Taxonomy of Genus Appias Hübner, 1819
Albatrosses, Puffins, Puffins and Albatrosses, Whites
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 4179
190570
2444470 | Families
Genera
Species |
Subclass
| Pterygota
Lang, 1888
Winged insects | 3816
189502
2439455 | Families
Genera
Species |
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 3399
186457
2415487 | Families
Genera
Species |
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 1891
142849
2048615 | Families
Genera
Species |
Order
| Lepidoptera
Linnaeus, 1758
Butterflies, Lepidopterans, Moths | 322
33641
568299 | Families
Genera
Species |
Suborder
| Glossata
Fabricius, 1775
Modern butterflies and moths | 208
32654
567261 | Families
Genera
Species |
Infraorder
| Heteroneura
Tillyard, 1918
| 197
32461
565431 | Families
Genera
Species | | Division | Ditrysia
Borner, 1925
Butterflies, Moths, Skippers | 186
32290
560864 | Families
Genera
Species |
Section
| Cossina
Herrich-Schäffer, 1856
| 74
20388
404639 | Families
Genera
Species |
Subsection
| Bombyces
Scopoli, 1763
| 66
18154
371101 | Families
Genera
Species |
Superfamily
| Papilionoidea
Latreille, 1802
Butterflies, Skippers, True butterflies | 13
3266
144998 | Families
Genera
Species | | Family | Pieridae
Duponchel, 1835
And Sulphurs, Brimstones, Orange-tips, Orangetips, Sulfurs | 230
13433 | Genera
Species | | Subfamily | Pierinae
Duponchel, 1835
And orangetips, Marbles, Marbles and Orangetips, Whites | 86
8651 | Genera
Species | | Tribe | Pierini
Swainson, 1820
Albatrosses, Cabbage Whites, Checkered Whites | 51
6623 | Genera
Species | | Subtribe | Appiadina
Kuznetsov, 1929
| 6
851 | Genera
Species | | Genus | Appias
Hübner, 1819
Albatrosses, Puffins, Puffins and Albatrosses, Whites | 713 | Species |
Subnodes of Appias Hübner, 1819 | Subgenus | Appias (Appias)
Hübner, 1819
Albatrosses, Puffins, Puffins and Albatrosses, Whites | 125 | Species | | Subgenus | Appias (Catophaga)
Hübner, 1819 ⇒ Tachyris
| 155 | Species | | Subgenus | Appias (Glutophrissa)
Butler, 1887
Albatrosses, Puffins, Whites | 49 | Species | | Subgenus | Appias (Hiposcritia)
Geyer, 1832 ⇒ Tachyris
| 9 | Species | | Subgenus | Appias (Phrissura)
Butler, 1870
| 10 | Species | | Subgenus | Appias (Tachyris)
Wallace, 1867 ⇒ Tachyris
| 1 | Species | | Species | Appias absyrtus Stoneham, 1957
| | | | Species | Appias abyssinica Talbot, 1932
| | | | Species | Appias acuminata Snellen, 1890
| | | | Species | Appias adamsoni Moore, 1905
| | | | Species | Appias adelpha Röber, 1892
| | | | Species | Appias adorabilis Fruhstorfer, 1910
| | | | Species | Appias aegina Fruhstorfer, 1910
| | | | Species | Appias aemilia Fruhstorfer, 1910
| | | | Species | Appias agar Fruhstorfer, 1910
| | | | Species | Appias agatha Staudinger, 1889
| | | | Species | Appias agave Felder, 1862
| | | | Species | Appias aglaia Stoneham, 1957
| | | | Species | Appias albata Hopffer, 1874
| | | | Species | Appias albida Mabille, 1887
| | | | Species | Appias alcesta Talbot, 1939
| | | | Species | Appias alope Wallace, 1867
| | | | Species | Appias amasene Boisduval, 1836
