Species list, Checklist of Superorder Holometabola (Holometabolous Insects) for Paleogene (Cenozoic)

Ants	Arachnids
Bees	Beetles
Booklice	Bugs
Bumblebees	Butterflies
Cicadas	Crane flies
Dragonflies	Earwigs
Flies	Isopods
Locusts	Mantises
Mayflies	Mosquitoes
Myriapods	Net-winged
Plant-parasitic	Scorpion flies
Termites	Wasps

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INSECTOIDGERENGDARK LIGHT

⌕

Checklist for Superorder Holometabola Heider, 1889 (Holometabolous Insects)
and region Paleogene (Cenozoic)

Class	Insecta Linnaeus, 1758 Insects, True insects	634 4230 9101	Families Genera Species	Taxom
Subclass	Pterygota Lang, 1888 Winged insects	631 4225 9075	Families Genera Species	Taxom
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	586 4092 8841	Families Genera Species	Taxom
Superorder	Holometabola Heider, 1889 Holometabolous Insects	372 3170 7222	Families Genera Species	Taxom

List of families for Paleogene
372 Families width 7222 Species and 36 Subspecies

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Acartophthalmidae Czerny, 1928	Acartophthalmid flies	2	Species	Taxom
Acroceridae Leach, 1815	Hunch-back flies	7	Species	Taxom
Adelidae Bruand, 1851	Adelid Moths	6	Species	Taxom
Aderidae Winkler, 1927	Ant-like leaf beetles	9	Species	Taxom
Agaonidae Walker, 1848	Fig wasp	2	Species	Taxom
Agromyzidae Fallen, 1823	Leaf miner flies	9	Species	Taxom
Agyrtidae Thomson, 1859	Primitive carrion beetles	4	Species	Taxom
Ampulicidae Shuckard, 1840	Ampulicid wasps	2	Species	Taxom
Andrenidae Webster, 1913	Andrenid bees	12	Species	Taxom
Angarosphecidae Rasnitsyn, 1975		1	Species	Taxom
Anisopodidae Knab, 1912	Window Midges	15	Species	Taxom
Anthicidae Latreille, 1819	Ant beetles	11	Species	Taxom
Anthomyiidae Robineau-Des..., 1830	Anthomyiid flies	9	Species	Taxom
Anthomyzidae Czerny, 1903	Anthomyzid flies	12	Species	Taxom
Anthribidae Billberg, 1820	Fungus Weevil	37	Species	Taxom
Apataniidae Wallengren, 1886	Early Smoky Wing Sedges	1	Species	Taxom
Aphelinidae Thomson, 1876	Aphelinid wasps	3	Species	Taxom
Apidae Latreille, 1802	Bumble bees	65	Species	Taxom
Arctiidae Leach, 1815	Footman moths	1	Species	Taxom
Argidae Konow, 1890	Argid flies	2	Species	Taxom
Artematopodidae Lacordaire, 1857	Soft bodied plant beetle	6	Species	Taxom
Asilidae Latreille, 1802	Assassin flies	58	Species	Taxom
Asteiidae Rondani, 1856	Asteiid Flies	1	Species	Taxom
Atelestidae Hennig, 1970		1	Species	Taxom
Athericidae Stuckenberg, 1973	Athericid flies	6	Species	Taxom
Attelabidae Billberg, 1820	Leaf Rolling Weevil	69	Species	Taxom
Aulacidae Shuckard, 1842	Aulacid wasps	11	Species	Taxom
Aulacigastridae Wollaston, 1864	Aulacigastrid flies	1	Species	Taxom
Autostichidae Le Marchand, 1947	Autostichid Moths	2	Species	Taxom
Belidae Schönherr, 1826	Belid Beetle	12	Species	Taxom
Beraeidae Wallengren, 1891	Small eastern caddisflies	3	Species	Taxom
Berendtimiridae Winkler, 1987		1	Species	Taxom
Bethylidae Halliday, 1839	Bethylid wasps	51	Species	Taxom
