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| | | | Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles)
and region Continents
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 2110
116133
1222976 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 2101
115931
1221959 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 1812
113780
1210168 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 929
84484
1007160 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 234
35522
419905 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 189
32170
363924 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 23
3594
50225 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 23
3594
50225 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 2963
42190 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 289
7423 | Genera
Species |
Taxom
|
Genera for Continents
289 Genera with 7423 Species #ABCDEFGHIJKLMNOPSTUVWXYZ
| Acanthonitis Janssens, 2012 | | 4 | Species | Taxom | | Afrodrepanus Krikken, 1921 | | 2 | Species | Taxom | | Agamopus Bates, 1928 | | 6 | Species | Taxom | | Aleiantus Olsoufieff, 1898 | | 4 | Species | Taxom | | Aliuscanthoniola Deschodt & S..., 2008 | | 1 | Species | Taxom | | Allogymnopleurus Janssens, 1941 | | 20 | Species | Taxom | | Allonitis Janssens, 1907 | | 1 | Species | Taxom | | Alloscelus Boucomont, 1923 | | 4 | Species | Taxom | | Amietina Cambefort, 1989 | | 4 | Species | Taxom | | Amphistomus van Lansberge, 1985 | | 25 | Species | Taxom | | Anachalcos Hope, 1901 | | 6 | Species | Taxom | | Anisocanthon Martinez & P..., 1868 | | 4 | Species | Taxom | | Anoctus Sharp, 1942 | | 5 | Species | Taxom | | Anomiopsis Westwood, 1944 | | 2 | Species | Taxom | | Anomiopsoides Blackwelder, 1945 | | 8 | Species | Taxom | | Anomiopus Westwood, 2006 | | 63 | Species | Taxom | | Anonthobium Paulian & Pl..., 1903 | | 6 | Species | Taxom | | Anonychonitis Janssens, 1950 | | 1 | Species | Taxom | | Anoplodrepanus Simonis, 1966 | | 2 | Species | Taxom | | Aphengium Harold, 1868 | | 4 | Species | Taxom | | Aphengoecus Péringuey, 1987 | | 2 | Species | Taxom | | Apotolamprus Olsofieff, 2008 | | 32 | Species | Taxom | | Aptenocanthon Matthews, 2000 | | 8 | Species | Taxom | | Apterepilissus Montreuil, 1958 | | 4 | Species | Taxom | | Aptychonitis Janssens, 1868 | | 2 | Species | Taxom | | Arachnodes Westwood, 2011 | | 63 | Species | Taxom | | Ateuchites Meunier, 1898 | | 1 | Species | Taxom | | Ateuchus Weber, 1996 | | 111 | Species | Taxom | | Attavicinus | | 1 | Species | Taxom | | Aulacopris White, 1859 | | 3 | Species | Taxom | | Baloghonthobium Paulian, 1986 | | 1 | Species | Taxom | | Bdelyropsis Pereira, Vul..., 1976 | | 3 | Species | Taxom | | Bdelyrus Harold, 1998 | | 28 | Species | Taxom | | Besourenga Vaz-de-Mello, 1939 | | 5 | Species | Taxom | | Bohepilissus Paulian, 1857 | | 2 | Species | Taxom | | Bolbites Harold, 1868 | | 1 | Species | Taxom | | Boletoscapter Matthews, 1974 | | 2 | Species | Taxom | | Boreocanthon Halffter, 1857 | | 13 | Species | Taxom | | Bradypodidium Vaz-de-Mello, 1993 | | 4 | Species | Taxom | | Bubas Mulsant, 1811 | | 3 | Species | Taxom | | Byrrhidium Harold, 1869 | | 3 | Species | Taxom | | Caccobius Thomson, 2014 | | 124 | Species | Taxom | | Caeconthobium Paulian, 1984 | | 1 | Species | Taxom | | Cambefortantus Paulian, 1975 | | 8 | Species | Taxom | | Cambefortius Branco, 1936 | | 8 | Species | Taxom | | Canthidium Erichson, 1886 | | 184 | Species | Taxom | | Canthochilum Chapin, 2008 | | 29 | Species | Taxom | | Canthodimorpha Davis, Schol..., 1999 | | 1 | Species | Taxom | | Canthon Hoffmannsegg, 1999 | Tumblebug | 260 | Species | Taxom | | Canthonella Chapin, 1867 | | 19 | Species | Taxom | | Canthonidia Paulian, 1846 | | 2 | Species | Taxom | | Canthonosoma MacLeay, 1871 | | 4 | Species | Taxom | | Canthotrypes Paulian, 1939 | | 1 | Species | Taxom | | Cassolus Sharp, 1885 | | 9 | Species | Taxom | | Catharsiocopris | | 1 | Species | Taxom | | Catharsius Hope, 1857 | | 118 | Species | Taxom | | Cephalodesmius Westwood, 1871 | | 3 | Species | Taxom | | Chalcocopris Burmeister, 2015 | | 2 | Species | Taxom | | Cheironitis van Lansberge, 1918 | | 28 | Species | Taxom | | Chironitis Lansberge, 1849 | | 5 | Species | Taxom | | Choeridium Serville, 1948 | | 6 | Species | Taxom | | Circellium Latreille, 1781 | | 1 | Species | Taxom | | Cleptocaccobius Cambefort, 1967 | | 26 | Species | Taxom | | Clypeodrepanus Krikken, 1953 | | 3 | Species | Taxom | | Copridaspidus Bloucomont, 1878 | | 1 | Species | Taxom | | Copris Geoffroy, 2009 | Mexican dung beetle | 364 | Species | Taxom | | Coproecus Reiche, 1844 | | 1 | Species | Taxom | | Coprophanaeus Olsoufieff, 2008 | | 55 | Species | Taxom | | Coptodactyla Burmeister, 1925 | | 17 | Species | Taxom | | Coptorhina Hope, 1857 | | 9 | Species | Taxom | | Cretonitis Nikolajev, 2007 | | 1 | Species | Taxom | | Cretorabaeus Nikolajev, 1995 | | 1 | Species | Taxom | | Cryptocanthon Balthasar, 2002 | | 45 | Species | Taxom | | Cyobius Sharp, 1875 | | 5 | Species | Taxom | | Cyptochirus Lesne, 1851 | | 4 | Species | Taxom | | Degallieridium Vaz-de-Mello, 2008 | | 1 | Species | Taxom | | Delopleurus Erichson, 1931 | | 11 | Species | Taxom | | Deltepilissus Silvério Per..., 1880 | | 2 | Species | Taxom | | Deltochilum Eschscholtz, 1939 | | 129 | Species | Taxom | | Deltorhinum Harold, 2010 | | 8 | Species | Taxom | | Demarziella Balthasar, 1986 | | 14 | Species | Taxom | | Dendropaemon Perty, 1831 | | 52 | Species | Taxom | | Diabroctis Gistl, 1990 | | 5 | Species | Taxom | | Diastellopalpus van Lansberge, 2000 | | 36 | Species | Taxom | | Dichotomius Hope, 1922 | | 225 | Species | Taxom | | Dicranocara | | 4 | Species | Taxom | | Digitonthophagus Balthasar, 2017 | | 16 | Species | Taxom | | Diorygopyx Matthews, 1871 | | 8 | Species | Taxom | | Disphysema Harold, 1873 | | 1 | Species | Taxom | | Dorbignyolus Branco, 1913 | | 1 | Species | Taxom | | Drepanocerus Kirby, 1851 | | 22 | Species | Taxom | | Drepanoplatynus Boucomont, 1921 | | 1 | Species | Taxom | | Drepanopodus Janssens, 1901 | | 2 | Species | Taxom | | Dwesasilvasedis | | 1 | Species | Taxom | | Endroedyolus Scholtz & Ho..., 1987 | | 1 | Species | Taxom | | Ennearabdus Lansberge, 1868 | | 1 | Species | Taxom | | Eodrepanus Barbero, Pal..., 1945 | | 9 | Species | Taxom | | Epactoides Olsoufieff, 1976 | | 39 | Species | Taxom | | Epilissus Blanchard, 2022 | | 43 | Species | Taxom | | Epirinus Reiche, 1901 | | 36 | Species | Taxom | | Eucranium Dejean, 1893 | | 7 | Species | Taxom | | Eudinopus Burmeister, 1802 | | 1 | Species | Taxom | | Euoniticellus Janssens, 1953 | | 19 | Species | Taxom | | Euonthophagus Balthasar, 1946 | | 38 | Species | Taxom | | Eurysternus Dalman, 1988 | | 58 | Species | Taxom | | Eusaproecius Branco, 1923 | | 7 | Species | Taxom | | Eutrichillum | | 3 | Species | Taxom | | Falsignambia Paulian, 1987 | | 1 | Species | Taxom | | Feeridium Vaz-de-Mello, 2008 | | 1 | Species | Taxom | | Frankenbergerius | | 9 | Species | Taxom | | Garreta Janssens, 1967 | | 33 | Species | Taxom | | Genieridium Vaz-de-Mello, 1998 | | 7 | Species | Taxom | | Gilletellus Janssens, 1857 | | 2 | Species | Taxom | | Glauconia Paulian, 0 | | 2 | Species | Taxom | | Glyphoderus Westwood, 1837 | | 3 | Species | Taxom | | Gromphas Brullé, 1891 | | 7 | Species | Taxom | | Gymnopleurus Illiger, 1843 | | 83 | Species | Taxom | | Hammondantus Cambefort, 1978 | | 1 | Species | Taxom | | Hansreia Halffter & M..., 2015 | | 6 | Species | Taxom | | Haroldius Boucomont, 1993 | | 40 | Species | Taxom | | Helictopleurus Orbigny, 1898 | | 79 | Species | Taxom | | Heliocopris Hope, 2013 | | 67 | Species | Taxom | | Heteroclitopus Péringuey, 1991 | | 10 | Species | Taxom | | Heteronitis Gillet, 1840 | | 5 | Species | Taxom | | Heterosyphus Paulian, 1975 | | 1 | Species | Taxom | | Holocanthon | | 3 | Species | Taxom | | Holocephalus Hope, 2001 | | 6 | Species | Taxom | Homalocopris Solsky, 1871 | | 0 | Species | Taxom | | Homalotarsus Janssens, 1932 | | 1 | Species | Taxom | | Homocopris Burmeister, 1859 | | 6 | Species | Taxom | | Hyalonthophagus Palestrini &..., 1910 | | 12 | Species | Taxom | | Hypocanthidium Balthasar, 1938 | | 1 | Species | Taxom | | Isocopris Pereira & Ma..., 2017 | | 7 | Species | Taxom | | Ixodina Roth, 0 | | 13 | Species | Taxom | | Janssensantus Paulian, 1987 | | 3 | Species | Taxom | | Janssensellus Cambefort, 1995 | | 2 | Species | Taxom | | Kheper Janssens, 1986 | | 33 | Species | Taxom | | Kolbeellus Jacobson, 1875 | | 1 | Species | Taxom | | Krikkenius Branco, 1982 | | 1 | Species | Taxom | | Labroma Sharp, 1974 | | 3 | Species | Taxom | | Larhodius Balthasar, 2003 | | 3 | Species | Taxom | | Latodrepanus Krikken, 2009 | | 6 | Species | Taxom | | Leotrichillum Vaz-de-Mello, 1998 | | 1 | Species | Taxom | | Lepanus Balthasar, 2019 | | 65 | Species | Taxom | | Liatongus Reitter, 1933 | | 49 | Species | Taxom | | Lithanomala Nikolajev, 1990 | | 4 | Species | Taxom | | Litocopris Waterhouse, 1869 | | 4 | Species | Taxom | | Lobateuchus Montreuil & al., 2010 | | 1 | Species | Taxom | | Lobidion Génier, 2010 | | 1 | Species | Taxom | | Lophodonitis Janssens, 2011 | | 2 | Species | Taxom | | Macroderes Westwood, 1880 | | 25 | Species | Taxom | | Macropanelus | | 3 | Species | Taxom | | Macropocopris Arrow, 1920 | | 1 | Species | Taxom | | Madateuchus Paulian, 1953 | | 1 | Species | Taxom | | Malagoniella Martínez, 1869 | | 21 | Species | Taxom | | Martinezidium Vaz-de-Mello, 2004 | | 4 | Species | Taxom | | Megalonitis Janssens, 1855 | | 3 | Species | Taxom | | Megaponerophilus Janssens, 1914 | | 1 | Species | Taxom | | Megatharsis Waterhouse, 1891 | | 1 | Species | Taxom | | Megathopa Eschscholtz, 1863 | | 6 | Species | Taxom | | Megathoposoma Balthasar, 1873 | | 1 | Species | Taxom | | Melanocanthon Halffter, 1915 | | 4 | Species | Taxom | | Mentophilus Blanchard, 1873 | | 2 | Species | Taxom | | Metacatharsius Paulian, 1965 | | 66 | Species | Taxom | | Milichus Péringuey, 1907 | | 17 | Species | Taxom | | Mimonthophagus Balthasar, 1902 | | 8 | Species | Taxom | | Mnematidium Ritsema, 1845 | | 2 | Species | Taxom | | Mnematium MacLeay, 1832 | | 2 | Species | Taxom | | Monoplistes Lansberge, 1923 | | 6 | Species | Taxom | | Namakwanus Scholtz & Ho..., 2005 | | 7 | Species | Taxom | | Nanos Westwood, 2011 | | 44 | Species | Taxom | Neateuchus Gillet, 1911 | | 0 | Species | Taxom | | Nebulasilvius Deschodt & C..., 2008 | | 2 | Species | Taxom | | Neocanthidium | | 2 | Species | Taxom | | Neonitis Péringuey, 1918 | | 2 | Species | Taxom | | Neosaproecius Branco, 1958 | | 2 | Species | Taxom | | Neosisyphus Müller, 1891 | | 27 | Species | Taxom | | Nesosisyphus Vinson, 1939 | | 4 | Species | Taxom | | Nesovinsonia Martinez & P..., 1939 | | 1 | Species | Taxom | | Nitiocellus Branco, 1921 | | 2 | Species | Taxom | | Nunoidium Vaz-de-Mello, 1913 | | 1 | Species | Taxom | | Ochicanthon Vaz-de-Mello, 2006 | | 53 | Species | Taxom | | Odontoloma Boheman, 1987 | | 20 | Species | Taxom | | Oficanthon Paulian, 1985 | | 1 | Species | Taxom | | Oniticellus Serville, 1934 | | 31 | Species | Taxom | | Onitis Fabricius, 2012 | | 188 | Species | Taxom | | Onoreidium Vaz-de-Mello, 1931 | | 4 | Species | Taxom | | Ontherus Erichson, 1930 | | 63 | Species | Taxom | | Onthobium Reiche, 1986 | | 14 | Species | Taxom | | Onthocharis Westwood, 1952 | | 6 | Species | Taxom | | Onthophagus Latreille, 1963 | | 2613 | Species | Taxom | | Onychothecus Boucomont, 1992 | | 4 | Species | Taxom | | Oruscatus Bates, 1870 | | 2 | Species | Taxom | | Outenikwanus Scholtz & Ho..., 1987 | | 1 | Species | Taxom | | Oxysternon Laporte, 1924 | | 17 | Species | Taxom | | Pachylomera Griffith & P..., 1874 | | 2 | Species | Taxom | | Pachysoma MacLeay, 1953 | | 19 | Species | Taxom | | Panelus Lewis, 0 | | 45 | Species | Taxom | | Paracanthon Balthasar, 1938 | | 16 | Species | Taxom | | Parachorius Harold, 1873 | | 22 | Species | Taxom | | Paracryptocanthon | | 2 | Species | Taxom | | Paragymnopleurus Shipp, 1997 | | 22 | Species | Taxom | | Paraphacosomoides | | 1 | Species | Taxom | | Paraphytus Harold, 1984 | | 14 | Species | Taxom | | Paroniticellus Balthasar, 1809 | | 1 | Species | Taxom | | Paronitis | | 1 | Species | Taxom | | Paronthobium Paulian, 1903 | | 23 | Species | Taxom | | Parvuhowdenius Deschodt & C..., 2008 | | 1 | Species | Taxom | | Peckolus Scholtz & Ho..., 2008 | | 3 | Species | Taxom | | Pedaria Laporte, 1939 | | 74 | Species | Taxom | | Pedaridium Harold, 1948 | | 10 | Species | Taxom | Peltecanthon Pereira, 1953 | | 0 | Species | Taxom | | Penalus Paulian, 1985 | | 1 | Species | Taxom | | Pereiraidium Vaz-de-Mello, 1946 | | 1 | Species | Taxom | | Phacosoma Boucomont, 1989 | | 2 | Species | Taxom | | Phaedotrogus Paulian, 1972 | | 1 | Species | Taxom | | Phalops Erichson, 2003 | | 40 | Species | Taxom | | Phanaeus MacLeay, 2017 | Burrowing Dung Beetles | 142 | Species | Taxom | | Pinacopodius Branco, 2007 | | 2 | Species | Taxom | | Pinacotarsus Harold, 1969 | | 2 | Species | Taxom | | Pinotus Erichson, 1929 | | 7 | Species | Taxom | | Platyonitis Janssens, 1974 | | 3 | Species | Taxom | | Pleuronitis Lansberge, 2011 | | 4 | Species | Taxom | | Ponerotrogus Silvestri, 1924 | | 1 | Species | Taxom | | Prionocephale Lin, 1980 | | 1 | Species | Taxom | | Proagoderus Lansberge, 2004 | | 132 | Species | Taxom | | Proteroscarabaeus Grabau, 1923 | | 6 | Species | Taxom | | Pseudignambia Paulian & Pl..., 1974 | | 2 | Species | Taxom | | Pseudocanthon Bates, 1846 | | 14 | Species | Taxom | | Pseudochironitis Ferreira, 1897 | | 3 | Species | Taxom | | Pseudocopris Ferreira, 1910 | | 2 | Species | Taxom | | Pseudonthobium Paulian, 2011 | | 9 | Species | Taxom | | Pseudosaproecius Balthasar, 1884 | | 12 | Species | Taxom | | Pseuduroxys Balthasar, 1938 | | 1 | Species | Taxom | | Pteronyx Lansberge, 1874 | | 1 | Species | Taxom | | Pycnopanelus Arrow, 1931 | | 3 | Species | Taxom | | Saphobiamorpha Brookes, 1944 | | 1 | Species | Taxom | | Saphobius Sharp, 1877 | | 15 | Species | Taxom | | Sarophorus Erichson, 1840 | | 12 | Species | Taxom | | Sauvagesinella Paulian, 1974 | | 4 | Species | Taxom | | Scaptocnemis Péringuey, 1901 | | 1 | Species | Taxom | | Scarabaeus Linnaeus, 1901 | | 169 | Species | Taxom | | Scatimus Erichson, 2020 | | 14 | Species | Taxom | | Scatonomus Erichson, 1954 | | 15 | Species | Taxom | | Scatrichus | | 3 | Species | Taxom | | Sceliages Westwood, 1844 | | 8 | Species | Taxom | | Scelocopris Zhang, 1989 | | 1 | Species | Taxom | | Scybalocanthon Martínez, 1949 | | 30 | Species | Taxom | | Scybalophagus Martínez, 1846 | | 5 | Species | Taxom | Sebasteos Westwood, 1845 | | 0 | Species | Taxom | | Silvaphilus | | 2 | Species | Taxom | | Silvinha Vaz-de-Mello, 2008 | | 1 | Species | Taxom | | Sinapisoma Boucomont, 1928 | | 2 | Species | Taxom | | Sinodrepanus Simonis, 2004 | | 9 | Species | Taxom | | Sisyphus Latreille, 1973 | | 120 | Species | Taxom | Sphaerocanthon Olsoufieff, 1947 | | 0 | Species | Taxom | | Stiptocnemis Branco, 1991 | | 2 | Species | Taxom | | Stiptopodius Harold, 1901 | | 15 | Species | Taxom | | Stiptotarsus Branco, 1963 | | 3 | Species | Taxom | | Strandius Balthasar, 1980 | | 1 | Species | Taxom | | Streblopus Lansberge, 1938 | | 2 | Species | Taxom | | Sukelus Branco, 1991 | | 1 | Species | Taxom | | Sulcodrepanus Krikken, 1840 | | 1 | Species | Taxom | | Sulcophanaeus Olsoufieff, 1856 | | 24 | Species | Taxom | | Sylvicanthon Halffter & M..., 1846 | | 17 | Species | Taxom | | Synapsis Forster, 1933 | | 28 | Species | Taxom | | Tanzanolus Scholtz & Ho..., 1960 | | 1 | Species | Taxom | | Temnoplectron Westwood, 2000 | | 21 | Species | Taxom | | Tesserodon Hope, 2017 | | 21 | Species | Taxom | | Tesserodoniella Vaz-de-Mello..., 2006 | | 2 | Species | Taxom | | Tetraechma Agassiz, 1939 | | 5 | Species | Taxom | | Tetramereia Klages, 1869 | | 1 | Species | Taxom | | Thyregis Blackburn, 1933 | | 4 | Species | Taxom | | Tibiodrepanus Krikken, 2011 | | 7 | Species | Taxom | | Tiniocellus Péringuey, 1851 | | 9 | Species | Taxom | | Tomogonus d´Orbigny, 1971 | | 11 | Species | Taxom | | Tragiscus Klug, 1855 | | 1 | Species | Taxom | | Trichillidium Vaz-de-Mello, 1932 | | 4 | Species | Taxom | | Trichillum Harold, 0 | | 18 | Species | Taxom | | Tropidonitis Janssens, 1857 | | 1 | Species | Taxom | | Unidentis | | 1 | Species | Taxom | | Uroxys Westwood, 1903 | | 65 | Species | Taxom | | Versicorpus Deschodt, Da..., 2005 | | 4 | Species | Taxom | | Vulcanocanthon Pereira & Ma..., 1867 | | 1 | Species | Taxom | | Walterantus Cambefort, 1960 | | 1 | Species | Taxom | | Xenocanthon Martínez, 1952 | | 2 | Species | Taxom | | Xinidium Harold, 1993 | | 4 | Species | Taxom | | Yvescambefortius Ochi & Kon, 1926 | | 1 | Species | Taxom | | Zonocopris Arrow, 2007 | | 2 | Species | Taxom |