| | | | Species | Appias ambigua Grose-Smith, 1895
| | | | Species | Appias andrea Bingham, 1907
| | | | Species | Appias antoniae Fruhstorfer, 1910
| | | | Species | Appias aornus Fruhstorfer, 1913
| | | | Species | Appias aperta Butler, 1886
| | | | Species | Appias arctimargo Hulstaert, 1924
| | | | Species | Appias ardens Butler, 1898
| | | | Species | Appias ares Swinhoe, 1885
| | | | Species | Appias argentifera Joicey & Talbot, 1928
| | | | Species | Appias argyridina Butler, 1885
| | | | Species | Appias arida Talbot, 1939
| | | | Species | Appias aristotemus
| | | | Species | Appias aristoxemus Fruhstorfer, 1908
| | | | Species | Appias aroa RIBBE, 1900
| | | | Species | Appias asteria Miskin, 1889
| | | | Species | Appias asterope Felder, 1862
| | | | Species | Appias aternia Fruhstorfer, 1910
| | | | Species | Appias athama Blanchard, 1848
Pacific albatross | 3 | Subspecies | | Species | Appias athena Fruhstorfer, 1902
| | | | Species | Appias aufidia Fruhstorfer, 1910
| | | | Species | Appias aurifera Fruhstorfer, 1910
| | | | Species | Appias auriflua Talbot, 1943
| | | | Species | Appias bachi Fabricius, 1775 ⇒ Cepora kotakii
| 1 | Subspecies | | Species | Appias balina Fruhstorfer, 1908
| | | | Species | Appias bandana Fruhstorfer, 1901
| | | | Species | Appias barea Fruhstorfer, 1910
| | | | Species | Appias baweanicus Fruhstorfer, 1906
| | | | Species | Appias bicolor Talbot, 1934
| | | | Species | Appias bouruensis Wallace, 1867
| | | | Species | Appias caledonica Felder, 1862
| | | | Species | Appias cana Fruhstorfer, 1910
| | | | Species | Appias canisia Hulstaert, 1924
| | | | Species | Appias caria Waterhouse & Lyell, 1914
| | | | Species | Appias cerussa Fruhstorfer, 1904
| | | | Species | Appias chelidon Fruhstorfer, 1905
| | | | Species | Appias chorasina Fruhstorfer, 1921
| | | | Species | Appias cilla Felder, 1865
| | | | Species | Appias citrina Fruhstorfer, 1910
| | | | Species | Appias citrinus Bergh, 1928
| | | | Species | Appias citronella Fruhstorfer, 1896
| | | | Species | Appias clavis Wallace, 1867
| | | | Species | Appias cneora Fabricius, 1793
| | | | Species | Appias coelita Fruhstorfer, 1899
| | | | Species | Appias colimba Moore, 1865
| | | | Species | Appias comorensis Talbot, 1943
| | | | Species | Appias confluens Hulstaert, 1924
| | | | Species | Appias confusa Fruhstorfer, 1910
| | | | Species | Appias coniata Butler, 1880
| | | | Species | Appias contracta Butler, 1888
| | | | Species | Appias cynisca Wallace, 1867
| | | | Species | Appias darada Felder, 1865
| | | | Species | Appias decolorata Hulstaert, 1924
| | | | Species | Appias defecta Gaede, 1916
| | | | Species | Appias deficiens Dufrane, 1947
| | | | Species | Appias delicata Butler, 1882
| | | | Species | Appias dido Talbot, 1943
| | | | Species | Appias dilutior Staudinger, 1889
| | | | Species | Appias distanti Moore, 1905
| | | | Species | Appias divisapex Hulstaert, 1924
| | | | Species | Appias dohertiella Butler, 1898
| | | | Species | Appias dohertyi Rothschild, 1892
| | | | Species | Appias dolorosa Fruhstorfer, 1910
