇒ Parahelice balssi K. Sakai, TÃ..., 2006 | | Pseudohelice daviei (K. Sakai, Tü..., 2006) | ⇒ Parahelice daviei K. Sakai, Tü..., 2006 | | Pseudohelice pilosa (K. Sakai, Tü..., 2006) | ⇒ Parahelice pilosa K. Sakai, Tü..., 2006 | | Pseudohelice quadrata Dana, 1851 | | | Pseudohelice subquadrata Dana, 1851 | | | Ptychognathus affinis De Man, 1895 | | | Ptychognathus altimanus Rathbun, 1914 | | | Ptychognathus barbatus A. Milne-Edw..., 1873 | | | Ptychognathus capillidigitatus Takeda, 1984 | | | Ptychognathus crassimanus Finnegan, 1931 | | | Ptychognathus demani Roux, 1917 | | | Ptychognathus dentatus De Man, 1892 | | | Ptychognathus easteranus Rathbun, 1907 | | | Ptychognathus glaber Stimpson, 1858 | | | Ptychognathus guijulugani Rathbun, 1914 | | | Ptychognathus hachijoensis Sakai, 1955 | | | Ptychognathus hachijyoensis (Sakai, 1955) | ⇒ Ptychognathus hachijoensis Sakai, 1955 | | Ptychognathus insolitus Osawa & N. K..., 2006 | | | Ptychognathus intermedius De Man, 1879 | | | Ptychognathus ishii Sakai, 1939 | | | Ptychognathus johannae Rathbun, 1914 | | | Ptychognathus lipkei N. K. Ng, 2010 | | | Ptychognathus makii Hsu & Shih, 2020 | | | Ptychognathus onyx Alcock, 1900 | | | Ptychognathus pilipes A. Milne-Edw..., 1868 | | | Ptychognathus pilosus De Man, 1892 | | | Ptychognathus polleni De Man, 1895 | | | Ptychognathus pusillus Heller, 1865 | | | Ptychognathus riedelii A. Milne-Edw..., 1868 | | | Ptychognathus sakaii Hsu, Shih & Li, 2022 | | | Ptychognathus spinicarpus Ortmann, 1894 | | | Ptychognathus stimpsoni Hsu & Shih, 2020 | | | Ptychognathus takahashii Sakai, 1939 | | | Ptychognathus takahasii Sakai, 1939 | | | Pyxidognathus deianira (Man, 1888) | ⇒ Parapyxidognathus deianira De Man, 1888 | | Pyxidognathus granulosus A. Milne-Edw..., 1879 | | | Pyxidognathus subglobosus Tesch, 1918 | | | Scutumara enodis Ng & Nakasone, 1993 | | | Scutumara laniger Tesch, 1918 | | | Scutumara miyakei Nakamura & T..., 1972 | | | Sestrostoma balssi Shen, 1932 | | | Sestrostoma depressum Sakai, 1965 | | | Sestrostoma toriumii Takeda, 1974 | | | Shurebus genoensis (Leach, 1847) | ⇒ Brachynotus sexdentatus Risso, 1827 | | Tetragrapsus jouyi Rathbun, 1894 | | | Thalassograpsus harpax Hilgendorf, 1892 | | | Utica barbimana A. Milne-Edw..., 1873 | | | Utica borneensis De Man, 1895 | | | Utica crassimana Haswell, 1882 | | | Utica glabra A. Milne-Edw..., 1873 | | | Utica gracilipes White, 1847 | | | Utica nausithoe De Man, 1895 | | | Utica setosipes Haswell, 1881 | | | Vanua linearius ♰ M. J. Rathbun, 1945 | | | Varuna altimanus (Rathbun, 1914) | ⇒ Ptychognathus altimanus Rathbun, 1914 | | Varuna angustifrons Karasawa, 1993 | | | Varuna litterata Fabricius, 1798 | Peregrine Crab | | Varuna tomentosa (Pfeffer, 1889) | ⇒ Varuna litterata Fabricius, 1798 | | Varuna yui Hwang & Takeda, 1986 | Sundaic Paddler Crab |
Subnodes of Varunidae with occurrences in Continents
Links, sources and further information Varunidae at artfakta.se
Varunidae at biodiversity.org.au
Varunidae at dyntaxa.se
Varunidae at gbif.org
Varunidae at irmng.org
Varunidae at itis.gov
Varunidae at marinespecies.org
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