Species list for Continents
7423 Species and 397 Subspecies | Acanthonitis bouyeri Moretto, 2010 | | | Acanthonitis nicolasi Cambefort, 1995 | | | Acanthonitis oberthuri Janssens, 1937 | | | Acanthonitis spissus Josso, 2012 | | | Afrodrepanus impressicollis Boheman, 1857 | | | Afrodrepanus marshalli Boucomont, 1921 | | | Agamopus castaneus Balthasar, 1938 | | | Agamopus convexus Balthasar, 1965 | | | Agamopus joker Costa-Silva,..., 2022 | | | Agamopus lampros Bates, 1887 | | | Agamopus unguicularis Harold, 1883 | | | Agamopus viridis Boucomont, 1928 | | | Aleiantus fiorii (Paulian, 1976) | ⇒ Epactoides fiorii Paulian, 1976 | | Aleiantus incertus (Lebis, 1953) | ⇒ Epactoides incertus Lebis, 1953 | | Aleiantus perinetanus (Paulian, 1975) | ⇒ Epactoides perinetanus Paulian, 1975 | | Aleiantus perrieri (Fairmaire, 1898) | ⇒ Epactoides perrieri Fairmaire, 1898 | | Aliuscanthoniola similaris Deschodt & S..., 2008 | | | Allogymnopleurus aeneus Harold, 1875 | | | Allogymnopleurus alluaudi Garreta, 1914 | | | Allogymnopleurus anthracinus Klug, 1845 | | | Allogymnopleurus chloris Klug, 1855 | | | Allogymnopleurus consocius Péringuey, 1901 | | | Allogymnopleurus histrio Balthasar, 1963 | | | Allogymnopleurus indigaceus Reiche, 1847 | | | Allogymnopleurus infranitens Fairmaire, 1887 | | | Allogymnopleurus jeanneli Garreta, 1914 | | | Allogymnopleurus maculosus MacLeay, 1821 | | | Allogymnopleurus olivieri Castelnau, 1840 | | | Allogymnopleurus sagnai Nicolas & Mo..., 2002 | | | Allogymnopleurus sericeicollis Frey, 1958 | | | Allogymnopleurus signaticollis Waterhouse, 1890 | | | Allogymnopleurus spilotus MacLeay, 1821 | | | Allogymnopleurus splendidus Bertoloni, 1849 | | | Allogymnopleurus thalassinus Klug, 1855 | | | Allogymnopleurus umbrinus Gerstaecker, 1871 | | | Allogymnopleurus youngai Endrödi, 1973 | | | Allogymnopleurus zavattarii Müller, 1941 | | | Allonitis nasutus Felsche, 1907 | | | Alloscelus combesi Paulian, 1949 | | | Alloscelus hirticomus d´Orbigny, 1913 | | | Alloscelus longipilis Frey, 1960 | | | Alloscelus paradoxus Boucomont, 1923 | | | Amietina cambeforti Branco, 1989 | | | Amietina eburnea Cambefort, 1981 | | | Amietina larrochei Cambefort, 1981 | | | Amietina nyassalandica Branco, 1989 | | | Amphistomus accidatus Matthews, 1974 | | | Amphistomus alfurorum Huijbregts &..., 2007 | | | Amphistomus calcaratus MacLeay, 1871 | | | Amphistomus complanatus Matthews, 1974 | | | Amphistomus cunninghamensis Matthews, 1974 | | | Amphistomus fasciculatus Gillet, 1927 | | | Amphistomus inermis Matthews, 1974 | | | Amphistomus kukali Paulian, 1985 | | | Amphistomus macphersonensis Matthews, 1974 | | | Amphistomus montanus Matthews, 1974 | | | Amphistomus monticola Paulian, 1985 | | | Amphistomus opacus Matthews, 1974 | | | Amphistomus palpebratus Matthews, 1974 | | | Amphistomus pectoralis Matthews, 1974 | | | Amphistomus primonactus Matthews, 1974 | | | Amphistomus pygmaeus Matthews, 1974 | | | Amphistomus respiciens Kolbe, 1905 | | | Amphistomus speculatus Gillet, 1927 | | | Amphistomus speculifer Matthews, 1974 | | | Amphistomus squalidus MacLeay, 1887 | | | Amphistomus storeyi Matthews, 1974 | | | Amphistomus trispiculatus Matthews, 1974 | | | Amphistomus tuberculatus Lansberge, 1885 | | | Amphistomus tuberosus Matthews, 1974 | | | Amphistomus uncinatus Paulian, 1985 | | | Anachalcos aurescens Bates, 1888 | | | Anachalcos convexus Boheman, 1857 | | | Anachalcos cupreus Fabricius, 1775 | | | Anachalcos mfwangani ♰ R. Paulian, 1976 | | | Anachalcos procerus Gerstaecker, 1871 | | | Anachalcos spectabilis (Péringuey, 1901) | ⇒ Chalconotus procerus Gerstaecker, 1871 | | Anisocanthon pygmaeus Gillet, 1911 | | | Anisocanthon sericinus Harold, 1868 | | | Anisocanthon strandi Balthasar, 1939 | | | Anisocanthon villosus Harold, 1868 | | | Anoctus laevis Sharp, 1875 | | | Anoctus myrmecophilus Arrow, 1907 | | | Anoctus rasilis Boucomont, 1921 | | | Anoctus scatonomoides Lansberge, 1885 | | | Anoctus siamensis Balthasar, 1942 | | | Anomiopsis bilobum schaefferi (Martínez, 1944) | ⇒ Anomiopsoides biloba Burmeister, 1861 | | Anomiopsis pereirai (Martínez, 1944) | ⇒ Anomiopsoides heteroclyta Blanchard, 1846 | | Anomiopsoides aberrans (Martínez, 1945) | ⇒ Anomiopsoides heteroclyta Blanchard, 1846 | | Anomiopsoides biloba Burmeister, 1861 | | | Anomiopsoides catamarcae (Martínez, 1945) | ⇒ Anomiopsoides heteroclyta Blanchard, 1846 | | Anomiopsoides cavifrons Burmeister, 1861 | | | Anomiopsoides fedemariai Ocampo, 2007 | | | Anomiopsoides heteroclyta Blanchard, 1846 | | | Anomiopsoides heteroclytum Blanchard, 1843 | | | Anomiopsoides xerophila Martínez, 1945 | | | Anomiopus aequalis Waterhouse, 1891 | | | Anomiopus alexandrei Canhedo, 2006 | | | Anomiopus andrei Canhedo, 2006 | | | Anomiopus apuskispay Figueroa & E..., 2015 | | | Anomiopus ataenioides Martínez, 1952 | | | Anomiopus batesi Waterhouse, 1891 | | | Anomiopus batesii Waterhouse, 1891 | | | Anomiopus birai Canhedo, 2006 | | | Anomiopus bonariensis Bruch, 1925 | | | Anomiopus brevipes Waterhouse, 1891 | | | Anomiopus cambeforti Canhedo, 2006 | | | Anomiopus caputipilus Canhedo, 2004 | | | Anomiopus chalceus Harold, 1867 | | | Anomiopus cirulito Cano, 2018 | | | Anomiopus cuprarius Harold, 1880 | | | Anomiopus edmondsi Canhedo, 2006 | | | Anomiopus foveicollis Canhedo, 2006 | | | Anomiopus galileoae Canhedo, 2006 | | | Anomiopus genieri Canhedo, 2006 | | | Anomiopus germari Harold, 1867 | | | Anomiopus gilli Canhedo, 2006 | | | Anomiopus globosus Canhedo, 2006 | | | Anomiopus gracilis Canhedo, 2006 | | | Anomiopus hirsutus Canhedo, 2006 | | | Anomiopus howdeni Canhedo, 2006 | | | Anomiopus idei Canhedo, 2006 | | | Anomiopus impartectus Canhedo, 2004 | | | Anomiopus intermedius Waterhouse, 1891 | | | Anomiopus juanae Martínez, 1952 | | | Anomiopus lacordairei Waterhouse, 1891 | | | Anomiopus laetus Waterhouse, 1891 | | | Anomiopus latistriatus Canhedo, 2006 | | | Anomiopus lauropalui Vulcano & Pe..., 1973 | | | Anomiopus lunatipes Canhedo, 2004 | | | Anomiopus mourai Canhedo, 2006 | | | Anomiopus myrmidon Westwood, 1842 | | | Anomiopus nigricans Westwood, 1842 | | | Anomiopus nigrocoeruleus Martínez, 1955 | | | Anomiopus octodentatus Canhedo, 2006 | | | Anomiopus palmispinus Canhedo, 2006 | | | Anomiopus panamensis Paulian, 1939 | | | Anomiopus paraensis Canhedo, 2004 | | | Anomiopus paraguaiensis Canhedo, 2004 | | | Anomiopus parallelus Harold, 1862 | | | Anomiopus pereirai Martínez, 1955 | | | Anomiopus pictus Harold, 1862 | | | Anomiopus pishtaco Edmonds & Fi..., 2013 | | | Anomiopus preissae Canhedo, 2004 | | | Anomiopus pumilius Canhedo, 2006 | | | Anomiopus puncticollis Harold, 1862 | | | Anomiopus quadridentatus Canhedo, 2004 | | | Anomiopus serranus Canhedo, 2006 | | | Anomiopus similis Canhedo, 2004 | | | Anomiopus simplex Waterhouse, 1891 | | | Anomiopus smaragdinus Westwood, 1842 | | | Anomiopus soledari Canhedo, 2004 | | | Anomiopus sulcaticollis Canhedo, 2006 | | | Anomiopus sulcatus Canhedo, 2006 | | | Anomiopus tuberculicollis Canhedo, 2006 | | | Anomiopus tuberifrons Canhedo, 2004 | | | Anomiopus validus Canhedo, 2006 | | | Anomiopus virescens Westwood, 1842 | | | Anomiopus zaguryi Canhedo, 2006 | | | Anonthobium fauveli Paulian, 1984 | | | Anonthobium micros Paulian, 1984 | | | Anonthobium monteithi Paulian, 1987 | | | Anonthobium moui Paulian, 1984 | | | Anonthobium muticum Fauvel, 1903 | | | Anonthobium tibiale Fauvel, 1903 | | | Anonychonitis freyi Janssens, 1950 | | | Anoplodrepanus pecki Howden, 1976 | | | Anoplodrepanus reconditus Matthews, 1966 | | | Aphengium cupreum Shipp, 1897 | | | Aphengium curtum Silva & Vaz-..., 2015 | | | Aphengium ibateguara Silva & Vaz-..., 2015 | | | Aphengium sordidum Harold, 1868 | | | Aphengoecus clypeatus Péringuey, 1901 | | | Aphengoecus multiserratus Scholtz & Ho..., 1987 | | | Apotolamprus ambohitsitondronensis Lebis, 1953 | | | Apotolamprus andranobeensis Montreuil & ..., 2012 | | | Apotolamprus antsingyensis Montreuil, 2008 | | | Apotolamprus bemarahaensis Montreuil, 2008 | | | Apotolamprus cyanescens Brancsik, 1892 | | | Apotolamprus darainaensis Montreuil, 2008 | | | Apotolamprus densepunctatus Montreuil, 2008 | | | Apotolamprus hagai Montreuil & ..., 2012 | | | Apotolamprus hanskii Montreuil, 2005 | | | Apotolamprus helenae Montreuil, 2004 | | | Apotolamprus latipennis Boucomont, 1937 | | | Apotolamprus mahajangaensis Montreuil, 2008 | | | Apotolamprus manjarivoloensis Montreuil, 2008 | | | Apotolamprus manomboensis Montreuil, 2008 | | | Apotolamprus marojejyensis Montreuil, 2005 | | | Apotolamprus metallicus Montreuil, 2008 | | | Apotolamprus milloti Lebis, 1953 | | | Apotolamprus nanoides Paulian, 1976 | | | Apotolamprus orangeaensis Montreuil, 2008 | | | Apotolamprus perinetensis Montreuil, 2008 | | | Apotolamprus peyrierasi Paulian, 1985 | | | Apotolamprus pseudomanomboensis Montreuil, 2008 | | | Apotolamprus pseudoquadrimaculatus Montreuil & ..., 2012 | | | Apotolamprus quadrimaculatus Lebis, 1953 | | | Apotolamprus quadrinotatus Boucomont, 1937 | | | Apotolamprus resplendens Montreuil, 2008 | | | Apotolamprus sahatezaensis Montreuil & ..., 2012 | | | Apotolamprus sericeus Paulian, 1976 | | | Apotolamprus sinuatotibialis Montreuil, 2008 | | | Apotolamprus sparsepunctatus Montreuil, 2008 | | | Apotolamprus vadoni Paulian, 1976 | | | Apotolamprus zombitsyensis Montreuil, 2008 | | | Aptenocanthon hopsoni Carter, 1936 | | | Aptenocanthon jimara Storey & Mon..., 2000 | | | Aptenocanthon kabura Storey & Mon..., 2000 | | | Aptenocanthon monteithi Storey, 1984 | | | Aptenocanthon rossi Matthews, 1974 | | | Aptenocanthon speewah Storey & Mon..., 2000 | | | Aptenocanthon winyar Storey & Mon..., 2000 | | | Aptenocanthon wollumbin Storey & Mon..., 2000 | | | Apterepilissus analavelonae Paulian, 1975 | | | Apterepilissus centralis Paulian, 1976 | | | Apterepilissus ovalis Felsche, 1911 | | | Apterepilissus sericeus Lebis, 1958 | | | Aptychonitis anomalus Gestro, 1895 | | | Aptychonitis sulcicollis Harold, 1868 | | | Arachnodes abadiei Lebis, 1953 | | | Arachnodes aeneoviridis Boucomont, 1937 | | | Arachnodes ampasindavae Paulian, 1976 | | | Arachnodes andriai Paulian, 1976 | | | Arachnodes angulosus Boucomont, 1937 | | | Arachnodes bicolor Montreuil, 2011 | | | Arachnodes biimpressus Lebis, 1953 | | | Arachnodes brunnipes Castelnau, 1840 | | | Arachnodes delaunayi Lebis, 1953 | | | Arachnodes delphinensis Lebis, 1953 | | | Arachnodes dichrous Paulian, 1976 | | | Arachnodes didiensis Lebis, 1953 | | | Arachnodes dieganus Paulian, 1976 | | | Arachnodes fairmairei Boucomont, 1937 | | | Arachnodes fulgens Kunckel, 1887 | | | Arachnodes fusconitens Fairmaire, 1899 | | | Arachnodes fuscovirens Fairmaire, 1889 | | | Arachnodes globuloides Paulian, 1976 | | | Arachnodes globulosus (Kunckel, 1887) | ⇒ Arachnodes globuloides Paulian, 1976 | | Arachnodes grossepunctatus Paulian, 1986 | | | Arachnodes hanskii Montreuil, 2003 | | | Arachnodes histeroides Castelnau, 1840 | | | Arachnodes insularis Paulian, 1976 | | | Arachnodes jeanneli Lebis, 1953 | | | Arachnodes kelifelyi Paulian, 1976 | | | Arachnodes luctuosus Boucomont, 1937 | | | Arachnodes mahafalyensis Montreuil, 2006 | | | Arachnodes manaitrai Montreuil, 2006 | | | Arachnodes manaitrai kirindyensis Montreuil, 2006 | | | Arachnodes manaitrai manaitrai | | | Arachnodes manomboensis Montreuil, 2006 | | | Arachnodes mantillerii Montreuil, 2003 | | | Arachnodes micans Lebis, 1953 | | | Arachnodes minutus Paulian, 1976 | | | Arachnodes morati Paulian, 1975 | | | Arachnodes morio Harold, 1879 | | | Arachnodes neonitidus Montreuil, 2011 | | | Arachnodes nitidus Castelnau, 1840 | | | Arachnodes nosybeensis Lebis, 1961 | | | Arachnodes oberthuri Boucomont, 1937 | | | Arachnodes philippi Lebis, 1953 | | | Arachnodes pierrettae Lebis, 1953 | | | Arachnodes pillula Paulian, 1976 | | | Arachnodes pusillus Lebis, 1953 | | | Arachnodes refulgens Fairmaire, 1901 | | | Arachnodes renaudi Lebis, 1958 | | | Arachnodes robinsoni Boucomont, 1937 | | | Arachnodes rubrolimbatus Boucomont, 1937 | | | Arachnodes rubrotinctus Lebis, 1953 | | | Arachnodes sakarahae Paulian, 1976 | | | Arachnodes saprinoides Fairmaire, 1889 | | | Arachnodes semichalceus Lebis, 1953 | | | Arachnodes seminitidus Lebis, 1953 | | | Arachnodes seyrigi Lebis, 1953 | | | Arachnodes soganus Paulian, 1976 | | | Arachnodes splendidus Lebis, 1953 | | | Arachnodes tibialis Lebis, 1956 | | | Arachnodes variolosus Lebis, 1953 | | | Arachnodes vicinus Castelnau, 1840 | | | Arachnodes viettei Montreuil, 2011 | | | Arachnodes vieui Paulian, 1976 | | | Arachnodes vigilans Lebis, 1953 | | | Arachnodes zombitsyensis Montreuil, 2011 | | | Ateuchites grandis ♰ Meunier, 1898 | | | Ateuchus aeneomicans Harold, 1868 | | | Ateuchus alipioi Pereira, 1954 | | | Ateuchus alutacius Kohlmann & S..., 2012 | | | Ateuchus alvarezi Martinez & H..., 1986 | | | Ateuchus ambiguus Martinez & M..., 1990 | | | Ateuchus apicatus Harold, 1867 | | | Ateuchus asperatus Harold, 1881 | | | Ateuchus balthasari Martínez, 1952 | | | Ateuchus bordoni Martinez & M..., 1990 | | | Ateuchus breve Harold, 1868 | | | Ateuchus calcaratus Harold, 1868 | | | Ateuchus candezei Harold, 1868 | | | Ateuchus carbonarius Harold, 1868 | | | Ateuchus carcavalloi Vulcano & Pe..., 1976 | | | Ateuchus carolinae Kohlmann, 1981 | | | Ateuchus cernyi Balthasar, 1939 | | | Ateuchus chrysopyge Bates, 1887 | | | Ateuchus colossus Moctezuma, S..., 2018 | | | Ateuchus columbianus Harold, 1868 | | | Ateuchus confusus Martinez & M..., 1987 | | | Ateuchus connexus Harold, 1868 | | | Ateuchus contractus Balthasar, 1939 | | | Ateuchus convexus (Hausmann, 1807) | ⇒ Scarabaeus convexus Hausmann, 1807 | | Ateuchus cujuchi Génier, 2015 | | | Ateuchus earthorus Kohlmann & S..., 2009 | | | Ateuchus ecuadorensis Boucomont, 1928 | | | Ateuchus euchalceus Balthasar, 1939 | | | Ateuchus fedescobari Montoya-Moli..., 2021 | | | Ateuchus femoratus Fabricius, 1801 | | | Ateuchus fetteri Kohlmann, 1996 | | | Ateuchus floridensis Génier, 2000 | | | Ateuchus freudei Balthasar, 1966 | | | Ateuchus frontale Boucomont, 1928 | | | Ateuchus fuscipes Blanchard, 1846 | | | Ateuchus fuscorubrus Blanchard, 1846 | | | Ateuchus genieri Montoya-Moli..., 2019 | | | Ateuchus gershensoni Kohlmann, 2000 | | | Ateuchus ginae Kohlmann, 1996 | | | Ateuchus globulus Boucomont, 1928 | | | Ateuchus granigerus Harold, 1875 | | | Ateuchus guatemalensis Bates, 1887 | | | Ateuchus halffteri Kohlmann, 1981 | | | Ateuchus hamatus Boucomont, 1928 | | | Ateuchus hendrichsi Kohlmann, 1996 | | | Ateuchus histeroides Weber, 1801 | | | Ateuchus histrio (Balthasar, 1939) | ⇒ Canthon septemmaculatus histrio Lepeletier &..., 1828 | | Ateuchus hoplopygus Harold, 1868 | | | Ateuchus hornai Balthasar, 1939 | | | Ateuchus howdeni Kohlmann, 1996 | | | Ateuchus hypocrita (Balthasar, 1939) | ⇒ Ateuchetus puncticollis Latreille, 1819 | | Ateuchus illaesum Harold, 1868 | | | Ateuchus illaesus Harold, 1868 | | | Ateuchus insulare Fleutiaux & ..., 1889 | | | Ateuchus irinus Balthasar, 1939 | | | Ateuchus klugi Harold, 1868 | | | Ateuchus laetitiae Kohlmann, 1981 | | | Ateuchus laevicolle Harold, 1868 | | | Ateuchus laterale Harold, 1868 | | | Ateuchus latus Boucomont, 1928 | | | Ateuchus lecontei Harold, 1868 | Leconte's dung beetle | | Ateuchus loricatus Boucomont, 1928 | | | Ateuchus luciae Matthews, 1966 | | | Ateuchus murrayi Harold, 1868 | | | Ateuchus mutilatus Harold, 1867 | | | Ateuchus myrmecophilus Boucomont, 1935 | | | Ateuchus nitidulus Castelnau, 1840 | | | Ateuchus oblongus Harold, 1883 | | | Ateuchus opacipennis Boucomont, 1928 | | | Ateuchus ovale Boucomont, 1928 | | | Ateuchus ovalis Boucomont, 1928 | | | Ateuchus parvus Balthasar, 1939 | | | Ateuchus pauki Balthasar, 1939 | | | Ateuchus pauperatus Germar, 1824 | | | Ateuchus perezvelai Kohlmann, 2000 | | | Ateuchus perpusillus Kohlmann, 2000 | | | Ateuchus persplendens Balthasar, 1939 | | | Ateuchus peruanus Balthasar, 1939 | | | Ateuchus pius (Illiger, 1803) | ⇒ Scarabaeus pius Illiger, 1803 | | Ateuchus procerus Harold, 1883 | | | Ateuchus pruneus Boucomont, 1928 | | | Ateuchus punctatissimus Génier, 2010 | | | Ateuchus puncticolle Harold, 1867 | | | Ateuchus pygidiale Harold, 1868 | | | Ateuchus pygidialis Harold, 1868 | | | Ateuchus rispolii Martínez, 1952 | | | Ateuchus robustus Harold, 1868 | | | Ateuchus rodriguezi Preudhomme d..., 1886 | | | Ateuchus romani Boucomont, 1927 | | | Ateuchus scatimoides Balthasar, 1939 | | | Ateuchus semicribratum Harold, 1868 | | | Ateuchus semicribratus Harold, 1868 | | | Ateuchus setulosus Balthasar, 1939 | | | Ateuchus