Bibionidae Fleming, 1821	Bibionid flies	245	Species	Taxom
Biphyllidae LeConte, 1861	Biphyllid beetles	1	Species	Taxom
Bittacidae Handlirsch, 1906	Hanging scorpionflies	11	Species	Taxom
Blasticotomidae Thomson, 1871	Blasticotomid sawflies	1	Species	Taxom
Blephariceridae Loew, 1861	Mountain midges	5	Species	Taxom
Bohartillidae Kinzelbach, 1969		1	Species	Taxom
Boleopsidae Kirejtshuk &..., 2013		1	Species	Taxom
Bombyliidae Latreille, 1802	Bee Flies	58	Species	Taxom
Bostrichidae Latreille, 1802	Auger beetles	6	Species	Taxom
Bothrideridae Erichson, 1845	Cocoon-forming beetles	4	Species	Taxom
Brachycentridae Ulmer, 1903	Humpless Casemaker Caddisflies	1	Species	Taxom
Brachyceritidae Hong, 2002		1	Species	Taxom
Brachypteridae (Erichson, 1845)	Short-winged Flower Beetles	1	Species	Taxom
Brachystomatidae Melander, 1908		2	Species	Taxom
Braconidae Latreille, 1829	Braconid wasps	230	Species	Taxom
Brentidae Billberg, 1820	Brenthidae	33	Species	Taxom
Buprestidae Leach, 1815	Flat-headed wood-borers	133	Species	Taxom
Byrrhidae Latreille, 1804	Moss beetles	6	Species	Taxom
Calamoceratidae Ulmer, 1905		3	Species	Taxom
Calliphoridae Brauer & Ber..., 1889	Blow flies	1	Species	Taxom
Camillidae Frey, 1921	Camillid flies	2	Species	Taxom
Cantharidae Imhoff, 1856	Cantharids	41	Species	Taxom
Carabidae Latreille, 1802	Carabid beetles	153	Species	Taxom
Caridae Thompson, 1992		5	Species	Taxom
Carnidae Newman, 1834	Bird flies	1	Species	Taxom
Castniidae Boisduval, 1828	Castniid moths	1	Species	Taxom
Cecidomyiidae Newman, 1834	Forest Midges	144	Species	Taxom
Cephidae Newman, 1834	Cephids	5	Species	Taxom
Cerambycidae Latreille, 1802	Long Horn beetles	89	Species	Taxom
Ceratopogonidae Newman, 1834	Biting midge	137	Species	Taxom
Chalcididae Latreille, 1817	Chalcidid wasps	7	Species	Taxom
Chamaemyiidae Hendel, 1910	Aphid flies	1	Species	Taxom
Chaoboridae Newman, 1834	Phantom midge	16	Species	Taxom
Chelonariidae Blanchard, 1845	Chelonarid beetles	1	Species	Taxom
Chimeromyiidae Grimaldi & C..., 2009		1	Species	Taxom
Chironomidae Erichson, 1842	Chironomids	204	Species	Taxom
Chloropidae Schoenherr, 1840	Chloropid flies	6	Species	Taxom
Choristidae Esben-Petersen, 1915		1	Species	Taxom
Chrysididae Latreille, 1802	Chrysidid wasps	9	Species	Taxom
Chrysomelidae Latreille, 1802	Casebearers	190	Species	Taxom
Chyromyidae Hendel, 1916	Chiromyid flies	1	Species	Taxom
Ciidae Leach, 1819	Minute tree fungus beetles	1	Species	Taxom
Cimbicidae Kirby, 1837	Cimbicid sawflies	5	Species	Taxom
Cimbrophlebiidae Willmann, 1977	Hangingfly	5	Species	Taxom
Cleridae Latreille, 1802	Checkered Beetle	26	Species	Taxom
Clusiidae Schönherr, 1840	Clusiid flies	4	Species	Taxom
Coccinellidae Latreille, 1807	Coccinelles	21	Species	Taxom
Coleophoridae Hübner, 1825	Case moths	2	Species	Taxom
Conopidae Latreille, 1802	Thick-headed Flies	2	Species	Taxom
Copromorphidae Meyrick, 1905	Fig Borers	1	Species	Taxom
Coptoclavidae Ponomarenko, 1961	Extinct family	1	Species	Taxom
Corioxenidae Kinzelbach, 1970		1	Species	Taxom
