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Checklist for Tribe Coprini Kolbe, 1805
and region Continents


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
2110
116133
1222976
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
2101
115931
1221959
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
1812
113780
1210168
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
929
84484
1007160
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
234
35522
419905
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
189
32170
363924
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
23
3594
50225
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
23
3594
50225
Families
Genera
Species
 
Taxom
 
FamilyScarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles
2963
42190
Genera
Species
 
Taxom
 
SubfamilyScarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles
289
7423
Genera
Species
 
Taxom
 
TribeCoprini
Kolbe, 1805
19
1188
Genera
Species
 
Taxom
 


Genera for Continents
19 Genera with 1188 Species
Canthidium Erichson, 1886184SpeciesTaxom
Catharsius Hope, 1857118SpeciesTaxom
Chalcocopris Burmeister, 20152SpeciesTaxom
Copridaspidus Bloucomont, 18781SpeciesTaxom
Copris Geoffroy, 2009Mexican dung beetle364SpeciesTaxom
Coptodactyla Burmeister, 192517SpeciesTaxom
Dichotomius Hope, 1922225SpeciesTaxom
Heliocopris Hope, 201367SpeciesTaxom
Holocephalus Hope, 20016SpeciesTaxom
Isocopris Pereira & Ma..., 20177SpeciesTaxom
Litocopris Waterhouse, 18694SpeciesTaxom
Macroderes Westwood, 188025SpeciesTaxom
Metacatharsius Paulian, 196566SpeciesTaxom
Ontherus Erichson, 193063SpeciesTaxom
Pseudocopris Ferreira, 19102SpeciesTaxom
Synapsis Forster, 193328SpeciesTaxom
Thyregis Blackburn, 19334SpeciesTaxom
Uroxys Westwood, 190365SpeciesTaxom
Xinidium Harold, 19934SpeciesTaxom


Species list for Continents
1188 Species and 64 Subspecies
Canthidium (Eucanthidium) laetum (Harold, 1867)⇒ Eucanthidium laetum Harold, 1867
Canthidium (Eucanthidium) macclevei Kohlmann & S..., 2006
Canthidium (Eucanthidium) pseudopuncticolle Solis & Kohl..., 2004
Canthidium abbreviatum Harold, 1867
Canthidium aeneolum Harold, 1867
Canthidium alvarezi Martinez & H..., 1986
Canthidium andersoni Kohlmann & S..., 2006
Canthidium angulicolle Balthasar, 1939
Canthidium angusticeps Bates, 1887
Canthidium annagabrielae Solis & Kohl..., 2004
Canthidium ardens Bates, 1887
Canthidium aterrimum Harold, 1867
Canthidium atomarium Balthasar, 1939
Canthidium atramentarium Balthasar, 1939
Canthidium atricolle Preudhomme d..., 1886
Canthidium aurichalceum Preudhomme d..., 1886
Canthidium auricolle Harold, 1867
Canthidium aurifex Bates, 1887
Canthidium ayri Carvalho de ..., 2019
Canthidium barbacenicum Preudhomme d..., 1886
Canthidium basale Harold, 1867
Canthidium basipunctatum Balthasar, 1939
Canthidium batesi Harold, 1867
Canthidium bicolor Boucomont, 1928
Canthidium bituberifrons Howden & Young, 1981
Canthidium bokermanni Martinez, Ha..., 1964
Canthidium bovinum Harold, 1867
Canthidium breve Germar, 1824
Canthidium caesareum Balthasar, 1939
Canthidium calidum Harold, 1880
Canthidium cavifrons Balthasar, 1939
Canthidium centrale Boucomont, 1928
Canthidium cereum Harold, 1868
Canthidium chabanaudi Boucomont, 1928
Canthidium chimalapense Moctezuma & ..., 2019
Canthidium chrysis Fabricius, 1801
Canthidium clypeale Harold, 1867
Canthidium coerulescens Balthasar, 1939
Canthidium cognatum Preudhomme d..., 1886
Canthidium collare Castelnau, 1840
Canthidium convexifrons Balthasar, 1939
Canthidium cupreum Blanchard, 1846
Canthidium cuprinum Harold, 1867
Canthidium darwini Kohlmann & S..., 2009
Canthidium decoratum Perty, 1830
Canthidium delgadoi Kohlmann & S..., 2006
Canthidium deplanatum Balthasar, 1939
Canthidium depressum Boucomont, 1928
Canthidium deyrollei Harold, 1867
Canthidium dimidiatum Lansberge, 1874
Canthidium discolor Harold, 1867
Canthidium discopygidiale Howden & Young, 1981
Canthidium dispar Harold, 1867
Canthidium dohrni Harold, 1867
Canthidium elegantulum Balthasar, 1939
Canthidium emoryi Solis & Kohl..., 2004
Canthidium epistomale Boucomont, 1928
Canthidium erythropterum Lucas, 1857
Canthidium escalerai Balthasar, 1939
Canthidium euchalceum Balthasar, 1939
Canthidium excisipes Balthasar, 1939
Canthidium feeri Carvalho de ..., 2019
Canthidium femoratum Boucomont, 1935
Canthidium flabellatum Harold, 1883
Canthidium flavicorne Blanchard, 1846
Canthidium flavipes Harold, 1867
Canthidium flavum Balthasar, 1939
Canthidium foveolatum Harold, 1867
Canthidium funebre Balthasar, 1939
Canthidium gemmingeri Harold, 1867
Canthidium gerstaeckeri Harold, 1867
Canthidium gigas Balthasar, 1939
Canthidium glabricolle Harold, 1867
Canthidium globulum Harold, 1867
Canthidium gracilipes Harold, 1867
Canthidium granuliferum Pereira, 1949
Canthidium guanacaste Howden & Gill, 1987
Canthidium guyanense Boucomont, 1928
Canthidium haagi Harold, 1867
Canthidium haroldi Preudhomme d..., 1886
Canthidium hespenheidei Howden & Young, 1981
Canthidium histrio Balthasar, 1939
Canthidium howdeni Kohlmann & S..., 2006
Canthidium humerale Germar, 1813
Canthidium hyla Balthasar, 1939
Canthidium hypocrita Balthasar, 1939
Canthidium imperiale Harold, 1876
Canthidium impressum Boucomont, 1928
Canthidium inerme Harold, 1867
Canthidium inoptatum Balthasar, 1939
Canthidium kelleri Martinez, Ha..., 1964
Canthidium kiesenwetteri Harold, 1867
Canthidium kirschi Harold, 1875
Canthidium kohlmanni Mora-Aguilar..., 2019
Canthidium korschefskyi Balthasar, 1939
Canthidium kraatzi Harold, 1867
Canthidium laevigatum Harold, 1867
Canthidium latipleurum Preudhomme d..., 1886
Canthidium latum Blanchard, 1846
Canthidium lebasi Harold, 1867
Canthidium lentum Erichson, 1847
Canthidium leucopterum Howden & Young, 1981
Canthidium lucidum Harold, 1867
Canthidium luteum Balthasar, 1939
Canthidium macroculare Howden & Gill, 1987
Canthidium magnum Harold, 1871
Canthidium manni Arrow, 1913
Canthidium margaritae Kohlmann & S..., 2006
Canthidium marianelae Solis & Kohl..., 2004
Canthidium marielae Solis & Kohl..., 2004
Canthidium marseuli Harold, 1867
Canthidium melanocephalum Olivier, 1789
Canthidium metallicum Harold, 1867
Canthidium minimum Harold, 1883
Canthidium miscellum Harold, 1883
Canthidium moestum Harold, 1867
Canthidium monoceros Harold, 1869
Canthidium moroni Kohlmann & S..., 2006
Canthidium multipunctatum Balthasar, 1939
Canthidium muticum Boheman, 1858
Canthidium nanum Harold, 1867
Canthidium nebularum Moctezuma, S..., 2019
Canthidium nigritum Preudhomme d..., 1886