simplex Lepeletier &..., 1828 | | | Ateuchus solisi Kohlmann, 1996 | | | Ateuchus spinipes Thunberg, 1818 | | | Ateuchus squalidus Fabricius, 1775 | | | Ateuchus steinbachi Boucomont, 1928 | | | Ateuchus striatulus Preudhomme d..., 1886 | | | Ateuchus subquadratus Harold, 1868 | | | Ateuchus substriatus Harold, 1868 | | | Ateuchus tenebrosus Balthasar, 1945 | | | Ateuchus texanus Robinson, 1948 | Notchfin blenny | | Ateuchus tona Lopera, Cham..., 2020 | | | Ateuchus tridenticeps Arrow, 1913 | | | Ateuchus tuza Kohlmann & V..., 2018 | | | Ateuchus viduum Blanchard, 1846 | | | Ateuchus viduus Blanchard, 1846 | | | Ateuchus vigilans Lansberge, 1874 | | | Ateuchus viridimicans Boucomont, 1935 | | | Ateuchus vividus Germar, 1824 | | | Ateuchus zoebischi Kohlmann, 1996 | | | Attavicinus monstrosus Bates, 1887 | | | Aulacopris matthewsi Storey, 1986 | Matthew's Flightless Dung Beetle | | Aulacopris maximus Matthews, 1974 | | | Aulacopris reichei White, 1859 | | | Baloghonthobium monommoides Paulian, 1986 | | | Bdelyropsis bowditchi Paulian, 1939 | | | Bdelyropsis newtoni Howden, 1971 | | | Bdelyropsis venezuelensis Howden, 1976 | | | Bdelyrus amazonensis Cook, 1998 | | | Bdelyrus apaporisae Cook, 1998 | | | Bdelyrus ashei Cook, 2000 | | | Bdelyrus boliviensis Cook, 1998 | | | Bdelyrus bowditchi (Paulian, 1939) | ⇒ Bdelyropsis bowditchi Paulian, 1939 | | Bdelyrus braziliensis Cook, 1998 | | | Bdelyrus bromeliatilis Cook, 1998 | | | Bdelyrus cochabambae Cook, 2000 | | | Bdelyrus ecuadorae Cook, 2000 | | | Bdelyrus geijskesi Huijbregts, 1984 | | | Bdelyrus genieri Cook, 1998 | | | Bdelyrus gilli Cook, 1998 | | | Bdelyrus grandis Cook, 1998 | | | Bdelyrus howdeni Cook, 1998 | | | Bdelyrus iauaretensis Cook, 1998 | | | Bdelyrus iquitosensis Cook, 2000 | | | Bdelyrus lagopus Harold, 1869 | | | Bdelyrus laplanadae Cook, 1998 | | | Bdelyrus leptomerus Cook, 1998 | | | Bdelyrus lobatus Cook, 1998 | | | Bdelyrus metaensis Cook, 1998 | | | Bdelyrus paraensis Cook, 1998 | | | Bdelyrus parvoculus Cook, 1998 | | | Bdelyrus parvus Cook, 1998 | | | Bdelyrus pecki Cook, 1998 | | | Bdelyrus peruviensis Cook, 1998 | | | Bdelyrus seminudus Bates, 1887 | | | Bdelyrus triangulus Cook, 1998 | | | Besourenga amarillai Aguilar-Julio, 2001 | | | Besourenga horacioi Martínez, 1967 | | | Besourenga minutum Saylor, 1935 | | | Besourenga minutus Saylor, 1935 | | | Besourenga vejdovskyi Balthasar, 1939 | | | Bohepilissus nitidus Balthasar, 1965 | | | Bohepilissus subtilis Boheman, 1857 | | | Bolbites onitoides Harold, 1868 | | | Boletoscapter cornutus MacLeay, 1887 | | | Boletoscapter furcatus Matthews, 1974 | | | Boreocanthon ateuchiceps Bates, 1887 | | | Boreocanthon coahuilensis Howden, 1966 | | | Boreocanthon depressipennis LeConte, 1859 | | | Boreocanthon ebenus Say, 1823 | | | Boreocanthon forreri Bates, 1887 | | | Boreocanthon halffteri Edmonds, 2022 | | | Boreocanthon integricollis Schaeffer, 1915 | | | Boreocanthon lecontei Harold, 1868 | | | Boreocanthon melanus Robinson, 1948 | | | Boreocanthon praticola LeConte, 1859 | | | Boreocanthon probus Germar, 1824 | | | Boreocanthon puncticollis LeConte, 1866 | | | Boreocanthon simplex LeConte, 1857 | | | Bradypodidium adisi Ratcliffe, 1980 | | | Bradypodidium bradyporum Boucomont, 1928 | | | Bradypodidium venezuelense Ferreira & G..., 1993 | | | Bradypodidium venezuelensis Ferreira & G..., 1993 | | | Bubas bison Linnaeus, 1767 | | | Bubas bubaloides Janssens, 1938 | | | Bubas bubalus Olivier, 1811 | | | Byrrhidium convexum Scholtz & Ho..., 1987 | | | Byrrhidium namaquensis Scholtz & Ho..., 1987 | | | Byrrhidium ovale Harold, 1869 | | | Caccobius (Caccobius) chujoi Paulian, 1942 | | | Caccobius (Caccophilus) scrofa (Balthasar, 1942) | ⇒ Cleptocaccobius scrofa Balthasar, 1942 | | Caccobius (Caccophilus) tortus Sharp, 1875 | | | Caccobius tortus matsudai Ochi & Araya, 1996 | | | Caccobius tortus tortus | | | Caccobius (Caccophilus) unicornis Fabricius, 1798 | | | Caccobius anthracites d´Orbigny, 1904 | | | Caccobius aterrimus Fabricius, 1798 | | | Caccobius atomarius (Balthasar, 1935) | ⇒ Caccobius brevis Waterhouse, 1875 | | Caccobius atratus Walter & Cam..., 1977 | | | Caccobius auberti d´Orbigny, 1902 | | | Caccobius balthasari (Tesar, 1969) | ⇒ Cleptocaccobius balthasarianus Cambefort, 1979 | | Caccobius bama Walter, 2014 | | | Caccobius bawangensis Ochi & Kon &..., 1997 | | | Caccobius bidentatus Boucomont, 1923 | | | Caccobius binodulus Harold, 1877 | | | Caccobius boucomonti Balthasar, 1935 | | | Caccobius brevis Waterhouse, 1875 | | | Caccobius cabellai Walter, 2019 | | | Caccobius callosifrons D Orbigny, 1905 | | | Caccobius castaneus (Klug, 1855) | ⇒ Caccocnemus castaneus Klug, 1855 | | Caccobius cavatus d´Orbigny, 1908 | | | Caccobius christophi Harold, 1879 | | | Caccobius convexifrons (Raffray, 1877) | ⇒ Caccophilus convexifrons Raffray, 1877 | | Caccobius corniceps d´Orbigny, 1913 | | | Caccobius cribrarius Boucomont, 1928 | | | Caccobius croceocinctus d´Orbigny, 1913 | | | Caccobius curvicornis Walter & Cam..., 1977 | | | Caccobius cuspidiger d´Orbigny, 1913 | | | Caccobius cyclotis Cambefort, 1984 | | | Caccobius demangei Boucomont, 1919 | | | Caccobius denticollis Harold, 1867 | | | Caccobius devagiriensis Nithya & Sabu, 2013 | | | Caccobius dickinsoni Masumoto, Oc..., 2022 | | | Caccobius diminutivus Walker, 1858 | | | Caccobius discrepans Péringuey, 1901 | | | Caccobius dorsalis (Harold, 1867) | ⇒ Caccophilus dorsalis Harold, 1867 | | Caccobius dybowskii d´Orbigny, 1902 | | | Caccobius elephantinus Balthasar, 1967 | | | Caccobius excavatus Bai, Zhang &..., 2007 | | | Caccobius ferrugineus Fahraeus, 1857 | | | Caccobius flavolimbatus Balthasar, 1942 | | | Caccobius foveolatus Frey, 1955 | | | Caccobius fukiensis Balthasar, 1942 | | | Caccobius fuliginosus (Roth, 1851) | ⇒ Caccophilus reticuliger d´Orbigny, 1904 | | Caccobius gallinus Arrow, 1907 | | | Caccobius gananensis d´Orbigny, 1904 | | | Caccobius genierorum Walter, 2014 | | | Caccobius gibbosulus d´Orbigny, 1915 | | | Caccobius gilleti Balthasar, 1933 | | | Caccobius globaticeps d´Orbigny, 1905 | | | Caccobius gonoderus Fairmaire, 1888 | | | Caccobius grossegranosus Balthasar, 1963 | | | Caccobius himalayanus Jekel, 1872 | | | Caccobius hirayamai (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccobius hirsutus Frey, 1958 | | | Caccobius histerinus Fahraeus, 1857 | | | Caccobius histeroides Ménétriés, 1832 | | | Caccobius histrio Balthasar, 1967 | | | Caccobius imitans Balthasar, 1933 | | | Caccobius inconspicuus (Fahraeus, 1857) | ⇒ Caccophilus fuliginosus Roth, 1851 | | Caccobius indicus Harold, 1867 | | | Caccobius inops Péringuey, 1901 | | | Caccobius ivorensis Cambefort, 1984 | | | Caccobius jessoensis Harold, 1867 | | | Caccobius jononis (Matsumura, 1936) | ⇒ Gibbonthophagus viduus Harold, 1875 | | Caccobius jossoi Walter, 2014 | | | Caccobius kelleri Olsoufieff, 1907 | | | Caccobius krikkeni Cambefort, 1980 | | | Caccobius lamottei (Cambefort, 1983) | ⇒ Cleptocaccobius lamottei Cambefort, 1983 | | Caccobius lateralis d´Orbigny, 1905 | | | Caccobius leleupi Balthasar, 1963 | | | Caccobius longipennis d´Orbigny, 1904 | | | Caccobius maruyamai Masumoto, Oc..., 2012 | | | Caccobius masumotoi Cambefort, 1990 | | | Caccobius megaponerae (Brauns, 1914) | ⇒ Megaponerophilus megaponerae Brauns, 1914 | | Caccobius meridionalis Boucomont, 1914 | | | Caccobius minor (Walter & Cam..., 1977) | ⇒ Cleptocaccobius minor Walter & Cam..., 1977 | | Caccobius mirabilepunctatus Cambefort, 1971 | | | Caccobius montreuilli Walter, 2019 | | | Caccobius morettoi Walter, 2014 | | | Caccobius mundus Ménétriés, 1838 | | | Caccobius nigritulus Klug, 1855 | | | Caccobius nikkoensis Lewis, 1895 | | | Caccobius obtusus Fahraeus, 1857 | | | Caccobius ocellipennis d´Orbigny, 1913 | | | Caccobius pantherinus Arrow, 1931 | | | Caccobius pentagonus d´Orbigny, 1908 | | | Caccobius pilosus Frey, 1958 | | | Caccobius polygonus d´Orbigny, 1911 | | | Caccobius postlulatus | | | Caccobius postlutatus d´Orbigny, 1905 | | | Caccobius pseudocorniceps Josso, 2018 | | | Caccobius pseudolaevis d´Orbigny, 1908 | | | Caccobius pulicarius Harold, 1875 | | | Caccobius pullus Jekel, 1872 | | | Caccobius punctatissimus Harold, 1867 | | | Caccobius reticuliger d´Orbigny, 1904 | | | Caccobius rufipennis Motschulsky, 1858 | | | Caccobius sappoensis Matsumura, 1936 | | | Caccobius sapporensis (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccobius scheuerni Cambefort, 1990 | | | Caccobius schreberi Linnaeus, 1767 | | | Caccobius semiaeneus d´Orbigny, 1905 | | | Caccobius semiluteus d´Orbigny, 1905 | | | Caccobius sericeus Frey, 1958 | | | Caccobius sericoides Balthasar, 1966 | | | Caccobius setifer d´Orbigny, 1905 | | | Caccobius signatipennis Harold, 1867 | | | Caccobius sordidus Harold, 1886 | | | Caccobius suzukii (Matsumura, 1936) | ⇒ Caccobius boucomonti Balthasar, 1935 | | Caccobius tai Cambefort, 1984 | | | Caccobius torticornis Arrow, 1931 | | | Caccobius tsunoellus (Matsumara, 1938) | ⇒ Caccophilus tortus Sharp, 1875 | | Caccobius tuberifrons d´Orbigny, 1902 | | | Caccobius ugandicus Frey, 1975 | | | Caccobius ultor Sharp, 1875 | | | Caccobius upembanus Frey, 1958 | | | Caccobius villiersi Cambefort, 1975 | | | Caccobius vulcanus Fabricius, 1801 | | | Caccobius yamauchii (Matsumura, 1936) | ⇒ Caccobius unicornis Fabricius, 1798 | | Caccobius yubariensis (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccobius zairensis Walter & Cam..., 1977 | | | Caccobius zambiensis Walter, 2014 | | | Caeconthobium parvulum Paulian, 1984 | | | Cambefortantus ankaratrae Paulian, 1975 | | | Cambefortantus blanci Paulian, 1975 | | | Cambefortantus helenae Montreuil, 2008 | | | Cambefortantus micros Paulian, 1986 | | | Cambefortantus myops Lebis, 1953 | | | Cambefortantus pauliani Montreuil, 2008 | | | Cambefortantus ranomafanensis Montreuil, 2008 | | | Cambefortantus ranomandryi Paulian, 1975 | | | Cambefortius bidentiger d´Orbigny, 1905 | | | Cambefortius cavipectus Branco, 1989 | | | Cambefortius dudleyi Branco, 1989 | | | Cambefortius eburneus Branco, 1989 | | | Cambefortius guineensis Branco, 1989 | | | Cambefortius lamizanai Cambefort, 1983 | | | Cambefortius teke Walter & Cam..., 1977 | | | Cambefortius turneri Boucomont, 1936 | | | Canthidium (Eucanthidium) laetum (Harold, 1867) | ⇒ Eucanthidium laetum Harold, 1867 | | Canthidium (Eucanthidium) macclevei Kohlmann & S..., 2006 | | | Canthidium (Eucanthidium) pseudopuncticolle Solis & Kohl..., 2004 | | | Canthidium abbreviatum Harold, 1867 | | | Canthidium aeneolum Harold, 1867 | | | Canthidium alvarezi Martinez & H..., 1986 | | | Canthidium andersoni Kohlmann & S..., 2006 | | | Canthidium angulicolle Balthasar, 1939 | | | Canthidium angusticeps Bates, 1887 | | | Canthidium annagabrielae Solis & Kohl..., 2004 | | | Canthidium ardens Bates, 1887 | | | Canthidium aterrimum Harold, 1867 | | | Canthidium atomarium Balthasar, 1939 | | | Canthidium atramentarium Balthasar, 1939 | | | Canthidium atricolle Preudhomme d..., 1886 | | | Canthidium aurichalceum Preudhomme d..., 1886 | | | Canthidium auricolle Harold, 1867 | | | Canthidium aurifex Bates, 1887 | | | Canthidium ayri Carvalho de ..., 2019 | | | Canthidium barbacenicum Preudhomme d..., 1886 | | | Canthidium basale Harold, 1867 | | | Canthidium basipunctatum Balthasar, 1939 | | | Canthidium batesi Harold, 1867 | | | Canthidium bicolor Boucomont, 1928 | | | Canthidium bituberifrons Howden & Young, 1981 | | | Canthidium bokermanni Martinez, Ha..., 1964 | | | Canthidium bovinum Harold, 1867 | | | Canthidium breve Germar, 1824 | | | Canthidium caesareum Balthasar, 1939 | | | Canthidium calidum Harold, 1880 | | | Canthidium cavifrons Balthasar, 1939 | | | Canthidium centrale Boucomont, 1928 | | | Canthidium cereum Harold, 1868 | | | Canthidium chabanaudi Boucomont, 1928 | | | Canthidium chimalapense Moctezuma & ..., 2019 | | | Canthidium chrysis Fabricius, 1801 | | | Canthidium clypeale Harold, 1867 | | | Canthidium coerulescens Balthasar, 1939 | | | Canthidium cognatum Preudhomme d..., 1886 | | | Canthidium collare Castelnau, 1840 | | | Canthidium convexifrons Balthasar, 1939 | | | Canthidium cupreum Blanchard, 1846 | | | Canthidium cuprinum Harold, 1867 | | | Canthidium darwini Kohlmann & S..., 2009 | | | Canthidium decoratum Perty, 1830 | | | Canthidium delgadoi Kohlmann & S..., 2006 | | | Canthidium deplanatum Balthasar, 1939 | | | Canthidium depressum Boucomont, 1928 | | | Canthidium deyrollei Harold, 1867 | | | Canthidium dimidiatum Lansberge, 1874 | | | Canthidium discolor Harold, 1867 | | | Canthidium discopygidiale Howden & Young, 1981 | | | Canthidium dispar Harold, 1867 | | | Canthidium dohrni Harold, 1867 | | | Canthidium elegantulum Balthasar, 1939 | | | Canthidium emoryi Solis & Kohl..., 2004 | | | Canthidium epistomale Boucomont, 1928 | | | Canthidium erythropterum Lucas, 1857 | | | Canthidium escalerai Balthasar, 1939 | | | Canthidium euchalceum Balthasar, 1939 | | | Canthidium excisipes Balthasar, 1939 | | | Canthidium feeri Carvalho de ..., 2019 | | | Canthidium femoratum Boucomont, 1935 | | | Canthidium flabellatum Harold, 1883 | | | Canthidium flavicorne Blanchard, 1846 | | | Canthidium flavipes Harold, 1867 | | | Canthidium flavum Balthasar, 1939 | | | Canthidium foveolatum Harold, 1867 | | | Canthidium funebre Balthasar, 1939 | | | Canthidium gemmingeri Harold, 1867 | | | Canthidium gerstaeckeri Harold, 1867 | | | Canthidium gigas Balthasar, 1939 | | | Canthidium glabricolle Harold, 1867 | | | Canthidium globulum Harold, 1867 | | | Canthidium gracilipes Harold, 1867 | | | Canthidium granuliferum Pereira, 1949 | | | Canthidium guanacaste Howden & Gill, 1987 | | | Canthidium guyanense Boucomont, 1928 | | | Canthidium haagi Harold, 1867 | | | Canthidium haroldi Preudhomme d..., 1886 | | | Canthidium hespenheidei Howden & Young, 1981 | | | Canthidium histrio Balthasar, 1939 | | | Canthidium howdeni Kohlmann & S..., 2006 | | | Canthidium humerale Germar, 1813 | | | Canthidium hyla Balthasar, 1939 | | | Canthidium hypocrita Balthasar, 1939 | | | Canthidium imperiale Harold, 1876 | | | Canthidium impressum Boucomont, 1928 | | | Canthidium inerme Harold, 1867 | | | Canthidium inoptatum Balthasar, 1939 | | | Canthidium kelleri Martinez, Ha..., 1964 | | | Canthidium kiesenwetteri Harold, 1867 | | | Canthidium kirschi Harold, 1875 | | | Canthidium kohlmanni Mora-Aguilar..., 2019 | | | Canthidium korschefskyi Balthasar, 1939 | | | Canthidium kraatzi Harold, 1867 | | | Canthidium laevigatum Harold, 1867 | | | Canthidium latipleurum Preudhomme d..., 1886 | | | Canthidium latum Blanchard, 1846 | | | Canthidium lebasi Harold, 1867 | | | Canthidium lentum Erichson, 1847 | | | Canthidium leucopterum Howden & Young, 1981 | | | Canthidium lucidum Harold, 1867 | | | Canthidium luteum Balthasar, 1939 | | | Canthidium macroculare Howden & Gill, 1987 | | | Canthidium magnum Harold, 1871 | | | Canthidium manni Arrow, 1913 | | | Canthidium margaritae Kohlmann & S..., 2006 | | | Canthidium marianelae Solis & Kohl..., 2004 | | | Canthidium marielae Solis & Kohl..., 2004 | | | Canthidium marseuli Harold, 1867 | | | Canthidium melanocephalum Olivier, 1789 | | | Canthidium metallicum Harold, 1867 | | | Canthidium minimum Harold, 1883 | | | Canthidium miscellum Harold, 1883 | | | Canthidium moestum Harold, 1867 | | | Canthidium monoceros Harold, 1869 | | | Canthidium moroni Kohlmann & S..., 2006 | | | Canthidium multipunctatum Balthasar, 1939 | | | Canthidium muticum Boheman, 1858 | | | Canthidium nanum Harold, 1867 | | | Canthidium nebularum Moctezuma, S..., 2019 | | | Canthidium nigritum Preudhomme d..., 1886 | | | Canthidium nitidum (Blanchard, 1846) | ⇒ Eucanthidium cupreum Blanchard, 1843 | | Canthidium nobile Harold, 1867 | | | Canthidium obscurum Harold, 1867 | | | Canthidium onitoides Perty, 1830 | | | Canthidium onthophagoides Martinez & H..., 1986 | | | Canthidium opacum Balthasar, 1939 | | | Canthidium orbiculatum (Lucas, 1857) | ⇒ Eucanthidium cupreum Blanchard, 1843 | | Canthidium pallidoalatum Howden & Young, 1981 | | | Canthidium paranum Harold, 1867 | | | Canthidium parvulum Harold, 1883 | | | Canthidium perceptibile Howden & Young, 1981 | | | Canthidium persplendens Balthasar, 1939 | | | Canthidium picipes Harold, 1867 | | | Canthidium pinotoides (Balthasar, 1939) | ⇒ Canthidium barbacenicum Preudhomme d..., 1886 | | Canthidium planovultum Howden & Young, 1981 | | | Canthidium prasinum Blanchard, 1846 | | | Canthidium priscillae Solis & Kohl..., 2004 | | | Canthidium pseudaurifex Balthasar, 1939 | | | Canthidium pseudoperceptibile Kohlmann & S..., 2006 | | | Canthidium pullus Felsche, 1910 | | | Canthidium punctatostriatum Mannerheim, 1829 | | | Canthidium punctatum