AntsAntsArachnidsArachnids
BeesBeesBeetlesBeetles
BookliceBookliceBugsBugs
BumblebeesBumblebeesButterfliesButterflies
CicadasCicadasCrane fliesCrane flies
DragonfliesDragonfliesEarwigsEarwigs
FliesFliesIsopodsIsopods
LocustsLocustsMantisesMantises
MayfliesMayfliesMosquitoesMosquitoes
MyriapodsMyriapodsNet-wingedNet-winged
Plant-parasiticPlant-parasiticScorpion fliesScorpion flies
TermitesTermitesWaspsWasps
INSECTOIDGERENGDARK LIGHT
Home Arthropoda Imprint Contact
TAXFAMGENSPESUB
⌕
Checklist for Order Coleoptera Linnaeus, 1758 (Beetles)
and region Raleigh County (West Virginia, USA)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
214
898
1250
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
214
898
1250
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
201
857
1175
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
155
731
1020
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
41
132
146
Families
Genera
Species
 
Taxom
 


List of families for Raleigh County (West Virginia)
41 Families width 146 Species and 1 Subspecies
#CBTEAMDSHNLGP
Carabidae Latreille, 1802Carabid beetles22SpeciesTaxom
Buprestidae Leach, 1815Flat-headed wood-borers6SpeciesTaxom
Tenebrionidae Latreille, 1802Darkling Beetle4SpeciesTaxom
Elateridae Leach, 1815Click Beetle7SpeciesTaxom
Cerambycidae Latreille, 1802Long Horn beetles16SpeciesTaxom
Anthicidae Latreille, 1819Ant beetles1SpeciesTaxom
Melyridae Leach, 1815Malachite Beetles1SpeciesTaxom
Dermestidae Hope, 1840Carpet beetles2SpeciesTaxom
Scarabaeidae Latreille, 1802Dung beetles7SpeciesTaxom
Hydrophilidae Latreille, 1802Silver Water Beetles3SpeciesTaxom
Brachypteridae Erichson, 1845Short-winged Flower Beetles1SpeciesTaxom
Nitidulidae Latreille, 1802Nitidules5SpeciesTaxom
Chrysomelidae Latreille, 1802Casebearers8SpeciesTaxom
Lucanidae Latreille, 1804Stag Beetle2SpeciesTaxom
Cantharidae Imhoff, 1856Cantharids0SpeciesTaxom
Coccinellidae Latreille, 1807Coccinelles12SpeciesTaxom
Curculionidae Latreille, 1802Ambrosia beetles7SpeciesTaxom
Latridiidae Erichson, 1842Minute brown scavenger beetle0SpeciesTaxom
Cupedidae Laporte, 1836Cupedids2SpeciesTaxom
Gyrinidae Latreille, 1802Whirligig beetle2SpeciesTaxom
Melandryidae Leach, 1815False Darkling Beetle2SpeciesTaxom
Histeridae Gyllenhal, 1808Clown beetle4SpeciesTaxom
Ptinidae Latreille, 1802Death-watch and Spider Beetles1SpeciesTaxom
Haliplidae Aubé, 1836Crawling Water Beetle0SpeciesTaxom
Dryopidae Billberg, 1820Long toed water beetle2SpeciesTaxom
Dytiscidae (Leach, 1815)Diving beetles1SpeciesTaxom
Derodontidae LeConte, 1861Tooth-neck fungus beetles2SpeciesTaxom
Leiodidae Fleming, 1821Round fungus beetles1SpeciesTaxom
Mycetophagidae Leach, 1815Hairy fungus beetle1SpeciesTaxom
Lampyridae Latreille, 1817Fireflies1SpeciesTaxom
Elmidae Curtis, 1830Drive beetles6SpeciesTaxom
Meloidae (Gyllenhal, 1810)Blister Beetle1SpeciesTaxom
Mordellidae Latreille, 1802Pintail beetles3SpeciesTaxom
Endomychidae Leach, 1815False Ladybirds1SpeciesTaxom
Silphidae Latreille, 1807Burying And Carrion Beetles3SpeciesTaxom
Staphylinidae Lameere, 1900Rove beetles4SpeciesTaxom
Scirtidae Fleming, 1821Marsh Beetle1SpeciesTaxom
Ptilodactylidae Laporte, 1836Ptilodactylid beetles1SpeciesTaxom
Erotylidae Latreille, 1802Pleasing Fungus Beetle1SpeciesTaxom
Throscidae Laporte, 1840False metallic wood-boring beetles1SpeciesTaxom
Thanerocleridae Chapin, 19211SpeciesTaxom


