Taxonomy of Genus Spialia Swinhoe, 1912
African grizzled skippers, Grizzled Skippers, Sandmen
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 4179
190570
2444470 | Families
Genera
Species |
Subclass
| Pterygota
Lang, 1888
Winged insects | 3816
189502
2439455 | Families
Genera
Species |
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 3399
186457
2415487 | Families
Genera
Species |
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 1891
142849
2048615 | Families
Genera
Species |
Order
| Lepidoptera
Linnaeus, 1758
Butterflies, Lepidopterans, Moths | 322
33641
568299 | Families
Genera
Species |
Suborder
| Glossata
Fabricius, 1775
Modern butterflies and moths | 208
32654
567261 | Families
Genera
Species |
Infraorder
| Heteroneura
Tillyard, 1918
| 197
32461
565431 | Families
Genera
Species | | Division | Ditrysia
Borner, 1925
Butterflies, Moths, Skippers | 186
32290
560864 | Families
Genera
Species | | Microorder | Apoditrysia
Minet, 1983
| 13
1792
34426 | Families
Genera
Species | | Phalanx | Obtectomera
Minet, 1986
| 8
1724
33194 | Families
Genera
Species |
Section
| Macrolepidoptera
Minet, 1991
Macro-moths | 6
1580
31274 | Families
Genera
Species |
Subsection
| Rhopalocera
Boisduval, 1836
Butterflies | 3
1099
24454 | Families
Genera
Species |
Superfamily
| Hesperioidea
Latreille, 1809
Skippers | 2
1096
24391 | Families
Genera
Species | | Family | Hesperiidae
Latreille, 1809
Indet. Skipper, Skipper, Skipper Butterflies, Skippers | 1096
24391 | Genera
Species | | Subfamily | Pyrginae
Burmeister, 1878
Dicot Skippers, Flats, Grizzled Skippers, Pyrgine skippers | 242
9165 | Genera
Species | | Tribe | Carcharodini
Verity, 1940
| 31
1255 | Genera
Species | | Genus | Spialia
Swinhoe, 1912
African grizzled skippers, Grizzled Skippers, Sandmen | 168 | Species |
Subnodes of Spialia Swinhoe, 1912 | Species | Spialia abscondita Plötz, 1884
Common Sandman, Diomus Grizzled Skipper | 1 | Subspecies | | Species | Spialia adenensis Butler, 1884 ⇒ Spialia doris
| | | | Species | Spialia aestiva Querci, 1932
| | | | Species | Spialia agylla Trimen, 1889
Grassveld Sandman | 2 | Subspecies | | Species | Spialia albodetersa Verity, 1929
| | | | Species | Spialia ali Oberthür, 1881
| | | | Species | Spialia alioides Verity, 1926
| | | | Species | Spialia alpina Erschov, 1874
| | | | Species | Spialia amenophis Reverdin, 1914
| | | | Species | Spialia anavalsae Coutsis, 1976
| | | | Species | Spialia aron
| | | | Species | Spialia asterodea
| | | | Species | Spialia asterodia Trimen, 1864
Asterodia Sandman, Star Sandman | | | | Species | Spialia aurivilli Shepard, 1935
| | | | Species | Spialia australis de Jong, 1978
| | | | Species | Spialia bamptoni Vári, 1976
| | | | Species | Spialia bettoni Butler, 1898 ⇒ Spialia kituina
| | | | Species | Spialia bifida Higgins, 1925
| | | | Species | Spialia carnea Reverdin, 1927
| 1 | Subspecies | | Species | Spialia catenata Jöst, 1957 ⇒ Spialia sertorius
| | | | Species | Spialia chenga Evans, 1957
| | | | Species | Spialia colotes Druce, 1875
Bushveld Sandman | 3 | Subspecies | | Species | Spialia confusa Higgins, 1925
Confusing Sandman | 2 | Subspecies | | Species | Spialia daphne Evans, 1949
| | | | Species | Spialia delagoae Trimen, 1898
Delagoa Grizzled Skipper, Delagoa Sandman | | | | Species | Spialia depauperata Strand, 1911
Deprived Grizzled Skipper, Wandering Sandman | 2 | Subspecies | | Species | Spialia diffusa Jöst, 1957 ⇒ Spialia sertorius
| | | | Species | Spialia diomus Hopffer, 1855
Common Sandman, Diomus Grizzled Skipper | 2 | Subspecies | | Species | Spialia doris Walker, 1870