| | | | Species | Appias domitia Felder, 1862
| | | | Species | Appias dopero Suffert, 1904
| | | | Species | Appias drosilla
Pure White | | | | Species | Appias dubia Aurivillius, 1898
| | | | Species | Appias dunaetha Fruhstorfer, 1915
| | | | Species | Appias durvasa Moore, 1857
| | | | Species | Appias eburnea Fruhstorfer, 1910
| | | | Species | Appias ega Boisduval, 1836
| | | | Species | Appias eleonora Boisduval, 1836
| | | | Species | Appias elis Fruhstorfer, 1910
| | | | Species | Appias ella Butler, 1898
| | | | Species | Appias emilia Fruhstorfer, 1902
| | | | Species | Appias enarete Boisduval, 1836
| | | | Species | Appias enaretina Fruhstorfer, 1900
| | | | Species | Appias enyo Boisduval, 1836
| | | | Species | Appias epaphioides Stoneham, 1957
| | | | Species | Appias epaphiopsis Gaede, 1916
| | | | Species | Appias epicoena Swinhoe, 1889
| | | | Species | Appias eudoxia Cramer, 1779
| | | | Species | Appias eumelis Boisduval, 1832
| | | | Species | Appias euphrosyne Stoneham, 1957
| | | | Species | Appias eurosundana Grose-Smith, 1895
| | | | Species | Appias eurynome Stoneham, 1957
| | | | Species | Appias eurytus Fruhstorfer, 1913
| | | | Species | Appias fabiola Fruhstorfer, 1905
| | | | Species | Appias falcidia Fruhstorfer, 1910
| | | | Species | Appias fasciata Fruhstorfer, 1910
| | | | Species | Appias fatima Snellen van Vollenhoven, 1866
| | | | Species | Appias festrada Fruhstorfer, 1910
| | | | Species | Appias figulina Butler, 1867
| | | | Species | Appias flaminia Fruhstorfer, 1910
| | | | Species | Appias flava RIBBE, 1886
| | | | Species | Appias flavida Trimen, 1889
| | | | Species | Appias flavius Grose-Smith, 1892
| | | | Species | Appias fleminius Fruhstorfer, 1911
| | | | Species | Appias florentia Grose-Smith, 1894
| | | | Species | Appias florentina RIBBE, 1898
| | | | Species | Appias floresiana Butler, 1898
| | | | Species | Appias formosana Wallace, 1866
| | | | Species | Appias furia Fruhstorfer, 1910
| | | | Species | Appias galathea Felder, 1862
| | | | Species | Appias galbana Fruhstorfer, 1910
| | | | Species | Appias galepsus Fruhstorfer, 1910
| | | | Species | Appias galerus Fruhstorfer, 1910
| | | | Species | Appias gellia Fruhstorfer, 1910
| | | | Species | Appias gisco Grose-Smith, 1895 ⇒ Saletara cycinna
| | | | Species | Appias grisea Moulton, 1923
| | | | Species | Appias griseoides Moulton, 1923
| | | | Species | Appias haendeli C. & R. Felder, 1865 ⇒ Appias (Glutophrissa) sabina
| 1 | Subspecies | | Species | Appias hagar Snellen van Vollenhoven, 1865
| | | | Species | Appias hainanensis Fruhstorfer, 1902
| | | | Species | Appias hecyra Mabille, 1880
| | | | Species | Appias helvola Fruhstorfer, 1910
| | | | Species | Appias hemichlora Mabille, 1897
| | | | Species | Appias herennia Fruhstorfer, 1910
| | | | Species | Appias hero Fabricius, 1793
| | | | Species | Appias higinia Godart, 1819
| | | | Species | Appias hippo Fruhstorfer, 1904
| | | | Species | Appias hippoides Moore, 1881 ⇒ Appias (Appias) lyncida
| 1 | Subspecies | | Species | Appias hippona Fruhstorfer, 1910