Canthidium nitidum (Blanchard, 1846)⇒ Eucanthidium cupreum Blanchard, 1843
Canthidium nobile Harold, 1867
Canthidium obscurum Harold, 1867
Canthidium onitoides Perty, 1830
Canthidium onthophagoides Martinez & H..., 1986
Canthidium opacum Balthasar, 1939
Canthidium orbiculatum (Lucas, 1857)⇒ Eucanthidium cupreum Blanchard, 1843
Canthidium pallidoalatum Howden & Young, 1981
Canthidium paranum Harold, 1867
Canthidium parvulum Harold, 1883
Canthidium perceptibile Howden & Young, 1981
Canthidium persplendens Balthasar, 1939
Canthidium picipes Harold, 1867
Canthidium pinotoides (Balthasar, 1939)⇒ Canthidium barbacenicum Preudhomme d..., 1886
Canthidium planovultum Howden & Young, 1981
Canthidium prasinum Blanchard, 1846
Canthidium priscillae Solis & Kohl..., 2004
Canthidium pseudaurifex Balthasar, 1939
Canthidium pseudoperceptibile Kohlmann & S..., 2006
Canthidium pullus Felsche, 1910
Canthidium punctatostriatum Mannerheim, 1829
Canthidium punctatum Schmidt, 1922
Canthidium puncticeps Harold, 1867
Canthidium puncticolle Harold, 1867
Canthidium puncticollis Harold, 1868
Canthidium quadridens Harold, 1867
Canthidium quercetorum Kohlmann, Ar..., 2018
Canthidium refulgens Boucomont, 1928
Canthidium riverai Kohlmann & S..., 2006
Canthidium ruficolle Germar, 1824
Canthidium rufinum Harold, 1867
Canthidium rufipes Harold, 1867
Canthidium rutilum Harold, 1867
Canthidium seladon Balthasar, 1939
Canthidium semicupreum Harold, 1868
Canthidium sladeni Arrow, 1903
Canthidium smaragdinum Harold, 1867
Canthidium smithi Bates, 1889
Canthidium splendidum Preudhomme d..., 1886
Canthidium stahli Harold, 1867
Canthidium stali Harold, 1867
Canthidium steinheili Harold, 1880
Canthidium stofeli Carvalho de ..., 2019
Canthidium subdopuncticolle Howden & Young, 1981
Canthidium sulcatum Perty, 1830
Canthidium sulcicolle Harold, 1868
Canthidium taurinum Harold, 1867
Canthidium tenebrosum Howden & Young, 1981
Canthidium thalassinum Erichson, 1847
Canthidium titschacki Balthasar, 1939
Canthidium tricolor Balthasar, 1939
Canthidium trinodosum Boheman, 1858
Canthidium tuberifrons Howden & Young, 1981
Canthidium variolosum Howden & Young, 1981
Canthidium versicolor Harold, 1867
Canthidium vespertinum Howden & Young, 1981
Canthidium violaceipenne Blanchard, 1846
Canthidium viride Lucas, 1857
Canthidium viridicolle Blanchard, 1846
Canthidium viridiobscurum Boucomont, 1928
Canthidium volxemi Preudhomme d..., 1886
Catharsius achates Olivier, 1789
Catharsius adamastor Gillet, 1932
Catharsius aegeus Génier, 2017
Catharsius aethiops Sharp, 1875
Catharsius africanus Ferreira, 1960
Catharsius alpheus Gillet, 1932
Catharsius approximans Kolbe, 1893
Catharsius balubanus Kolbe, 1893
Catharsius bellus Gillet, 1918
Catharsius belus Gillet, 1918
Catharsius biconifer Felsche, 1907
Catharsius bicornutus Kolbe, 1893
Catharsius birmanensis Lansberge, 1874
Catharsius borneensis (Paulian, 1936)⇒ Catharsius molossus Linnaeus, 1758
Catharsius bradshawi Lansberge, 1887
Catharsius brevicornis Felsche, 1907
Catharsius brittoni Ferreira, 1963
Catharsius brutus Harold, 1880
Catharsius calaharicus Kolbe, 1893
Catharsius capucinus Fabricius, 1781
Catharsius cassius Kolbe, 1893
Catharsius chinai Ferreira, 1960
Catharsius congolensis Ferreira, 1962
Catharsius coptorhinaformis Josso & Prév..., 2009
Catharsius crassicornis Gillet, 1911
Catharsius dayacus Lansberge, 1886
Catharsius disseptus (Péringuey, 1901)⇒ Catharsius calaharicus Kolbe, 1893
Catharsius dubius (Péringuey, 1901)⇒ Catharsius molossus Linnaeus, 1758
Catharsius duciformis Ferreira, 1959
Catharsius dux Harold, 1878
Catharsius erechtheus Felsche, 1907
Catharsius eteocles (Laporte, 1840)⇒ Catharsius alpheus Gillet, 1932
Catharsius fastidiosus Thomson, 1858
Catharsius folicornis Josso & Prév..., 2019
Catharsius furcillatus Guérin-Ménev..., 1847
Catharsius furcillatus furcillatus Guérin, 1847
Catharsius furcillatus laeviplaga G. Müller, 1941
Catharsius garambae Ferreira, 1962
Catharsius genieri Moretto, 2014
Catharsius gorilla Thomson, 1858
Catharsius gorilloides Felsche, 1907
Catharsius granulatus Sharp, 1875
Catharsius guineensis Ferreira, 1963
Catharsius haafi Ferreira, 1960
Catharsius haroldi Takano, 2021
Catharsius harpagus Harold, 1877
Catharsius heros Boheman, 1860
Catharsius hertli Ferreira, 1964
Catharsius jacksoni Waterhouse, 1891
Catharsius javanus Lansberge, 1886
Catharsius juheli Josso, 2011
Catharsius kangeanus (Paulian, 1936)⇒ Catharsius molossus Linnaeus, 1758
Catharsius karlwerneri Moretto, 2008
Catharsius kolbei Felsche, 1907
Catharsius laticeps Boheman, 1857
Catharsius latifossa Müller, 1941
Catharsius lepineyi Boucomont, 1934
Catharsius longiceps Gillet, 1907
Catharsius luluensis Ferreira, 1960
Catharsius lycaon Kolbe, 1893
Catharsius machadoi Ferreira, 1960
Catharsius marcellus Kolbe, 1893
Catharsius melancholicus (Boheman, 1860)⇒ Catharsius longiceps Gillet, 1907
Catharsius minutus (Lansberge, 1882)⇒ Metacatharsius minutus Lansberge, 1882
Catharsius mirabilis Felsche, 1901
Catharsius molossus Linnaeus, 1758
Catharsius morettoi Josso & Prév..., 2009
Catharsius mossambicanus Ferreira, 1960
Catharsius mulanjeensis Josso & Prév..., 2019
Catharsius nathani Frey, 1956
Catharsius neptunus Kolbe, 1893
Catharsius ninus Gillet, 1918
Catharsius oedipus Felsche, 1907
Catharsius omoensis Müller, 1941
Catharsius oryx Frey, 1974
Catharsius pallas Génier, 2017
Catharsius pandion Harold, 1877
Catharsius parafastidiosus Ferreira, 1971
Catharsius peleus (Olivier, 1789)⇒ Metacatharsius peleus Olivier, 1789
Catharsius phidias (Olivier, 1789)⇒ Catharsius platicornis Ferreira, 1960
Catharsius philus Kolbe, 1893
Catharsius pithecius Fabricius, 1775
Catharsius platicornis Ferreira, 1960
Catharsius platycerus Klug, 1855
Catharsius platynotus Felsche, 1907
Catharsius platypus Sharp, 1875
Catharsius polynices Kolbe, 1893
Catharsius pseudocongolensis Ferreira, 1963
Catharsius pseudolycaon Ferreira, 1960
Catharsius pseudooedipus Ferreira, 1960
Catharsius pseudoplaticornis Josso & Prév..., 2019
Catharsius pseudovitulus Müller, 1941
Catharsius punctatus Josso & Prév..., 2019
Catharsius quadridentatus Lansberge, 1885
Catharsius renaudpauliani Ochi & Kon, 1996
Catharsius rhinoceros Klug, 1855
Catharsius saegeri Ferreira, 1962
Catharsius sagax Quenstedt, 1806
Catharsius satyrus Kolbe, 1893
Catharsius scopas Felsche, 1907
Catharsius semirubidus Fairmaire, 1891
Catharsius sesostris Waterhouse, 1888
Catharsius severini Gillet, 1907
Catharsius simillimus Ferreira, 1960
Catharsius somalicus Gillet, 1918
Catharsius spectabilis Gillet, 1918