Schmidt, 1922 | | | Canthidium puncticeps Harold, 1867 | | | Canthidium puncticolle Harold, 1867 | | | Canthidium puncticollis Harold, 1868 | | | Canthidium quadridens Harold, 1867 | | | Canthidium quercetorum Kohlmann, Ar..., 2018 | | | Canthidium refulgens Boucomont, 1928 | | | Canthidium riverai Kohlmann & S..., 2006 | | | Canthidium ruficolle Germar, 1824 | | | Canthidium rufinum Harold, 1867 | | | Canthidium rufipes Harold, 1867 | | | Canthidium rutilum Harold, 1867 | | | Canthidium seladon Balthasar, 1939 | | | Canthidium semicupreum Harold, 1868 | | | Canthidium sladeni Arrow, 1903 | | | Canthidium smaragdinum Harold, 1867 | | | Canthidium smithi Bates, 1889 | | | Canthidium splendidum Preudhomme d..., 1886 | | | Canthidium stahli Harold, 1867 | | | Canthidium stali Harold, 1867 | | | Canthidium steinheili Harold, 1880 | | | Canthidium stofeli Carvalho de ..., 2019 | | | Canthidium subdopuncticolle Howden & Young, 1981 | | | Canthidium sulcatum Perty, 1830 | | | Canthidium sulcicolle Harold, 1868 | | | Canthidium taurinum Harold, 1867 | | | Canthidium tenebrosum Howden & Young, 1981 | | | Canthidium thalassinum Erichson, 1847 | | | Canthidium titschacki Balthasar, 1939 | | | Canthidium tricolor Balthasar, 1939 | | | Canthidium trinodosum Boheman, 1858 | | | Canthidium tuberifrons Howden & Young, 1981 | | | Canthidium variolosum Howden & Young, 1981 | | | Canthidium versicolor Harold, 1867 | | | Canthidium vespertinum Howden & Young, 1981 | | | Canthidium violaceipenne Blanchard, 1846 | | | Canthidium viride Lucas, 1857 | | | Canthidium viridicolle Blanchard, 1846 | | | Canthidium viridiobscurum Boucomont, 1928 | | | Canthidium volxemi Preudhomme d..., 1886 | | | Canthochilum alleni Tarasov & Va..., 2016 | | | Canthochilum anacaona Zayas & Matt..., 1966 | | | Canthochilum andyi Chapin, 1935 | | | Canthochilum baracutey Zayas & Matt..., 1966 | | | Canthochilum boringuensis | | | Canthochilum borinquensis Matthews, 1965 | | | Canthochilum brodzinskyorum Philips & Ivie, 2008 | | | Canthochilum brozinskyorum | | | Canthochilum cemi Zayas & Matt..., 1966 | | | Canthochilum ciboney Matthews, 1969 | | | Canthochilum darlingtoni Matthews, 1969 | | | Canthochilum guayca Zayas & Matt..., 1966 | | | Canthochilum gundlachi Harold, 1868 | | | Canthochilum hispidum Chapin, 1934 | | | Canthochilum hispidum hispidum | | | Canthochilum hispidum iunceanum Matthews, 1965 | | | Canthochilum hispidum serropunctae Matthews, 1965 | | | Canthochilum histeroides Harold, 1868 | | | Canthochilum magnum Philips & Ivie, 2008 | | | Canthochilum matthewsi Philips & Ivie, 2008 | | | Canthochilum mimicum Philips & Ivie, 2008 | | | Canthochilum nebulonemi Philips & Ivie, 2008 | | | Canthochilum oakleyi Chapin, 1934 | | | Canthochilum philipsivieorum Tarasov & Va..., 2016 | | | Canthochilum pijirigua Zayas & Matt..., 1966 | | | Canthochilum platycnemis Philips & Ivie, 2008 | | | Canthochilum taino Matthews, 1966 | | | Canthochilum tureyra Zayas & Matt..., 1966 | | | Canthochilum xericum Philips & Ivie, 2008 | | | Canthodimorpha lawrencei Davis, Schol..., 1999 | | | Canthon (Boreocanthon) coahuilensis Howden, 1966 | Rockpool blenny | | Canthon (Boreocanthon) depressipennis LeConte, 1859 | | | Canthon (Boreocanthon) ebenus Say, 1823 | | | Canthon (Boreocanthon) forreri Bates, 1887 | | | Canthon (Boreocanthon) lecontei Harold, 1868 | | | Canthon (Boreocanthon) melanus Robinson, 1948 | | | Canthon (Boreocanthon) mixtus Robinson, 1948 | | | Canthon (Boreocanthon) praticola Le Conte, 1859 | | | Canthon (Boreocanthon) probus Germar, 1824 | | | Canthon (Boreocanthon) puncticollis (LeConte, 1866) | ⇒ Canthon punctatus Schmidt, 1922 | | Canthon puncticollis integricollis Schaeffer, 1915 | | | Canthon (Boreocanthon) simplex Le Conte, 1857 | | | Canthon (Canthon) chalcites Haldeman, 1843 | | | Canthon (Canthon) cyanellus LeConte, 1859 | | | Canthon cyanellus sallei (Harold, 1863) | ⇒ Canthon cyanellus LeConte, 1859 | | Canthon (Canthon) humectus Say, 1831 | Longhorn blenny | | Canthon humectus alvarengai Halffter, 1961 | | | Canthon humectus assimilis Robinson, 1946 | | | Canthon humectus blumei Halffter & H..., 2003 | | | Canthon humectus hidalgoensis Bates, 1887 | | | Canthon humectus humectus Say, 1832 | | | Canthon humectus incisus Robinson, 1948 | | | Canthon humectus riverai Halffter & H..., 2003 | | | Canthon humectus sayi Robinson, 1948 | | | Canthon (Canthon) imitator Brown, 1946 | | | Canthon imitator floridanus Brown, 1946 | | | Canthon (Canthon) indigaceus (LeConte, 1866) | ⇒ Canthon humectus Say, 1831 | | Canthon indigaceus chevrolati Harold, 1868 | | | Canthon indigaceus chiapas Robinson, 1948 | | | Canthon indigaceus indigaceus | | | Canthon (Canthon) obliquus Horn, 1894 | | | Canthon (Canthon) pilularius Linnaeus, 1758 | Dull tumblebug | | Canthon (Canthon) vigilans LeConte, 1858 | Kalia | | Canthon (Glaphyrocanthon) femoralis Chevrolat, 1834 | | | Canthon (Glaphyrocanthon) viridis (Palisot de B..., 1805) | ⇒ Epilissus viridis Klug, 1832 | | Canthon aberrans Harold, 1868 | | | Canthon acutiformis Balthasar, 1939 | | | Canthon acutoides (Schmidt, 1922) | ⇒ Glaphyrocanthon acutoides Schmidt, 1922 | | Canthon acutus Harold, 1868 | | | Canthon aequinoctialis Harold, 1868 | | | Canthon angularis Harold, 1868 | | | Canthon angularis angularis | | | Canthon angularis obenbergeri Balthasar, 1940 | | | Canthon angustatus Harold, 1867 | | | Canthon antoniomartinezi Rivera-Cerva..., 1999 | | | Canthon arcabuquensis Molano & Medina, 2010 | | | Canthon arriagadai Vieira, Vaz-..., 2019 | | | Canthon ateuchiceps Bates, 1887 | | | Canthon ateuchipes Bates, 1887 | | | Canthon auricollis Redtenbacher, 1867 | | | Canthon balachowskyi Martinez & H..., 1972 | | | Canthon balteatus Boheman, 1858 | | | Canthon bicolor Castelnau, 1840 | | | Canthon bifurcatus (Robinson, 1948) | ⇒ Canthon aberrans Harold, 1868 | | Canthon bimaculatus (Schmidt, 1922) | ⇒ Glaphyrocanthon bimaculatus Schmidt, 1922 | | Canthon bisignatus Balthasar, 1939 | | | Canthon bispinatus (Robinson, 1941) | ⇒ Melanocanthon bispinatus Robinson, 1941 | | Canthon bispinus Germar, 1824 | | | Canthon blumei Halffter & H..., 2003 | | | Canthon bonaerensis Vieira, Vaz-..., 2019 | | | Canthon brunneus (Schmidt, 1922) | ⇒ Glaphyrocanthon brunneus Schmidt, 1922 | | Canthon brunnipennis Schmidt, 1922 | | | Canthon caelius Bates, 1887 | | | Canthon callosus Harold, 1868 | | | Canthon candens Gistel, 1857 | | | Canthon carbonarius (Harold, 1868) | ⇒ Francmonrosia carbonarius Harold, 1868 | | Canthon championi (Bates, 1887) | ⇒ Glaphyrocanthon championi Bates, 1887 | | Canthon chiapas (Robinson, 1948) | ⇒ Canthon indigaceus chiapas Robinson, 1948 | | Canthon chiriguano Martinez & H..., 1972 | | | Canthon cinctellus Germar, 1824 | | | Canthon circulatus (Harold, 1868) | ⇒ Glaphyrocanthon circulatus Harold, 1868 | | Canthon cleidecostae Vaz-de-Mello..., 2020 | | | Canthon cobosi Pereira & Ma..., 1960 | | | Canthon coeruleicollis Blanchard, 1846 | | | Canthon coerulescens Schmidt, 1922 | | | Canthon coloratus (Schmidt, 1922) | ⇒ Glaphyrocanthon coloratus Schmidt, 1922 | | Canthon columbianus (Schmidt, 1920) | ⇒ Glaphyrocanthon columbianus Schmidt, 1920 | | Canthon corporaali Balthasar, 1939 | | | Canthon corpulentus Harold, 1868 | | | Canthon corruscus Castelnau, 1840 | | | Canthon curvipes Harold, 1868 | | | Canthon curvodilatatus Schmidt, 1920 | | | Canthon daguerrei Martínez, 1951 | | | Canthon darlingtoni (Paulian, 1939) | ⇒ Scybalocanthon darlingtoni Paulian, 1939 | | Canthon delgadoi Rivera-Cerva..., 1999 | | | Canthon delicatulus Balthasar, 1939 | | | Canthon deloyai Rivera-Cerva..., 1999 | | | Canthon delpontei Martinez & H..., 1972 | | | Canthon denticulatus Schmidt, 1922 | | | Canthon dentiger Harold, 1868 | | | Canthon deplanatus Harold, 1868 | | | Canthon deyrollei Harold, 1868 | | | Canthon dives Harold, 1868 | | | Canthon divinator Gistel, 1857 | | | Canthon ebeneus Say, 1823 | | | Canthon edentulus Harold, 1868 | | | Canthon edentulus granuliceps Felsche, 1910 | | | Canthon edmondsi Rivera-Cerva..., 1999 | | | Canthon enkerlini Martinez, Ha..., 1964 | | | Canthon euryscelis Bates, 1887 | | | Canthon fallax Harold, 1868 | | | Canthon floridanus Brown, 1946 | | | Canthon forcipatus Harold, 1868 | | | Canthon formosus Harold, 1868 | | | Canthon foveiventris (Schmidt, 1920) | ⇒ Sylvicanthon foveiventris Schmidt, 1920 | | Canthon fulgidus Redtenbacher, 1867 | | | Canthon fulgidus fulgidus | | | Canthon fulgidus martinezi Nunes, Nunes..., 2018 | | | Canthon fulgidus pereirai Nunes, Nunes..., 2018 | | | Canthon fuscipes Erichson, 1847 | | | Canthon gagatinus amethystinus (Harold, 1863) | ⇒ Canthon humectus Say, 1831 | | Canthon gemellatus Erichson, 1847 | | | Canthon granuliceps Felsche, 1910 | | | Canthon granulifer (Schmidt, 1920) | ⇒ Melanocanthon granulifer Schmidt, 1920 | | Canthon haroldi Nunes, Nunes..., 2020 | | | Canthon hartmanni Howden & Gill, 1987 | | | Canthon helleri (Schmidt, 1922) | ⇒ Glaphyrocanthon helleri Schmidt, 1922 | | Canthon hendrichsi Halffter & M..., 1968 | | | Canthon heyrovskyi Balthasar, 1939 | | | Canthon histrio (Lepeletier &..., 1828) | ⇒ Canthon septemmaculatus Latreille, 1812 | | Canthon honsi Balthasar, 1939 | | | Canthon hudsonias Forster, 1771 | | | Canthon humboldti Solis & Kohl..., 2002 | | | Canthon humeralis (Horn, 1870) | ⇒ Boreocanthon simplex LeConte, 1857 | | Canthon ibarragrassoi Martínez, 1952 | | | Canthon incisus | | | Canthon integricollis (Schaeffer, 1915) | ⇒ Boreocanthon puncticollis LeConte, 1866 | | Canthon inusitatus Kohlmann & S..., 2006 | | | Canthon janthinus Blanchard, 1846 | | | Canthon juanae (Martínez, 1949) | ⇒ Canthon aberrans Harold, 1868 | | Canthon juvencus Harold, 1868 | | | Canthon laesus (Erichson, 1847) | ⇒ Glaphyrocanthon rubrescens Blanchard, 1843 | | Canthon lafargei Drapiez, 1820 | | | Canthon laminatus Balthasar, 1939 | | | Canthon lamprimus Bates, 1887 | | | Canthon lamproderes Redtenbacher, 1867 | | | Canthon latipes (Blanchard, 1843) | ⇒ Francmonrosia latipes Blanchard, 1843 | | Canthon latipes atricornis Blanchard, 1846 | | | Canthon latipes latipes | | | Canthon latipes viridans Balthasar, 1939 | | | Canthon leechi Martinez, Ha..., 1964 | | | Canthon lituratus Germar, 1813 | | | Canthon lividus Blanchard, 1846 | | | Canthon lividus lividus | | | Canthon lividus seminitens Harold, 1868 | | | Canthon lucreciae Halffter & H..., 2009 | | | Canthon luctuosus Harold, 1868 | | | Canthon lunatus (Schmidt, 1922) | ⇒ Pseudepilissus lunatus Schmidt, 1922 | | Canthon lunatus tibialis Schmidt, 1922 | | | Canthon luteicollis Erichson, 1847 | | | Canthon machadoi Pereira & Ma..., 1967 | | | Canthon magnus Molano & Par..., 2015 | | | Canthon maldonadoi Martínez, 1951 | | | Canthon manantlanensis Rivera-Cerva..., 1999 | | | Canthon marii (Brèthes, 1925) | ⇒ Scatonomus insignis Harold, 1867 | | Canthon marmoratus Pereira & Ma..., 1956 | | | Canthon matthewsi Martinez & H..., 1972 | | | Canthon melancholicus Harold, 1868 | | | Canthon meridionalis Martinez, Ha..., 1964 | | | Canthon militaris (Horn, 1870) | ⇒ Boreocanthon simplex LeConte, 1857 | | Canthon modestus Harold, 1867 | | | Canthon moniliatus (Bates, 1887) | ⇒ Scybalocanthon moniliatus Bates, 1887 | | Canthon monilifer Blanchard, 1846 | | | Canthon montanus Rivera-Cerva..., 1999 | | | Canthon moroni Rivera-Cerva..., 1999 | | | Canthon morsei Howden, 1966 | | | Canthon mutabilis Lucas, 1857 | | | Canthon muticus Harold, 1867 | | | Canthon muticus hendrichsi Halffter & M..., 1968 | | | Canthon nigellus (Schmidt, 1922) | ⇒ Scybalocanthon nigellus Schmidt, 1922 | | Canthon nigricornis Say, 1823 | | | Canthon nigripennis Lansberge, 1874 | | | Canthon nyctelius Bates, 1887 | | | Canthon obscuriellus Schmidt, 1922 | | | Canthon obscurus (Schmidt, 1920) | ⇒ Sylvicanthon obscurus Schmidt, 1920 | | Canthon occidentalis Halffter & R..., 2015 | | | Canthon octodentatus Schmidt, 1920 | | | Canthon ohausi (Blackwelder, 1944) | ⇒ Canthon angustatus Harold, 1867 | | Canthon oliverioi Pereira & Ma..., 1956 | | | Canthon orfilai Martínez, 1949 | | | Canthon ornatus Redtenbacher, 1867 | | | Canthon ornatus bipunctatus Burmeister, 1873 | | | Canthon ornatus ornatus | | | Canthon ornatus thoracicus Harold, 1868 | | | Canthon pacificus Rivera-Cerva..., 1999 | | | Canthon pallidus (Schmidt, 1922) | ⇒ Glaphyrocanthon pallidus Schmidt, 1922 | | Canthon perplexum (LeConte, 1847) | ⇒ Pseudocanthon perplexus LeConte, 1847 | | Canthon perplexus LeConte, 1847 | | | Canthon perseverans Matthews, 1966 | | | Canthon pilularium Linnaeus, 1758 | | | Canthon piluliformis Blanchard, 1846 | | | Canthon plagiatus Harold, 1880 | | | Canthon planus (Lucas, 1857) | ⇒ Pseudepilissus planus Lucas, 1857 | | Canthon podagricus Harold, 1868 | | | Canthon politus Harold, 1868 | | | Canthon principalis Burmeister, 1873 | | | Canthon punctaticollis (Schaeffer, 1915) | ⇒ Melanocanthon punctaticollis Schaeffer, 1915 | | Canthon punctatus Schmidt, 1922 | | | Canthon pygidialis (Schmidt, 1922) | ⇒ Scybalocanthon pygidialis Schmidt, 1922 | | Canthon quadratus Blanchard, 1846 | | | Canthon quadriguttatus Olivier, 1789 | | | Canthon quadripunctatus Redtenbacher, 1867 | | | Canthon quinquemaculatus Castelnau, 1840 | | | Canthon reichei Felsche, 1910 | | | Canthon reyesi Martinez & H..., 1972 | | | Canthon riverai Halffter & H..., 2003 | | | Canthon rubrescens Blanchard, 1846 | | | Canthon rutilans (Castelnau, 1840) | ⇒ Francmonrosia rutilans Castelnau, 1840 | | Canthon rutilans cyanescens Harold, 1868 | | | Canthon rutilans rutilans | | | Canthon rzedowskii Rivera-Cerva..., 1999 | | | Canthon seminulus Harold, 1867 | | | Canthon semiopacus Harold, 1868 | | | Canthon septemmaculatus Latreille, 1812 | | | Canthon septemmaculatus cincticollis Lucas, 1857 | | | Canthon septemmaculatus histrio Lepeletier &..., 1828 | | | Canthon septemmaculatus septemmaculatus | | | Canthon sericatus Schmidt, 1922 | | | Canthon signifer (Harold, 1868) | ⇒ Nesocanthon signifer Harold, 1868 | | Canthon silvaticus Solis & Kohl..., 2002 | | | Canthon simulans Martínez, 1950 | | | Canthon smaragdulus Fabricius, 1781 | | | Canthon smaragdulus smaragdulus | | | Canthon smaragdulus subviridis Schmidt, 1922 | | | Canthon sordidus (Harold, 1868) | ⇒ Trichocanthon sordidus Harold, 1868 | | Canthon splendidus Schmidt, 1922 | | | Canthon staigi Pereira, 1953 | | | Canthon steinheili Harold, 1880 | | | Canthon subcyaneaus Erichson, 1848 | | | Canthon subcyaneus Erichson, 1848 | | | Canthon subhyalinus Harold, 1867 | | | Canthon subhyalinus subhyalinoides Balthasar, 1939 | | | Canthon subhyalinus subhyalinus | | | Canthon substriatus Harold, 1868 | | | Canthon sulcatus Castelnau, 1840 | | | Canthon terciae Nunes, Nunes..., 2020 | | | Canthon tetraodon Blanchard, 1846 | | | Canthon tibialis (Schmidt, 1922) | ⇒ Pseudepilissus lunatus Schmidt, 1922 | | Canthon triangularis Drury, 1770 | | | Canthon trimaculatus (Schmidt, 1922) | ⇒ Scybalocanthon trimaculatus Schmidt, 1922 | | Canthon unicolor Blanchard, 1846 | | | Canthon unicolor fortemarginatus Balthasar, 1939 | | | Canthon unicolor tristis Harold, 1862 | | | Canthon unicolor unicolor | | | Canthon variabilis Martínez, 1948 | | | Canthon vazquezae Martinez, Ha..., 1964 | | | Canthon velutinus Harold, 1868 | | | Canthon vidaurrei Vieira, Vaz-..., 2019 | | | Canthon viduus Harold, 1868 | | | Canthon violaceus Olivier, 1789 | | | Canthon virens Mannerheim, 1829 | | | Canthon virens chalybaeus Blanchard, 1846 | | | Canthon virens paraguayanus Balthasar, 1939 | | | Canthon virens scrutator Balthasar, 1939 | | | Canthon virens surinamensis Endrödi, 1962 | | | Canthon virens virens | | | Canthon vulcanoae Pereira & Ma..., 1956 | | | Canthon xanthopus (Blanchard, 1846) | ⇒ Sylvicanthon xanthopus Blanchard, 1846 | | Canthon ziggy Vieira, Vaz-..., 2019 | | | Canthon zuninoi Rivera-Cerva..., 1999 | | | Canthonella amazonica Ratcliffe & ..., 1999 | | | Canthonella baorucensis Ivie & Philips, 1990 | | | Canthonella barrerai Halffter & M..., 1968 | | | Canthonella catharinensis Pereira & Ma..., 1956 | | | Canthonella constans Matthews, 1966 | | | Canthonella gomezi Halffter & M..., 1968 | | | Canthonella howdeni Ivie & Philips, 1990 | | | Canthonella instriata Boucomont, 1928 | | | Canthonella isabellae Matthews, 1966 | | | Canthonella jarmilae Ivie & Philips, 2008 | | | Canthonella lenkoi Pereira & Ma..., 1956 | | | Canthonella leptoculata Ratcliffe & ..., 1999 | | | Canthonella parva Chapin, 1930 | | | Canthonella parva luquillensis Matthews, 1965 | | | Canthonella parva parva | | | Canthonella pygmaea Harold, 1869 | | | Canthonella quesqueya Ivie & Philips, 2008 | | | Canthonella sikesi Ivie & Philips, 2008 | | | Canthonella silphoides