Corylophidae LeConte, 1852	Hooded beetles	1	Species	Taxom
Cossidae Leach, 1815	Borers	5	Species	Taxom
Crabronidae Latreille, 1802	Bee Wolves	42	Species	Taxom
Cryptochetidae Brues & Mela..., 1932	Cryptochetid Flies	1	Species	Taxom
Cryptophagidae Kirby, 1937	Mould beetles	11	Species	Taxom
Ctenophthalmidae Rothschild, 1915		4	Species	Taxom
Cucujidae Latreille, 1802	Flat bark beetles	10	Species	Taxom
Culicidae Meigen, 1818	Mosquito	21	Species	Taxom
Cupedidae Laporte, 1836	Cupedids	15	Species	Taxom
Curculionidae Latreille, 1802	Ambrosia beetles	452	Species	Taxom
Curtonotidae Duda, 1934	Curtonotid flies	1	Species	Taxom
Cylindrotomidae (Mulsant, 1842)	Cylindrotomid Crane Flies	9	Species	Taxom
Cynipidae Latreille, 1802	Cynipid gall wasps	12	Species	Taxom
Cypselosomatidae Hennig, 1958	Cypselosomatid flies	1	Species	Taxom
Dascillidae Guérin-Ménev..., 1843	Orchid beetles	2	Species	Taxom
Dermestidae Hope, 1840	Carpet beetles	30	Species	Taxom
Derodontidae LeConte, 1861	Tooth-neck fungus beetles	1	Species	Taxom
Diapriidae Latreille, 1796	Diapriid wasps	20	Species	Taxom
Diastatidae Hendel, 1917	Bog flies	4	Species	Taxom
Dinopanorpidae Carpenter, 1972		7	Species	Taxom
Diopsidae Billberg, 1820	Stalk-eyed flies	3	Species	Taxom
Diprionidae Rohwer, 1911	Conifer sawflies	2	Species	Taxom
Dipseudopsidae Ulmer, 1904		4	Species	Taxom
Ditomyiidae Keilin, 1919		1	Species	Taxom
Dixidae Schiner, 1868	Dixid Midges	12	Species	Taxom
Dolichopodidae Latreille, 1809	Long legged flies	135	Species	Taxom
Drosophilidae Rondani, 1856	Fruit flies	3	Species	Taxom
Dryinidae Haliday, 1840	Dryinid wasps	21	Species	Taxom
Dryomyzidae Schiner, 1862	Dryomyzid flies	3	Species	Taxom
Dryophthoridae (Schönherr, 1825)	Palm weevils	1	Species	Taxom
Dryopidae Billberg, 1820	Long toed water beetle	7	Species	Taxom
Dytiscidae Leach, 1815	Diving beetles	34	Species	Taxom
Ecnomidae Ulmer, 1903		20	Species	Taxom
Elachistidae Bruand, 1851	Elachistid moths	12	Species	Taxom
Elateridae Leach, 1815	Click Beetle	103	Species	Taxom
Electrotomidae Rasnitsyn, 1977		1	Species	Taxom
Elmidae Curtis, 1830	Drive beetles	4	Species	Taxom
Embolemidae Förster, 1856	Embolemid wasps	3	Species	Taxom
Empididae Latreille, 1809	Balloon flies	95	Species	Taxom
Encyrtidae Walker, 1837	Encyrtid wasps	11	Species	Taxom
Endomychidae Leach, 1815	False Ladybirds	3	Species	Taxom
Eomeropidae Cockerell, 1909		4	Species	Taxom
Eophlebomyiidae Cockerell, 1925		1	Species	Taxom
Eorpidae Archibald & al., 2013		3	Species	Taxom
Eostephanitidae Hong, 2002		1	Species	Taxom
Ephydridae Zetterstedt, 1837	Brine flies	8	Species	Taxom
Erebidae Leach, 1815	Erebid Moths	4	Species	Taxom
Erotylidae Latreille, 1802	Pleasing Fungus Beetle	11	Species	Taxom
Eucharitidae Latreille, 1809	Eucharitid wasp	1	Species	Taxom
Eucnemidae Eschscholtz, 1829	Click Beetles	20	Species	Taxom
Eulophidae Westwood, 1829	Eulophid wasps	6	Species	Taxom
Eupelmidae Walker, 1833	Eupelmid wasps	6	Species	Taxom
Eurytomidae Walker, 1832	Eurytomid wasps	8	Species	Taxom