Catharsius straeleni Ferreira, 1963
Catharsius stricticornis Josso & Prév..., 2019
Catharsius stuhlmanni Kolbe, 1893
Catharsius takanoi Josso & Prév..., 2019
Catharsius timorensis (Lansberge, 1879)⇒ Catharsius molossus Linnaeus, 1758
Catharsius tricornutus DeGeer, 1778
Catharsius tridens Josso & Prév..., 2009
Catharsius ugandicus Gillet, 1918
Catharsius ulysses Boheman, 1857
Catharsius upembanus Ferreira, 1962
Catharsius vansoni Ferreira, 1960
Catharsius vitulus Boheman, 1857
Chalcocopris hesperus Olivier, 1789
Chalcocopris inexpectatus Rossini & Va..., 2015
Copridaspidus peregrinus Harold, 1878
Copris (Copris) cambodiensis Ochi, Kon & ..., 2008
Copris cambodiensis cambodiensis
Copris cambodiensis masumotoi Ochi, Kon & ..., 2011
Copris (Copris) fukiensis Balthasar, 1952
Copris (Copris) kachinensis Ochi, Kon & ..., 2008
Copris (Copris) kartlinus O. N. Kabakov, 1988
Copris (Copris) punctatus Gillet, 1910
Copris punctatus formosanus Gillet, 1922
Copris punctatus punctatus
Copris (Copris) schoolmeestersi Ochi & Kon, 2010
Copris (Copris) tripartitus Waterhouse, 1875
Copris (Microcopris) propinquus Felsche, 1910
Copris (Paracopris) cariniceps Felsche, 1910
Copris (Sinocopris) barclayi Ochi, Kon & Bai, 2009
Copris (Sinocopris) pecuarius Lewis, 1884
Copris pecuarius continentalis Ochi & Kon, 2010
Copris (Sinocopris) uenoi Hanboonsong ..., 2003
Copris achates (Péringuey, 1901)⇒ Copris cornifrons Boheman, 1860
Copris acutidens Motschulsky, 1860
Copris afgoi Cambefort, 1992
Copris agnus Sharp, 1875
Copris algol Nguyen-phung, 1989
Copris amabilis Kolbe, 1914
Copris amazonicus Darling & Gé..., 2018
Copris amyntor (Klug, 1855)⇒ Copris harrisi Waterhouse, 1891
Copris anceus Olivier, 1789
Copris andrewesi Waterhouse, 1891
Copris angusticornis Arrow, 1933
Copris angustus Nguyen-phung..., 1986
Copris anomiopseoides (Boucomont, 1924)⇒ Paracopris anomiopseoides Boucomont, 1924
Copris antares Bacescu & Ca..., 1947
Copris apicepunctatus Balthasar, 1942
Copris arcturus Gillet, 1907
Copris ares Zelenka, 1993
Copris arimotoi Ochi & Araya, 1994
Copris arizonensis Schaeffer, 1906
Copris armatus Harold, 1869
Copris armeniacus Falderman, 1835
Copris armiger Gillet, 1910
Copris arrowi Felsche, 1910
Copris aspericollis Gillet, 1910
Copris atropolitus Gillet, 1910
Copris bartolozzi Ochi, Kon & ..., 2019
Copris basilewskyi Ferreira, 1962
Copris basipunctatus Balthasar, 1942
Copris bellator Chevrolat, 1844
Copris bengalensis Shiino, 1978
Copris bidens (Pfeffer, 1888)⇒ Liatongus femoratus Illiger, 1800
Copris bihamatus Balthasar, 1965
Copris binodis Thunberg, 1818
Copris bootes Klug, 1855
Copris boucardi Harold, 1869
Copris bovinus Gillet, 1908
Copris brachypterus Nomura, 1964
Copris brachypterus brachypterus
Copris brachypterus toguchii Ochi & Kawahara, 2012
Copris brevicornis Darling & Gé..., 2018
Copris brittoni Ferreira, 1962
Copris caelatus Fabricius, 1794
Copris caliginosus Kohlmann, Ca..., 2003
Copris calvus Sharp, 1875
Copris cambeforti Nguyen-phung, 1988
Copris camerunus Felsche, 1904
Copris caobangensis Bui, Dumack ..., 2018
Copris capensis Waterhouse, 1891
Copris carinicus Gillet, 1910
Copris carmelita Fabricius, 1801
Copris cartwrighti (Robinson, 1941)⇒ Copris fricator cartwrighti Robinson, 1941
Copris cassius Péringuey, 1901
Copris celebensis Lansberge, 1886
Copris cheni Ochi, Kon & Bai, 2007
Copris chimalapensis Mora-Aguilar..., 2015
Copris chinmontanus Ochi, Kon & ..., 2019
Copris ciliatus
Copris colmanti Gillet, 1908
Copris complexus Nguyen-phung, 1989Robber Baron Cave meshweaver
Copris compressipennis (Gillet, 1910)⇒ Paracopris compressipennis Gillet, 1910
Copris confucius Harold, 1877
Copris continentalis Ochi & Kon, 2010
Copris coriarius Gillet, 1907
Copris corniculatus (Mulsant, 1842)⇒ Copris lunaris Linnaeus, 1758
Copris cornifrons Boheman, 1860
Copris corniger (Sahlberg, 1823)⇒ Onthophagus taurus Schreber, 1759
Copris corpulentus Gillet, 1910
Copris costaricensis Gahan, 1894
Copris costaricensis costaricensis
Copris costaricensis dolichocerus Matthews, 1961
Copris costariensis
Copris crassus Deschodt & D..., 2015
Copris cribratus Gillet, 1927
Copris cribricollis (Gillet, 1910)⇒ Catharsius pithecius Fabricius, 1775
Copris curlettii Moretto, 2011
Copris davidi Darling & Gé..., 2018
Copris davidis Deyrolle, 1878
Copris davisi Nguyen-phung..., 1986
Copris davisoni Waterhouse, 1891
Copris decellei Cambefort, 1992
Copris delicatus Arrow, 1931
Copris denticulatus Nguyen-phung, 1988
Copris diversus Waterhouse, 1891
Copris doriae Harold, 1877
Copris draco Arrow, 1906
Copris dracunculus Fereira, 1959
Copris druidum O. Heer, 1862
Copris dudleyi Cambefort, 1992
Copris eburneus Cambefort, 1992
Copris elpheneroides Felsche, 1910
Copris elphenor Klug, 1855
Copris erratus Lansberge, 1886
Copris erratus erratus
Copris erratus masayukii Ochi & Kon, 2004
Copris evanidus Klug, 1855
Copris excisus Waterhouse, 1891
Copris fallaciosus Gillet, 1907Kenyan dung beetle
Copris fallax Felsche, 1910
Copris felschei (Reitter, 1893)⇒ Copris armeniacus Falderman, 1835
Copris fidius Olivier, 1789
Copris frankenbergerianus Balthasar, 1958
Copris freyi Ferreira, 1961
Copris fricator Fabricius, 1787
Copris fricator cartwrighti Robinson, 1941
Copris fricator fricator
Copris fujiokai Ochi & Kon, 1996
Copris fujiokai poringensis Ochi & Kon, 2005
Copris furciceps Felsche, 1910
Copris garambae Cambefort, 1992
Copris gazellarum Gillet, 1918
Copris gibbulus Lansberge, 1886
Copris gibbulus borneensis Ochi & Kon, 2005
Copris gibbulus gibbulus
Copris gilleti Kolbe, 1907
Copris gladiator Arrow, 1933
Copris gopheri Hubbard, 1894Copris tortoise commensal scarab
Copris gracilis Waterhouse, 1891
Copris graueri (Kolbe, 1914)⇒ Copris camerunus Felsche, 1904
Copris halffteri Matthews, 1959
Copris harrisi Waterhouse, 1891
Copris hartli Frey, 1975
Copris hatayamai Ochi & Araya, 1992
Copris hidakai Ochi & Kon, 1996
Copris hintoni (Saylor, 1933)⇒ Copris klugi Harold, 1869
Copris hispanus Linnaeus, 1764
Copris hispanus cavolinii Petagna, 1792
Copris hispanus hispanus Linné, 1764
Copris hispanus persianus Miksic, 1958
Copris hosakai Ochi & Kon, 2014
Copris howdeni Matthews & H..., 1959
Copris humilis Gillet, 1908
Copris hybridus Gillet, 1910
Copris igualensis Warner, 1990
Copris illotus Balthasar, 1942
Copris imitans Felsche, 1910
Copris inaequabilis Zhang, 1997
Copris incertus Say, 1835Black dung beetle
Copris indicus (Gillet, 1910)⇒ Copris fricator Fabricius, 1787
Copris inemarginatus Blatchley, 1918
Copris inhalatus Quedenfeldt, 1884
Copris inhalatus grandis Nguyen-phung..., 1986