| | | | Species | Appias horatia Fruhstorfer, 1910
| | | | Species | Appias hostilia Fruhstorfer, 1910
| | | | Species | Appias hypatia Drury, 1782
| | | | Species | Appias icilia Fruhstorfer, 1910
| | | | Species | Appias ida Lucas, 1852
| | | | Species | Appias imbecilis Moore, 1884
| | | | Species | Appias imbeilis
| | | | Species | Appias inanis Van Eecke, 1913
| | | | Species | Appias indigis Weymer, 1887
| | | | Species | Appias indroides Honrath, 1889
| | | | Species | Appias infralimbalis Strand, 1912
| | | | Species | Appias infuscata Fruhstorfer, 1910
| | | | Species | Appias inornata Moore, 1878
| | | | Species | Appias intermedia Dufrane, 1948
| | | | Species | Appias invitabilis Fruhstorfer, 1910
| | | | Species | Appias ione Stoneham, 1957
| | | | Species | Appias iria Fruhstorfer, 1910
| | | | Species | Appias irvini Swinhoe, 1890
| | | | Species | Appias isokani Grose-Smith, 1889
| | | | Species | Appias iwasakii Matsumura, 1919
| | | | Species | Appias jacquinotii Lucas, 1852
| | | | Species | Appias kabiraensis Murayama, 1970
| | | | Species | Appias kawakamii Matsumura, 1909
| | | | Species | Appias klossi Corbet, 1941
| | | | Species | Appias korridona Grose-Smith, 1894
| | | | Species | Appias lagai Dufrane, 1947
| | | | Species | Appias lagela Moore, 1874
| | | | Species | Appias lageloides Crowley, 1900
| | | | Species | Appias lalasis Grose-Smith, 1887
| 1 | Subspecies | | Species | Appias lankapura Moore, 1879
| | | | Species | Appias larissa Felder, 1865
| | | | Species | Appias latifasciata Moore, 1881 ⇒ Appias (Appias) lyncida
| | | | Species | Appias latimarginata Gaede, 1916
| | | | Species | Appias lepidana Fruhstorfer, 1910
| | | | Species | Appias leptis Felder, 1865 ⇒ Appias (Appias) indra
| 5 | Subspecies | | Species | Appias leucosticta Butler, 1898
| | | | Species | Appias leytensis Fruhstorfer, 1911
| | | | Species | Appias libitina Godart, 1819
| | | | Species | Appias lileia Fruhstorfer, 1910
| | | | Species | Appias limbata Fruhstorfer, 1910
| | | | Species | Appias limbophora Strand, 1913
| | | | Species | Appias limia Fruhstorfer, 1910
| | | | Species | Appias lucasi
| | | | Species | Appias lucasii Wallace, 1867
| | | | Species | Appias lurida Fruhstorfer, 1910
| | | | Species | Appias lutatia Fruhstorfer, 1910
| | | | Species | Appias luvuensis Joicey & Talbot, 1927
| | | | Species | Appias lycaste Felder, 1865
| | | | Species | Appias lynceola Felder, 1865
| | | | Species | Appias maccina Fruhstorfer, 1911
| | | | Species | Appias maculata Staudinger, 1884
| | | | Species | Appias mahana Moore, 1887
| | | | Species | Appias mahoboides
| | | | Species | Appias majungana Grose-Smith, 1891
| | | | Species | Appias malatha Boisduval, 1833
| | | | Species | Appias manaia Hopkins, 1927
| | | | Species | Appias margarita Cramer, 1777 ⇒ Appias (Glutophrissa) drusilla
| 1 | Subspecies | | Species | Appias marginata Fruhstorfer, 1910
| | | | Species | Appias masabae Stoneham, 1957
| | | | Species | Appias massilia Fruhstorfer, 1910
| | | | Species | Appias matsumurai Sonan, 1930
| | | | Species | Appias matuta Doubleday, 1847
