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Checklist for Order Coleoptera Linnaeus, 1758 (Beetles)
and region Grave (North Brabant, Netherlands)


Class
 		Insecta
Linnaeus, 1758
Insects, True insects		193
856
1188		Families
Genera
Species		 
Taxom
 

Subclass
 		Pterygota
Lang, 1888
Winged insects		192
855
1187		Families
Genera
Species		 
Taxom
 

Infraclass
 		Neoptera
van der Wulp, 1890
Modern Wing-folding insects		180
827
1136		Families
Genera
Species		 
Taxom
 

Superorder
 		Holometabola
Heider, 1889
Holometabolous Insects		133
657
910		Families
Genera
Species		 
Taxom
 

Order
 		Coleoptera
Linnaeus, 1758
Beetles, Coleopterans		27
130
161		Families
Genera
Species		 
Taxom
 


List of families for Grave
27 Families width 161 Species and 2 Subspecies
#DCSBEHMANGP
Dytiscidae Leach, 1815Diving beetles12SpeciesTaxom
Carabidae Latreille, 1802Carabid beetles21SpeciesTaxom
Coccinellidae Latreille, 1807Coccinelles27SpeciesTaxom
Cerambycidae Latreille, 1802Long Horn beetles18SpeciesTaxom
Chrysomelidae Latreille, 1802Casebearers11SpeciesTaxom
Scarabaeidae Latreille, 1802Dung beetles11SpeciesTaxom
Buprestidae Leach, 1815Flat-headed wood-borers2SpeciesTaxom
Elateridae Leach, 1815Click Beetle2SpeciesTaxom
Hydrophilidae Latreille, 1802Silver Water Beetles13SpeciesTaxom
Melyridae Leach, 1815Malachite Beetles2SpeciesTaxom
Attelabidae Billberg, 1820Leaf Rolling Weevil2SpeciesTaxom
Cantharidae Imhoff, 1856Cantharids1SpeciesTaxom
Curculionidae Latreille, 1802Ambrosia beetles14SpeciesTaxom
Nitidulidae Latreille, 1802Nitidules1SpeciesTaxom
Silphidae (Latreille, 1807)Burying And Carrion Beetles4SpeciesTaxom
Dryopidae (Billberg, 1820)Long toed water beetle1SpeciesTaxom
Endomychidae Leach, 1815False Ladybirds1SpeciesTaxom
Gyrinidae Latreille, 1802Whirligig beetle1SpeciesTaxom
Haliplidae Aubé, 1836Crawling Water Beetle5SpeciesTaxom
Histeridae Gyllenhal, 1808Clown beetle1SpeciesTaxom
Meloidae (Gyllenhal, 1810)Blister Beetle1SpeciesTaxom
Noteridae C. G. Thomson, 1860Burrowing water beetle2SpeciesTaxom
Staphylinidae Lameere, 1900Rove beetles3SpeciesTaxom
Elmidae Curtis, 1830Drive beetles2SpeciesTaxom
Ptinidae Latreille, 1802Death-watch and Spider Beetles1SpeciesTaxom
Pyrochroidae (Latreille, 1807)Cardinal beetles1SpeciesTaxom
Silvanidae (Kirby, 1837)Flat bark beetles1SpeciesTaxom