Copris inhalatus inhalatus
Copris inhalatus ngotto Moretto, 2011
Copris inhalatus perturbator Péringuey, 1901
Copris inhalatus sanctaeluciae Nguyen-phung..., 1986
Copris insidiosus Péringuey, 1901
Copris integer (Reiche, 1847)⇒ Copris fallaciosus Gillet, 1907
Copris interioris Kolbe, 1897
Copris iris Sharp, 1875
Copris jacchoides Nguyen-phung..., 1987
Copris jacchus Lang, 1971
Copris jahi Nguyen-phung..., 1986
Copris javanensis Bui, Ziegler..., 2019
Copris johkii Ochi & Kon, 2014
Copris juanmorai Mora-Aguilar..., 2015
Copris jucundus Gillet, 1932
Copris kasagii Ochi & Kon, 1996
Copris kasanka Josso & Prév..., 2016
Copris katangae Cambefort & ..., 1996
Copris keralensis Gill, 1986
Copris kimioi Ochi & Kon, 2021
Copris kiriromensis Ochi & Kon, 2021
Copris kiuchii Masumoto, 1989
Copris klapperichi Balthasar, 1942
Copris klugi Harold, 1869
Copris klugi sierrensis (Matthews, 1961)⇒ Copris sierrensis Matthews, 1961
Copris kusakabei Ochi, Kon & ..., 2005
Copris laeviceps Harold, 1869
Copris laevigatus (Gillet, 1927)⇒ Onthophagus laevigatus Fabricius, 1798
Copris laiiformis Nguyen-phung..., 1987
Copris laioides Boucomont, 1932
Copris laius Harold, 1868
Copris laius angusticornis Nguyen-phung..., 1987
Copris laius laius
Copris lannathai Hanboonsong ..., 2003
Copris laosensis Ochi & Kon, 2021
Copris leakeyorum R. Paulian, 1976
Copris lecontei Matthews, 1961
Copris lecontei isthmiensis Matthews, 1961
Copris lecontei lecontei Matthews, 1962
Copris leruei Josso & Prév..., 2017
Copris lugubris Boheman, 1858
Copris lunaris Linnaeus, 1758Horned Dung Beetle
Copris macacus Lansberge, 1886
Copris macclevei Warner, 1990
Copris macer Péringuey, 1901
Copris maesi Ratcliffe, 1998
Copris magicus Harold, 1881
Copris manni Ochi, Kon & Bai, 2009
Copris marcoberrai Ochi, Kon & ..., 2019
Copris marcus Gillet, 1921
Copris martinae Nguyen-phung, 1988
Copris martinae martinae
Copris martinae ntchisi Moretto, 2011
Copris martinezi Matthews & H..., 1968Streaked blenny
Copris matthewsi Delgado & Ko..., 2001
Copris matthewsi matthewsi
Copris matthewsi pacificus Delgado & Ko..., 2001
Copris medogensis Y.-W. Zhang, 1988
Copris megaceratoides Waterhouse, 1891
Copris megasoma Matthews & H..., 1959
Copris mesacanthus Harold, 1878
Copris mesacanthus mesacanthus
Copris mesacanthus transvaalensis Nguyen-phung, 1988
Copris mexicanus Matthews & H..., 1959
Copris micros Ochi & Kon, 2021
Copris mindorensis Ochi, 1992
Copris minutus Drury, 1773Small Black Dung Beetle
Copris misellus Péringuey, 1901
Copris miyakei Ochi & Kon, 2004
Copris moecha LeConte, 1854
Copris moechus LeConte, 1854
Copris moffartsi Gillet, 1907
Copris mongkhoni Hanboonsong ..., 2003
Copris montivagus (Gillet, 1908)⇒ Copris harrisi Waterhouse, 1891
Copris montreuili Ochi, Kon & ..., 2019
Copris morettoi Josso & Prév..., 2020
Copris moroni Darling & Gé..., 2018
Copris morphaeus Gillet, 1932
Copris mourgliai Zelenka, 1993
Copris muticus (Boheman, 1857)⇒ Litocopris muticus Boheman, 1857
Copris neglectus Moxey, 1963
Copris neotridens Génier & Pré..., 2017
Copris nepos Gillet, 1908
Copris nevinsoni Waterhouse, 1891Warton meshweaver
Copris nonarmatus Ochi & Kon, 2021
Copris novaki Zelenka, 1992
Copris nubilosus Kohlmann & C..., 2003
Copris numa Lansberge, 1886Braken Bat Cave meshweaver
Copris obenbergeri Balthasar, 1933Government Canyon Bat Cave meshweaver
Copris obesus Boheman, 1857
Copris ochus Motschulsky, 1860
Copris olivieri Illiger, 1800
Copris orion Klug, 1835
Copris orion caffer Gillet, 1908
Copris orion centralis Gillet, 1908
Copris orion orion
Copris orion overlaeti Balthasar, 1961
Copris orphanus (Guérin-Ménev..., 1847)⇒ Copris evanidus Klug, 1855
Copris parapecuarius Ochi, Kon & ..., 2008
Copris parasonensis Ochi, Kon & ..., 2019
Copris paraszechouanicus Ochi, Kon & ..., 2018
Copris pauliani Nguyen-phung..., 1986
Copris peculiariprotibialis Ochi & Kon, 2023
Copris pedarioides Lansberge, 1886
Copris perturbator (Péringuey, 1901)⇒ Copris inhalatus perturbator Péringuey, 1901
Copris phungae Cambefort, 1992
Copris phungae gabonicus Cambefort, 1992
Copris phungae phungae
Copris phylax Gillet, 1908
Copris poggii Ochi & Kon, 2005
Copris popaensis Ochi, Kon & ..., 2019
Copris potanini Semenov, 1891
Copris prevosti Josso, 2011
Copris pristinus W. D. Pierce, 1946
Copris pseudobootes Ferreira, 1962
Copris pseudochus Ochi & Kon, 2004
Copris pseudochus basipunctulatus Ochi, Kon & ..., 2020
Copris pseudochus chuduraziensis Ochi, Kon & ..., 2019
Copris pseudochus pseudochus
Copris pseudomoffartsi Moretto, 2011
Copris pseudosinicus Balthasar, 1958
Copris pueli Mollandin De..., 1905
Copris puncticollis Boheman, 1857
Copris punctipennis Boucomont, 1914
Copris punctiventris Waterhouse, 1891
Copris punctulatus Wiedemann, 1823
Copris punctulatus punctulatus
Copris punctulatus septentrionalis Ochi, Kon & ..., 2019
Copris punjabensis Gillet, 1921
Copris quadridens (Say, 1835)⇒ Phanaeus quadridens Say, 1835
Copris quasilaevigatus Ochi, Kon & Bai, 2007
Copris ramosiceps Gillet, 1921
Copris rebouchei Harold, 1869
Copris reflexus (Fabricius, 1787)⇒ Copris minutus Drury, 1773
Copris remotus LeConte, 1866
Copris remotus dicyrtus Matthews & H..., 1959
Copris remotus remotus LeConte, 1866
Copris renwarti Nguyen-phung..., 1986
Copris repertus Walker, 1858
Copris ribbei Lansberge, 1886
Copris riekoae Ochi & Kon, 2003
Copris ritsemae Harold, 1875
Copris ruandanus Balthasar, 1961
Copris rugosus (Gillet, 1908)⇒ Copris inhalatus Quedenfeldt, 1884
Copris ruricola Balthasar, 1933
Copris sabinus Gillet, 1910
Copris sacontala Redtenbacher, 1848
Copris sallei Harold, 1869
Copris sarpedon Harold, 1868
Copris saundersi Harold, 1869
Copris scorpio Zelenka, 1993
Copris serius Nguyen-phung, 1987
Copris sexdentatus (Thunberg, 1818)⇒ Copris sarpedon Harold, 1868
Copris siamensis Gillet, 1910
Copris siangensis Biswas, 1980
Copris sierrensis Matthews, 1961
Copris signatus Walker, 1858
Copris simonettai Cambefort, 1992
Copris simplex (Harold, 1869)⇒ Dichotomius simplex Taschenberg, 1870
Copris simulator Nguyen-phung..., 1986
Copris singularis Nguyen-phung..., 1986
Copris sinicus Hope, 1842
Copris sinon Olivier, 1789
Copris sodalis Walker, 1858
Copris sonensis Bui, Dumack ..., 2018
Copris sorex Balthasar, 1942
Copris sphaeropterus Harold, 1877
Copris spinator Harold, 1881
Copris subdolus Balthasar, 1958
Copris subpunctatus Gillet, 1910
Copris subsidens Péringuey, 1901
Copris subterranea O. Heer, 1862
Copris subterraneus Heer, 1862
Copris sumatrensis Gillet, 1921
Copris surdus Arrow, 1931
Copris susanae Darling & Gé..., 2018