Genera for Raleigh County (West Virginia)
132 Genera with 146 Species
#ABCDEFGHILMNOPSTUVZ
Agonum Bonelli, 18232SpeciesTaxom
Agrilus Curtis, 19632SpeciesTaxom
Alaetrinus Iwan, 18051SpeciesTaxom
Ampedus Dejean, 18383SpeciesTaxom
Anelaphus Linsley, 1793Oak twig pruners1SpeciesTaxom
Anisodactylus Dejean, 17981SpeciesTaxom
Anthicus Paykull, 18481SpeciesTaxom
Anthocomus Erichson, 17811SpeciesTaxom
Anthrenus Geoffroy, 1767Carpet beetles2SpeciesTaxom
Aphodius Hellwig, 17741SpeciesTaxom
Ataenius Harold, 18482SpeciesTaxom
Athous Eschscholtz, 18371SpeciesTaxom
Berosus Leach, 17971SpeciesTaxom
Brachypterus Kugelann, 17921SpeciesTaxom
Brachysomida Casey, 18241SpeciesTaxom
Bradycellus Erichson, 1837
0SpeciesTaxom
Brassicogethes Audisio and ..., 17751SpeciesTaxom
Buprestis Linnaeus, 1841Jewel Beetle1SpeciesTaxom
Calligrapha Chevrolat, 1758Calligrapha Beetle1SpeciesTaxom
Calodromius Reitter, 19331SpeciesTaxom
Carabus Linnaeus, 1801Ground beetle1SpeciesTaxom
Cassida Linnaeus, 1776Tortoise Beetles1SpeciesTaxom
Ceruchus MacLeay, 18011SpeciesTaxom
Chaetocnema Stephens, 18801SpeciesTaxom
Chalcophora Dejean, 17701SpeciesTaxom
Chauliognathus Hentz, 1829
Soldier Beetle0SpeciesTaxom
Chilocorus Leach, 1835Red-spotted lady beetles1SpeciesTaxom
Chlaenius Bonelli, 1771Vivid Metallic Ground Beetle2SpeciesTaxom
Chrysochus Chevrolat, 17751SpeciesTaxom
Cicindela Linnaeus, 1873Common Tiger Beetles7SpeciesTaxom
Clytus Laicharting, 17951SpeciesTaxom
Coccinella Linnaeus, 17581SpeciesTaxom
Coleomegilla Timberlake, 17751SpeciesTaxom
Conotrachelus Schönherr, 18762SpeciesTaxom
Corticarina Reitter, 1881
0SpeciesTaxom
Crioceris Geoffroy, 1758Asparagus beetle1SpeciesTaxom
Cryptolaemus Mulsant, 18531SpeciesTaxom
Cupes Fabricius, 18011SpeciesTaxom
Cyclocephala Dejean, 1911Masked Chafers1SpeciesTaxom
Cycloneda Crotch, 1835Spotless Lady Beetles1SpeciesTaxom
Cymbiodyta Bedel, 18691SpeciesTaxom
Diabrotica Dejean, 1843Corn rootworms1SpeciesTaxom
Dicerca Eschscholtz, 18231SpeciesTaxom
Dichotomius Hope, 17671SpeciesTaxom
Dineutus MacLeay, 1838whirligig beetle2SpeciesTaxom
Diomus Mulsant, 18351SpeciesTaxom
Dircaea Fabricius, 18661SpeciesTaxom
Disonycha Dejean, 17921SpeciesTaxom
Enochrus Thomson, 18441SpeciesTaxom
Epilachna Dejean, 1850Epilachna Lady Beetles1SpeciesTaxom
Epuraea Erichson, 1843
0SpeciesTaxom
Euspilotus Lewis, 18111SpeciesTaxom
Fabogethes Audisio and ..., 18301SpeciesTaxom
Gaurotes Le Conte, 18472SpeciesTaxom
Glischrochilus Reitter, 1790Beer bugs2SpeciesTaxom
Goes LeConte, 18471SpeciesTaxom
Hadrobregmus Thomson, 18251SpeciesTaxom
Haliplus Latreille, 1802
0SpeciesTaxom
Harmonia Mulsant, 1773Greater Lady Beetles1SpeciesTaxom
Helichus Erichson, 18242SpeciesTaxom
Hemicrepidius Germar, 18061SpeciesTaxom
Hippodamia Dejean, 1842Black-spotted lady beetles1SpeciesTaxom
Hister Linnaeus, 18251SpeciesTaxom