Harold, 1867 | | | Canthonidia hirsuta Paulian, 1939 | | | Canthonidia rubromaculata Blanchard, 1846 | | | Canthonosoma castelnaui Harold, 1868 | | | Canthonosoma macleayi Harold, 1868 | | | Canthonosoma mastersi MacLeay, 1871 | | | Canthonosoma mastersii MacLeay, 1871 | | | Canthotrypes oberthuri Paulian, 1939 | | | Cassolus fukiensis Balthasar, 1960 | | | Cassolus gotoi Masumoto, 1986 | | | Cassolus humeralis Arrow, 1907 | | | Cassolus javanus Boucomont, 1914 | | | Cassolus minutus Boucomont, 1914 | | | Cassolus nudus Sharp, 1875 | | | Cassolus peninsularis Arrow, 1907 | | | Cassolus pongchaii Masumoto, 1989 | | | Cassolus sumatranus Lansberge, 1885 | | | Catharsiocopris quadrituberculatus Balthasar, 1967 | | | Catharsius achates Olivier, 1789 | | | Catharsius adamastor Gillet, 1932 | | | Catharsius aegeus Génier, 2017 | | | Catharsius aethiops Sharp, 1875 | | | Catharsius africanus Ferreira, 1960 | | | Catharsius alpheus Gillet, 1932 | | | Catharsius approximans Kolbe, 1893 | | | Catharsius balubanus Kolbe, 1893 | | | Catharsius bellus Gillet, 1918 | | | Catharsius belus Gillet, 1918 | | | Catharsius biconifer Felsche, 1907 | | | Catharsius bicornutus Kolbe, 1893 | | | Catharsius birmanensis Lansberge, 1874 | | | Catharsius borneensis (Paulian, 1936) | ⇒ Catharsius molossus Linnaeus, 1758 | | Catharsius bradshawi Lansberge, 1887 | | | Catharsius brevicornis Felsche, 1907 | | | Catharsius brittoni Ferreira, 1963 | | | Catharsius brutus Harold, 1880 | | | Catharsius calaharicus Kolbe, 1893 | | | Catharsius capucinus Fabricius, 1781 | | | Catharsius cassius Kolbe, 1893 | | | Catharsius chinai Ferreira, 1960 | | | Catharsius congolensis Ferreira, 1962 | | | Catharsius coptorhinaformis Josso & Prév..., 2009 | | | Catharsius crassicornis Gillet, 1911 | | | Catharsius dayacus Lansberge, 1886 | | | Catharsius disseptus (Péringuey, 1901) | ⇒ Catharsius calaharicus Kolbe, 1893 | | Catharsius dubius (Péringuey, 1901) | ⇒ Catharsius molossus Linnaeus, 1758 | | Catharsius duciformis Ferreira, 1959 | | | Catharsius dux Harold, 1878 | | | Catharsius erechtheus Felsche, 1907 | | | Catharsius eteocles (Laporte, 1840) | ⇒ Catharsius alpheus Gillet, 1932 | | Catharsius fastidiosus Thomson, 1858 | | | Catharsius folicornis Josso & Prév..., 2019 | | | Catharsius furcillatus Guérin-Ménev..., 1847 | | | Catharsius furcillatus furcillatus Guérin, 1847 | | | Catharsius furcillatus laeviplaga G. Müller, 1941 | | | Catharsius garambae Ferreira, 1962 | | | Catharsius genieri Moretto, 2014 | | | Catharsius gorilla Thomson, 1858 | | | Catharsius gorilloides Felsche, 1907 | | | Catharsius granulatus Sharp, 1875 | | | Catharsius guineensis Ferreira, 1963 | | | Catharsius haafi Ferreira, 1960 | | | Catharsius haroldi Takano, 2021 | | | Catharsius harpagus Harold, 1877 | | | Catharsius heros Boheman, 1860 | | | Catharsius hertli Ferreira, 1964 | | | Catharsius jacksoni Waterhouse, 1891 | | | Catharsius javanus Lansberge, 1886 | | | Catharsius juheli Josso, 2011 | | | Catharsius kangeanus (Paulian, 1936) | ⇒ Catharsius molossus Linnaeus, 1758 | | Catharsius karlwerneri Moretto, 2008 | | | Catharsius kolbei Felsche, 1907 | | | Catharsius laticeps Boheman, 1857 | | | Catharsius latifossa Müller, 1941 | | | Catharsius lepineyi Boucomont, 1934 | | | Catharsius longiceps Gillet, 1907 | | | Catharsius luluensis Ferreira, 1960 | | | Catharsius lycaon Kolbe, 1893 | | | Catharsius machadoi Ferreira, 1960 | | | Catharsius marcellus Kolbe, 1893 | | | Catharsius melancholicus (Boheman, 1860) | ⇒ Catharsius longiceps Gillet, 1907 | | Catharsius minutus (Lansberge, 1882) | ⇒ Metacatharsius minutus Lansberge, 1882 | | Catharsius mirabilis Felsche, 1901 | | | Catharsius molossus Linnaeus, 1758 | | | Catharsius morettoi Josso & Prév..., 2009 | | | Catharsius mossambicanus Ferreira, 1960 | | | Catharsius mulanjeensis Josso & Prév..., 2019 | | | Catharsius nathani Frey, 1956 | | | Catharsius neptunus Kolbe, 1893 | | | Catharsius ninus Gillet, 1918 | | | Catharsius oedipus Felsche, 1907 | | | Catharsius omoensis Müller, 1941 | | | Catharsius oryx Frey, 1974 | | | Catharsius pallas Génier, 2017 | | | Catharsius pandion Harold, 1877 | | | Catharsius parafastidiosus Ferreira, 1971 | | | Catharsius peleus (Olivier, 1789) | ⇒ Metacatharsius peleus Olivier, 1789 | | Catharsius phidias (Olivier, 1789) | ⇒ Catharsius platicornis Ferreira, 1960 | | Catharsius philus Kolbe, 1893 | | | Catharsius pithecius Fabricius, 1775 | | | Catharsius platicornis Ferreira, 1960 | | | Catharsius platycerus Klug, 1855 | | | Catharsius platynotus Felsche, 1907 | | | Catharsius platypus Sharp, 1875 | | | Catharsius polynices Kolbe, 1893 | | | Catharsius pseudocongolensis Ferreira, 1963 | | | Catharsius pseudolycaon Ferreira, 1960 | | | Catharsius pseudooedipus Ferreira, 1960 | | | Catharsius pseudoplaticornis Josso & Prév..., 2019 | | | Catharsius pseudovitulus Müller, 1941 | | | Catharsius punctatus Josso & Prév..., 2019 | | | Catharsius quadridentatus Lansberge, 1885 | | | Catharsius renaudpauliani Ochi & Kon, 1996 | | | Catharsius rhinoceros Klug, 1855 | | | Catharsius saegeri Ferreira, 1962 | | | Catharsius sagax Quenstedt, 1806 | | | Catharsius satyrus Kolbe, 1893 | | | Catharsius scopas Felsche, 1907 | | | Catharsius semirubidus Fairmaire, 1891 | | | Catharsius sesostris Waterhouse, 1888 | | | Catharsius severini Gillet, 1907 | | | Catharsius simillimus Ferreira, 1960 | | | Catharsius somalicus Gillet, 1918 | | | Catharsius spectabilis Gillet, 1918 | | | Catharsius straeleni Ferreira, 1963 | | | Catharsius stricticornis Josso & Prév..., 2019 | | | Catharsius stuhlmanni Kolbe, 1893 | | | Catharsius takanoi Josso & Prév..., 2019 | | | Catharsius timorensis (Lansberge, 1879) | ⇒ Catharsius molossus Linnaeus, 1758 | | Catharsius tricornutus DeGeer, 1778 | | | Catharsius tridens Josso & Prév..., 2009 | | | Catharsius ugandicus Gillet, 1918 | | | Catharsius ulysses Boheman, 1857 | | | Catharsius upembanus Ferreira, 1962 | | | Catharsius vansoni Ferreira, 1960 | | | Catharsius vitulus Boheman, 1857 | | | Cephalodesmius armiger Westwood, 1842 | | | Cephalodesmius laticollis Pascoe, 1863 | | | Cephalodesmius quadridens MacLeay, 1871 | | | Chalcocopris hesperus Olivier, 1789 | | | Chalcocopris inexpectatus Rossini & Va..., 2015 | | | Cheironitis arrowi Janssens, 1937 | | | Cheironitis asbenicus Gillet, 1909 | | | Cheironitis audens Péringuey, 1901 | | | Cheironitis candezei Lansberge, 1875 | | | Cheironitis dentifemoralis Ferreira, 1976 | | | Cheironitis eumenes Gebler, 1860 | | | Cheironitis flabellatus Boucomont, 1923 | | | Cheironitis furcifer Rossi, 1792 | | | Cheironitis haroldi Ballion, 1870 | | | Cheironitis hoplosternus Harold, 1868 | | | Cheironitis hungaricus Herbst, 1789 | | | Cheironitis imitator Balthasar, 1963 | | | Cheironitis indicus (Lansberge, 1875) | ⇒ Cheironitis arrowi Janssens, 1937 | | Cheironitis irroratus Rossi, 1790 | | | Cheironitis klapperichi (Balthasar, 1955) | ⇒ Cheironitis furcifer Rossi, 1792 | | Cheironitis moeris Pallas, 1781 | | | Cheironitis muelleri Janssens, 1943 | | | Cheironitis osiridis Reiche, 1856 | | | Cheironitis pamphilus Ménétriés, 1849 | | | Cheironitis petraensis Coppo, 2021 | | | Cheironitis phoebus Reitter, 1892 | | | Cheironitis scabrosus Fabricius, 1776 | | | Cheironitis socotranus Gahan, 1900 | | | Cheironitis sterculius Ballion, 1870 | | | Cheironitis ungaricus Herbst, 1789 | | | Cheironitis ungaricus irroratus (Rossi, 1790) | ⇒ Cheironitis irroratus Rossi, 1790 | | Cheironitis ungaricus ungaricus Herbst, 1789 | | | Cheironitis viridicans Gillet, 1918 | | | Chironitis furcifer Rossi, 1792 | | | Chironitis hungaricus (Herbst, 1789) | ⇒ Cheironitis ungaricus Herbst, 1789 | | Chironitis irroratus Rossi, 1790 | | | Chironitis moeris Pallas, 1781 | | | Chironitis pamphilus Ménétriése, 1849 | | | Choeridium ebeninum ♰ G. H. Horn, 1876 | | | Choeridium insulare (Fleutiaux & ..., 1889) | ⇒ Ateuchus illaesus Harold, 1868 | | Choeridium loricatum (Boucomont, 1928) | ⇒ Ateuchus loricatus Boucomont, 1928 | | Choeridium punctatus (Robinson, 1948) | ⇒ Ateuchus histeroides Weber, 1801 | | Choeridium steinbachi (Boucomont, 1928) | ⇒ Ateuchus steinbachi Boucomont, 1928 | | Choeridium texanus (Robinson, 1948) | ⇒ Ateuchus texanus Robinson, 1948 | | Circellium bacchus Fabricius, 1781 | Flightless dung beetle | | Cleptocaccobius arrowi Cambefort, 1985 | | | Cleptocaccobius balthasarianus Cambefort, 1979 | | | Cleptocaccobius biceps d´Orbigny, 1905 | | | Cleptocaccobius boucomonti Cambefort, 1985 | | | Cleptocaccobius clementi Cambefort, 1971 | | | Cleptocaccobius convexifrons Raffray, 1877 | | | Cleptocaccobius dorbignyi Cambefort, 1984 | | | Cleptocaccobius durantoni Cambefort, 1985 | | | Cleptocaccobius humilis d´Orbigny, 1902 | | | Cleptocaccobius inermis Arrow, 1931 | | | Cleptocaccobius khatimae Cambefort, 1985 | | | Cleptocaccobius lamottei Cambefort, 1983 | | | Cleptocaccobius minor Walter & Cam..., 1977 | | | Cleptocaccobius morettoi Josso & Prév..., 2006 | | | Cleptocaccobius postlutatus d´Orbigny, 1905 | | | Cleptocaccobius schaedlei d´Orbigny, 1902 | | | Cleptocaccobius scrofa Balthasar, 1942 | | | Cleptocaccobius semiluteus d´Orbigny, 1905 | | | Cleptocaccobius seminulum Klug, 1855 | | | Cleptocaccobius signaticollis d´Orbigny, 1902 | | | Cleptocaccobius simplex Boucomont, 1923 | | | Cleptocaccobius simplex meridionalis Cambefort, 1985 | | | Cleptocaccobius simplex simplex | | | Cleptocaccobius uniseries d´Orbigny, 1905 | | | Cleptocaccobius viridicollis Fahraeus, 1857 | | | Cleptocaccobius youngai Balthasar, 1967 | | | Clypeodrepanus digitatus Krikken, 2009 | | | Clypeodrepanus striatus Boucomont, 1921 | | | Clypeodrepanus strigatus Janssens, 1953 | | | Copridaspidus peregrinus Harold, 1878 | | | Copris (Copris) cambodiensis Ochi, Kon & ..., 2008 | | | Copris cambodiensis cambodiensis | | | Copris cambodiensis masumotoi Ochi, Kon & ..., 2011 | | | Copris (Copris) fukiensis Balthasar, 1952 | | | Copris (Copris) kachinensis Ochi, Kon & ..., 2008 | | | Copris (Copris) kartlinus ♰ O. N. Kabakov, 1988 | | | Copris (Copris) punctatus Gillet, 1910 | | | Copris punctatus formosanus Gillet, 1922 | | | Copris punctatus punctatus | | | Copris (Copris) schoolmeestersi Ochi & Kon, 2010 | | | Copris (Copris) tripartitus Waterhouse, 1875 | | | Copris (Microcopris) propinquus Felsche, 1910 | | | Copris (Paracopris) cariniceps Felsche, 1910 | | | Copris (Sinocopris) barclayi Ochi, Kon & Bai, 2009 | | | Copris (Sinocopris) pecuarius Lewis, 1884 | | | Copris pecuarius continentalis Ochi & Kon, 2010 | | | Copris (Sinocopris) uenoi Hanboonsong ..., 2003 | | | Copris achates (Péringuey, 1901) | ⇒ Copris cornifrons Boheman, 1860 | | Copris acutidens Motschulsky, 1860 | | | Copris afgoi Cambefort, 1992 | | | Copris agnus Sharp, 1875 | | | Copris algol Nguyen-phung, 1989 | | | Copris amabilis Kolbe, 1914 | | | Copris amazonicus Darling & Gé..., 2018 | | | Copris amyntor (Klug, 1855) | ⇒ Copris harrisi Waterhouse, 1891 | | Copris anceus Olivier, 1789 | | | Copris andrewesi Waterhouse, 1891 | | | Copris angusticornis Arrow, 1933 | | | Copris angustus Nguyen-phung..., 1986 | | | Copris anomiopseoides (Boucomont, 1924) | ⇒ Paracopris anomiopseoides Boucomont, 1924 | | Copris antares Bacescu & Ca..., 1947 | | | Copris apicepunctatus Balthasar, 1942 | | | Copris arcturus Gillet, 1907 | | | Copris ares Zelenka, 1993 | | | Copris arimotoi Ochi & Araya, 1994 | | | Copris arizonensis Schaeffer, 1906 | | | Copris armatus Harold, 1869 | | | Copris armeniacus Falderman, 1835 | | | Copris armiger Gillet, 1910 | | | Copris arrowi Felsche, 1910 | | | Copris aspericollis Gillet, 1910 | | | Copris atropolitus Gillet, 1910 | | | Copris bartolozzi Ochi, Kon & ..., 2019 | | | Copris basilewskyi Ferreira, 1962 | | | Copris basipunctatus Balthasar, 1942 | | | Copris bellator Chevrolat, 1844 | | | Copris bengalensis Shiino, 1978 | | | Copris bidens (Pfeffer, 1888) | ⇒ Liatongus femoratus Illiger, 1800 | | Copris bihamatus Balthasar, 1965 | | | Copris binodis Thunberg, 1818 | | | Copris bootes Klug, 1855 | | | Copris boucardi Harold, 1869 | | | Copris bovinus Gillet, 1908 | | | Copris brachypterus Nomura, 1964 | | | Copris brachypterus brachypterus | | | Copris brachypterus toguchii Ochi & Kawahara, 2012 | | | Copris brevicornis Darling & Gé..., 2018 | | | Copris brittoni Ferreira, 1962 | | | Copris caelatus Fabricius, 1794 | | | Copris caliginosus Kohlmann, Ca..., 2003 | | | Copris calvus Sharp, 1875 | | | Copris cambeforti Nguyen-phung, 1988 | | | Copris camerunus Felsche, 1904 | | | Copris caobangensis Bui, Dumack ..., 2018 | | | Copris capensis Waterhouse, 1891 | | | Copris carinicus Gillet, 1910 | | | Copris carmelita Fabricius, 1801 | | | Copris cartwrighti (Robinson, 1941) | ⇒ Copris fricator cartwrighti Robinson, 1941 | | Copris cassius Péringuey, 1901 | | | Copris celebensis Lansberge, 1886 | | | Copris cheni Ochi, Kon & Bai, 2007 | | | Copris chimalapensis Mora-Aguilar..., 2015 | | | Copris chinmontanus Ochi, Kon & ..., 2019 | | | Copris ciliatus | | | Copris colmanti Gillet, 1908 | | | Copris complexus Nguyen-phung, 1989 | Robber Baron Cave meshweaver | | Copris compressipennis (Gillet, 1910) | ⇒ Paracopris compressipennis Gillet, 1910 | | Copris confucius Harold, 1877 | | | Copris continentalis Ochi & Kon, 2010 | | | Copris coriarius Gillet, 1907 | | | Copris corniculatus (Mulsant, 1842) | ⇒ Copris lunaris Linnaeus, 1758 | | Copris cornifrons Boheman, 1860 | | | Copris corniger (Sahlberg, 1823) | ⇒ Onthophagus taurus Schreber, 1759 | | Copris corpulentus Gillet, 1910 | | | Copris costaricensis Gahan, 1894 | | | Copris costaricensis costaricensis | | | Copris costaricensis dolichocerus Matthews, 1961 | | | Copris costariensis | | | Copris crassus Deschodt & D..., 2015 | | | Copris cribratus Gillet, 1927 | | | Copris cribricollis (Gillet, 1910) | ⇒ Catharsius pithecius Fabricius, 1775 | | Copris curlettii Moretto, 2011 | | | Copris davidi Darling & Gé..., 2018 | | | Copris davidis Deyrolle, 1878 | | | Copris davisi Nguyen-phung..., 1986 | | | Copris davisoni Waterhouse, 1891 | | | Copris decellei Cambefort, 1992 | | | Copris delicatus Arrow, 1931 | | | Copris denticulatus Nguyen-phung, 1988 | | | Copris diversus Waterhouse, 1891 | | | Copris doriae Harold, 1877 | | | Copris draco Arrow, 1906 | | | Copris dracunculus Fereira, 1959 | | | Copris druidum ♰ O. Heer, 1862 | | | Copris dudleyi Cambefort, 1992 | | | Copris eburneus Cambefort, 1992 | | | Copris elpheneroides Felsche, 1910 | | | Copris elphenor Klug, 1855 | | | Copris erratus Lansberge, 1886 | | | Copris erratus erratus | | | Copris erratus masayukii Ochi & Kon, 2004 | | | Copris evanidus Klug, 1855 | | | Copris excisus Waterhouse, 1891 | | | Copris fallaciosus Gillet, 1907 | Kenyan dung beetle | | Copris fallax Felsche, 1910 | | | Copris felschei (Reitter, 1893) | ⇒ Copris armeniacus Falderman, 1835 | | Copris fidius Olivier, 1789 | | | Copris frankenbergerianus Balthasar, 1958 | | | Copris freyi Ferreira, 1961 | | | Copris fricator Fabricius, 1787 | | | Copris fricator cartwrighti Robinson, 1941 | | | Copris fricator fricator | | | Copris fujiokai Ochi & Kon, 1996 | | | Copris fujiokai poringensis Ochi & Kon, 2005 | | | Copris furciceps Felsche, 1910 | | | Copris garambae Cambefort, 1992 | | | Copris gazellarum Gillet, 1918 | | | Copris gibbulus Lansberge, 1886 | | | Copris gibbulus borneensis Ochi & Kon, 2005 | | | Copris gibbulus gibbulus | | | Copris gilleti Kolbe, 1907 | | | Copris gladiator Arrow, 1933 | | | Copris gopheri Hubbard, 1894 | Copris tortoise commensal scarab | | Copris gracilis Waterhouse, 1891 | | | Copris graueri (Kolbe, 1914) | ⇒ Copris camerunus Felsche, 1904 | | Copris halffteri Matthews, 1959 | | | Copris harrisi Waterhouse, 1891 | | | Copris hartli Frey, 1975 | | | Copris hatayamai Ochi & Araya, 1992 | | | Copris hidakai Ochi & Kon, 1996 | | | Copris hintoni (Saylor, 1933) | ⇒ Copris klugi Harold, 1869 | | Copris hispanus Linnaeus, 1764 | | | Copris hispanus cavolinii Petagna, 1792 | | | Copris hispanus hispanus Linné, 1764 | | | Copris hispanus persianus Miksic, 1958 | | | Copris hosakai Ochi & Kon, 2014 | | | Copris howdeni Matthews & H..., 1959 | | | Copris humilis Gillet, 1908 | | | Copris hybridus Gillet, 1910 | | | Copris igualensis Warner, 1990 | | | Copris illotus Balthasar, 1942 | | | Copris imitans Felsche, 1910 | | | Copris inaequabilis Zhang, 1997 | | | Copris incertus Say, 1835 | Black dung beetle | | Copris indicus (Gillet, 1910) | ⇒ Copris fricator Fabricius, 1787 | | Copris inemarginatus Blatchley, 1918 | | | Copris inhalatus Quedenfeldt, 1884 | | | Copris inhalatus grandis Nguyen-phung..., 1986 | | | Copris inhalatus inhalatus | | | Copris inhalatus ngotto Moretto, 2011 | | | Copris inhalatus perturbator Péringuey, 1901 | | | Copris inhalatus sanctaeluciae Nguyen-phung..., 1986 | | | Copris insidiosus Péringuey, 1901 | | | Copris integer (Reiche, 1847) | ⇒ Copris fallaciosus Gillet, 1907 | | Copris interioris Kolbe, 1897 | | | Copris iris Sharp, 1875 | | | Copris jacchoides Nguyen-phung..., 1987 | | | Copris jacchus Lang, 1971 | | | Copris jahi Nguyen-phung..., 1986 | | | Copris javanensis Bui, Ziegler..., 2019 | | | Copris johkii Ochi & Kon, 2014 | | | Copris juanmorai Mora-Aguilar..., 2015 | | | Copris jucundus Gillet, 1932 | | | Copris kasagii Ochi & Kon, 1996 | | | Copris kasanka Josso & Prév..., 2016 | | | Copris katangae Cambefort & ..., 1996 | | | Copris keralensis Gill, 1986 | | | Copris kimioi Ochi & Kon, 2021 | | | Copris kiriromensis Ochi & Kon, 2021 | | | Copris kiuchii Masumoto, 1989 | | | Copris klapperichi Balthasar, 1942 | | | Copris klugi Harold, 1869 | | | Copris klugi sierrensis (Matthews, 1961) | ⇒ Copris sierrensis Matthews, 1961 | | Copris kusakabei Ochi, Kon & ..., 2005 | | | Copris laeviceps Harold, 1869 | | | Copris laevigatus (Gillet, 1927) | ⇒ Onthophagus laevigatus Fabricius, 1798 | | Copris laiiformis Nguyen-phung..., 1987 | | | Copris laioides Boucomont, 1932 | | | Copris laius Harold, 1868 | | | Copris laius angusticornis Nguyen-phung..., 1987 | | | Copris laius laius | | | Copris lannathai Hanboonsong ..., 2003 | | | Copris laosensis Ochi & Kon, 2021 | | | Copris leakeyorum ♰ R. Paulian, 1976 | | | Copris lecontei Matthews, 1961 | | | Copris lecontei isthmiensis Matthews, 1961 | | | Copris lecontei lecontei Matthews, 1962 | | | Copris leruei Josso & Prév..., 2017 | | | Copris lugubris Boheman, 1858 | | | Copris lunaris Linnaeus, 1758 | Horned Dung Beetle | | Copris macacus Lansberge, 1886 | | | Copris macclevei Warner, 1990 | | | Copris macer Péringuey, 1901 | | | Copris maesi Ratcliffe, 1998 | | | Copris magicus Harold, 1881 | | | Copris manni Ochi, Kon & Bai, 2009 | | | Copris marcoberrai Ochi, Kon & ..., 2019 | | | Copris marcus Gillet, 1921 | | | Copris martinae Nguyen-phung, 1988 | | | Copris martinae martinae | | | Copris martinae ntchisi Moretto, 2011 | | | Copris martinezi Matthews & H..., 1968 | Streaked blenny | | Copris matthewsi Delgado & Ko..., 2001 | | | Copris matthewsi matthewsi | | | Copris matthewsi pacificus Delgado & Ko..., 2001 | | | Copris medogensis Y.