Copris szechouanicus Balthasar, 1958
Copris takanoi Ochi, Kon & Bai, 2020
Copris teramachii Ochi, Kon & Bai, 2020
Copris tetraodon Gillet, 1910
Copris tricorniprothoracicus Ochi, Kon & ..., 2020
Copris tridens (Felsche, 1901)⇒ Copris hispanus hispanus Linné, 1764
Copris tridentatus Solis & Kohl..., 2003
Copris truncatus Felsche, 1901
Copris tsukamotoi Ochi, Kon & ..., 2008
Copris tullius Olivier, 1789
Copris typhoeus Gerstaecker, 1884
Copris umbilicatus Abeille de P..., 1901
Copris urus (Boheman, 1857)⇒ Catharsius molossus Linnaeus, 1758
Copris usambaricus Gillet, 1908
Copris vankhaii Nguyen-phung, 1988
Copris vankhaii genopunctatus Nguyen-phung, 1988
Copris vankhaii vankhaii
Copris vankhalii Nguyen-phung, 1988
Copris victorini (Boheman, 1857)⇒ Copris ritsemae Harold, 1875
Copris vietnamicus Kabakov, 1994
Copris vietnamimontanus Ochi, Kon & ..., 2018
Copris vilhenai Ferreira, 1962
Copris vitalisi Gillet, 1921
Copris vrydaghi Ferreira, 1962
Copris warneri Mccleve & Ko..., 2005
Copris wiesei Kolbe, 1914
Copris xingkeyangi Ochi, Kon & Bai, 2020
Copris yangi Ochi, Kon & Bai, 2007
Copris youngai Balthasar, 1967
Copris zhangi Ochi, Kon & Bai, 2009
Coptodactyla brooksi Matthews, 1976
Coptodactyla depressa Paulian, 1933
Coptodactyla ducalis Blackburn, 1903
Coptodactyla glabricollis Hope, 1842
Coptodactyla lesnei Paulian, 1933
Coptodactyla matthewsi Reid, 2000
Coptodactyla merdeka Reid, 2000
Coptodactyla meridionalis Matthews, 1976
Coptodactyla monstrosa Felsche, 1909
Coptodactyla nitida (Paulian, 1933)⇒ Coptodactyla subaenea Harold, 1877
Coptodactyla onitoides Gillet, 1925
Coptodactyla papua Lansberge, 1885
Coptodactyla stereocera Gillet, 1911
Coptodactyla storeyi Reid, 2000
Coptodactyla subaenea Harold, 1877
Coptodactyla torresica Mathews, 1976
Coptodactyla tuberculata Gillet, 1925
Dichotomius (Homocanthonides) smaragdinus Perty, 1830
Dichotomius (Luederwaldtinia) affinis Felsche, 1910
Dichotomius (Luederwaldtinia) agenor Harold, 1869
Dichotomius (Luederwaldtinia) alvarengai Maldaner, N..., 2015
Dichotomius (Luederwaldtinia) amplicollis Harold, 1869
Dichotomius (Luederwaldtinia) angeloi Nunes, Carv..., 2016
Dichotomius (Luederwaldtinia) apicalis Luederwaldt, 1931
Dichotomius (Luederwaldtinia) assifer Eschscholtz, 1822
Dichotomius (Luederwaldtinia) batesi Harold, 1869
Dichotomius (Luederwaldtinia) bechynei Martínez, 1973
Dichotomius (Luederwaldtinia) belus Harold, 1880
Dichotomius (Luederwaldtinia) benesi Vaz-de-Mello..., 2016
Dichotomius (Luederwaldtinia) bosqui Pereira, 1941
Dichotomius (Luederwaldtinia) carbonarius Mannerheim, 1829
Dichotomius (Luederwaldtinia) catimbau Valois, Vaz-..., 2017
Dichotomius (Luederwaldtinia) centralis Harold, 1869
Dichotomius (Luederwaldtinia) crinicollis Germar, 1824
Dichotomius (Luederwaldtinia) cuprinus Felsche, 1901
Dichotomius (Luederwaldtinia) danieli (Kohlmann & S..., 1997)⇒ Luederwaldtinia danieli Kohlmann & S..., 1997
Dichotomius (Luederwaldtinia) deyrollei Harold, 1869
Dichotomius (Luederwaldtinia) eucranioides (Pereira & An..., 1955)⇒ Luederwaldtinia eucranioides Pereira & An..., 1955
Dichotomius (Luederwaldtinia) fallax Harold, 1880
Dichotomius (Luederwaldtinia) favi (Kohlmann & S..., 1997)⇒ Luederwaldtinia favi Kohlmann & S..., 1997
Dichotomius (Luederwaldtinia) femoratus (Howden & Young, 1981)⇒ Luederwaldtinia femoratus Howden & Young, 1981
Dichotomius (Luederwaldtinia) fimbriatus Harold, 1869
Dichotomius (Luederwaldtinia) fortepunctatus Luederwaldt, 1923
Dichotomius (Luederwaldtinia) fortestriatus Luederwaldt, 1924
Dichotomius (Luederwaldtinia) gamboaensis (Howden & Young, 1981)⇒ Luederwaldtinia gamboaensis Howden & Young, 1981
Dichotomius (Luederwaldtinia) geminatus Arrow, 1913
Dichotomius (Luederwaldtinia) gilletti Valois, Vaz-..., 2017
Dichotomius (Luederwaldtinia) glaucus Harold, 1869
Dichotomius (Luederwaldtinia) globulus Felsche, 1901
Dichotomius (Luederwaldtinia) guaribensis Valois, Vaz-..., 2017
Dichotomius (Luederwaldtinia) hempeli Pereira, 1942
Dichotomius (Luederwaldtinia) iannuzziae Valois, Vaz-..., 2017
Dichotomius (Luederwaldtinia) inachoides Felsche, 1901
Dichotomius (Luederwaldtinia) inachus Erichson, 1847
Dichotomius (Luederwaldtinia) interstitialis Luederwaldt, 1931
Dichotomius (Luederwaldtinia) irinus Harold, 1867
Dichotomius (Luederwaldtinia) laevicollis Felsche, 1901
Dichotomius (Luederwaldtinia) louzadai Nunes, Carv..., 2016
Dichotomius (Luederwaldtinia) lucasi Harold, 1869
Dichotomius (Luederwaldtinia) luctuosus Harold, 1869
Dichotomius (Luederwaldtinia) lycas Felsche, 1901
Dichotomius (Luederwaldtinia) machadoi Martinez & P..., 1967
Dichotomius (Luederwaldtinia) malyi Maldaner, N..., 2015
Dichotomius (Luederwaldtinia) micans Luederwaldt, 1924
Dichotomius (Luederwaldtinia) muticus Luederwaldt, 1922
Dichotomius (Luederwaldtinia) mysticus Luederwaldt, 1935
Dichotomius (Luederwaldtinia) nisus Olivier, 1789
Dichotomius (Luederwaldtinia) nobilis Waterhouse, 1891
Dichotomius (Luederwaldtinia) ocellatopunctatus Felsche, 1901
Dichotomius (Luederwaldtinia) opacipennis Luederwaldt, 1931
Dichotomius opacipennis fornicatus Luederwaldt, 1931
Dichotomius (Luederwaldtinia) opalescens Felsche, 1910
Dichotomius (Luederwaldtinia) parcepunctatus Felsche, 1901
Dichotomius (Luederwaldtinia) problematicus Luederwaldt, 1924
Dichotomius problematicus planus Luederwaldt, 1924
Dichotomius (Luederwaldtinia) punctatostriatus (Felsche, 1901)⇒ Luederwaldtinia punctatostriatus Felsche, 1901
Dichotomius (Luederwaldtinia) rodrigoi (Kohlmann & S..., 1997)⇒ Luederwaldtinia rodrigoi Kohlmann & S..., 1997
Dichotomius (Luederwaldtinia) rugatus (Luederwaldt, 1935)⇒ Luederwaldtinia rugatus Luederwaldt, 1935
Dichotomius (Luederwaldtinia) schiffleri Vaz-de-Mello..., 2001
Dichotomius (Luederwaldtinia) semisquamosus Curtis, 1845
Dichotomius (Luederwaldtinia) sericeus Harold, 1867
Dichotomius (Luederwaldtinia) sexdentatus Luederwaldt, 1925
Dichotomius (Luederwaldtinia) simplicicornis Luederwaldt, 1935
Dichotomius (Luederwaldtinia) speciosus Waterhouse, 1891
Dichotomius (Luederwaldtinia) superbus Felsche, 1901
Dichotomius (Luederwaldtinia) texanus (Felsche, 1901)⇒ Luederwaldtinia texanus Felsche, 1901
Dichotomius (Luederwaldtinia) triquetrus Luederwaldt, 1924
Dichotomius (Luederwaldtinia) tristis Luederwaldt, 1924
Dichotomius (Luederwaldtinia) validipilosus Luederwaldt, 1931
Dichotomius (Luederwaldtinia) vidaurrei Nunes & Vaz-..., 2013
Dichotomius (Luederwaldtinia) yucatanus Bates, 1887
Dichotomius (Luederwaldtinia) ziliolii Gandini & Ag..., 2009