Hololepta Paykull, 1825Clown Beetles1SpeciesTaxom
Hoplia Illiger, 1823Monkey Beetles1SpeciesTaxom
Hydroporus Clairville, 18231SpeciesTaxom
Hylobius Germar, 1797Pine Weevil1SpeciesTaxom
Hymenorus Mulsant, 1851
0SpeciesTaxom
Hyperaspis Chevrolat, 1837
Sigil Lady Beetles0SpeciesTaxom
Hypulus Paykull, 18381SpeciesTaxom
Ithycerus Schönherr, 17711SpeciesTaxom
Laricobius Rosenhauer, 18612SpeciesTaxom
Latridius Herbst, 1793
0SpeciesTaxom
Lebia Latreille, 1823colorful foliage ground beetle1SpeciesTaxom
Leiodes Latreille, 19931SpeciesTaxom
Lilioceris Reitter, 17631SpeciesTaxom
Litargus Erichson, 18561SpeciesTaxom
Litomeces Murray, 19221SpeciesTaxom
Loricera Latreille, 17751SpeciesTaxom
Lucanus Scopoli, 17581SpeciesTaxom
Lucidota Laporte, 17901SpeciesTaxom
Macronychus Mueller, 18251SpeciesTaxom
Madarellus Casey, 18241SpeciesTaxom
Mecyclothorax Sharp, 20151SpeciesTaxom
Melanotus Schwarz, 1837Click Beetle1SpeciesTaxom
Meloe Linnaeus, 1815Oil Beetle1SpeciesTaxom
Meracantha Kirby, 18051SpeciesTaxom
Monochamus Dejean, 1824Sawyer beetles1SpeciesTaxom
Mordellina Schilsky, 18451SpeciesTaxom
Mordellistena Costa, 19171SpeciesTaxom
Mulsantina Weise, 18991SpeciesTaxom
Mycetina Mulsant, 18381SpeciesTaxom
Necrophila Kirby & Spence, 17581SpeciesTaxom
Neomida Mulsant, 17771SpeciesTaxom
Nicrophorus Fabricius, 1825Burying Beetle1SpeciesTaxom
Oiceoptoma Leach, 17811SpeciesTaxom
Ontholestes Ganglbauer, 18021SpeciesTaxom
Optioservus Sanderson, 18632SpeciesTaxom
Pallodes Erichson, 18051SpeciesTaxom
Panscopus Schönherr, 18311SpeciesTaxom
Paramordellaria Ermisch, 18241SpeciesTaxom
Paratachys Casey, 1918
0SpeciesTaxom
Paratenetus Spinola, 18441SpeciesTaxom
Philonthus Stephens, 18302SpeciesTaxom
Photuris Dejean, 1833
Femme fatale fireflies0SpeciesTaxom
Pidonia Mulsant, 18241SpeciesTaxom
Platynus Bonelli, 18481SpeciesTaxom
Plegaderus Erichson, 18251SpeciesTaxom
Prionocyphon Redtenbacher, 18251SpeciesTaxom
Propylaea Mulsant, 17581SpeciesTaxom
Pseudanostirus Dolin, 18341SpeciesTaxom
Psyllobora Dejean, 1824Fungus-eating Lady Beetles1SpeciesTaxom
Ptilodactyla Illiger, 18231SpeciesTaxom
Ptosima Dejean, 18231SpeciesTaxom
Sarosesthes Thomson, 17751SpeciesTaxom
Selenophorus Dejean, 18631SpeciesTaxom
Stenelmis Dufour, 19383SpeciesTaxom
Stenolophus Dejean, 1823Seedcorn Beetle2SpeciesTaxom
Stenus Latreille, 18741SpeciesTaxom
Strangalepta Casey, 18241SpeciesTaxom
Strangalia Serville, 17751SpeciesTaxom
Strophiona Casey, 17711SpeciesTaxom
Synaphaeta Thomson, 18521SpeciesTaxom
Tenomerga Neboiss, 18311SpeciesTaxom
Tetracha Hope, 1767Metallic Tiger Beetles1SpeciesTaxom
Tritoma Fabricius, 18261SpeciesTaxom
Trixagus Kugelann, 18591SpeciesTaxom
Tylotus Schonherr, 17971SpeciesTaxom
Typocerus LeConte, 17951SpeciesTaxom
Uracanthus Hope, 18921SpeciesTaxom
Valgus Scriba, 17891SpeciesTaxom
Zenodosus Wolcott, 18351SpeciesTaxom