Genera for Grave
130 Genera with 161 Species
#ABCDEGHILMNOPRSTUV
Acilius Leach, 1758Small Flat Diving Beetles2SpeciesTaxom
Acupalpus Latreille, 18291SpeciesTaxom
Adalia Mulsant, 17582SpeciesTaxom
Agapanthia Serville, 17751SpeciesTaxom
Agelastica Dejean, 17581SpeciesTaxom
Agonum Bonelli, 17841SpeciesTaxom
Agrilinus Mulsant & Rey, 17741SpeciesTaxom
Agrilus Curtis, 19631SpeciesTaxom
Agrypnus Eschscholtz, 17581SpeciesTaxom
Amara Bonelli, 1825Seed-Eating Ground Beetles1SpeciesTaxom
Ampedus Dejean, 17841SpeciesTaxom
Anacaena Thomson, 18292SpeciesTaxom
Anaglyptus Mulsant, 17581SpeciesTaxom
Anatis Mulsant, 1758Giant Lady Beetles1SpeciesTaxom
Anchomenus Bonelli, 17631SpeciesTaxom
Anisosticta Chevrolat, 17581SpeciesTaxom
Anomala Samouelle, 1763Shining Leaf Chafer1SpeciesTaxom
Anthocomus Erichson, 17841SpeciesTaxom
Aphidecta Weise, 17581SpeciesTaxom
Attelabus Linnaeus, 17631SpeciesTaxom
Bembidion Latreille, 18251SpeciesTaxom
Brumus Mulsant, 17581SpeciesTaxom
Calathus Bonelli, 18501SpeciesTaxom
Calosoma Weber, 1758Caterpillar hunters1SpeciesTaxom
Calvia Mulsant, 17672SpeciesTaxom
Cantharis Linnaeus, 17831SpeciesTaxom
Carabus Linnaeus, 1764Ground beetle1SpeciesTaxom
Cercyon Leach, 17905SpeciesTaxom
Chaetarthria Stephens, 17971SpeciesTaxom
Chilocorus Leach, 1790Red-spotted lady beetles2SpeciesTaxom
Chlorophanus C.R. Sahlberg, 17851SpeciesTaxom
Chrysolina Motschulsky, 17631SpeciesTaxom
Chrysomela Linnaeus, 17583SpeciesTaxom
Cicindela Linnaeus, 1758Common Tiger Beetles2SpeciesTaxom
Cionus Clairville, 17631SpeciesTaxom
Clytus Laicharting, 17581SpeciesTaxom
Coccidula Gyllenhal, 17831SpeciesTaxom
Coccinella Linnaeus, 17583SpeciesTaxom
Cortodera Mulsant, 17831SpeciesTaxom
Crioceris Geoffroy, 1758Asparagus beetle2SpeciesTaxom
Cybister Curtis, 17741SpeciesTaxom
Cychramus Kugelann, 17871SpeciesTaxom
Dendroxena Motschulsky, 17711SpeciesTaxom
Deporaus Leach / Samo..., 17581SpeciesTaxom
Deronectes Sharp, 1882
0SpeciesTaxom
Dicheirotrichus Jacquelin du..., 18271SpeciesTaxom
Dorytomus Germar, 1787Willow Weevils2SpeciesTaxom
Dryops Olivier, 18471SpeciesTaxom
Duvalius Delarouzee, 19131SpeciesTaxom
Dytiscus Linnaeus, 17751SpeciesTaxom
Endomychus Panzer, 17581SpeciesTaxom
Enochrus Thomson, 17942SpeciesTaxom
Galerucella Crotch, 1758Water-lily Beetle1SpeciesTaxom
Grammoptera Serville, 17811SpeciesTaxom
Graptodytes Seidlitz, 17871SpeciesTaxom
Gyrinus Geoffroy, 1826Whirligig Beetle1SpeciesTaxom
Hadroplontus C. G. Thomson, 17751SpeciesTaxom
Haliplus Latreille, 18025SpeciesTaxom
Halyzia Mulsant, 17581SpeciesTaxom
Harmonia Mulsant, 1763Greater Lady Beetles2SpeciesTaxom
Helophorus Fabricius, 17612SpeciesTaxom
Hippodamia Dejean, 1777Black-spotted lady beetles2SpeciesTaxom
Hololepta Paykull, 1776Clown Beetles1SpeciesTaxom