Dichotomius (Selenocopris) acuticornis Luederwaldt, 1930
Dichotomius (Selenocopris) aielloae Nunes, 2016
Dichotomius (Selenocopris) ascanius Harold, 1869
Dichotomius ascanius aceratus Pereira, 1944
Dichotomius ascanius piceus Luederwaldt, 1930
Dichotomius (Selenocopris) bicuspis Germar, 1824
Dichotomius (Selenocopris) darwini Nunes & Vaz-..., 2016
Dichotomius (Selenocopris) enrietti (Vernalha, 1952)⇒ Selenocopris enrietti Vernalha, 1952
Dichotomius (Selenocopris) fissiceps Felsche, 1901
Dichotomius (Selenocopris) fissus Harold, 1867
Dichotomius (Selenocopris) fonsecae Luederwaldt, 1925
Dichotomius (Selenocopris) guaranii Gandini & Ag..., 2009
Dichotomius (Selenocopris) inflaticollis Felsche, 1910
Dichotomius (Selenocopris) ingens Luederwaldt, 1935
Dichotomius (Selenocopris) manni Nunes & Vaz-..., 2016
Dichotomius (Selenocopris) mundus Harold, 1869
Dichotomius (Selenocopris) parisi Nunes & Vaz-..., 2016
Dichotomius (Selenocopris) pauloensis Luederwaldt, 1925
Dichotomius (Selenocopris) periotoi Nunes & Vaz-..., 2016
Dichotomius (Selenocopris) punctulatipennis Luederwaldt, 1930
Dichotomius (Selenocopris) pygidialis Luederwaldt, 1922
Dichotomius (Selenocopris) quadraticeps Felsche, 1901
Dichotomius (Selenocopris) rugosipennis Luederwaldt, 1930
Dichotomius (Selenocopris) simplex (Taschenberg, 1870)⇒ Selenocopris simplex Taschenberg, 1870
Dichotomius (Selenocopris) simulans Luederwaldt, 1925
Dichotomius (Selenocopris) singularis Felsche, 1907
Dichotomius (Selenocopris) socius Luederwaldt, 1929
Dichotomius (Selenocopris) spadiceus Luederwaldt, 1928
Dichotomius achamas Harold, 1867
Dichotomius adrastus Harold, 1875
Dichotomius agesilaus Waterhouse, 1891
Dichotomius alutaceus Felsche, 1901
Dichotomius alyattes Harold, 1880
Dichotomius amicitiae Kohlmann & S..., 1997
Dichotomius andresi Sarmiento-Ga..., 2014
Dichotomius annae Linnaeus, 1767
Dichotomius anthrax Felsche, 1901
Dichotomius arnaudi Nunes & Vaz-..., 2019
Dichotomius asenjoi Nunes & Vaz-..., 2019
Dichotomius australis Luederwaldt, 1931
Dichotomius baiano Nunes & Vaz-..., 2019
Dichotomius barbarae Nunes & Vaz-..., 2019
Dichotomius berthalutzae Valois, Silv..., 2022
Dichotomius bicornis Waterhouse, 1891
Dichotomius bitiensis Gillet, 1911
Dichotomius blancoi Nunes & Vaz-..., 2019
Dichotomius boreus Olivier, 1789
Dichotomius borgmeieiri Pereira & Ma..., 1959
Dichotomius borgmeieri (Pereira & Ma..., 1959)⇒ Dichotomius borgmeieiri Pereira & Ma..., 1959
Dichotomius bos Blanchard, 1846
Dichotomius bucki (Pereira, 1953)⇒ Luederwaldtinia bucki Pereira, 1953
Dichotomius buqueti Lucas, 1857
Dichotomius calcaratus Arrow, 1913
Dichotomius camargoi Martínez, 1956
Dichotomius camposeabrai Martínez, 1974
Dichotomius candango Nunes & Vaz-..., 2019
Dichotomius carinatus Luederwaldt, 1925
Dichotomius carolinus Linnaeus, 1767Carolina copris
Dichotomius clypeatus Luederwaldt, 1929
Dichotomius coenosus Erichson, 1848
Dichotomius colonicus (Say, 1835)⇒ Dichotomius carolinus Linnaeus, 1767
Dichotomius comarapensis Génier, 2000
Dichotomius compressicollis Luederwaldt, 1929
Dichotomius conicollis Blanchard, 1846
Dichotomius costaricensis Luederwaldt, 1935
Dichotomius cotopaxi Guérin-Ménev..., 1855
Dichotomius dahli Landin, 1956
Dichotomius depressicollis Harold, 1867
Dichotomius diabolicus Harold, 1875
Dichotomius divergens Luederwaldt, 1923
Dichotomius dynastus Gandini & Ag..., 2009
Dichotomius edmondsi Nunes & Vaz-..., 2019
Dichotomius emas Nunes & Vaz-..., 2019
Dichotomius enioi Montoya-Moli..., 2021
Dichotomius feeri Nunes & Vaz-..., 2019
Dichotomius fernandosilvai Nunes & Vaz-..., 2019
Dichotomius filgueirasi Nunes & Vaz-..., 2019
Dichotomius fornicatus Luederwaldt, 1931
Dichotomius foveicollis Kirsch, 1871
Dichotomius frizzasae Nunes & Vaz-..., 2019
Dichotomius furtadoi Nunes & Vaz-..., 2019
Dichotomius gandinii Rossini & Va..., 2020
Dichotomius gardneri Nunes & Vaz-..., 2019
Dichotomius geraldimi Maldaner & V..., 2021
Dichotomius haroldi Waterhouse, 1891
Dichotomius henripittieri Montoya-Moli..., 2021
Dichotomius horridus Felsche, 1911
Dichotomius horvathi Balthasar, 1970
Dichotomius imitator Felsche, 1901
Dichotomius itatiaiaensis Nunes & Vaz-..., 2019
Dichotomius joelus Gandini & Ag..., 2009
Dichotomius latilobatus Boilly & Vaz..., 2021
Dichotomius latistriatus (Luederwaldt, 1935)⇒ Luederwaldtinia latistriatus Luederwaldt, 1935
Dichotomius longiceps Taschenberg, 1870
Dichotomius lucianomourai Nunes & Vaz-..., 2019
Dichotomius luctuosioides Luederwaldt, 1922
Dichotomius mamillatus Felsche, 1901
Dichotomius marcoscarvalhoi Nunes & Vaz-..., 2019
Dichotomius maya Peraza & Deloya, 2006
Dichotomius melzeri Luederwaldt, 1922
Dichotomius missionus Martínez, 1947
Dichotomius monstrosus Harold, 1875
Dichotomius mormon Ljungh, 1799
Dichotomius motai Pereira, 1942
Dichotomius nemoricola Pereira, 1942
Dichotomius nevermanni Luederwaldt, 1935
Dichotomius nimuendaju Luederwaldt, 1925
Dichotomius nimuendajui Luederwaldt, 1925
Dichotomius nitidissimus Waterhouse, 1891
Dichotomius nutans Harold, 1867
Dichotomius ohausi Luederwaldt, 1923
Dichotomius paschoali Nunes & Vaz-..., 2019
Dichotomius planicollis Gillet, 1911
Dichotomius podalirius Felsche, 1901
Dichotomius prietoi Martinez & M..., 1982
Dichotomius protectus Harold, 1867
Dichotomius provisorius Luederwaldt, 1925
Dichotomius puncticollis Luederwaldt, 1935
Dichotomius quadrilobatus Chamorro, Lo..., 2021
Dichotomius quadrinodosus Felsche, 1901
Dichotomius quinquedens Felsche, 1910
Dichotomius quinquelobatus Felsche, 1901
Dichotomius rafanunezi Montoya-Moli..., 2021
Dichotomius reclinatus Felsche, 1901
Dichotomius reichei Harold, 1869
Dichotomius ribeiroi Pereira, 1954
Dichotomius ricardosilvai Nunes & Vaz-..., 2019
Dichotomius robustus Luederwaldt, 1935Apex mesh weaver
Dichotomius rondoniaensis Nunes & Vaz-..., 2019
Dichotomius rotundatus Burmeister, 1874
Dichotomius rugosicollis Luederwaldt, 1935
Dichotomius sagittarius Harold, 1869
Dichotomius satanas Harold, 1867
Dichotomius satanas angustus Luederwaldt, 1923
Dichotomius satanas satanas
Dichotomius seag Nunes & Vaz-..., 2019
Dichotomius semiaeneus Germar, 1824
Dichotomius semicircularis Luederwaldt, 1929
Dichotomius subaeneus Castelnau, 1840
Dichotomius telamon Harold, 1869
Dichotomius tissianiae Nunes & Vaz-..., 2019
Dichotomius torulosus Luederwaldt, 1925
Dichotomius travassosi (Martínez, 1955)⇒ Dichotomius nitidissimus Waterhouse, 1891
Dichotomius triangulariceps Blanchard, 1846
Dichotomius valoisae Silva, Moura..., 2020
Dichotomius vargasae Nunes & Vaz-..., 2019
Dichotomius virescens Luederwaldt, 1935