Aden Skipper, Desert Grizzled Skipper | 4 | Subspecies | | Species | Spialia driomus
| | | | Species | Spialia dromus Plötz, 1884
Dromus Grizzled Skipper, Forest Sandman | | | | Species | Spialia elongata Higgins, 1925 ⇒ Spialia dromus
| | | | Species | Spialia eucrate Ochsenheimer, 1808
| | | | Species | Spialia eupator Hemming, 1932
| | | | Species | Spialia evanidus Butler, 1880
| | | | Species | Spialia extrema Pfeiffer, 1938
| | | | Species | Spialia fasciata Higgins, 1925 ⇒ Spialia confusa
| | | | Species | Spialia ferax Wallengren, 1863
Ferax Grizzled Skipper | | | | Species | Spialia galba Fabricius, 1793
Indian Grizzled, Indian Grizzled Skipper, Indian Skipper | 3 | Subspecies | | Species | Spialia gavarniensis Warren, 1926
| | | | Species | Spialia gecko Evans, 1949
| | | | Species | Spialia geron Watson, 1893
Aden Skipper, Baluchi Skipper | 2 | Subspecies | | Species | Spialia gracilis Verity, 1919
| | | | Species | Spialia guadarramensis Warren, 1925
| | | | Species | Spialia hebra
| 1 | Subspecies | | Species | Spialia hellas de Nicéville, 1889
| | | | Species | Spialia hermona Evans, 1957
| | | | Species | Spialia hibiscae Hoffmannsegg, 1804 ⇒ Spialia sertorius
Red-underwing Skipper | 2 | Subspecies | | Species | Spialia higginsi Evans, 1937
| | | | Species | Spialia hilaris Staudinger, 1901
| | | | Species | Spialia infraurata Verity, 1929
| | | | Species | Spialia ioan Warren, 1926
| | | | Species | Spialia jason Oberthür, 1912
| | | | Species | Spialia kempnyi Schawerda, 1919
| | | | Species | Spialia kiki Higgins, 1974
| | | | Species | Spialia kituina Karsch, 1896
Kitui Grizzled Skipper | | | | Species | Spialia lacreuzei Oberthür, 1912
| | | | Species | Spialia leucomelas Oberthür, 1912 ⇒ Spialia dromus
| | | | Species | Spialia lugens Staudinger, 1886
| | | | Species | Spialia machacoana Butler, 1899 ⇒ Spialia diomus
| | | | Species | Spialia maculatus BREMER & GREY, 1853
| | | | Species | Spialia mafa Trimen, 1870
Mafa grizzled skipper, Mafa Sandman | 2 | Subspecies | | Species | Spialia mangana Rebel, 1899
Arabian Grizzled Skipper | | | | Species | Spialia medioalbotersa Verity, 1940 ⇒ Spialia sertorius
| | | | Species | Spialia melaleuca Oberthür, 1912 ⇒ Spialia dromus
| | | | Species | Spialia meridionalis Higgins, 1925 ⇒ Spialia dromus
| | | | Species | Spialia minor Rebel, 1910
| | | | Species | Spialia minuscula Reverdin, 1933
| | | | Species | Spialia murasaki Sugitani, 1936
| | | | Species | Spialia nana Trimen, 1889
| | | | Species | Spialia nanus Trimen, 1889
Dwarf Sandman | | | | Species | Spialia nora Plötz, 1884 ⇒ Spialia colotes
| | | | Species | Spialia oberthuri Aurivillius, 1925 ⇒ Spialia mafa
| | | | Species | Spialia obscura Higgins, 1925
| | | | Species | Spialia occidentalis de Jong, 1977
| | | | Species | Spialia orbifer Hübner, 1823
Brick Skipper, Hungarian Skipper | 5 | Subspecies | | Species | Spialia osthelderi Pfeiffer, 1932
| 2 | Subspecies | | Species | Spialia parvula Verity, 1921
| | | | Species | Spialia paula Higgins, 1925
Mite Sandman | | | | Species | Spialia phlomidis Herrich-Schäffer, 1845
Balochi Skipper, Persian Skipper | 3 | Subspecies | | Species | Spialia phlonidis
| | | | Species | Spialia ploetzi Aurivillius, 1891
Forest Grizzled Skipper, Froest Grizzled Skipper | 2 | Subspecies | | Species | Spialia poggei Lederer, 1858
| | | | Species | Spialia postesselloides Verity, 1938 ⇒ Spialia sertorius
| | | | Species | Spialia rebeli Aurivillius, 1891 ⇒ Spialia ploetzi
Forest Grizzled Skipper | 1 | Subspecies | | Species | Spialia rehfoussi Oberthür, 1913