Hydroporus Clairville, 17611SpeciesTaxom
Hygrotus Stephens, 17832SpeciesTaxom
Hyphydrus Illiger, 17611SpeciesTaxom
Ilybius Erichson, 17741SpeciesTaxom
Laccobius Erichson, 18291SpeciesTaxom
Laccophilus Leach, 17741SpeciesTaxom
Leptinotarsa Chevrolat, 1824Potato Beetles1SpeciesTaxom
Leptura Linnaeus, 17581SpeciesTaxom
Litomeces Murray, 19221SpeciesTaxom
Loricera Latreille, 17751SpeciesTaxom
Malachius Fabricius, 17581SpeciesTaxom
Melinopterus Mulsant, 17981SpeciesTaxom
Meloe Linnaeus, 1758Oil Beetle1SpeciesTaxom
Melolontha Fabricius, 1758Billy witch1SpeciesTaxom
Myrrha Mulsant, 17581SpeciesTaxom
Nedyus Schönherr, 17581SpeciesTaxom
Nicrophorus Fabricius, 1783Burying Beetle1SpeciesTaxom
Noterus Clairville, 17762SpeciesTaxom
Oberea Dejean, 17581SpeciesTaxom
Odacantha Paykull, 17661SpeciesTaxom
Oenopia Mulsant, 17581SpeciesTaxom
Oiceoptoma Leach, 17581SpeciesTaxom
Ontholestes Ganglbauer, 17851SpeciesTaxom
Oodes Bonelli, 17921SpeciesTaxom
Oryctes Hellwig, 1758Coconut rhinoceros beetles1SpeciesTaxom
Otiorhynchus Germar, 1824Strawberry root weevil4SpeciesTaxom
Oulimnius des Gozis, 18062SpeciesTaxom
Oxypselaphus Chaudoir, 17841SpeciesTaxom
Oxythyrea Mulsant, 17611SpeciesTaxom
Pachytodes Pic, 17811SpeciesTaxom
Panagaeus Latreille, 17581SpeciesTaxom
Paracorymbia Miroshnikov, 17751SpeciesTaxom
Paradromius Fowler, 17951SpeciesTaxom
Paranchus Lindroth, 17961SpeciesTaxom
Phyllopertha Stephens, 17581SpeciesTaxom
Phymatodes Mulsant, 17581SpeciesTaxom
Plagiodera Laicharting, 17811SpeciesTaxom
Platambus C.G. Thomson, 17581SpeciesTaxom
Platynus Bonelli, 17901SpeciesTaxom
Pogonocherus Dejean, 17581SpeciesTaxom
Propylaea Mulsant, 17581SpeciesTaxom
Pseudovadonia Lobanov, Mur..., 17761SpeciesTaxom
Psyllobora Dejean, 1758Fungus-eating Lady Beetles1SpeciesTaxom
Pterostichus Bonelli, 1796Woodland Ground Beetle1SpeciesTaxom
Ptinomorphus Mulsant & Rey, 17671SpeciesTaxom
Pyrochroa Geoffroy, 1761cardinal beetle1SpeciesTaxom
Pyrrhalta Joannis, 17991SpeciesTaxom
Rhinoncus Schönherr, 17581SpeciesTaxom
Rhysothorax Bedel, 17921SpeciesTaxom
Rhyzobius Stephens, 17872SpeciesTaxom
Rutpela Nakane & Ohb..., 17611SpeciesTaxom
Scabrostomus Gordon and S..., 19911SpeciesTaxom
Scaphidium Olivier, 17901SpeciesTaxom
Serica MacLeay, 17581SpeciesTaxom
Silpha Linnaeus, 17581SpeciesTaxom
Stenurella Villiers, 17922SpeciesTaxom
Stictoleptura Casey, 17581SpeciesTaxom
Strangalia Serville, 17581SpeciesTaxom
Strophosoma Billberg, 17751SpeciesTaxom
Subcoccinella Huber, 17581SpeciesTaxom
Tanymecus Germar, 17871SpeciesTaxom
Trachys Fabricius, 17581SpeciesTaxom
Tunicopterus Tikhomirova, 19681SpeciesTaxom
Tylotus Schonherr, 17971SpeciesTaxom
Tytthaspis Crotch, 17581SpeciesTaxom
Uleiota Latreille, 17611SpeciesTaxom
Valgus Scriba, 17581SpeciesTaxom