-W. Zhang, 1988 | | | Copris megaceratoides Waterhouse, 1891 | | | Copris megasoma Matthews & H..., 1959 | | | Copris mesacanthus Harold, 1878 | | | Copris mesacanthus mesacanthus | | | Copris mesacanthus transvaalensis Nguyen-phung, 1988 | | | Copris mexicanus Matthews & H..., 1959 | | | Copris micros Ochi & Kon, 2021 | | | Copris mindorensis Ochi, 1992 | | | Copris minutus Drury, 1773 | Small Black Dung Beetle | | Copris misellus Péringuey, 1901 | | | Copris miyakei Ochi & Kon, 2004 | | | Copris moecha LeConte, 1854 | | | Copris moechus LeConte, 1854 | | | Copris moffartsi Gillet, 1907 | | | Copris mongkhoni Hanboonsong ..., 2003 | | | Copris montivagus (Gillet, 1908) | ⇒ Copris harrisi Waterhouse, 1891 | | Copris montreuili Ochi, Kon & ..., 2019 | | | Copris morettoi Josso & Prév..., 2020 | | | Copris moroni Darling & Gé..., 2018 | | | Copris morphaeus Gillet, 1932 | | | Copris mourgliai Zelenka, 1993 | | | Copris muticus (Boheman, 1857) | ⇒ Litocopris muticus Boheman, 1857 | | Copris neglectus Moxey, 1963 | | | Copris neotridens Génier & Pré..., 2017 | | | Copris nepos Gillet, 1908 | | | Copris nevinsoni Waterhouse, 1891 | Warton meshweaver | | Copris nonarmatus Ochi & Kon, 2021 | | | Copris novaki Zelenka, 1992 | | | Copris nubilosus Kohlmann & C..., 2003 | | | Copris numa Lansberge, 1886 | Braken Bat Cave meshweaver | | Copris obenbergeri Balthasar, 1933 | Government Canyon Bat Cave meshweaver | | Copris obesus Boheman, 1857 | | | Copris ochus Motschulsky, 1860 | | | Copris olivieri Illiger, 1800 | | | Copris orion Klug, 1835 | | | Copris orion caffer Gillet, 1908 | | | Copris orion centralis Gillet, 1908 | | | Copris orion orion | | | Copris orion overlaeti Balthasar, 1961 | | | Copris orphanus (Guérin-Ménev..., 1847) | ⇒ Copris evanidus Klug, 1855 | | Copris parapecuarius Ochi, Kon & ..., 2008 | | | Copris parasonensis Ochi, Kon & ..., 2019 | | | Copris paraszechouanicus Ochi, Kon & ..., 2018 | | | Copris pauliani Nguyen-phung..., 1986 | | | Copris peculiariprotibialis Ochi & Kon, 2023 | | | Copris pedarioides Lansberge, 1886 | | | Copris perturbator (Péringuey, 1901) | ⇒ Copris inhalatus perturbator Péringuey, 1901 | | Copris phungae Cambefort, 1992 | | | Copris phungae gabonicus Cambefort, 1992 | | | Copris phungae phungae | | | Copris phylax Gillet, 1908 | | | Copris poggii Ochi & Kon, 2005 | | | Copris popaensis Ochi, Kon & ..., 2019 | | | Copris potanini Semenov, 1891 | | | Copris prevosti Josso, 2011 | | | Copris pristinus ♰ W. D. Pierce, 1946 | | | Copris pseudobootes Ferreira, 1962 | | | Copris pseudochus Ochi & Kon, 2004 | | | Copris pseudochus basipunctulatus Ochi, Kon & ..., 2020 | | | Copris pseudochus chuduraziensis Ochi, Kon & ..., 2019 | | | Copris pseudochus pseudochus | | | Copris pseudomoffartsi Moretto, 2011 | | | Copris pseudosinicus Balthasar, 1958 | | | Copris pueli Mollandin De..., 1905 | | | Copris puncticollis Boheman, 1857 | | | Copris punctipennis Boucomont, 1914 | | | Copris punctiventris Waterhouse, 1891 | | | Copris punctulatus Wiedemann, 1823 | | | Copris punctulatus punctulatus | | | Copris punctulatus septentrionalis Ochi, Kon & ..., 2019 | | | Copris punjabensis Gillet, 1921 | | | Copris quadridens (Say, 1835) | ⇒ Phanaeus quadridens Say, 1835 | | Copris quasilaevigatus Ochi, Kon & Bai, 2007 | | | Copris ramosiceps Gillet, 1921 | | | Copris rebouchei Harold, 1869 | | | Copris reflexus (Fabricius, 1787) | ⇒ Copris minutus Drury, 1773 | | Copris remotus LeConte, 1866 | | | Copris remotus dicyrtus Matthews & H..., 1959 | | | Copris remotus remotus LeConte, 1866 | | | Copris renwarti Nguyen-phung..., 1986 | | | Copris repertus Walker, 1858 | | | Copris ribbei Lansberge, 1886 | | | Copris riekoae Ochi & Kon, 2003 | | | Copris ritsemae Harold, 1875 | | | Copris ruandanus Balthasar, 1961 | | | Copris rugosus (Gillet, 1908) | ⇒ Copris inhalatus Quedenfeldt, 1884 | | Copris ruricola Balthasar, 1933 | | | Copris sabinus Gillet, 1910 | | | Copris sacontala Redtenbacher, 1848 | | | Copris sallei Harold, 1869 | | | Copris sarpedon Harold, 1868 | | | Copris saundersi Harold, 1869 | | | Copris scorpio Zelenka, 1993 | | | Copris serius Nguyen-phung, 1987 | | | Copris sexdentatus (Thunberg, 1818) | ⇒ Copris sarpedon Harold, 1868 | | Copris siamensis Gillet, 1910 | | | Copris siangensis Biswas, 1980 | | | Copris sierrensis Matthews, 1961 | | | Copris signatus Walker, 1858 | | | Copris simonettai Cambefort, 1992 | | | Copris simplex (Harold, 1869) | ⇒ Dichotomius simplex Taschenberg, 1870 | | Copris simulator Nguyen-phung..., 1986 | | | Copris singularis Nguyen-phung..., 1986 | | | Copris sinicus Hope, 1842 | | | Copris sinon Olivier, 1789 | | | Copris sodalis Walker, 1858 | | | Copris sonensis Bui, Dumack ..., 2018 | | | Copris sorex Balthasar, 1942 | | | Copris sphaeropterus Harold, 1877 | | | Copris spinator Harold, 1881 | | | Copris subdolus Balthasar, 1958 | | | Copris subpunctatus Gillet, 1910 | | | Copris subsidens Péringuey, 1901 | | | Copris subterranea ♰ O. Heer, 1862 | | | Copris subterraneus Heer, 1862 | | | Copris sumatrensis Gillet, 1921 | | | Copris surdus Arrow, 1931 | | | Copris susanae Darling & Gé..., 2018 | | | Copris szechouanicus Balthasar, 1958 | | | Copris takanoi Ochi, Kon & Bai, 2020 | | | Copris teramachii Ochi, Kon & Bai, 2020 | | | Copris tetraodon Gillet, 1910 | | | Copris tricorniprothoracicus Ochi, Kon & ..., 2020 | | | Copris tridens (Felsche, 1901) | ⇒ Copris hispanus hispanus Linné, 1764 | | Copris tridentatus Solis & Kohl..., 2003 | | | Copris truncatus Felsche, 1901 | | | Copris tsukamotoi Ochi, Kon & ..., 2008 | | | Copris tullius Olivier, 1789 | | | Copris typhoeus Gerstaecker, 1884 | | | Copris umbilicatus Abeille de P..., 1901 | | | Copris urus (Boheman, 1857) | ⇒ Catharsius molossus Linnaeus, 1758 | | Copris usambaricus Gillet, 1908 | | | Copris vankhaii Nguyen-phung, 1988 | | | Copris vankhaii genopunctatus Nguyen-phung, 1988 | | | Copris vankhaii vankhaii | | | Copris vankhalii Nguyen-phung, 1988 | | | Copris victorini (Boheman, 1857) | ⇒ Copris ritsemae Harold, 1875 | | Copris vietnamicus Kabakov, 1994 | | | Copris vietnamimontanus Ochi, Kon & ..., 2018 | | | Copris vilhenai Ferreira, 1962 | | | Copris vitalisi Gillet, 1921 | | | Copris vrydaghi Ferreira, 1962 | | | Copris warneri Mccleve & Ko..., 2005 | | | Copris wiesei Kolbe, 1914 | | | Copris xingkeyangi Ochi, Kon & Bai, 2020 | | | Copris yangi Ochi, Kon & Bai, 2007 | | | Copris youngai Balthasar, 1967 | | | Copris zhangi Ochi, Kon & Bai, 2009 | | | Coproecus hemisphaericus Guérin-Ménev..., 1844 | | | Coprophanaeus (Coprophanaeus) pluto Harold, 1863 | | | Coprophanaeus pluto nogueirai (Arnaud, 2002) | ⇒ Coprophanaeus pluto Harold, 1863 | | Coprophanaeus (Megaphanaeus) bellicosus Olivier, 1789 | | | Coprophanaeus (Megaphanaeus) bonariensis Gory, 1844 | | | Coprophanaeus (Megaphanaeus) ensifer Germar, 1821 | | | Coprophanaeus (Megaphanaeus) lancifer Linnaeus, 1767 | | | Coprophanaeus (Metallophanaeus) horus Waterhouse, 1891 | | | Coprophanaeus (Metallophanaeus) machadoi Pereira & An..., 1955 | | | Coprophanaeus (Metallophanaeus) pertyi Olsufieff, 1924 | | | Coprophanaeus (Metallophanaeus) pessoai Pereira, 1949 | | | Coprophanaeus (Metallophanaeus) punctatus Olsufieff, 1924 | | | Coprophanaeus (Metallophanaeus) saphirinus Sturm, 1826 | | | Coprophanaeus saphirinus chabrillaci Thomson, 1857 | | | Coprophanaeus saphirinus saphirinus | | | Coprophanaeus (Metallophanaeus) thalassinus Perty, 1830 | | | Coprophanaeus (Metallophanaeus) vazdemelloi Arnaud, 2002 | | | Coprophanaeus abas MacLeay, 1819 | | | Coprophanaeus acrisius MacLeay, 1819 | | | Coprophanaeus boucardi Nevinson, 1891 | | | Coprophanaeus callegarii Arnaud, 2002 | | | Coprophanaeus caroliae Edmonds, 2008 | | | Coprophanaeus cerberus Harold, 1875 | | | Coprophanaeus chiriquensis Olsufieff, 1924 | | | Coprophanaeus christophorowi Olsufieff, 1924 | | | Coprophanaeus conocephalus Olsufieff, 1924 | | | Coprophanaeus corythus Harold, 1863 | | | Coprophanaeus cyanescens Olsufieff, 1924 | | | Coprophanaeus dardanus MacLeay, 1819 | | | Coprophanaeus degallieri Arnaud, 1997 | | | Coprophanaeus edmondsi (Arnaud, 1997) | ⇒ Coprophanaeus conocephalus Olsufieff, 1924 | | Coprophanaeus gamezi Arnaud, 2002 | | | Coprophanaeus gephyra Kohlmann & S..., 2012 | | | Coprophanaeus gilli Arnaud, 1997 | | | Coprophanaeus ignecinctus Felsche, 1909 | | | Coprophanaeus jasius Olivier, 1789 | | | Coprophanaeus jasius acrisius (MacLeay, 1819) | ⇒ Coprophanaeus acrisius MacLeay, 1819 | | Coprophanaeus kohlmanni (Arnaud, 2002) | ⇒ Coprophanaeus morenoi Arnaud, 1982 | | Coprophanaeus larseni Arnaud, 2002 | | | Coprophanaeus lecromi Arnaud, 2002 | | | Coprophanaeus lichyi Arnaud, 2002 | | | Coprophanaeus magnoi Arnaud, 2002 | | | Coprophanaeus milon Blanchard, 1846 | | | Coprophanaeus morenoi Arnaud, 1982 | | | Coprophanaeus ohausi Felsche, 1911 | | | Coprophanaeus parvulus Olsufieff, 1924 | | | Coprophanaeus pecki Howden & Young, 1981 | | | Coprophanaeus rigoutorum Arnaud, 2002 | | | Coprophanaeus solisi Arnaud, 1997 | | | Coprophanaeus spitzi Pessôa, 1934 | | | Coprophanaeus strandi Balthasar, 1939 | | | Coprophanaeus suredai Arnaud, 1996 | | | Coprophanaeus telamon Erichson, 1847 | | | Coprophanaeus telamon nevinsoni (Arnaud & Gamez, 2002) | ⇒ Coprophanaeus corythus Harold, 1863 | | Coprophanaeus terrali Arnaud, 2002 | | | Coprophanaeus uhleri (Maly & Pokorny, 2008) | ⇒ Coprophanaeus chiriquensis Olsufieff, 1924 | | Coptodactyla brooksi Matthews, 1976 | | | Coptodactyla depressa Paulian, 1933 | | | Coptodactyla ducalis Blackburn, 1903 | | | Coptodactyla glabricollis Hope, 1842 | | | Coptodactyla lesnei Paulian, 1933 | | | Coptodactyla matthewsi Reid, 2000 | | | Coptodactyla merdeka Reid, 2000 | | | Coptodactyla meridionalis Matthews, 1976 | | | Coptodactyla monstrosa Felsche, 1909 | | | Coptodactyla nitida (Paulian, 1933) | ⇒ Coptodactyla subaenea Harold, 1877 | | Coptodactyla onitoides Gillet, 1925 | | | Coptodactyla papua Lansberge, 1885 | | | Coptodactyla stereocera Gillet, 1911 | | | Coptodactyla storeyi Reid, 2000 | | | Coptodactyla subaenea Harold, 1877 | | | Coptodactyla torresica Mathews, 1976 | | | Coptodactyla tuberculata Gillet, 1925 | | | Coptorhina angolensis Arrow, 1906 | | | Coptorhina auspicata Péringuey, 1901 | | | Coptorhina davidi Janssens, 1939 | | | Coptorhina excavata Frolov, Akhm..., 2008 | | | Coptorhina klugi Hope, 1833 | | | Coptorhina klugii Hope, 1833 | | | Coptorhina nanus Péringuey, 1888 | | | Coptorhina nitefacta Gillet, 1932 | | | Coptorhina nitidipennis Boheman, 1857 | | | Cretonitis copripes ♰ Nikolajev, 2007 | | | Cretorabaeus elongatus ♰ G. V. Nikolajev, 1995 | | | Cryptocanthon altus Howden, 1976 | | | Cryptocanthon andersoni Cook, 2002 | | | Cryptocanthon andradei Arias & Medina, 2014 | | | Cryptocanthon bochilae Cook, 2002 | | | Cryptocanthon brevisetosus Howden, 1973 | | | Cryptocanthon buriticaorum Arias & Medina, 2014 | | | Cryptocanthon campbellorum Howden, 1973 | | | Cryptocanthon cephalopunctatus Martínez-Rev..., 2020 | | | Cryptocanthon chimalapensis Mora-Aguilar..., 2018 | | | Cryptocanthon chiriquinus Howden & Gill, 1987 | | | Cryptocanthon cristobalensis Howden, 1973 | | | Cryptocanthon curticrinis Cook, 2002 | | | Cryptocanthon denticulatum | | | Cryptocanthon denticulum Cook, 2002 | | | Cryptocanthon encenillensis Martínez-Rev..., 2020 | | | Cryptocanthon escobari Cook, 2002 | | | Cryptocanthon foveatus Cook, 2002 | | | Cryptocanthon galbao Cook, 2002 | | | Cryptocanthon genieri Cook, 2002 | | | Cryptocanthon gilli Cook, 2002 | | | Cryptocanthon howdeni Cook, 2002 | | | Cryptocanthon humidus Howden, 1973 | | | Cryptocanthon iskaypachak Giraldo-Mendoza, 2022 | | | Cryptocanthon lindemanae Howden & Gill, 1987 | | | Cryptocanthon lobatus Howden, 1973 | | | Cryptocanthon lobipygus Cook, 2002 | | | Cryptocanthon mailinae Martínez-Rev..., 2020 | | | Cryptocanthon medinae Cook, 2002 | | | Cryptocanthon montebello Cook, 2002 | | | Cryptocanthon napoensis Cook, 2002 | | | Cryptocanthon nebulinus Howden, 1973 | | | Cryptocanthon newtoni Howden, 1976 | | | Cryptocanthon ocosingo Cook, 2002 | | | Cryptocanthon osaensis Cook, 2002 | | | Cryptocanthon otonga Cook, 2002 | | | Cryptocanthon paradoxus Balthasar, 1942 | | | Cryptocanthon parvus Howden, 1973 | | | Cryptocanthon peckorum Howden, 1973 | | | Cryptocanthon pulidae | | | Cryptocanthon pulidoae Arias & Medina, 2014 | | | Cryptocanthon pumilus Cook, 2002 | | | Cryptocanthon punctatus Cook, 2002 | | | Cryptocanthon rayonensis Cook, 2002 | | | Cryptocanthon solisi Cook, 2002 | | | Cryptocanthon urguensis Cook, 2002 | | | Cyobius brendelli Ochi, Kon & ..., 2010 | | | Cyobius cheyi Ochi, Kon & ..., 2006 | | | Cyobius mendeli Ochi, Kon & ..., 2010 | | | Cyobius tsukamotoi Ochi, Kon & ..., 2008 | | | Cyobius wallacei Sharp, 1875 | | | Cyptochirus ambiguus (Kirby, 1828) | ⇒ Drepanocerus trogiformis Roth, 1851 | | Cyptochirus decellei Simonis & Zu..., 1980 | | | Cyptochirus distinctus Janssens, 1953 | | | Cyptochirus trogiformis Roth, 1851 | | | Degallieridium lilliputanum Vaz-de-Mello, 2008 | | | Delopleurus darrenmanni Frolov, 2014 | | | Delopleurus fossatus Frolov, 2014 | | | Delopleurus giletti Janssens, 1939 | | | Delopleurus gilletti Janssens, 1939 | | | Delopleurus krikkeni Frolov, 2014 | | | Delopleurus mencli Kral, 2014 | | | Delopleurus naviauxi Frolov & Cam..., 2014 | | | Delopleurus parvus Sharp, 1875 | | | Delopleurus pubescens Frolov, 2014 | | | Delopleurus pullus Boheman, 1857 | | | Delopleurus striatus Arrow, 1931 | | | Deltepilissus diabolicus Harold, 1880 | | | Deltepilissus infernalis Harold, 1880 | | | Deltochilum abdominalis Martínez, 1947 | | | Deltochilum acanthus Kohlmann & S..., 2012 | | | Deltochilum acropyge Bates, 1887 | | | Deltochilum aequinoctiale Buquet, 1844 | | | Deltochilum alpercata Silva, Louza..., 2015 | | | Deltochilum amandaarcanjoae Silva, Louza..., 2015 | | | Deltochilum amazonicum Bates, 1887 | | | Deltochilum arrowi Paulian, 1939 | | | Deltochilum arturoi Silva, Louza..., 2015 | | | Deltochilum aspericolle Bates, 1870 | | | Deltochilum aureopilosum Paulian, 1939 | | | Deltochilum barbipes Bates, 1870 | | | Deltochilum batesi Paulian, 1938 | | | Deltochilum bezdeki González-Alv..., 2014 | | | Deltochilum bolivariensis González-Alv..., 2021 | | | Deltochilum bordoni Halffter & M..., 1976 | | | Deltochilum brasiliense Castelnau, 