Species list for Raleigh County (West Virginia)
146 Species and 1 Subspecies
Agonum (Europhilus) palustre Goulet, 1969Dogtoothed goby
Agonum (Olisares) extensicolle Say, 1823
Agrilus (Agrilus) ruficollis Fabricius, 1787Red-necked Cane Borer
Agrilus syrphus Descarpentri..., 1963
Alaetrinus minimus Palisot de B..., 1805
Ampedus (Ampedus) linteus Say, 1839
Ampedus (Ampedus) protervus LeConte, 1853
Ampedus (Ampedus) semicinctus Randall, 1838
Anelaphus villosus Fabricius, 1793Oak pruner
Anisodactylus (Anadaptus) sanctaecrucis Fabricius, 1798
Anthicus (Anthicus) cervinus LaFerté-Séne..., 1848Ant-like flower beetle
Anthocomus (Celidus) equestris Fabricius, 1781Black-Banded Soft-Winged Flower Beetle
Anthrenus (Florilinus) museorum (Linnaeus, 1761)⇒ Anthrenus festivus Erichson, 1848
Anthrenus (Florilinus) verbasci Linnaeus, 1767Varied carpet beetle
Aphodius (Aphodius) pedellus (DeGeer, 1774)⇒ Aphodius fimetarius Linnaeus, 1758
Ataenius gracilis Melsheimer, 1845
Ataenius spretulus Haldeman, 1848Black turfgrass ataenius
Athous (Athous) brightwelli Kirby, 1837
Berosus (Berosus) peregrinus Herbst, 1797
Brachypterus urticae Fabricius, 1792Nettle Pollen Beetle
Brachysomida bivittata (Say, 1824)⇒ 
Brassicogethes aeneus Fabricius, 1775Blossom beetle
Buprestis (Buprestis) consularis (Gory, 1841)⇒ Buprestis nuttalli Kirby, 1837
Calligrapha (Calligrapha) philadelphica Linnaeus, 1758Dogwood Calligrapha
Calodromius sphex Andrewes, 1933
Carabus (Lichnocarabus) vinctus Weber, 1801
Cassida (Cassida) rubiginosa O.F. Müller, 1776Black crowberry
Ceruchus piceus (Weber, 1801)⇒ Ceruchus chrysomelinus Hochenwarth, 1785
Chaetocnema pulicaria F. E. Melshe..., 1880Clover Flea Beetle
Chalcophora virginiensis Drury, 1770Large flat-headed pine heartwood borer
Chilocorus stigma Say, 1835Twice-stabbed lady beetle
Chlaenius (Chlaeniellus) tricolor Dejean, 1826Prickly scorpion's-tail
Chlaenius (Chlaenius) sericeus Forster, 1771Green Pubescent Ground Beetle
Chrysochus auratus Fabricius, 1775Dogbane beetle
Cicindela (Cicindela) ancocisconensis T.W.Harris, 1852Appalachian Tiger Beetle
Cicindela (Cicindela) duodecimguttata Dejean, 1825Twelve-spot Tiger Beetle
Cicindela (Cicindela) repanda Dejean, 1825Bronze Tiger Beetle
Cicindela (Cicindela) scutellaris Say, 1823Chartreuse Tiger Beetle
Cicindela (Cicindela) sexguttata Fabricius, 1775Six-spotted Green Tiger Beetle
Cicindela (Cicindela) tranquebarica Herbst, 1806Diffracted Tiger Beetle
Cicindela pamphila (LeConte, 1873)⇒ Habroscelimorpha pamphila LeConte, 1873
Clytus ruricola Olivier, 1795Round-necked Longhorn Beetle
Coccinella (Coccinella) septempunctata Linnaeus, 17587-spot Ladybird
Coleomegilla maculata DeGeer, 1775Pink spotted lady beetle
Conotrachelus anaglypticus Say, 1831Cambium Curculio
Conotrachelus juglandis LeConte, 1876Butternut curculio