Species list for Grave
161 Species and 2 Subspecies
Acilius (Acilius) canaliculatus Nicolai, 1822
Acilius (Acilius) sulcatus Linnaeus, 1758
Acupalpus (Acupalpus) exiguus Dejean, 1829
Adalia (Adalia) bipunctata Linnaeus, 17582-spot Ladybird
Adalia (Adalia) decempunctata Linnaeus, 175810-spot Ladybird
Agapanthia (Epoptes) villosoviridescens (DeGeer, 1775)⇒ Agapanthia daurica daurica Ganglbauer, 1883
Agelastica alni Linnaeus, 1758Alder leaf beetle
Agonum (Agonum) muelleri Herbst, 1784Müller'S Harp Ground Beetle
Agrilinus ater De Geer, 1774
Agrilus syrphus Descarpentri..., 1963
Agrypnus murinus Linnaeus, 1758Mousegrey click beetle
Amara (Amara) montivaga Sturm, 1825
Ampedus (Ampedus) pomorum Herbst, 1784
Anacaena limbata Fabricius, 1792
Anacaena lutescens Stephens, 1829
Anaglyptus (Anaglyptus) mysticus Linnaeus, 1758Grey-coated longhorn beetle
Anatis ocellata Linnaeus, 1758Eye-spotted lady beetle
Anchomenus (Anchomenus) dorsalis Pontoppidan, 1763
Anisosticta novemdecimpunctata Linnaeus, 175819-spot ladybird
Anomala (Anomala) dubia (Scopoli, 1763)⇒ Anomala undulata Melsheimer, 1845
Anthocomus (Anthocomus) rufus Herbst, 1784
Aphidecta obliterata Linnaeus, 1758Ladybird beetle
Attelabus (Attelabus) nitens Scopoli, 1763Oak leaf-roller
Bembidion (Notaphus) obliquum Sturm, 1825
Brumus quadripustulatus (Linnaeus, 1758)⇒ Harmonia axyridis Pallas, 1773
Calathus (Neocalathus) cinctus Motschulsky, 1850
Calosoma (Acalosoma) inquisitor Linnaeus, 1758Caterpillar hunter
Calvia (Anisocalvia) quatuordecimguttata Linnaeus, 1758Cream Spotted Lady-beetle
Calvia (Calvia) decemguttata Linnaeus, 176710 spot ladybird
Cantharis coccinea Schaller, 1783
Carabus (Archicarabus) nemoralis Müller, 1764Bronze Ground Beetle
Cercyon (Cercyon) bifenestratus K?ster, 1851
Cercyon (Cercyon) convexiusculus Stephens, 1829Jumping-bean sapium
Cercyon (Cercyon) marinus Thomson, 1853
Cercyon (Cercyon) sternalis Sharp, 1918
Cercyon (Dicyrtocercyon) ustulatus (Preyssler, 1790)⇒ Cercyon nigriceps Marsham, 1802
Chaetarthria seminulum Herbst, 1797
Chilocorus bipustulatus Linnaeus, 1758Heather Lady Beetle
Chilocorus renipustulatus Scriba, 1790Kidney-spot Ladybird
Chlorophanus viridis (Linnaeus, 1785)⇒ Chlorophanus flavescens Schoenherr, 1826
Chrysolina (Fastuolina) fastuosa Scopoli, 1763Dead-nettle leaf beetle
Chrysomela (Chrysomela) populi Linnaeus, 1758Balsam poplar leaf beetle
Chrysomela elongata (Linnaeus, 1758)⇒ Tillus elongatus Linnaeus, 1758
Chrysomela tanaceti (Linnæus, 1758)⇒ Galeruca tanaceti Linnaeus, 1758
Cicindela (Cicindela) campestris (Linnaeus, 1758)⇒ Cicindela hybrida hybrida Linnaeus, 1758
Cicindela (Cicindela) hybrida Linnaeus, 1758Northern dune tiger beetle
Cionus tuberculosus Scopoli, 1763
Clytus arietis Linnaeus, 1758Wasp beetle
Coccidula rufa Herbst, 1783
Coccinella (Coccinella) quinquepunctata Linnaeus, 17585-spot Ladybird
Coccinella (Coccinella) septempunctata Linnaeus, 17587-spot Ladybird
Coccinella (Spilota) undecimpunctata Linnaeus, 175811-spot Ladybird
Cortodera humeralis Schaller, 1783
Crioceris (Crioceris) asparagi Linnaeus, 1758Asparagus beetle
Crioceris (Crioceris) duodecimpunctata Linnaeus, 1758Spotted asparagus beetle
Cybister (Scaphinectes) lateralimarginalis De Geer, 1774Grand Junction camissonia