Dichotomius woodruffi Solis & Kohl..., 2022
Dichotomius worontzowi Pereira, 1942
Dichotomius zikani Luederwaldt, 1922
Heliocopris alatus Felsche, 1910
Heliocopris anadematus Gillet, 1908
Heliocopris andersoni Bates, 1868
Heliocopris anguliceps Janssens, 1943
Heliocopris antenor (Olivier, 1789)⇒ Heliocopris colossus Bates, 1868
Heliocopris antiquus I. Fujiyama, 1968
Heliocopris ares Moretto & Mi..., 2022
Heliocopris atropos Boheman, 1860
Heliocopris beccarii Harold, 1871
Heliocopris biimpressus Kolbe, 1893
Heliocopris bucephalus Fabricius, 1775
Heliocopris camerunus Pokorny & Zidek, 2009
Heliocopris colossus Bates, 1868
Heliocopris corniculatus Janssens, 1939
Heliocopris coronatus Felsche, 1901
Heliocopris cuneifer Lesne, 1906
Heliocopris densissa Roth, 1851
Heliocopris dianae (Hope, 1842)⇒ Heliocopris mutabilis Kolbe, 1893
Heliocopris dilloni Guérin-Ménev..., 1847
Heliocopris dolosus Janssens, 1939
Heliocopris dominus Bates, 1868
Heliocopris erycoides Felsche, 1907
Heliocopris eryx (Fabricius, 1801)⇒ Heliocopris neptunus Boheman, 1857
Heliocopris eryx eryx
Heliocopris eryx ikelenge Moretto, 2017
Heliocopris eryx mutabilis (Kolbe, 1895)⇒ Heliocopris mutabilis Kolbe, 1893
Heliocopris eryx tridentatus Moretto, 2017
Heliocopris faunus Boheman, 1857
Heliocopris felschei Kolbe, 1904
Heliocopris fischeri Schäfer, 2019
Heliocopris fonsecai Ferreira, 1967
Heliocopris furcithorax Müller, 1941
Heliocopris gigas Linnaeus, 1758
Heliocopris gigas erythraeus Müller, 1941
Heliocopris gigas gigas
Heliocopris hamadryas Fabricius, 1775
Heliocopris hamifer Harold, 1878
Heliocopris haroldi Kolbe, 1893
Heliocopris helleri Felsche, 1907
Heliocopris hermes Gillet, 1911
Heliocopris hunteri Waterhouse, 1891
Heliocopris isidis (Boheman, 1857)⇒ Heliocopris andersoni Bates, 1868
Heliocopris japetus Klug, 1855
Heliocopris kolbei Felsche, 1901
Heliocopris letiranti Moretto, 2021
Heliocopris maou Moretto, 2014
Heliocopris marshalli Péringuey, 1901
Heliocopris midas Fabricius, 1775
Heliocopris mimus Janssens, 1939
Heliocopris minos Gillet, 1907
Heliocopris mutabilis Kolbe, 1893
Heliocopris myrmidon Kolbe, 1893
Heliocopris neptuniformis Felsche, 1907
Heliocopris neptunoides Janssens, 1939
Heliocopris neptunus Boheman, 1857
Heliocopris pauliani Janssens, 1939
Heliocopris pirmal Fabricius, 1798
Heliocopris quinqueangulatus Janssens, 1939
Heliocopris samson Harold, 1878
Heliocopris sirius (Gillet, 1925)⇒ Heliocopris hamifer Harold, 1878
Heliocopris smithi Moretto, 2017
Heliocopris solitarius Kolbe, 1893
Heliocopris staudingeri Kolbe, 1893
Heliocopris stroehlei Moretto, 2014
Heliocopris sylvanus Gillet, 1925
Heliocopris tyrannus Thomson, 1859
Heliocopris werneri Moretto & Mi..., 2013
Holocephalus cristatus Gillet, 1907
Holocephalus eridanus Olivier, 1789
Holocephalus julieni Smith & Génier, 2001
Holocephalus pokornyi Nunes & Vaz-..., 2016
Holocephalus sculptus Gillet, 1907
Holocephalus simoni Smith & Génier, 2001
Isocopris foveolatus Luederwaldt, 1931
Isocopris hypocrita Lucas, 1859
Isocopris imitator Felsche, 1901
Isocopris inhiatus Germar, 1824
Isocopris nitidus Luederwaldt, 1922
Isocopris tarsalis Luederwaldt, 1931
Isocopris xacriaba Rossini & Va..., 2017
Litocopris muticus Boheman, 1857
Litocopris punctiventris Waterhouse, 1891
Litocopris ruandanus Balthasar, 1961
Litocopris simplex Harold, 1869
Macroderes amplior Frolov & Sch..., 2004
Macroderes arrowi Janssens, 1939
Macroderes bias Olivier, 1789
Macroderes cederbergensis Abdalla & De..., 2018
Macroderes cornutus Frolov & Sch..., 2004
Macroderes dubius Péringuey, 1901
Macroderes endroedyi Frolov & Sch..., 2004
Macroderes fornicatus Sharp, 1880
Macroderes foveatus Frolov & Sch..., 2004
Macroderes gifboomi Abdalla & De..., 2018
Macroderes greeni Kirby, 1818
Macroderes leipoldti Abdalla & De..., 2018
Macroderes minutus Frolov & Sch..., 2004
Macroderes mutilans Kolbe, 1908
Macroderes mutilatus Kolbe, 1908
Macroderes namakwanus Frolov & Sch..., 2004
Macroderes nitidus Harold, 1877
Macroderes oreatus Abdalla & De..., 2018
Macroderes politulus (Preudhomme d..., 1880)⇒ Macroderes bias Olivier, 1789
Macroderes porselinus Abdalla & De..., 2018
Macroderes pristinus Sharp, 1880
Macroderes soleiana Abdalla & De..., 2018
Macroderes spectabilis Péringuey, 1901
Macroderes tortuosus Abdalla & De..., 2018
Macroderes undulatus Preudhomme d..., 1880
Metacatharsius abortivus Fairmaire, 1891
Metacatharsius anderseni Waterhouse, 1891
Metacatharsius anomalus Felsche, 1910
Metacatharsius auberti Fairmaire, 1891
Metacatharsius basilewskyi Balthasar, 1961
Metacatharsius bateke Walter, 1976
Metacatharsius benadirensis Müller, 1947
Metacatharsius bidentatus Ferreira, 1964
Metacatharsius brevicosta Müller, 1947
Metacatharsius clypeolatus Balthasar, 1940
Metacatharsius convexiusculus Shipp, 1897
Metacatharsius darfurensis Balthasar, 1965
Metacatharsius dentinum Ferreira, 1964
Metacatharsius duplicatus Müller, 1941
Metacatharsius exiguiformis Ferreira, 1964
Metacatharsius exiguus Boheman, 1860
Metacatharsius ferreirae Balthasar, 1965
Metacatharsius ferrugineus Olivier, 1789
Metacatharsius freyi Ferreira, 1964
Metacatharsius gardoensis Balthasar, 1941
Metacatharsius holyi Balthasar, 1965
Metacatharsius imitator Balthasar, 1965
Metacatharsius inermis Castelnau, 1840
Metacatharsius janssensi Frey, 1963
Metacatharsius kindiai Paulian, 1939
Metacatharsius latifrons Harold, 1868
Metacatharsius lomii Müller, 1941
Metacatharsius marani Balthasar, 1940
Metacatharsius minutus Lansberge, 1882
Metacatharsius modestus Balthasar, 1965
Metacatharsius murati Paulian, 1939
Metacatharsius nubiensis Harold, 1878
Metacatharsius omoensis Müller, 1941
Metacatharsius opacus Waterhouse, 1891
Metacatharsius ovulum Balthasar, 1940
Metacatharsius parcepunctatus Balthasar, 1965
Metacatharsius patrizii Müller, 1947
Metacatharsius peleus Olivier, 1789
Metacatharsius pernitidus Müller, 1941
Metacatharsius perpunctatus Balthasar, 1965
Metacatharsius petrovitzi Balthasar, 1965
Metacatharsius politus Paulian, 1934
Metacatharsius pollicatus Harold, 1878
Metacatharsius pseudoopacus Ferreira, 1965
Metacatharsius pumilioniformis Ferreira, 1964
Metacatharsius pumilionis Wallengren, 1881
Metacatharsius pusio Kolbe, 1908
Metacatharsius renaudi Frey, 1960
Metacatharsius rochai Ferreira, 1964
Metacatharsius rosinae Balthasar, 1970
Metacatharsius rubidus Fairmaire, 1891
Metacatharsius rugosipennis Frey, 1975
Metacatharsius rusingae R. Paulian, 1976
Metacatharsius seminulum Balthasar, 1940
Metacatharsius simulator Balthasar, 1939