Evaniidae Latreille, 1802	Ensign wasp	6	Species	Taxom
Expansicornidae Hong, 2002		1	Species	Taxom
Figitidae Thomson, 1862	Figitid wasps	4	Species	Taxom
Formicidae Latreille, 1809	Ants	545	Species	Taxom
Fungivoritidae Rohdendorf, 1957		1	Species	Taxom
Geometridae Leach, 1815	Cankerworm moths	3	Species	Taxom
Geotrupidae Latreille, 1802	Dor beetles	5	Species	Taxom
Glaphyridae MacLeay, 1819	Bumble bee scarab beetle	2	Species	Taxom
Glaphyropteridae Pongracz, 1935		14	Species	Taxom
Glossinidae Theobald, 1903	Tik-tik flies	4	Species	Taxom
Glossosomatidae Wallengren, 1891	Little Black Caddisflies	4	Species	Taxom
Goeridae Ulmer, 1903		4	Species	Taxom
Gracillariidae Stainton, 1854	Blotch leaf miner moths	2	Species	Taxom
Gyrinidae Latreille, 1802	Whirligig beetle	5	Species	Taxom
Halictidae Thomson, 1869	Flower bees	10	Species	Taxom
Heleomyzidae (Westwood, 1840)	Heleomyzid flies	18	Species	Taxom
Helicopsychidae Ulmer, 1912	Snail Case caddisflies	12	Species	Taxom
Heloridae Förster, 1856	Helorid wasps	1	Species	Taxom
Helotidae Reitter, 1876		1	Species	Taxom
Hepialidae Stephens, 1829	Ghost Moths	1	Species	Taxom
Hesperiidae Latreille, 1809	Skipper	7	Species	Taxom
Hesperinidae Hendel, 1928		4	Species	Taxom
Hippoboscidae Samouelle, 1819	Keds	3	Species	Taxom
Histeridae Gyllenhal, 1808	Clown beetle	5	Species	Taxom
Hoffeinsmyiidae Michelsen, 2009		1	Species	Taxom
Huaxiasciaritidae (Hong, 2002)		26	Species	Taxom
Hybosoridae Erichson, 1847	Pill scarab beetles	1	Species	Taxom
Hybotidae Meigen, 1820	Dance flies	25	Species	Taxom
Hydraenidae Mulsant, 1844	Cascade Beetles	2	Species	Taxom
Hydrobiosidae Ulmer, 1905		6	Species	Taxom
Hydrophilidae Latreille, 1802	Silver Water Beetles	49	Species	Taxom
Hydropsychidae Curtis, 1835	Net Spinning caddisflies	11	Species	Taxom
Hydroptilidae Stephens, 1836	Micro caddisflies	52	Species	Taxom
Hygrobiidae Régimbart, 1878	Screech beetles	1	Species	Taxom
Ibaliidae Thomson, 1862	Ibaliid wasps	3	Species	Taxom
Ichneumonidae Latreille, 1802	Ichneumon flies	189	Species	Taxom
Incurvariidae Spuler, 1898	Fairy moths	1	Species	Taxom
Ithyceridae (Schönherr, 1826)	New York weevil	1	Species	Taxom
Jacobsoniidae Heller, 1926	Jacobson's beetles	1	Species	Taxom
Kalophryganeidae Haupt, 1956		1	Species	Taxom
Kateretidae Kirkby, 1837	Pollen Beetles	2	Species	Taxom
Keroplatidae Rondani, 1856	Fungus gnats	40	Species	Taxom
Lampyridae Latreille, 1817	Fireflies	6	Species	Taxom
Latridiidae Erichson, 1842	Minute brown scavenger beetle	23	Species	Taxom
Lauxaniidae Macquart, 1835	Beach flies	4	Species	Taxom
Leiodidae Fleming, 1821	Round fungus beetles	11	Species	Taxom
Lepidostomatidae Ulmer, 1903	Bizarre Caddisflies	16	Species	Taxom
Leptoceridae Leach, 1815	Long Horned caddisflies	25	Species	Taxom
Limnephilidae Kolenati, 1848	Northern caddisflies	25	Species	Taxom
Limnichidae Erichson, 1846	Minute marsh loving beetle	1	Species	Taxom
Limoniidae (Speiser, 1909)	Craneflies	42	Species	Taxom
Lonchaeidae Loew, 1861	Lance Flies	2	Species	Taxom