| | | | Species | Appias melanides Fruhstorfer, 1910
| | | | Species | Appias menandrus Fruhstorfer, 1910
| | | | Species | Appias micromalayana Fruhstorfer, 1910
| | | | Species | Appias minato Fruhstorfer, 1898
| | | | Species | Appias mindanaensis Butler, 1883
| | | | Species | Appias minuta Ungemach, 1932
| | | | Species | Appias moira Fruhstorfer, 1913
| | | | Species | Appias monna Fruhstorfer, 1902
| | | | Species | Appias montana Rothschild, 1896
| | | | Species | Appias montanus Rothschild, 1896 ⇒ Appias (Appias) phoebe
| | | | Species | Appias mutina Fruhstorfer, 1910
| | | | Species | Appias myndus Fruhstorfer, 1913
| | | | Species | Appias nagare Grose-Smith, 1889
| | | | Species | Appias narendra Moore, 1877
| | | | Species | Appias natalensis Von Neustetter, 1927
| | | | Species | Appias nebo Grose-Smith, 1894
| | | | Species | Appias neombo Boisduval, 1836
| | | | Species | Appias nerissa Semper, 1875
| | | | Species | Appias neronis Fruhstorfer, 1910
| | | | Species | Appias nerva Fruhstorfer, 1910
| | | | Species | Appias nicobarica Moore, 1905
| | | | Species | Appias nigricantemarginatus Goeze, 1779
| | | | Species | Appias nigrocincta Martin, 1919
| | | | Species | Appias nikomedeia Fruhstorfer, 1910
| | | | Species | Appias norma Evans, 1924
| | | | Species | Appias nyasana Butler, 1889
| | | | Species | Appias obscurior Fruhstorfer, 1910
| | | | Species | Appias ochracea Moulton, 1914
| | | | Species | Appias ochraciens Talbot, 1943
| | | | Species | Appias ochrea Bartel, 1905
| | | | Species | Appias olferna Swinhoe, 1890
Appias olferna libythea, Eastern Striped Albatross | 2 | Subspecies | | Species | Appias orbona Boisduval, 1833
| | | | Species | Appias ozolia Fruhstorfer, 1910
| | | | Species | Appias palawanica Staudinger, 1889
| | | | Species | Appias pancheia Fruhstorfer, 1910
| | | | Species | Appias panda Fruhstorfer, 1903
Nicobar Albatross | 2 | Subspecies | | Species | Appias papissa Fruhstorfer, 1910
| | | | Species | Appias paula Röber, 1892
| | | | Species | Appias paulinia Snellen van Vollenhoven, 1865
| | | | Species | Appias peducaea Fruhstorfer, 1910
| | | | Species | Appias perakana Fruhstorfer, 1902
| | | | Species | Appias pertineus
| | | | Species | Appias phestus Westwood, 1888
| | | | Species | Appias phileris Boisduval, 1833
| | | | Species | Appias pietersi Kalis, 1933
| | | | Species | Appias pione Fruhstorfer, 1910
| | | | Species | Appias placidea
| 1 | Subspecies | | Species | Appias plaethoria Fruhstorfer, 1910
| | | | Species | Appias plana Butler, 1879
| | | | Species | Appias plotina Fruhstorfer, 1902
| | | | Species | Appias poppea Lucas, 1835
| | | | Species | Appias principalis Fruhstorfer, 1910
| | | | Species | Appias pseudola
| 1 | Subspecies | | Species | Appias pseudolalage Moore, 1879
| | | | Species | Appias pseudoleis Fruhstorfer, 1910
| | | | Species | Appias psyche Felder, 1862
| | | | Species | Appias pulonus Fruhstorfer, 1906
| | | | Species | Appias pulverobasalis Joicey & Noakes, 1916
| | | | Species | Appias punctata Fruhstorfer, 1910