Cychramus luteus Fabricius, 1787
Dendroxena quadrimaculata Scopoli, 1771Carrion beetle
Deporaus (Deporaus) betulae Linnaeus, 1758Birch Leaf Roller
Dicheirotrichus (Trichocellus) placidus (Gyllenhal, 1827)⇒ Trichocellus placidus Gyllenhal, 1827
Dorytomus (Dorytomus) longimanus Forster, 1771
Dorytomus (Dorytomus) tremulae Fabricius, 1787
Dryops (Dryops) luridus (Erichson, 1847)⇒ Dryops gracilis Karsh, 1881
Duvalius paroecus dryops Bokor, 1913
Dytiscus hermanni (Fabricius, 1775)⇒ Hygrobia hermanni Fabricius, 1775
Endomychus coccineus Linnaeus, 1758False Ladybird
Enochrus (Lumetus) testaceus Fabricius, 1801
Enochrus (Methydrus) affinis Thunberg, 1794
Galerucella (Galerucella) nymphaeae Linnaeus, 1758Leaf Beetle
Grammoptera (Grammoptera) ruficornis Fabricius, 1781
Graptodytes pictus Fabricius, 1787
Gyrinus (Gyrinus) substriatus Sturm, 1826Whirligig Beetle
Hadroplontus litura Fabricius, 1775Canadian Thistle Stem Weevil
Haliplus (Haliplidius) varius Nicolai, 1822
Haliplus (Haliplinus) fluviatilis Aubé, 1836
Haliplus (Haliplinus) heydeni Wehncke, 1875
Haliplus (Liaphlus) laminatus Schaller, 1783
Haliplus (Neohaliplus) lineatocollis Marsham, 1802
Halyzia sedecimguttata Linnaeus, 175816-Spot Orange
Harmonia axyridis Pallas, 1773Asian lady beetle
Harmonia quadripunctata Pontoppidan, 1763Cream-streaked lady beetle
Helophorus (Rhopalohelophorus) brevipalpis Bedel, 1881
Helophorus (Rhopalohelophorus) granularis Linnaeus, 1761
Hippodamia (Hemisphaerica) tredecimpunctata Linnaeus, 175813-spot ladybird
Hippodamia (Hippodamia) variegata Goeze, 1777Adonis' Ladybird
Hololepta (Hololepta) plana Sulzer, 1776
Hydroporus (Hydroporus) palustris (Linnaeus, 1761)⇒ Hydroporus notatus Sturm, 1835
Hygrotus (Hygrotus) inaequalis Fabricius, 1777
Hygrotus (Hygrotus) versicolor Schaller, 1783
Hyphydrus ovatus Linnaeus, 1761
Ilybius (Ilybius) ater De Geer, 1774
Laccobius (Laccobius) colon Stephens, 1829
Laccophilus hyalinus De Geer, 1774
Leptinotarsa decemlineata Say, 1824Colorado beetle
Leptura (Leptura) quadrifasciata (Linnaeus, 1758)⇒ Strangalia attenuata Linnaeus, 1758
Litomeces gracilis sphecodes Schmidt, 1922
Loricera (Loricera) pilicornis Fabricius, 1775Bigtooth aspen
Malachius (Malachius) bipustulatus Linnaeus, 1758Malachite beetle
Melinopterus sphacelatus Panzer, 1798
Meloe (Meloe) proscarabaeus (Linnaeus, 1758)⇒ Mordella aradasiana Patti, 1840
Melolontha (Melolontha) melolontha Linnaeus, 1758Cockchafer
Myrrha (Myrrha) octodecimguttata Linnaeus, 175818-spot ladybird
Nedyus quadrimaculatus Linnaeus, 1758Small Nettle Weevil
Nicrophorus vespilloides Herbst, 1783Boreal Carrion Beetle
Noterus clavicornis (De Geer, 1774)⇒ Noterus crassicornis O.F.Müller, 1776
Noterus crassicornis O.F.Müller, 1776Smaller Noterus
Oberea (Oberea) oculata Linnaeus, 1758Eyed longhorn beetle
Odacantha (Odacantha) melanura Linnaeus, 1766
Oenopia conglobata Linnaeus, 1758Ladybird beetle
Oiceoptoma thoracicum Linnaeus, 1758Red-breasted Carrion Beetle
Ontholestes tessellatus Geoffroy, 1785
Oodes helopioides Fabricius, 1792
Oryctes (Oryctes) nasicornis Linnaeus, 1758European rhinoceros beetle
Otiorhynchus (Dorymerus) sulcatus Fabricius, 1775Black vine beetle
Otiorhynchus (Otiorhynchus) aurifer Boheman, 1843