Lucanidae Latreille, 1804	Stag Beetle	10	Species	Taxom
Lycaenidae Leach, 1815	Blue butterflies	4	Species	Taxom
Lycidae Laporte, 1836	Lycid beetles	5	Species	Taxom
Lygistorrhinidae Hendel, 1936	Long beaked fungus gnat	1	Species	Taxom
Lymexylidae Fleming, 1821	Ship-timber beetle	3	Species	Taxom
Lyonetiidae Stainton, 1854	Lyonet Moths	2	Species	Taxom
Megachilidae Schenk, 1860	And Resin Bees	25	Species	Taxom
Megalyridae Schletterer, 1889	Megalyrid wasps	8	Species	Taxom
Megamerinidae Hendel, 1913	Megamerinid flies	1	Species	Taxom
Megaspilidae Ashmead, 1893	Megaspilid wasps	5	Species	Taxom
Melandryidae Leach, 1815	False Darkling Beetle	15	Species	Taxom
Melittidae Schenk, 1860	Melittid bees	6	Species	Taxom
Meloidae Gyllenhal, 1810	Blister Beetle	8	Species	Taxom
Melyridae Leach, 1815	Malachite Beetles	15	Species	Taxom
Mengeidae Pierce, 1908		2	Species	Taxom
Mesokristenseniidae Huang, Nel &..., 2010		1	Species	Taxom
Micromalthidae Barber, 1913	Micromalthid beetles	2	Species	Taxom
Micropezidae Illiger, 1798	Stilt Flies	4	Species	Taxom
Micropterigidae Herrich-Schä..., 1855	Mandibulate Archaic Moths	6	Species	Taxom
Milichiidae Schiner, 1862	Filth flies	2	Species	Taxom
Molannidae Wallengren, 1891	Hood Casemaker Caddisflies	2	Species	Taxom
Monotomidae Laporte, 1840	Minute Clubbed Beetles	2	Species	Taxom
Mordellidae Latreille, 1802	Pintail beetles	27	Species	Taxom
Muscidae Latreille, 1802	Face flies	7	Species	Taxom
Mutillidae André, 1907	Cow ant	2	Species	Taxom
Mycetophagidae Leach, 1815	Hairy fungus beetle	4	Species	Taxom
Mycetophilidae Newman, 1834	Fungus Gnat	358	Species	Taxom
Mycteridae Blanchard, 1845	Mycterid beetles	3	Species	Taxom
Mydidae Bezzi, 1903	Mydas flies	1	Species	Taxom
Mymaridae Haliday, 1833	Fairflies	22	Species	Taxom
Mymarommatidae Debauche, 1948	False fairy wasp	2	Species	Taxom
Myrmecolacidae Saunders, 1872		15	Species	Taxom
Myrmosidae Fox, 1894	Myrmosid Wasps	8	Species	Taxom
Mythicomyiidae Melander, 1902	Micro Bee Flies	7	Species	Taxom
Nemestrinidae Macquart, 1834	Tangle-veined flies	5	Species	Taxom
Nemonychidae Bedel, 1882	Nemonychid weevils	2	Species	Taxom
Nepticulidae Stainton, 1854	Leaf miners	3	Species	Taxom
Neurochaetidae Mcalpine, 1978	Upside-Down Flies	2	Species	Taxom
Nitidulidae Latreille, 1802	Nitidules	35	Species	Taxom
Noctuidae Latreille, 1809	Armyworms	13	Species	Taxom
Nosodendridae Erichson, 1846	Nosodendrid beetles	4	Species	Taxom
Notodontidae Stephens, 1829	Bag Shelter Moths	2	Species	Taxom
Nymphalidae Rafinesque, 1815	Admirals	33	Species	Taxom
Nymphomyiidae Tokunaga, 1932	Nymphomyiid crane flies	1	Species	Taxom
Ochodaeidae Mulsant & Rey, 1870	Sand-loving scarab beetle	1	Species	Taxom
Odiniidae Hendel, 1920	Odiniid flies	2	Species	Taxom
Odontoceridae Wallengren, 1891	Mortarjoint Casemaker Caddisflies	7	Species	Taxom
Oecophoridae Bruand, 1851	Concealer Moths	16	Species	Taxom
Oedemeridae Latreille, 1810	False Blister Beetle	4	Species	Taxom
Oestridae Leach, 1815	Bot flies	4	Species	Taxom