| | | | Species | Appias purana Fruhstorfer, 1910
| | | | Species | Appias raksasa Kalis, 1946
| | | | Species | Appias ramosa Fruhstorfer, 1898
| | | | Species | Appias retexta Swinhoe, 1890
| | | | Species | Appias reversa Stoneham, 1957
| | | | Species | Appias rhodanus Ward, 1871
| | | | Species | Appias rhodope Suffert, 1904 ⇒ Glutophrissa sylvia
| 1 | Subspecies | | Species | Appias rhodopiana Strand, 1913
| | | | Species | Appias ribbei Fruhstorfer, 1902
| | | | Species | Appias roepstorffi Moore, 1884
| | | | Species | Appias rouxii Boisduval, 1836
| | | | Species | Appias saba Hopffer, 1862
| | | | Species | Appias saina Grose-Smith, 1894
| | | | Species | Appias saumlakiana Kalis, 1933
| | | | Species | Appias sawela Fruhstorfer, 1896
| | | | Species | Appias saweloides Fruhstorfer, 1902
| | | | Species | Appias sekarensis RIBBE, 1886
| | | | Species | Appias semiepaphia Strand, 1911
| | | | Species | Appias semiflava Fruhstorfer, 1910
| | | | Species | Appias semperi Moore, 1905
| | | | Species | Appias shima Sonan, 1930
| | | | Species | Appias shiva Swinhoe, 1885
| | | | Species | Appias simplicior Strand, 1913
| | | | Species | Appias sithonia Fruhstorfer, 1911
| | | | Species | Appias solis Joicey & Noakes, 1915
| | | | Species | Appias solstitialis Butler, 1898
| | | | Species | Appias sopara Fruhstorfer, 1910
| | | | Species | Appias soramus Fruhstorfer, 1910
| | | | Species | Appias sosias Fruhstorfer, 1913
| | | | Species | Appias spec Wallace, 1867
| | | | Species | Appias statilia Fruhstorfer, 1910
| | | | Species | Appias subochracea Fruhstorfer, 1910
| | | | Species | Appias subtuslutea Roepke, 1935
| | | | Species | Appias sudanensis Talbot, 1932
| | | | Species | Appias sufflava Fruhstorfer, 1910
| | | | Species | Appias sulana Fruhstorfer, 1899
| | | | Species | Appias sulanorum Fruhstorfer, 1902
| | | | Species | Appias swinhoei Moore, 1905
| | | | Species | Appias taihokuna Nomura, 1932
| | | | Species | Appias tambora Fruhstorfer, 1902
| | | | Species | Appias tamibia Strand, 1912
| | | | Species | Appias taprobana Moore, 1879
| | | | Species | Appias tenimberensis Joicey & Talbot, 1922
| | | | Species | Appias tenuifasciata Von Neustetter, 1926
| | | | Species | Appias terentilia Fruhstorfer, 1910
| | | | Species | Appias thalia Stonehma, 1957
| | | | Species | Appias thasia Fruhstorfer, 1902
| | | | Species | Appias thrasea Fruhstorfer, 1910
| | | | Species | Appias thronion Fruhstorfer, 1910
| | | | Species | Appias thyre Fruhstorfer, 1911
| | | | Species | Appias thyria Godart, 1819
| | | | Species | Appias tibericus Fruhstorfer, 1910
| | | | Species | Appias timorensis Butler, 1898
| | | | Species | Appias tombugensis Fruhstorfer, 1902
| | | | Species | Appias tristitia Fruhstorfer, 1910
| | | | Species | Appias tsurui Matsumura, 1909
| | | | Species | Appias udei C. & R. Felder, 1865 ⇒ Appias (Glutophrissa) sabina
| 1 | Subspecies | | Species | Appias umbratilis Fruhstorfer, 1896
| | | | Species | Appias uranides Fruhstorfer, 1910
| | | | Species | Appias vacans Hampson, 1888
| | | | Species | Appias vadus Fruhstorfer, 1910