Crioceris (Crioceris) asparagi Linnaeus, 1758Asparagus beetle
Cryptolaemus montrouzieri Mulsant, 1853Mealy Bug Destroyer Larvae
Cupes capitata Fabricius, 1801
Cyclocephala borealis Arrow, 1911Northern masked chafer
Cycloneda munda Say, 1835Polished Lady Beetle
Cymbiodyta semistriata Zimmermann, 1869
Diabrotica undecimpunctata Mannerheim, 1843Arizona cottonwood
Dicerca (Dicerca) divaricata (Say, 1823)⇒ Dicerca moesta Fabricius, 1793
Dichotomius carolinus Linnaeus, 1767Carolina copris
Dineutus (Cyclinus) assimilis Kirby, 1837
Dineutus (Cyclinus) discolor Aubé, 1838
Diomus terminatus Say, 1835Firefly Duskyling
Dircaea liturata LeConte, 1866
Disonycha discoidea Fabricius, 1792Passionflower Flea Beetle
Enochrus (Methydrus) ochraceus Melsheimer, 1844
Epilachna varivestis Mulsant, 1850Ladybird Beetle
Euspilotus (Hesperosaprinus) assimilis Paykull, 1811
Fabogethes nigrescens Stephens, 1830Black Pollen Beetle
Gaurotes (Gaurotes) cyanipennis Say, 1824
Gaurotes (Gaurotes) thoracica Haldeman, 1847
Glischrochilus (Librodor) fasciatus Olivier, 1790Picnic Beetle
Glischrochilus (Librodor) sanguinolentus Olivier, 1790
Goes pulverulentus Haldeman, 1847Beech borer
Hadrobregmus notatus Say, 1825
Harmonia axyridis Pallas, 1773Asian lady beetle
Helichus basalis LeConte, 1852
Helichus lithophilus Germar, 1824
Hemicrepidius (Hemicrepidius) memnonius Herbst, 1806
Hippodamia (Hippodamia) convergens Guérin-Ménev..., 1842Convergent Lady Beetle
Hister depurator Say, 1825
Hololepta (Hololepta) aequalis Say, 1825
Hoplia (Hoplia) trifasciata Say, 1823Three lined Hoplia
Hydroporus niger (Say, 1823)⇒ Hydroporus memnonius Nicolai, 1822
Hylobius (Hylobitelus) pales Herbst, 1797Pales weevil
Hypulus simulator Newman, 1838
Ithycerus noveboracensis J.R. Forster, 1771New York weevil
Laricobius nigrinus Fender, 1945
Laricobius rubidus LeConte, 1861
Lebia (Lebia) viridis Say, 1823Flower Lebia Beetle
Leiodes impressa Baranowski, 1993Longtail tuna
Lilioceris (Lilioceris) lilii Scopoli, 1763Lily Beetle
Litargus (Alitargus) balteatus LeConte, 1856Aquatic oligochaete worm
Litomeces gracilis sphecodes Schmidt, 1922
Loricera (Loricera) pilicornis Fabricius, 1775Bigtooth aspen
Lucanus (Lucanus) cervus Linnaeus, 1758Stag beetle
Lucidota atra Olivier, 1790Black Firefly
Macronychus glabratus Say, 1825
Madarellus undulatus Say, 1824
Mecyclothorax scarites Liebherr, 2015
Melanotus (Melanotus) similis Kirby, 1837
Meloe (Meloe) americanus (Leach, 1815)⇒ Meloe impressa Kirby, 1837
Meracantha contracta Beauvois, 1805
Monochamus (Monochamus) scutellatus Say, 1824Oregon fir sawyer
Mordellina pustulata Melsheimer, 1845
Mordellistena (Mordellistena) coelioxys Lea, 1917
Mulsantina hudsonica Casey, 1899Hudsonian Lady Beetle
Mycetina perpulchra Newman, 1838
Necrophila americana Linnaeus, 1758American Carrion Beetle