Omalisidae Lacordaire, 1857		1	Species	Taxom
Omethidae LeConte, 1861	False fireflies	1	Species	Taxom
Ommatidae Sharp & Muir, 1912		1	Species	Taxom
Opetiidae Rondani, 1856	Flat-footed flies	1	Species	Taxom
Opomyzidae (Fallen, 1820)	Opomyzid flies	2	Species	Taxom
Orussidae Newman, 1834	Orussid wasps	1	Species	Taxom
Ostomatidae Reitter, 1882	Bark-gnawing beetles	1	Species	Taxom
Otitidae (Macquart, 1835)	Picture-winged flies	3	Species	Taxom
Paleomelittidae Engel, 2001		1	Species	Taxom
Pallopteridae Loew, 1862	Flutter flies	3	Species	Taxom
Pamphiliidae Cameron, 1890	Leaf-rolling sawflies	4	Species	Taxom
Panorpidae Linnaeus, 1758	Common scorpionflies	11	Species	Taxom
Panorpodidae Byers, 1965	Short-faced scorpionflies	5	Species	Taxom
Papilionidae Latreille, 1802	Apollos	9	Species	Taxom
Passalidae Leach, 1815	Bess beetle	1	Species	Taxom
Pediciidae Osten Sacken, 1860	Hairy-Eyed Crane Flies	3	Species	Taxom
Pelecinidae Haliday, 1839	Ancestral And Modern Pelecinid Wasps	2	Species	Taxom
Peradeniidae Naumann & Ma..., 1985		1	Species	Taxom
Pergidae Rohwer, 1911	Pergid flies	1	Species	Taxom
Perilampidae Latreille, 1809	Perilampid waps	2	Species	Taxom
Periscelididae Oldenberg, 1914	Periscelidid Flies	2	Species	Taxom
Phaeomyiidae (Verbeke, 1950)	Marsh flies	1	Species	Taxom
Phalacridae Leach, 1815	Shining flower beetles	3	Species	Taxom
Philopotamidae Stephens, 1829	Finger Net caddisflies	17	Species	Taxom
Phoridae Curtis, 1833	Coffin Flies	80	Species	Taxom
Phryganeidae (Leach, 1815)	Giant casemaker	23	Species	Taxom
Pieridae Duponchel, 1835	Brimstones	6	Species	Taxom
Piophilidae Macquart, 1835	Cheese flies	1	Species	Taxom
Pipunculidae Walker, 1834	Big headed flies	12	Species	Taxom
Platygastridae Haliday, 1833	Platygastrid wasps	7	Species	Taxom
Platypezidae Fallen, 1815	Flat footed flies	6	Species	Taxom
Plutellidae Guenée, 1845	Diamond-back moths	4	Species	Taxom
Polycentropodidae Ulmer, 1903	Tube Maker Caddisflies	80	Species	Taxom
Pompilidae Latreille, 1805	Pompilid wasps	19	Species	Taxom
Prionoceridae Lacaordaire, 1857		1	Species	Taxom
Proctotrupidae Latreille, 1802	Proctotrupid wasps	14	Species	Taxom
Proneottiophilidae Hennig, 1969		1	Species	Taxom
Proterhinidae (Fauvel, 1891)		2	Species	Taxom
Protoxenidae Pohl, Beutel..., 2005		1	Species	Taxom
Psephenidae Lacordaire, 1854	Water pennies	3	Species	Taxom
Pseudopomyzidae (Mcalpine, 1966)	Banana flies	2	Species	Taxom
Pseudostenolestidae Garrouste & Nel, 2015		1	Species	Taxom
Psilidae Macquart, 1835	Carrot flies	1	Species	Taxom
Psychidae Boisduval, 1828	Bagmoths	4	Species	Taxom
Psychodidae Newman, 1834	Drain flies	37	Species	Taxom
Psychomyiidae Walker, 1852	Net Tube Caddisflies	9	Species	Taxom
Pteromalidae Dalman, 1820	Pteromalid wasps	26	Species	Taxom
Pterophoridae Zeller, 1841	Plume Moths	1	Species	Taxom
Ptiliidae Heer, 1843	Feather-wing beetles	8	Species	Taxom
Ptilodactylidae Laporte, 1836	Ptilodactylid beetles	2	Species	Taxom
Ptinidae Latreille, 1802	Death-watch and Spider Beetles	49	Species	Taxom