Metacatharsius tchadensis Balthasar, 1961
Metacatharsius tenellus Müller, 1947
Metacatharsius togoensis Kolbe, 1893
Metacatharsius transvaalensis Balthasar, 1968
Metacatharsius trianguliceps Balthasar, 1965
Metacatharsius tricarenatus Ferreira, 1964
Metacatharsius troglodytes (Boheman, 1857)⇒ Metacatharsius anderseni Waterhouse, 1891
Metacatharsius tuberifrons Fairmaire, 1884
Metacatharsius useramus Kolbe, 1893
Metacatharsius wittei Ferreira, 1964
Metacatharsius zuluanus Balthasar, 1965
Ontherus (Caelontherus) mexicanus Harold, 1868
Ontherus aequatorius Bates, 1891
Ontherus alexis Blanchard, 1846
Ontherus amplector Génier, 1996
Ontherus androgynus Génier, 1996
Ontherus aphodioides Burmeister, 1874
Ontherus appendiculatus Mannerheim, 1829
Ontherus ashei Génier, 1996
Ontherus atlantidis Génier, 1996
Ontherus azteca Harold, 1869
Ontherus brevicollis Kirsch, 1870
Ontherus brevipennis Harold, 1867
Ontherus bridgesi Waterhouse, 1891
Ontherus cambeforti Génier, 1996
Ontherus carinicollis Luederwaldt, 1930
Ontherus carinifrons Luederwaldt, 1930
Ontherus cephalotes Harold, 1869
Ontherus compressicornis Luederwaldt, 1931
Ontherus contractus (Burmeister, 1874)⇒ Ontherus appendiculatus Mannerheim, 1829
Ontherus dentatus Luederwaldt, 1930
Ontherus diabolicus Génier, 1996
Ontherus didymus (Erichson, 1847)⇒ Ontherus alexis Blanchard, 1846
Ontherus digitatus Harold, 1868
Ontherus edentulus Génier, 1996
Ontherus elegans Luederwaldt, 1930
Ontherus erosioides Luederwaldt, 1930
Ontherus erosus Harold, 1875
Ontherus felicitae González & M..., 2015
Ontherus gilli Génier, 1996
Ontherus gladiator Génier, 1998
Ontherus grandis Luederwaldt, 1931
Ontherus hadros Génier, 1996
Ontherus howdeni Génier, 1996
Ontherus incisus Kirsch, 1870
Ontherus insolitus Génier, 1996
Ontherus irinus Balthasar, 1938
Ontherus kirschii Harold, 1867
Ontherus laminifer Balthasar, 1938
Ontherus lichyi Martínez, 1947
Ontherus lobifrons Génier, 1996
Ontherus lunicollis Génier, 1996
Ontherus magnus Génier, 1996
Ontherus monilistriatus Génier, 1996
Ontherus obliquus Génier, 1996
Ontherus pilatus Génier, 1996
Ontherus planus Génier, 1996
Ontherus podiceps Harold, 1868
Ontherus politus Génier, 1996
Ontherus pseudodidymus Génier, 1996
Ontherus pubens Génier, 1996
Ontherus raptor Génier, 1996
Ontherus rectangulidens Génier, 1996
Ontherus rectus Génier, 1996
Ontherus sanctaemartae Génier, 1996
Ontherus sextuberculatus Génier, 1996
Ontherus stridulator Génier, 1996
Ontherus strius (Howden & Young, 1981)⇒ Ontherus azteca Harold, 1869
Ontherus sulcator Fabricius, 1775
Ontherus tenuistriatus Génier, 1996
Ontherus trituberculatus Balthasar, 1938
Ontherus ulcopygus Génier, 1996
Ontherus virescens Lucas, 1859
Ontherus zikani Luederwaldt, 1930
Pseudocopris lunaticeps Gillet, 1910
Pseudocopris tubericollis Felsche, 1910
Synapsis aquisgranensis Forster, 1869
Synapsis birmanicus Gillet, 1907
Synapsis boonlongi Hanboonsong ..., 1999
Synapsis brahminus Hope, 1831
Synapsis cambeforti (Ochi, Kon & ..., 2008)⇒ Synapsis ritsemae Lansberge, 1874
Synapsis davidi (Fairmaire, 1878)⇒ Synapsis davidis Fairmaire, 1878
Synapsis davidis Fairmaire, 1878
Synapsis dickinsoni Hanboonsong ..., 1999
Synapsis gilleti Arrow, 1931
Synapsis horaki Zidek & Pokorny, 2010
Synapsis kiuchii Hanboonsong ..., 1999
Synapsis masumotoi Ochi, 1992
Synapsis naxiorum Kral & Rejsek, 2000
Synapsis ochii Masumoto, 1995
Synapsis ovalis Boucomont, 1920
Synapsis puluongensis Bui & Bonkowski, 2018
Synapsis punctatus Ochi, Kon & ..., 2008
Synapsis ritsemae Lansberge, 1874
Synapsis roslihashimi Ochi, Kon & ..., 2008
Synapsis satoi Ochi & Kon, 2007
Synapsis simplex Sharp, 1875
Synapsis strnadi Kral, 2002
Synapsis thoas (Sharp, 1875)⇒ Synapsis ritsemae Lansberge, 1874
Synapsis tmolus Fischer von ..., 1821
Synapsis tridens Sharp, 1881
Synapsis vietnamicus Ochi, Kon & ..., 2020
Synapsis yama Gillet, 1911
Synapsis yunnanus Arrow, 1933
Thyregis kershawi Blackburn, 1904
Thyregis monteithi Matthews, 1976
Thyregis relictus Matthews, 1976
Thyregis tarsatus Paulian, 1933
Uroxys angulatus Harold, 1868
Uroxys angulicollis Boheman, 1858
Uroxys aterrimus Harold, 1867
Uroxys bahianus Boucomont, 1927
Uroxys batesi Harold, 1868
Uroxys besti Ratcliffe, 1980
Uroxys bidentis (Howden & Young, 1981)⇒ Uroxys boneti Pereira & Ha..., 1961
Uroxys boneti Pereira & Ha..., 1961
Uroxys bonplandi Kohlmann, So..., 2019
Uroxys brachialis Arrow, 1933
Uroxys brevis Waterhouse, 1891
Uroxys catharinensis Balthasar, 1966
Uroxys caucanus Arrow, 1933
Uroxys chichanich Delgado & Ko..., 2007
Uroxys coarctatus Harold, 1867
Uroxys corniculatus Harold, 1880
Uroxys corporaali Balthasar, 1940
Uroxys cuprescens Westwood, 1842
Uroxys deavilai Delgado & Ko..., 2007
Uroxys depressifrons (Howden & Young, 1981)⇒ Uroxys pauliani Balthasar, 1940
Uroxys dilaticollis Blanchard, 1846
Uroxys dureti (Martínez, 1947)⇒ Uroxys epipleurale Boucomont, 1928
Uroxys dybasi Howden & Young, 1981
Uroxys elongatus Harold, 1868
Uroxys epipleurale Boucomont, 1928
Uroxys frankenbergeri Balthasar, 1940
Uroxys gatunensis Howden & Young, 1981
Uroxys gorgon Arrow, 1933
Uroxys histeroides Arrow, 1933
Uroxys inconspicuus Harold, 1868
Uroxys kratochvili Balthasar, 1940
Uroxys laevipennis Kirsch, 1870
Uroxys latesulcatus Bates, 1891
Uroxys latus Arrow, 1933
Uroxys lobata
Uroxys lojanus Arrow, 1933
Uroxys macrocularis (Howden & Young, 1981)⇒ Uroxys boneti Pereira & Ha..., 1961
Uroxys magnus Balthasar, 1940
Uroxys metagorgon Howden & Young, 1981
Uroxys metallescens Harold, 1868
Uroxys microcularis Howden & Young, 1981
Uroxys micros Bates, 1887
Uroxys minutus Harold, 1868
Uroxys monstrosus Balthasar, 1940
Uroxys nebulinus Howden & Gill, 1987
Uroxys ohausi Balthasar, 1938
Uroxys pauliani Balthasar, 1940
Uroxys peruanus Balthasar, 1940
Uroxys platypyga Howden & Young, 1981
Uroxys productus Arrow, 1933
Uroxys pygmaeus Harold, 1883
Uroxys rugatus Boucomont, 1928
Uroxys simplex Waterhouse, 1891
Uroxys spaethi Balthasar, 1940
Uroxys striatus Harold, 1868
Uroxys sulai Balthasar, 1940
Uroxys sulcicollis Harold, 1880
Uroxys tacanensis Delgado & Ko..., 2007
Uroxys terminalis Waterhouse, 1891
Uroxys thoracalis Balthasar, 1940
Uroxys transversifrons Howden & Gill, 1987
Uroxys trinitatis Arrow, 1933
Uroxys tuberculatus Lansberge, 1874
Uroxys variabilis Robinson, 1951
Uroxys vincentiae Arrow, 1903
Xinidium davisi Cambefort, 1985
Xinidium dentilabris Harold, 1869
Xinidium dewitzi Harold, 1878
Xinidium howdeni Cambefort, 1993


Links, sources and further information
Coprini at artfakta.se
Coprini at dyntaxa.se
Coprini at itis.gov