Otiorhynchus (Pendragon) ovatus Linnaeus, 1758Strawberry root weevil
Otiorhynchus (Pocodalemes) crataegi Germar, 1824Privet weevil
Oulimnius rivularis Rosenhauer, 1856
Oulimnius tuberculatus Müller, 1806Seychelles Magpie-Robin
Oxypselaphus obscurus Herbst, 1784
Oxythyrea funesta Poda, 1761
Pachytodes cerambyciformis (Schrank, 1781)⇒ Judolia cordifera Olivier, 1795
Panagaeus (Panagaeus) cruxmajor Linnaeus, 1758Crucifix ground beetle
Paracorymbia fulva DeGeer, 1775
Paradromius (Manodromius) linearis Olivier, 1795
Paranchus albipes Fabricius, 1796White-legged harp ground beetle
Phyllopertha horticola (Linnaeus, 1758)⇒ Phyllopertha intermixta Arrow, 1913
Phymatodes (Phymatodes) testaceus Linnaeus, 1758Long-Horned Beetle
Plagiodera (Plagiodera) versicolora Laicharting, 1781Imported poplar and willow beetle
Platambus maculatus Linnaeus, 1758
Platynus (Platynus) assimilis Paykull, 1790
Pogonocherus (Pogonocherus) hispidus (Linnaeus, 1758)⇒ Pogonocherus hispidulus Piller & Mit..., 1783
Propylaea quatuordecimpunctata Linnaeus, 175814-Spotted Ladybird Beetle
Pseudovadonia livida Fabricius, 1776Fairy-ring longhorn beetle
Psyllobora vigintiduopunctata Linnaeus, 175822-spot ladybird
Pterostichus (Argutor) vernalis Panzer, 1796Spring Blackclock
Ptinomorphus imperialis Linnaeus, 1767
Pyrochroa serraticornis (Scopoli, 1761)⇒ Trichalus serraticornis Fabricius, 1775
Pyrrhalta viburni Paykull, 1799Guelder-Rose Leaf Beetle
Rhinoncus (Rhinoncus) pericarpius Linnaeus, 1758
Rhysothorax rufus Fabricius, 1792Red sand scarab beetle
Rhyzobius chrysomeloides Herbst, 1792
Rhyzobius litura Fabricius, 1787
Rutpela maculata (Poda, 1761)⇒ Judolia antecurrens H. F. Wickham, 1913
Scabrostomus tanytarsus Skelley & Wo..., 1991Masked stargazer
Scaphidium quadrimaculatum Olivier, 1790
Serica brunna Linnaeus, 1758
Silpha atrata (Linnæus, 1758)⇒ Phosphuga atrata atrata Linnaeus, 1758
Stenurella (Nigrostenurella) nigra Linnaeus, 1758
Stenurella melanura Fabricius, 1792
Stictoleptura (Stictoleptura) rubra (Linnaeus, 1758)⇒ Aredolpona rubra rubra Linné, 1758
Strangalia (Strangalina) attenuata Linnaeus, 1758
Strophosoma (Strophosoma) capitatum DeGeer, 1775
Subcoccinella vigintiquatuorpunctata Linnaeus, 175824-spot Ladybird
Tanymecus (Tanymecus) palliatus Fabricius, 1787Beet leaf weevil
Trachys minutus Linnaeus, 1758Metallic wood-boring beetle
Tunicopterus sigara ♰ Tikhomirova, 1968
Tylotus chrysops Herbst, 1797
Tytthaspis (Tytthaspis) sedecimpunctata Linnaeus, 175816 spot ladybird
Uleiota planatus (Linnaeus, 1761)⇒ Uleiota planata Linnaeus, 1761
Valgus hemipterus Linnaeus, 1758Bug flower chafer

Subnodes of Coleoptera with occurrences in Grave
SuborderAdephaga
Schellenberg, 1806
Adephagans, Ground Beetles and Water Beetles, Predaceous Diving and Whirligig Beetles		5
34
41		Families
Genera
Species
Taxom

SuborderPolyphaga
Emery, 1886
Polyphags		22
96
120		Families
Genera
Species
Taxom



Coleoptera (Linnaeus 1758)
Coleoptera (Linnaeus 1758)


Links, sources and further information
Coleoptera at szmn.sbras.ru
Coleoptera at papua-insects.nl
Coleoptera at artfakta.se
Coleoptera at dyntaxa.se
Coleoptera at gbif.org
Coleoptera at irmng.org
Coleoptera at itis.gov
Coleoptera at kaefer-der-welt.de
Coleoptera at marinespecies.org