Ptychopteridae Osten Sacken, 1862	Phantom crane flies	6	Species	Taxom
Pyralidae Latreille, 1802	Flour Moths	4	Species	Taxom
Pyrochroidae Latreille, 1807	Cardinal beetles	2	Species	Taxom
Pythidae Solier, 1834	Dead log bark beetles	1	Species	Taxom
Rangomaramidae Jaschhof & D..., 2002		3	Species	Taxom
Rhagionidae Latreille, 1802	Snipe flies	26	Species	Taxom
Rhipiphoridae Gemminger & ..., 1870	Hump back beetles	3	Species	Taxom
Rhopalosomatidae Ashmead, 1896	Rhopalosomatid wasps	2	Species	Taxom
Rhynchitidae Gistel, 1848	Leaf-rolling weevils	3	Species	Taxom
Richardiidae (Snow, 1896)	Picture-winged flies	3	Species	Taxom
Riodinidae Grote, 1895	Metalmark Butterflies	5	Species	Taxom
Ripiphoridae Gemminger & ..., 1870	Rhipiphoridae	3	Species	Taxom
Roproniidae Bradley, 1905	Roproniid wasps	1	Species	Taxom
Salpingidae Leach, 1815	Narrow-waisted bark beetle	3	Species	Taxom
Saturniidae Boisduval, 1837	And Giant Silk Moths	1	Species	Taxom
Scarabaeidae Latreille, 1802	Dung beetles	65	Species	Taxom
Scathophagidae Robineau-Des..., 1830	Dung flies	4	Species	Taxom
Scatopsidae Newman, 1834	Dung midges	9	Species	Taxom
Scelionidae Haliday, 1839	Scelionid wasps	54	Species	Taxom
Scenopinidae Burmeister, 1835	Window Flies	1	Species	Taxom
Schizopodidae (LeConte, 1859)	False Jewel Beetles	1	Species	Taxom
Sciaridae Billberg, 1820	Black fungus gnats	6	Species	Taxom
Sciomyzidae Fallen, 1820	Marsh Flies	13	Species	Taxom
Scirtidae Fleming, 1821	Marsh Beetle	33	Species	Taxom
Scolebythidae Evans, 1963	Scolebythid wasps	5	Species	Taxom
Scoliidae Leach, 1815	Mammoth wasps	2	Species	Taxom
Scraptiidae Mulsant, 1856	Antlike leaf beetles	9	Species	Taxom
Scydmaenidae (Leach, 1815)	Ant-like stone beetles	21	Species	Taxom
Sepsidae Walker, 1833	Black scavenger flies	5	Species	Taxom
Sericostomatidae Stephens, 1836	Bushtailed Caddisflies	6	Species	Taxom
Sesiidae Boisduval, 1828	Clear Wings	2	Species	Taxom
Silphidae Latreille, 1807	Burying And Carrion Beetles	7	Species	Taxom
Silvanidae Kirby, 1837	Flat bark beetles	3	Species	Taxom
Simuliidae Newman, 1834	Black flies	13	Species	Taxom
Sinisilvanidae (Hong, 2002)		1	Species	Taxom
Siricidae Billberg, 1820	Horntail	5	Species	Taxom
Smicripidae Horn, 1879	Palmetto beetle	2	Species	Taxom
Sphaeroceridae Macquart, 1835	Dung flies	1	Species	Taxom
Sphecidae Latreille, 1802	Digger & Sand wasps	10	Species	Taxom
Sphingidae Latreille, 1802	Hawk Moths	1	Species	Taxom
Staphylinidae Lameere, 1900	Rove beetles	206	Species	Taxom
Stenopsychidae Martynov, 1924		1	Species	Taxom
Stephanidae Leach, 1815	Crown wasp	7	Species	Taxom
Stratiomyidae Latreille, 1802	Soldier flies	19	Species	Taxom
Stylopidae Kirby, 1813	Strepsiptera	1	Species	Taxom
Syrphidae Latreille, 1802	Drone flies	94	Species	Taxom
Tabanidae Latreille, 1802	Breeze flies	17	Species	Taxom
Tachinidae (Robineau-Des..., 1830)	Bristle Flies	4	Species	Taxom
Tanaostigmatidae Howard, 1890	Tanaostigmatid wasps	1	Species	Taxom
Tanyderidae Alexander, 1919	Primitive crane flies	3	Species	Taxom