| | | | Species | Spialia retrograda Verity, 1940
| | | | Species | Spialia rosae Hernández-Roldán, Dapporto, Dincă, Vicente & Vila, 2016
| | | | Species | Spialia sao Hübner, 1803 ⇒ Spialia sertorius
| | | | Species | Spialia sataspes Trimen, 1864
Boland Sandman | | | | Species | Spialia secessa Trimen, 1891
| | | | Species | Spialia secessus Trimen, 1891
Wolkberg Sandman | | | | Species | Spialia secunda Hübner, 1823 ⇒ Spialia orbifer
Hungarian Skipper, Orbed Red-underwing Skipper | 1 | Subspecies | | Species | Spialia semiconfluens de Jong, 1978
| | | | Species | Spialia sertorius Hoffmannsegg, 1804
Brick Skipper, Red Underwing Skipper | 8 | Subspecies | | Species | Spialia shanta Evans, 1957
| | | | Species | Spialia spec Fabricius, 1793
| | | | Species | Spialia spio Linnaeus, 1764
Mountain Sandman, Spio Grizzled Skipper | | | | Species | Spialia struvei Püngeler, 1914
| 4 | Subspecies | | Species | Spialia subgracilis Verity, 1921
| | | | Species | Spialia subtusbrunnea Reverdin, 1922
| | | | Species | Spialia superna Moore, 1865
| | | | Species | Spialia tesselloides Hübner, 1823 ⇒ Spialia orbifer
Hungarian Skipper, Orbed Red-underwing Skipper | 1 | Subspecies | | Species | Spialia therapne Rambur, 1832
Corsican Red-underwing Skipper | | | | Species | Spialia therapnoides Oberthür, 1910
| | | | Species | Spialia transvaaliae Trimen, 1889
| | | | Species | Spialia trimeni Aurivillius, 1925 ⇒ Spialia secessa
| | | | Species | Spialia vindex Cramer, 1782 ⇒ Spialia spio
| | | | Species | Spialia wrefordi Evans, 1951
Wreford´s Grizzled Skipper | | | | Species | Spialia zaira Plötz, 1884 ⇒ Spialia dromus
| | | | Species | Spialia zebra Butler, 1888 ⇒ Spialia ploetzi
Zebra Grizzled Skipper, Zebra Skipper | 2 | Subspecies | | Species | Spialia zona Mabille, 1875
| | |
Synonyms Native, vernacular names and spellings of Spialia | English | African grizzled skippers, Grizzled Skippers, Sandmen |
Distribution of Spialia Swinhoe, 1912 Afghanistan
| Albania
| Algeria
| Andorra
| Angola
| Argentina
| Armenia
| Australia
| Austria
| Azerbaijan
| Bangladesh
| Belgium
| Belize
| Benin
| Bhutan
| Bolivia
| Bosnia and Herzegovina
| Botswana
| Brazil
2 Species | Bulgaria
| Burkina Faso
| Cambodia
| Cameroon
| Canada
| China
| Colombia
| Costa Rica
| Croatia
| Cuba
| Czech Republic
| Democratic Republic of the Congo
| Djibouti
| Ecuador
| Egypt
| Ethiopia
| France
| Gabon
| Gambia
| Germany
| Ghana
| Greece
| Guinea
| Guinea-Bissau
| Hungary
| India
| Indonesia
| Iran
| Iraq
| Israel
| Italy
| Ivory Coast
| Japan
| Jordan
| Kazakhstan
| Kenya
| Kyrgyzstan
| Laos
| Lebanon
| Lesotho
| Liberia
| Liechtenstein
| Luxembourg
| Madagascar
| Malawi
| Mali
| Malta
| Mauritania
| Mauritius
| Mexico
4 Species | Moldova
| Mongolia
| Montenegro
| Morocco
| Mozambique
| Myanmar
| Namibia
| Nepal
| Netherlands
| New Zealand
| Nicaragua
| Niger
| Nigeria
| Norway
| Oman
| Pakistan
| Palestine
| Panama
| Papua New Guinea
| Peru
2 Species | Poland
| Portugal
| Republic of the Congo
| Republic South Africa
| Romania
| Russia
| Rwanda
| Saint Lucia
| Saudi Arabia
| Senegal
| Serbia
| Sierra Leone
| Slovakia
| Slovenia
| Somalia
| Somaliland
| South Sudan
| Spain
| Sri Lanka
| Sudan
| Switzerland
| Syria
| Tajikistan
| Tanzania
| Thailand
| Togo
| Tunisia
| Turkey
| Turkmenistan
| Uganda
| Ukraine
| United Arab Emirates
| United Kingdom
| USA
| Uzbekistan
| Vietnam
| Yemen
| Zambia
| Zimbabwe
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Links, sources and further information
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