1840 | | | Deltochilum burmeisteri (Harold, 1867) | ⇒ Calhyboma mexicanum Burmeister, 1848 | | Deltochilum calcaratum Bates, 1870 | | | Deltochilum cangalha Silva, Louza..., 2015 | | | Deltochilum carinatum Westwood, 1837 | | | Deltochilum carrilloi González-Alv..., 2014 | | | Deltochilum costalimai Pereira & D´..., 1955 | | | Deltochilum crenulipes Paulian, 1938 | | | Deltochilum cristatum Paulian, 1938 | | | Deltochilum cristinae Martínez, 1991 | | | Deltochilum cupreicolle Blanchard, 1846 | | | Deltochilum cupreicolle viridescens Martínez, 1948 | | | Deltochilum dentipes Eschscholtz, 1822 | | | Deltochilum diringshofeni Pereira & Ma..., 1956 | | | Deltochilum elevatum Castelnau, 1840 | | | Deltochilum elongatum Felsche, 1907 | | | Deltochilum enceladus Kolbe, 1893 | | | Deltochilum eurymedon Génier, 2012 | | | Deltochilum feeri Silva, Louza..., 2015 | | | Deltochilum femorale Bates, 1870 | | | Deltochilum finestriatum Silva, Louza..., 2015 | | | Deltochilum furcatum Castelnau, 1840 | | | Deltochilum fuscocupreum Bates, 1870 | | | Deltochilum genieri González-Alv..., 2021 | | | Deltochilum gibbosum Fabricius, 1775 | Humpback Dung Beetle | | Deltochilum gibbosum gibbosum Fabricius, 1775 | | | Deltochilum gibbosum panamensis Howden, 1966 | | | Deltochilum gibbosum sublaeve Bates, 1887 | | | Deltochilum gibbosus Fabricius, 1775 | | | Deltochilum gigante Silva & Vaz-..., 2014 | | | Deltochilum gilli González-Alv..., 2021 | | | Deltochilum granulatum Bates, 1870 | | | Deltochilum granulosum Paulian, 1933 | | | Deltochilum guildingii Westwood, 1835 | | | Deltochilum guyanense Paulian, 1933 | | | Deltochilum howdeni Martínez, 1955 | | | Deltochilum hypponum Buquet, 1844 | | | Deltochilum icariforme Paulian, 1938 | | | Deltochilum icaroides Balthasar, 1939 | | | Deltochilum icarus Olivier, 1789 | | | Deltochilum inaequale Balthasar, 1939 | | | Deltochilum inesae González-Alv..., 2021 | | | Deltochilum irroratum Castelnau, 1840 | | | Deltochilum jocelynae González-Alv..., 2021 | | | Deltochilum kolbei Paulian, 1938 | | | Deltochilum kolleri Silva, Louza..., 2015 | | | Deltochilum komareki Balthasar, 1939 | | | Deltochilum laetiusculum Bates, 1870 | | | Deltochilum laevigatum (Balthasar, 1939) | ⇒ Deltochilum peruanum Paulian, 1938 | | Deltochilum larseni Silva, Louza..., 2015 | | | Deltochilum lindemannae Balthasar, 1967 | | | Deltochilum lobipes Bates, 1887 | | | Deltochilum longiceps Paulian, 1938 | | | Deltochilum loperae González & M..., 2010 | | | Deltochilum louzadai González-Alv..., 2014 | | | Deltochilum luederwaldti Pereira & D´..., 1955 | | | Deltochilum mariafernandae Silva, Vaz-d..., 2017 | | | Deltochilum mexicanum Burmeister, 1848 | | | Deltochilum molanoi González-Alv..., 2014 | | | Deltochilum morbillosum Burmeister, 1848 | | | Deltochilum mourei Pereira, 1949 | | | Deltochilum multicolor Balthasar, 1939 | | | Deltochilum nonstriatum González-Alv..., 2021 | | | Deltochilum orbiculare Lansberge, 1874 | | | Deltochilum orbignyi Blanchard, 1846 | | | Deltochilum orbignyi amazonicum Bates, 1887 | | | Deltochilum orbignyi orbignyi | | | Deltochilum panamensis Howden, 1966 | | | Deltochilum paresi Silva, Louza..., 2015 | | | Deltochilum parile Bates, 1887 | | | Deltochilum parile parile | | | Deltochilum parile tolimensis Paulian, 1938 | | | Deltochilum peruanum Paulian, 1938 | | | Deltochilum plebejum Balthasar, 1939 | | | Deltochilum pretiosum Harold, 1875 | | | Deltochilum pseudoicarus Balthasar, 1939 | | | Deltochilum pseudoparile Paulian, 1938 | | | Deltochilum punctatum Harold, 1880 | | | Deltochilum quasistriatum González-Alv..., 2021 | | | Deltochilum riehli Harold, 1868 | | | Deltochilum ritamourae Silva, Louza..., 2015 | | | Deltochilum robustus Molano & Gon..., 2010 | | | Deltochilum rosamariae Martínez, 1991 | | | Deltochilum rubripenne Gory, 1831 | | | Deltochilum scabriusculum Bates, 1887 | Rockpool blenny | | Deltochilum schefflerorum Silva, Louza..., 2015 | | | Deltochilum sculpturatum Felsche, 1907 | | | Deltochilum septemstriatum Paulian, 1938 | | | Deltochilum sericeum Paulian, 1938 | | | Deltochilum sextuberculatum Bates, 1870 | | | Deltochilum silphoides Balthasar, 1939 | | | Deltochilum speciosissimum Balthasar, 1939 | | | Deltochilum spinipes Paulian, 1938 | | | Deltochilum streblopodum Silva, Louza..., 2015 | | | Deltochilum sublaeve Bates, 1887 | | | Deltochilum submetallicum Castelnau, 1840 | | | Deltochilum subrubrum Silva, Louza..., 2015 | | | Deltochilum susanae González-Alv..., 2021 | | | Deltochilum tenuistriatum González-Alv..., 2021 | | | Deltochilum tessellatum Bates, 1870 | | | Deltochilum titovidaurrei Silva, Louza..., 2015 | | | Deltochilum trisignatum Harold, 1880 | | | Deltochilum tumidum Howden, 1966 | Rockpool blenny | | Deltochilum valgum Burmeister, 1873 | | | Deltochilum valgum acropyge Bates, 1887 | | | Deltochilum valgum longiceps Paulian, 1938 | | | Deltochilum variolosum Burmeister, 1873 | | | Deltochilum verruciferum Felsche, 1911 | | | Deltochilum violaceum Paulian, 1938 | | | Deltochilum violetae Martínez, 1991 | | | Deltochilum viridescens Martinez, 1948 | | | Deltochilum viridicatum Silva, Louza..., 2015 | | | Deltochilum viridicupreum Balthasar, 1939 | | | Deltorhinum armatum Génier, 2010 | | | Deltorhinum batesi Harold, 1867 | | | Deltorhinum bilobatum Génier, 2010 | | | Deltorhinum genieri Montoya-Moli..., 2019 | | | Deltorhinum guyanensis Génier, 2010 | | | Deltorhinum kempffmercadoi Génier, 2010 | | | Deltorhinum robustum Génier, 2010 | | | Deltorhinum vazdemelloi Génier, 2010 | | | Demarziella alternata Lea, 1923 | | | Demarziella eungella Matthews, 1976 | | | Demarziella geminata MacLeay, 1871 | | | Demarziella imitatrix Balthasar, 1967 | | | Demarziella interrupta Carter, 1936 | | | Demarziella metallica Carter, 1936 | | | Demarziella mirifica Balthasar, 1961 | | | Demarziella planitarsis Petrovitz, 1971 | | | Demarziella pratensis Matthews, 1976 | | | Demarziella scarpensis Matthews, 1976 | | | Demarziella storeyi Matthews, 1976 | | | Demarziella sylvestris Matthews, 1976 | | | Demarziella tropicalis Matthews, 1976 | | | Demarziella yorkensis Matthews & S..., 1986 | | | Dendropaemon (Coprophanaeoides) compressipennis Génier & Arn..., 2016 | | | Dendropaemon (Coprophanaeoides) cribrosus Génier & Arn..., 2016 | | | Dendropaemon (Coprophanaeoides) renatii Olsoufieff, 1924 | | | Dendropaemon (Dendropaemon) aenigmaticus Génier & Arn..., 2016 | | | Dendropaemon (Dendropaemon) amazonicus Génier & Arn..., 2016 | | | Dendropaemon (Dendropaemon) amyntas Harold, 1868 | | | Dendropaemon amyntas waterhousi Olsufieff, 1924 | | | Dendropaemon (Dendropaemon) angustulus Génier & Arn..., 2016 | | | Dendropaemon (Dendropaemon) ater (Laporte, 1832) | ⇒ Dendropaemon telephus Waterhouse, 1891 | | Dendropaemon (Dendropaemon) flechtmanni Génier & Arn..., 2016 | | | Dendropaemon (Dendropaemon) larseni Génier & Arn..., 2016 | | | Dendropaemon (Dendropaemon) piceus Perty, 1830 | | | Dendropaemon (Dendropaemon) telephus Waterhouse, 1891 | | | Dendropaemon (Dendropaemon) viridis Perty, 1830 | | | Dendropaemon (Glaphyropaemon) angustipennis (Harold, 1869) | ⇒ Dendropaemon silvanus Blut, 1939 | | Dendropaemon (Glaphyropaemon) bahianus (Harold, 1868) | ⇒ Coprophanaeoides bahianus Harold, 1868 | | Dendropaemon (Glaphyropaemon) inemarginatus Génier & Arn..., 2016 | | | Dendropaemon (Nigropaemon) nigritulus Génier & Arn..., 2016 | | | Dendropaemon (Onthoecus) attalus Génier & Arn..., 2016 | | | Dendropaemon (Onthoecus) lydiae Génier & Arn..., 2016 | | | Dendropaemon (Onthoecus) morettoi Génier & Arn..., 2016 | | | Dendropaemon (Paradendropaemon) ganglbaueri Felsche, 1909 | | | Dendropaemon (Paradendropaemon) vazdemelloi Génier & Arn..., 2016 | | | Dendropaemon (Rutilopaemon) refulgens (Waterhouse, 1891) | ⇒ Coprophanaeoides refulgens Waterhouse, 1891 | | Dendropaemon refulgens olsufieffi Blut, 1939 | | | Dendropaemon (Streblopaemon) fractipes Felsche, 1909 | | | Dendropaemon (Sulcopaemon) fasces Blut, 1939 | | | Dendropaemon (Sulcopaemon) haroldi Olsoufieff, 1924 | | | Dendropaemon (Sulcopaemon) latistriatus Génier & Arn..., 2016 | | | Dendropaemon (Sulcopaemon) nitidicollis Olsoufieff, 1924 | | | Dendropaemon (Titthopaemon) denticollis Felsche, 1909 | | | Dendropaemon denticollis lividus Blut, 1939 | | | Dendropaemon batrachites Blut, 1939 | | | Dendropaemon bluti Génier & Arn..., 2016 | | | Dendropaemon bonariensis (Bruch, 1925) | ⇒ Anomiopus caputipilus Canhedo, 2004 | | Dendropaemon carinifer Génier & Arn..., 2016 | | | Dendropaemon convexus (Harold, 1869) | ⇒ Tetramereia convexa Harold, 1869 | | Dendropaemon crenatostriatus Felsche, 1909 | | | Dendropaemon fredericki Klages, 1906 | | | Dendropaemon furtadoi Génier & Arn..., 2016 | | | Dendropaemon hirticollis (Olsoufieff, 1924) | ⇒ Coprophanaeoides hirticollis Olsufieff, 1924 | | Dendropaemon inflatus Génier & Arn..., 2016 | | | Dendropaemon lobatus (Waterhouse, 1891) | ⇒ Coprophanaeoides bahianus Harold, 1868 | | Dendropaemon pauliani Martinez & P..., 1960 | | | Dendropaemon pilosissimus Génier & Arn..., 2016 | | | Dendropaemon quadratus Laporte, 1832 | | | Dendropaemon silvanus Blut, 1939 | | | Dendropaemon similis Blut, 1939 | | | Dendropaemon smaragdinus Waterhouse, 1891 | | | Dendropaemon subcylindricus Blut, 1939 | | | Dendropaemon tenuitarsis Felsche, 1909 | | | Dendropaemon viridipennis Laporte, 1831 | | | Diabroctis cadmus Harold, 1868 | | | Diabroctis mimas Linnaeus, 1758 | | | Diabroctis mirabilis Harold, 1877 | | | Diabroctis pseudomimas Valois, Hara..., 2019 | | | Diabroctis venezuelensis Martinez & C..., 1990 | | | Diastellopalpus anthonyi Walter, 1989 | | | Diastellopalpus astorgisorum Walter, 2013 | | | Diastellopalpus balubanus Kolbe, 1893 | | | Diastellopalpus basilobatus d´Orbigny, 1902 | | | Diastellopalpus cambeforti Josso & Pré..., 2000 | | | Diastellopalpus camiadei Moretto, 1999 | | | Diastellopalpus conradti d´Orbigny, 1902 | | | Diastellopalpus drumonti Moretto, 1999 | | | Diastellopalpus dudleyi Cambefort, 1980 | | | Diastellopalpus fuelleborni Kolbe, 1901 | | | Diastellopalpus gilleti d´Orbigny, 1908 | | | Diastellopalpus hassoni Moretto, 1999 | | | Diastellopalpus ikelenge Josso & Pré..., 2014 | | | Diastellopalpus infernalis Lansberge, 1887 | | | Diastellopalpus jirouxi Josso & Pré..., 2000 | | | Diastellopalpus johnstoni Waterhouse, 1885 | | | Diastellopalpus laevibasis d´Orbigny, 1902 | | | Diastellopalpus lamellicollis Quedenfeldt, 1884 | | | Diastellopalpus legrandi Josso & Pré..., 2000 | | | Diastellopalpus metriogonus (d´Orbigny, 1914) | ⇒ Diastellopalpus neavei d´Orbigny, 1914 | | Diastellopalpus monapoides Bates, 1888 | | | Diastellopalpus murrayi Harold, 1868 | | | Diastellopalpus neavei d´Orbigny, 1914 | | | Diastellopalpus nigerrimus Kolbe, 1893 | | | Diastellopalpus noctis Thomson, 1858 | | | Diastellopalpus pluton d´Orbigny, 1902 | | | Diastellopalpus pseudoschultzei Josso & Pré..., 2000 | | | Diastellopalpus quinquedens Bates, 1888 | | | Diastellopalpus schultzei Gillet, 1927 | | | Diastellopalpus semirubidus d´Orbigny, 1913 | | | Diastellopalpus sulciger Kolbe, 1893 | | | Diastellopalpus thomsoni Bates, 1888 | | | Diastellopalpus thomsoni malawiensis Josso & Pré..., 2000 | | | Diastellopalpus thomsoni thomsoni | | | Diastellopalpus tridens (Fairmaire, 1891) | ⇒ Diastellopalpus thomsoni Bates, 1888 | | Diastellopalpus walteri Josso & Pré..., 2000 | | | Dichotomius (Homocanthonides) smaragdinus Perty, 1830 | | | Dichotomius (Luederwaldtinia) affinis Felsche, 1910 | | | Dichotomius (Luederwaldtinia) agenor Harold, 1869 | | | Dichotomius (Luederwaldtinia) alvarengai Maldaner, N..., 2015 | | | Dichotomius (Luederwaldtinia) amplicollis Harold, 1869 | | | Dichotomius (Luederwaldtinia) angeloi Nunes, Carv..., 2016 | | | Dichotomius (Luederwaldtinia) apicalis Luederwaldt, 1931 | | | Dichotomius (Luederwaldtinia) assifer Eschscholtz, 1822 | | | Dichotomius (Luederwaldtinia) batesi Harold, 1869 | | | Dichotomius (Luederwaldtinia) bechynei Martínez, 1973 | | | Dichotomius (Luederwaldtinia) belus Harold, 1880 | | | Dichotomius (Luederwaldtinia) benesi Vaz-de-Mello..., 2016 | | | Dichotomius (Luederwaldtinia) bosqui Pereira, 1941 | | | Dichotomius (Luederwaldtinia) carbonarius Mannerheim, 1829 | | | Dichotomius (Luederwaldtinia) catimbau Valois, Vaz-..., 2017 | | | Dichotomius (Luederwaldtinia) centralis Harold, 1869 | | | Dichotomius (Luederwaldtinia) crinicollis Germar, 1824 | | | Dichotomius (Luederwaldtinia) cuprinus Felsche, 1901 | | | Dichotomius (Luederwaldtinia) danieli (Kohlmann & S..., 1997) | ⇒ Luederwaldtinia danieli Kohlmann & S..., 1997 | | Dichotomius (Luederwaldtinia) deyrollei Harold, 1869 | | | Dichotomius (Luederwaldtinia) eucranioides (Pereira & An..., 1955) | ⇒ Luederwaldtinia eucranioides Pereira & An..., 1955 | | Dichotomius (Luederwaldtinia) fallax Harold, 1880 | | | Dichotomius (Luederwaldtinia) favi (Kohlmann & S..., 1997) | ⇒ Luederwaldtinia favi Kohlmann & S..., 1997 | | Dichotomius (Luederwaldtinia) femoratus (Howden & Young, 1981) | ⇒ Luederwaldtinia femoratus Howden & Young, 1981 | | Dichotomius (Luederwaldtinia) fimbriatus Harold, 1869 | | | Dichotomius (Luederwaldtinia) fortepunctatus Luederwaldt, 1923 | | | Dichotomius (Luederwaldtinia) fortestriatus Luederwaldt, 1924 | | | Dichotomius (Luederwaldtinia) gamboaensis (Howden & Young, 1981) | ⇒ Luederwaldtinia gamboaensis Howden & Young, 1981 | | Dichotomius (Luederwaldtinia) geminatus Arrow, 1913 | | | Dichotomius (Luederwaldtinia) gilletti Valois, Vaz-..., 2017 | | | Dichotomius (Luederwaldtinia) glaucus Harold, 1869 | | | Dichotomius (Luederwaldtinia) globulus Felsche, 1901 | | | Dichotomius (Luederwaldtinia) guaribensis Valois, Vaz-..., 2017 | | | Dichotomius (Luederwaldtinia) hempeli Pereira, 1942 | | | Dichotomius (Luederwaldtinia) iannuzziae Valois, Vaz-..., 2017 | | | Dichotomius (Luederwaldtinia) inachoides Felsche, 1901 | | | Dichotomius (Luederwaldtinia) inachus Erichson, 1847 | | | Dichotomius (Luederwaldtinia) interstitialis Luederwaldt, 1931 | | | Dichotomius (Luederwaldtinia) irinus Harold, 1867 | | | Dichotomius (Luederwaldtinia) laevicollis Felsche, 1901 | | | Dichotomius (Luederwaldtinia) louzadai Nunes, Carv..., 2016 | | | Dichotomius (Luederwaldtinia) lucasi Harold, 1869 | | | Dichotomius (Luederwaldtinia) luctuosus Harold, 1869 | | | Dichotomius (Luederwaldtinia) lycas Felsche, 1901 | | | Dichotomius (Luederwaldtinia) machadoi Martinez & P..., 1967 | | | Dichotomius (Luederwaldtinia) malyi Maldaner, N..., 2015 | | | Dichotomius (Luederwaldtinia) micans Luederwaldt, 1924 | | | Dichotomius (Luederwaldtinia) muticus Luederwaldt, 1922 | | | Dichotomius (Luederwaldtinia) mysticus Luederwaldt, 1935 | | | Dichotomius (Luederwaldtinia) nisus Olivier, 1789 | | | Dichotomius (Luederwaldtinia) nobilis Waterhouse, 1891 | | | Dichotomius (Luederwaldtinia) ocellatopunctatus Felsche, 1901 | | | Dichotomius (Luederwaldtinia) opacipennis Luederwaldt, 1931 | | | Dichotomius opacipennis fornicatus Luederwaldt, 1931 | | | Dichotomius (Luederwaldtinia) opalescens Felsche, 1910 | | | Dichotomius (Luederwaldtinia) parcepunctatus Felsche, 1901 | | | Dichotomius (Luederwaldtinia) problematicus Luederwaldt, 1924 | | | Dichotomius problematicus planus Luederwaldt, 1924 | | | Dichotomius (Luederwaldtinia) punctatostriatus (Felsche, 1901) | ⇒ Luederwaldtinia punctatostriatus Felsche, 1901 | | Dichotomius (Luederwaldtinia) rodrigoi (Kohlmann & S..., 1997) | ⇒ Luederwaldtinia rodrigoi Kohlmann & S..., 1997 | | Dichotomius (Luederwaldtinia) rugatus (Luederwaldt, 1935) | ⇒ Luederwaldtinia rugatus Luederwaldt, 1935 | | Dichotomius (Luederwaldtinia) schiffleri Vaz-de-Mello..., 2001 | | | Dichotomius (Luederwaldtinia) semisquamosus Curtis, 1845 | | | Dichotomius (Luederwaldtinia) sericeus Harold, 1867 | | | Dichotomius (Luederwaldtinia) sexdentatus Luederwaldt, 1925 | | | Dichotomius (Luederwaldtinia) simplicicornis Luederwaldt, 1935 | | | Dichotomius (Luederwaldtinia) speciosus Waterhouse, 1891 | | | Dichotomius (Luederwaldtinia) superbus Felsche, 1901 | | | Dichotomius (Luederwaldtinia) texanus (Felsche, 1901) | ⇒ Luederwaldtinia texanus Felsche, 1901 | | Dichotomius (Luederwaldtinia) triquetrus Luederwaldt, 1924 | | | Dichotomius (Luederwaldtinia) tristis Luederwaldt, 1924 | | | Dichotomius (Luederwaldtinia) validipilosus Luederwaldt, 1931 | | | Dichotomius (Luederwaldtinia) vidaurrei Nunes & Vaz-..., 2013 | | | Dichotomius (Luederwaldtinia) yucatanus Bates, 1887 | | | Dichotomius (Luederwaldtinia) ziliolii Gandini & Ag..., 2009 | | | Dichotomius (Selenocopris) acuticornis Luederwaldt, 1930 | | | Dichotomius (Selenocopris) aielloae Nunes, 2016 | | | Dichotomius (Selenocopris) ascanius Harold, 1869 | | | Dichotomius ascanius aceratus