Tenebrionidae Latreille, 1802	Darkling Beetle	59	Species	Taxom
Tenthredinidae Latreille, 1804	Common sawflies	52	Species	Taxom
Tephritidae Newman, 1834	Fruit flies	1	Species	Taxom
Tetracampidae Förster, 1856	Tetracampid wasps	2	Species	Taxom
Tetratomidae Billberg, 1820	Polypore fungus beetle	1	Species	Taxom
Therevidae Newman, 1834	Stiletto flies	9	Species	Taxom
Throscidae Laporte, 1840	False metallic wood-boring beetles	12	Species	Taxom
Tineidae Latreille, 1810	& Allies	21	Species	Taxom
Tiphiidae Leach, 1815	Flower wasps	8	Species	Taxom
Tipulidae Latreille, 1802	Crane Flies	326	Species	Taxom
Tortricidae Latreille, 1803	Bell Moths	8	Species	Taxom
Torymidae Walker, 1833	Apple seed chalcids	12	Species	Taxom
Trachypachidae C. G. Thomson, 1857	False ground beetle	1	Species	Taxom
Trichoceridae Rondani, 1841	Winter Crane Flies	17	Species	Taxom
Trichogrammatidae Haliday, 1851	Trichogrammatid wasps	4	Species	Taxom
Trigonalidae (Cresson, 1887)	Trigonalid sawflies	1	Species	Taxom
Trigonalyidae		1	Species	Taxom
Tritarsidae Hong, 2002		1	Species	Taxom
Trogidae (MacLeay, 1819)	Gnawer beetles	2	Species	Taxom
Trogossitidae Latreille, 1802	Bark gnawing beetle	14	Species	Taxom
Tshekardocoleidae Rohdendorf, 1944		3	Species	Taxom
Uraniidae Blanchard, 1845	Swallowtail Moths	1	Species	Taxom
Urolibellulidae Zeiri & al., 2015		1	Species	Taxom
Vespidae Leach, 1815	And Allies	35	Species	Taxom
Xyelidae Newman, 1835	Sawfly	9	Species	Taxom
Xylomyidae Verrall, 1901	Wood soldier flies	3	Species	Taxom
Xylophagidae Fallen, 1810	Awl-flies	8	Species	Taxom
Yantarocentridae Ivanov & Mel..., 2015		1	Species	Taxom
Yponomeutidae Stephens, 1829	Ermine Moths	6	Species	Taxom
Zopheridae Solier, 1834	Cylindrical bark beetles	11	Species	Taxom
Zygaenidae Latreille, 1809	Burnet Forester	2	Species	Taxom

Subnodes of Holometabola with occurrences in Paleogene

Order	Coleoptera Linnaeus, 1758 Beetles, Coleopterans	111 1198 2377	Families Genera Species	Taxom
Order	Diptera Linnaeus, 1758 Flies, Gnats, hoverflies, Mosquitoes, True flies, Two-Winged Flies	106 904 2561	Families Genera Species	Taxom
Order	Hymenoptera Linnaeus, 1758 Ants, Bees, Sawflies, Wasps	74 778 1667	Families Genera Species	Taxom
Order	Lepidoptera Linnaeus, 1758 Butterflies, Lepidopterans, Moths	38 162 203	Families Genera Species	Taxom
Order	Mecoptera Hyatt & Arms, 1891 Hanging flies, Hangingflies, Scorpion flies, Scorpionflies, snow scorpionflies, Snowflies	9 17 51	Families Genera Species	Taxom
Order	Strepsiptera Kirby, 1813 Parasitic insects, Strepsipterans, Stylops, Twisted-wing insects, Twisted-wing parasites, Twisted-winged Insects, twisted-winged parasite, Twisted-winged parasites	6 10 23	Families Genera Species	Taxom
Order	Trichoptera Kirby, 1813 Caddis-Flies, Caddisflies, Caddisfly, Cased Caddis, Cased Caddis-Flies, Caseless Caddis, Caseless Caddis-Flies, Rail-flies, Sedge-flies	28 101 340	Families Genera Species	Taxom

Links, sources and further information

Holometabola at itis.gov