| | | | Species | Appias vana Fruhstorfer, 1902
| | | | Species | Appias vasava Fruhstorfer, 1910
| | | | Species | Appias venusta Moore, 1881
| | | | Species | Appias virginea Hulstaert, 1923
| | | | Species | Appias virilis Fruhstorfer, 1910
| | | | Species | Appias wallacei Butler, 1898
| | | | Species | Appias waltraude
| | | | Species | Appias weberi C. & R. Felder, 1865 ⇒ Appias (Glutophrissa) sabina
| 1 | Subspecies | | Species | Appias weiskei RIBBE, 1934
| | | | Species | Appias whiteheadi Grose-Smith, 1887
| | | | Species | Appias xenia Fruhstorfer, 1902
| | | | Species | Appias yaksha Fruhstorfer, 1910
| | | | Species | Appias yamazakii Sonan, 1936
| | | | Species | Appias yambo Neave, 1904 ⇒ Dixeia dixeyi
| | | | Species | Appias yayeyamana Matsumura, 1909
| | | | Species | Appias yunnana Mell, 1943
| | | | Species | Appias zamboanga Felder, 1862
| | | | Species | Appias zamora Felder, 1862
| | | | Species | Appias zelmira Stoll, 1780
| | | | Species | Appias zoe Snellen van Vollenhoven, 1865
| | | | Species | Appias zondervani Toxopeus, 1950
| | |
Synonyms | Andopodum (Scudder, 1875) | | Andropodum (Scudder, 1875) | | Appius (Brown, 1942) | | Catophaga (Hubner, 1819) | | Glutophrissa (Butler, 1887) | | Hiposcrita (Geyer, 1832) | | Hiposcritia (Geyer, 1832) | | Hyposcritia (Butler, 1898) | | Lade (Nicéville, 1898) | | Lade de (Nicéville, 1898) | | Tachyris (Wallace, 1867) | | Trigonia (Geyer, 1837) |
Native, vernacular names and spellings of Appias | English | Albatrosses, Puffins, Puffins and Albatrosses, Whites |
Distribution of Appias Hübner, 1819 Algeria, Angola, Argentina, Armenia, Australia, Bangladesh, Belgium, Belize, Benin, Bhutan, Botswana, Brazil, Brunei, Burkina Faso, Burundi, Cambodia, Cameroon, Cape Verde, Central African Republic, Chile, China, Colombia, Comoros, Cook Islands, Costa Rica, Cuba, Democratic Republic of the Congo, Denmark, Dominica, Dominican Republic, East Timor, Ecuador, Egypt, Equatorial Guinea, Ethiopia, Fiji, France, Gabon, Gambia, Germany, Ghana, Greece, Guatemala, Guinea, Haiti, Honduras, Hungary, Iceland, India, Indonesia, Iran, Ireland, Italy, Ivory Coast, Jamaica, Japan, Jordan, Kenya, Kiribati, Laos, Liberia, Madagascar, Malawi, Malaysia, Maldives, Mali, Malta, Marshall Islands, Mexico, Micronesia, Morocco, Mozambique, Myanmar, Namibia, Nepal, Netherlands, New Zealand, Nicaragua, Niger, Nigeria, Norway, Oman, Pakistan, Palau, Panama, Papua New Guinea, Paraguay, Peru, Philippines, Portugal, Republic of the Congo, Republic South Africa, Romania, Rwanda, Samoa, Saudi Arabia, Senegal, Sierra Leone, Singapore, Solomon Islands, South Sudan, Spain, Sri Lanka, Sudan, Sweden, Taiwan, Tanzania, Thailand, Togo, Tonga, Trinidad and Tobago, Tunisia, Uganda, United Kingdom, Uruguay, USA, Vanuatu, Vietnam, Yemen, Zambia, Zimbabwe
Distribution on islands Alor Archipelago, Barat Daya Islands, Kai Islands, Langkawi, Murray Islands, Obi Islands, Orkney Islands, Schouten Islands, Shetland Islands
Links, sources and further information
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