Neomida bicornis Fabricius, 1777Two-Horned Darkling Beetle
Nicrophorus orbicollis Say, 1825Roundneck Sexton Beetle
Oiceoptoma inaequale Fabricius, 1781Ridged Carrion Beetle
Ontholestes cingulatus Gravenhorst, 1802Gold and Brown Rove Beetle
Optioservus (Optioservus) fastiditus LeConte, 1850Fastidious Riffle Beetle
Optioservus (Optioservus) ovalis LeConte, 1863
Pallodes pallidus Beauvois, 1805
Panscopus erinaceus Say, 1831
Paramordellaria triloba Say, 1824
Paratenetus punctatus Spinola, 1844
Philonthus (Philonthus) cyanipennis Fabricius, 1793Staph Beetle
Philonthus (Trionthus) caeruleipennis Mannerheim, 1830
Pidonia (Pidonia) ruficollis Say, 1824
Platynus (Platynus) tenuicollis LeConte, 1848
Plegaderus (Plegaderus) transversus Say, 1825
Prionocyphon discoideus Say, 1825
Propylaea quatuordecimpunctata Linnaeus, 175814-Spotted Ladybird Beetle
Pseudanostirus hamatus Say, 1834
Psyllobora vigintimaculata Say, 1824Ladybird Beetle
Ptilodactyla serricollis Say, 1823
Ptosima gibbicollis Say, 1823Redbud Borer
Sarosesthes fulminans Fabricius, 1775
Selenophorus (Selenophorus) opalinus LeConte, 1863
Stenelmis bicarinata LeConte, 1852
Stenelmis crenata Say, 1824
Stenelmis mera Sanderson, 1938Puff
Stenolophus (Agonoderus) comma Fabricius, 1775
Stenolophus (Stenolophus) ochropezus Say, 1823
Stenus cicindela Sharp, 1874
Strangalepta abbreviata Germar, 1824Strangalepta Flower Longhorn Beetle
Strangalia (Strangalina) luteicornis Fabricius, 1775Yellow-Horned Flower Longhorn Beetle
Strophiona nitens Forster, 1771Chestnut bark borer
Synaphaeta guexi LeConte, 1852Spotted tree borer
Tenomerga cinerea Say, 1831
Tetracha (Tetracha) virginica Linnaeus, 1767Virginia Big-headed Tiger Beetle
Tritoma pulchra Say, 1826Handsome Tritoma
Trixagus chevrolati Bonvouloir, 1859
Tylotus chrysops Herbst, 1797
Typocerus velutinus Olivier, 1795Banded flower longhorn beetle
Uracanthus cryptophagus Olliff, 1892Australian citrus branch borer
Valgus canaliculatus Olivier, 1789
Zenodosus sanguineus Say, 1835Blood-Colored Checkered Beetle

Subnodes of Coleoptera with occurrences in Raleigh County (West Virginia)
SuborderAdephaga
Schellenberg, 1806
Adephagans, Ground Beetles and Water Beetles, Predaceous Diving and Whirligig Beetles
4
18
25
Families
Genera
Species

Taxom

SuborderArchostemata
Kolbe, 1908
Cupedids, Ommatids, Reticulated Beetles and Telephone-pole Beetles
1
2
2
Family
Genera
Species

Taxom

SuborderPolyphaga
Emery, 1886
Polyphags
36
112
119
Families
Genera
Species

Taxom



Coleoptera (Linnaeus 1758)
Coleoptera (Linnaeus 1758)


Links, sources and further information
Coleoptera at szmn.sbras.ru
Coleoptera at papua-insects.nl
Coleoptera at artfakta.se
Coleoptera at dyntaxa.se
Coleoptera at gbif.org
Coleoptera at irmng.org
Coleoptera at itis.gov
Coleoptera at kaefer-der-welt.de
Coleoptera at marinespecies.org