Pereira, 1944 | | | Dichotomius ascanius piceus Luederwaldt, 1930 | | | Dichotomius (Selenocopris) bicuspis Germar, 1824 | | | Dichotomius (Selenocopris) darwini Nunes & Vaz-..., 2016 | | | Dichotomius (Selenocopris) enrietti (Vernalha, 1952) | ⇒ Selenocopris enrietti Vernalha, 1952 | | Dichotomius (Selenocopris) fissiceps Felsche, 1901 | | | Dichotomius (Selenocopris) fissus Harold, 1867 | | | Dichotomius (Selenocopris) fonsecae Luederwaldt, 1925 | | | Dichotomius (Selenocopris) guaranii Gandini & Ag..., 2009 | | | Dichotomius (Selenocopris) inflaticollis Felsche, 1910 | | | Dichotomius (Selenocopris) ingens Luederwaldt, 1935 | | | Dichotomius (Selenocopris) manni Nunes & Vaz-..., 2016 | | | Dichotomius (Selenocopris) mundus Harold, 1869 | | | Dichotomius (Selenocopris) parisi Nunes & Vaz-..., 2016 | | | Dichotomius (Selenocopris) pauloensis Luederwaldt, 1925 | | | Dichotomius (Selenocopris) periotoi Nunes & Vaz-..., 2016 | | | Dichotomius (Selenocopris) punctulatipennis Luederwaldt, 1930 | | | Dichotomius (Selenocopris) pygidialis Luederwaldt, 1922 | | | Dichotomius (Selenocopris) quadraticeps Felsche, 1901 | | | Dichotomius (Selenocopris) rugosipennis Luederwaldt, 1930 | | | Dichotomius (Selenocopris) simplex (Taschenberg, 1870) | ⇒ Selenocopris simplex Taschenberg, 1870 | | Dichotomius (Selenocopris) simulans Luederwaldt, 1925 | | | Dichotomius (Selenocopris) singularis Felsche, 1907 | | | Dichotomius (Selenocopris) socius Luederwaldt, 1929 | | | Dichotomius (Selenocopris) spadiceus Luederwaldt, 1928 | | | Dichotomius achamas Harold, 1867 | | | Dichotomius adrastus Harold, 1875 | | | Dichotomius agesilaus Waterhouse, 1891 | | | Dichotomius alutaceus Felsche, 1901 | | | Dichotomius alyattes Harold, 1880 | | | Dichotomius amicitiae Kohlmann & S..., 1997 | | | Dichotomius andresi Sarmiento-Ga..., 2014 | | | Dichotomius annae Linnaeus, 1767 | | | Dichotomius anthrax Felsche, 1901 | | | Dichotomius arnaudi Nunes & Vaz-..., 2019 | | | Dichotomius asenjoi Nunes & Vaz-..., 2019 | | | Dichotomius australis Luederwaldt, 1931 | | | Dichotomius baiano Nunes & Vaz-..., 2019 | | | Dichotomius barbarae Nunes & Vaz-..., 2019 | | | Dichotomius berthalutzae Valois, Silv..., 2022 | | | Dichotomius bicornis Waterhouse, 1891 | | | Dichotomius bitiensis Gillet, 1911 | | | Dichotomius blancoi Nunes & Vaz-..., 2019 | | | Dichotomius boreus Olivier, 1789 | | | Dichotomius borgmeieiri Pereira & Ma..., 1959 | | | Dichotomius borgmeieri (Pereira & Ma..., 1959) | ⇒ Dichotomius borgmeieiri Pereira & Ma..., 1959 | | Dichotomius bos Blanchard, 1846 | | | Dichotomius bucki (Pereira, 1953) | ⇒ Luederwaldtinia bucki Pereira, 1953 | | Dichotomius buqueti Lucas, 1857 | | | Dichotomius calcaratus Arrow, 1913 | | | Dichotomius camargoi Martínez, 1956 | | | Dichotomius camposeabrai Martínez, 1974 | | | Dichotomius candango Nunes & Vaz-..., 2019 | | | Dichotomius carinatus Luederwaldt, 1925 | | | Dichotomius carolinus Linnaeus, 1767 | Carolina copris | | Dichotomius clypeatus Luederwaldt, 1929 | | | Dichotomius coenosus Erichson, 1848 | | | Dichotomius colonicus (Say, 1835) | ⇒ Dichotomius carolinus Linnaeus, 1767 | | Dichotomius comarapensis Génier, 2000 | | | Dichotomius compressicollis Luederwaldt, 1929 | | | Dichotomius conicollis Blanchard, 1846 | | | Dichotomius costaricensis Luederwaldt, 1935 | | | Dichotomius cotopaxi Guérin-Ménev..., 1855 | | | Dichotomius dahli Landin, 1956 | | | Dichotomius depressicollis Harold, 1867 | | | Dichotomius diabolicus Harold, 1875 | | | Dichotomius divergens Luederwaldt, 1923 | | | Dichotomius dynastus Gandini & Ag..., 2009 | | | Dichotomius edmondsi Nunes & Vaz-..., 2019 | | | Dichotomius emas Nunes & Vaz-..., 2019 | | | Dichotomius enioi Montoya-Moli..., 2021 | | | Dichotomius feeri Nunes & Vaz-..., 2019 | | | Dichotomius fernandosilvai Nunes & Vaz-..., 2019 | | | Dichotomius filgueirasi Nunes & Vaz-..., 2019 | | | Dichotomius fornicatus Luederwaldt, 1931 | | | Dichotomius foveicollis Kirsch, 1871 | | | Dichotomius frizzasae Nunes & Vaz-..., 2019 | | | Dichotomius furtadoi Nunes & Vaz-..., 2019 | | | Dichotomius gandinii Rossini & Va..., 2020 | | | Dichotomius gardneri Nunes & Vaz-..., 2019 | | | Dichotomius geraldimi Maldaner & V..., 2021 | | | Dichotomius haroldi Waterhouse, 1891 | | | Dichotomius henripittieri Montoya-Moli..., 2021 | | | Dichotomius horridus Felsche, 1911 | | | Dichotomius horvathi Balthasar, 1970 | | | Dichotomius imitator Felsche, 1901 | | | Dichotomius itatiaiaensis Nunes & Vaz-..., 2019 | | | Dichotomius joelus Gandini & Ag..., 2009 | | | Dichotomius latilobatus Boilly & Vaz..., 2021 | | | Dichotomius latistriatus (Luederwaldt, 1935) | ⇒ Luederwaldtinia latistriatus Luederwaldt, 1935 | | Dichotomius longiceps Taschenberg, 1870 | | | Dichotomius lucianomourai Nunes & Vaz-..., 2019 | | | Dichotomius luctuosioides Luederwaldt, 1922 | | | Dichotomius mamillatus Felsche, 1901 | | | Dichotomius marcoscarvalhoi Nunes & Vaz-..., 2019 | | | Dichotomius maya Peraza & Deloya, 2006 | | | Dichotomius melzeri Luederwaldt, 1922 | | | Dichotomius missionus Martínez, 1947 | | | Dichotomius monstrosus Harold, 1875 | | | Dichotomius mormon Ljungh, 1799 | | | Dichotomius motai Pereira, 1942 | | | Dichotomius nemoricola Pereira, 1942 | | | Dichotomius nevermanni Luederwaldt, 1935 | | | Dichotomius nimuendaju Luederwaldt, 1925 | | | Dichotomius nimuendajui Luederwaldt, 1925 | | | Dichotomius nitidissimus Waterhouse, 1891 | | | Dichotomius nutans Harold, 1867 | | | Dichotomius ohausi Luederwaldt, 1923 | | | Dichotomius paschoali Nunes & Vaz-..., 2019 | | | Dichotomius planicollis Gillet, 1911 | | | Dichotomius podalirius Felsche, 1901 | | | Dichotomius prietoi Martinez & M..., 1982 | | | Dichotomius protectus Harold, 1867 | | | Dichotomius provisorius Luederwaldt, 1925 | | | Dichotomius puncticollis Luederwaldt, 1935 | | | Dichotomius quadrilobatus Chamorro, Lo..., 2021 | | | Dichotomius quadrinodosus Felsche, 1901 | | | Dichotomius quinquedens Felsche, 1910 | | | Dichotomius quinquelobatus Felsche, 1901 | | | Dichotomius rafanunezi Montoya-Moli..., 2021 | | | Dichotomius reclinatus Felsche, 1901 | | | Dichotomius reichei Harold, 1869 | | | Dichotomius ribeiroi Pereira, 1954 | | | Dichotomius ricardosilvai Nunes & Vaz-..., 2019 | | | Dichotomius robustus Luederwaldt, 1935 | Apex mesh weaver | | Dichotomius rondoniaensis Nunes & Vaz-..., 2019 | | | Dichotomius rotundatus Burmeister, 1874 | | | Dichotomius rugosicollis Luederwaldt, 1935 | | | Dichotomius sagittarius Harold, 1869 | | | Dichotomius satanas Harold, 1867 | | | Dichotomius satanas angustus Luederwaldt, 1923 | | | Dichotomius satanas satanas | | | Dichotomius seag Nunes & Vaz-..., 2019 | | | Dichotomius semiaeneus Germar, 1824 | | | Dichotomius semicircularis Luederwaldt, 1929 | | | Dichotomius subaeneus Castelnau, 1840 | | | Dichotomius telamon Harold, 1869 | | | Dichotomius tissianiae Nunes & Vaz-..., 2019 | | | Dichotomius torulosus Luederwaldt, 1925 | | | Dichotomius travassosi (Martínez, 1955) | ⇒ Dichotomius nitidissimus Waterhouse, 1891 | | Dichotomius triangulariceps Blanchard, 1846 | | | Dichotomius valoisae Silva, Moura..., 2020 | | | Dichotomius vargasae Nunes & Vaz-..., 2019 | | | Dichotomius virescens Luederwaldt, 1935 | | | Dichotomius woodruffi Solis & Kohl..., 2022 | | | Dichotomius worontzowi Pereira, 1942 | | | Dichotomius zikani Luederwaldt, 1922 | | | Dicranocara deschodti Frolov & Sch..., 2003 | | | Dicranocara inexpectata Deschodt & S..., 2007 | | | Dicranocara tatasensis Deschodt & S..., 2007 | | | Dicranocara vandersmisseni Moretto, 2016 | | | Digitonthophagus aksumensis Génier, 2017 | | | Digitonthophagus biflagellatus Génier, 2017 | | | Digitonthophagus bonasus Fabricius, 1775 | | | Digitonthophagus catta Fabricius, 1787 | Beetle | | Digitonthophagus dilatatus Génier, 2017 | | | Digitonthophagus eucatta Génier, 2017 | | | Digitonthophagus falciger Génier, 2017 | | | Digitonthophagus fimator Génier, 2017 | | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Digitonthophagus lusinganus Orbigny, 1905 | | | Digitonthophagus namaquensis Génier, 2017 | | | Digitonthophagus petilus Génier, 2017 | | | Digitonthophagus sahelicus Moretto, 2017 | | | Digitonthophagus uks Génier, 2017 | | | Digitonthophagus ulcerosus Génier, 2017 | | | Digitonthophagus viridicollis Génier, 2017 | | | Diorygopyx asciculifer Matthews, 1974 | | | Diorygopyx cuspidatus Matthews, 1974 | | | Diorygopyx duplodentatus Matthews, 1974 | | | Diorygopyx incomptus Matthews, 1974 | | | Diorygopyx incrassatus Matthews, 1974 | | | Diorygopyx niger Matthews, 1974 | | | Diorygopyx simpliciclunis Matthews, 1974 | | | Diorygopyx tibialis MacLeay, 1871 | | | Disphysema candezei Harold, 1873 | | | Dorbignyolus mirecribratus d´Orbigny, 1913 | | | Drepanocerus abyssinicus (Roth, 1851) | ⇒ Ixodina abyssinica Roth, 1851 | | Drepanocerus ambiguus Kirby, 1828 | | | Drepanocerus arrowi (Bathasar, 1932) | ⇒ Sinodrepanus rex Boucomont, 1912 | | Drepanocerus bechynei (Janssens, 1953) | ⇒ Eodrepanus bechynei Janssens, 1953 | | Drepanocerus endroedy Endrodi, 1976 | | | Drepanocerus fastiditus (Péringuey, 1901) | ⇒ Eodrepanus fastiditus Péringuey, 1901 | | Drepanocerus impressicollis (Boheman, 1857) | ⇒ Afrodrepanus impressicollis Boheman, 1857 | | Drepanocerus kirbyi Kirby, 1828 | | | Drepanocerus laticollis (Fahraeus, 1857) | ⇒ Latodrepanus laticollis Fahraeus, 1857 | | Drepanocerus marshalli (Boucomont, 1921) | ⇒ Afrodrepanus marshalli Boucomont, 1921 | | Drepanocerus orientalis Krikken, 2009 | | | Drepanocerus parallelus (Raffray, 1877) | ⇒ Eodrepanus parallelus Raffray, 1877 | | Drepanocerus patrizii Boucomont, 1923 | | | Drepanocerus planus Fabricius, 1801 | | | Drepanocerus rex (Boucomont, 1912) | ⇒ Sinodrepanus rex Boucomont, 1912 | | Drepanocerus roconditus | | | Drepanocerus runicus (Arrow, 1909) | ⇒ Ixodina runica Arrow, 1909 | | Drepanocerus sinicus (Harold, 1868) | ⇒ Tibiodrepanus sinicus Harold, 1868 | | Drepanocerus striatus (Boucomont, 1921) | ⇒ Clypeodrepanus striatus Boucomont, 1921 | | Drepanocerus strigatus (Janssens, 1953) | ⇒ Clypeodrepanus strigatus Janssens, 1953 | | Drepanocerus sulcicollis (Castelnau, 1840) | ⇒ Sulcodrepanus sulcicollis Castelnau, 1840 | | Drepanocerus trogiformis Roth, 1851 | | | Drepanoplatynus gilleti Boucomont, 1921 | | | Drepanopodus costatus Wiedemann, 1823 | | | Drepanopodus proximus Péringuey, 1901 | | | Dwesasilvasedis medinae Deschodt & S..., 2008 | | | Endroedyolus paradoxus Scholtz & Ho..., 1987 | | | Ennearabdus lobocephalus Harold, 1868 | | | Eodrepanus bechynei Janssens, 1953 | | | Eodrepanus coopei Barbero, Pal..., 2009 | | | Eodrepanus fastiditus Péringuey, 1901 | | | Eodrepanus integriceps Janssens, 1953 | | | Eodrepanus liuchungloi Kryzhanovsky..., 1966 | | | Eodrepanus morgani Barbero, Pal..., 2009 | | | Eodrepanus paolae Barbero, Pal..., 2009 | | | Eodrepanus parallelus Raffray, 1877 | | | Eodrepanus striatulus Paulian, 1945 | | | Epactoides andringitrae Paulian, 1975 | | | Epactoides ankasokae Paulian, 1975 | | | Epactoides basilewskyi Balthasar, 1960 | | | Epactoides betschi Paulian, 1975 | | | Epactoides costatus Paulian, 1992 | | | Epactoides emkeni Montreuil, 2017 | | | Epactoides femoralis Montreuil, 2005 | | | Epactoides fiorii Paulian, 1976 | | | Epactoides frontalis Montreuil, 2003 | | | Epactoides giganteus Rossini, Vaz..., 2021 | | | Epactoides hanskii Montreuil, 2005 | | | Epactoides helenae Montreuil, 2005 | | | Epactoides humberti Paulian, 1975 | | | Epactoides humeralis Paulian, 1976 | | | Epactoides hyphydroides Lebis, 1953 | | | Epactoides incertus Lebis, 1953 | | | Epactoides jounii Wirta & Mont..., 2008 | | | Epactoides lacustris Paulian, 1976 | | | Epactoides lissus Lebis, 1953 | | | Epactoides madecassus Paulian, 1936 | | | Epactoides mahaboi Paulian, 1976 | | | Epactoides major Paulian, 1991 | | | Epactoides mananarae Paulian, 1976 | | | Epactoides mangabeensis Wirta & Mont..., 2008 | | | Epactoides masoalae Paulian, 1976 | | | Epactoides olsoufieffi Lebis, 1953 | | | Epactoides paradoxus Paulian, 1976 | | | Epactoides pauliani Lebis, 1953 | | | Epactoides perinetanus Paulian, 1975 | | | Epactoides perrieri Fairmaire, 1898 | | | Epactoides rahagai Montreuil, 2005 | | | Epactoides sambavae Paulian, 1976 | | | Epactoides semiaeneus Lebis, 1953 | | | Epactoides spinicollis Montreuil, 2003 | | | Epactoides tiinae Montreuil, 2005 | | | Epactoides vadoni Montreuil, 2005 | | | Epactoides vagecarinatus Lebis, 1953 | | | Epactoides viridicollis Montreuil, 2005 | | | Epactoides vondrozoi Paulian, 1976 | | | Epilissus alluaudi Lebis, 1953 | | | Epilissus andranobeensis Montreuil, 2011 | | | Epilissus antoetrae Paulian, 1975 | | | Epilissus apotolamproides Lebis, 1961 | | | Epilissus armipes Lebis, 1953 | | | Epilissus bemarahaensis Montreuil & ..., 2022 | | | Epilissus colasi Paulian, 1935 | | | Epilissus cuprarius Fairmaire, 1889 | | | Epilissus cuprarius discretus Montreuil, 2006 | | | Epilissus cuprarius serindicus Paulian, 1976 | | | Epilissus delphinensis Lebis, 1953 | | | Epilissus discretus Montreuil, 2006 | | | Epilissus emmae Lebis, 1953 | | | Epilissus emmae emmae Lebis, 1953 | | | Epilissus emmae obscuripennis Montreuil, 2006 | | | Epilissus fantamattii Montreuil, 2011 | | | Epilissus fuscovirens (Fairmaire, 1889) | ⇒ Arachnodes fuscovirens Fairmaire, 1889 | | Epilissus genieri Montreuil, 2006 | | | Epilissus guillaumeti Paulian, 1975 | | | Epilissus hova Künckel d’..., 1887 | | | Epilissus mantasoae Paulian, 1976 | | | Epilissus micheli Lebis, 1953 | | | Epilissus morio (Harold, 1879) | ⇒ Arachnodes morio Harold, 1879 | | Epilissus morio hova Künckel d’He..., 1887 | | | Epilissus morio morio Harold, 1879 | | | Epilissus morio serindicus Paulian, 1976 | | | Epilissus niger (Lansberge, 1874) | ⇒ Lepanus niger Lansberge, 1874 | | Epilissus planatus Montreuil, 2006 | | | Epilissus prasinus Klug, 1832 | | | Epilissus robustus Montreuil & ..., 2022 | | | Epilissus ruteri Lebis, 1953 | | | Epilissus semicribrosus (Fairmaire, 1898) | ⇒ Nanos semicribrosus Fairmaire, 1898 | | Epilissus semilissus Montreuil & ..., 2022 | | | Epilissus sikorai Paulian, 1977 | | | Epilissus splendidus (Fairmaire, 1889) | ⇒ Arachnodes splendidus Lebis, 1953 | | Epilissus splendidus delphinensis Lebis, 1953 | | | Epilissus splendidus descarpentriesi Lebis, 1953 | | | Epilissus splendidus splendidus Fairmaire, 1889 | | | Epilissus striatoides Paulian, 1976 | | | Epilissus striatus Castelnau, 1840 | | | Epilissus superbus Montreuil & ..., 2022 | | | Epilissus viridis Klug, 1832 | | | Epilissus vohemarensis Montreuil & ..., 2022 | | | Epirinus aeneus Wiedemann, 1823 | | | Epirinus aquilus Medina & Sch..., 2005 | | | Epirinus asper Péringuey, 1901 | | | Epirinus bentoi Ferreira, 1964 | | | Epirinus comosus Péringuey, 1901 | | | Epirinus convexus Scholtz & Ho..., 1987 | | | Epirinus davisi Scholtz & Ho..., 1987 | | | Epirinus drakomontanus Scholtz & Ho..., 1987 | | | Epirinus flagellatus Fabricius, 1775 | | | Epirinus granulatus Scholtz & Ho..., 1987 | | | Epirinus gratus Péringuey, 1901 | | | Epirinus hilaris Péringuey, 1901 | | | Epirinus hluhluwensis Medina & Sch..., 2005 | | | Epirinus hopei Castelnau, 1840 | | | Epirinus inparrugosus Deschodt & D..., 2019 | | | Epirinus jacobsae Deschodt & D..., 2019 | | | Epirinus minimus Medina & Sch..., 2005 | | | Epirinus montanus Scholtz & Ho..., 1987 | | | Epirinus mucrodentatus Scholtz & Ho..., 1987 | | | Epirinus muellerae Deschodt & D..., 2019 | | | Epirinus ngomae Medina & Sch..., 2005 | | | Epirinus obtusus Boheman, 1857 | | | Epirinus pseudorelictus Deschodt & D..., 2019 | | | Epirinus pseudorugosus Medina & Sch..., 2005 | | | Epirinus punctatus Scholtz & Ho..., 1987 | | | Epirinus pygidialis Scholtz & Ho..., 1987 | | | Epirinus pygidialus Scholtz & Ho..., 1987 | | | Epirinus relictus Scholtz & Ho..., 1987 | | | Epirinus rugosus Scholtz & Ho..., 1987 | | | Epirinus schoolmeestersi Deschodt & D..., 2019 | | | Epirinus scrobiculatus Harold, 1880 | | | Epirinus sebastiani Medina & Sch..., 2005 | | | Epirinus silvestris Cambefort, 1978 | | | Epirinus striatus Scholtz & Ho..., 1987 | | | Epirinus sulcipennis Boheman, 1857 | | | Epirinus validus Péringuey, 1901 | | | Eucranium arachnoides Brullé, 1837 | | | Eucranium belenae Ocampo, 2010 | | | Eucranium cyclosoma Burmeister, 1861 | | | Eucranium dentifrons Guérin-Ménev..., 1838 | | | Eucranium planicolle Burmeister, 1861 | | | Eucranium pulvinatum (Burmeister, 1873) | ⇒ Eucranium arachnoides Brullé, 1837 | | Eucranium simplicifrons Fairmaire, 1893 | | | Eudinopus dytiscoides Schreibers, 1802 | | | Euoniticellus africanus Harold, 1873 | Greater sandy dung beetle | | Euoniticellus cambeforti Masumoto, 1987 | | | Euoniticellus capnus Cambefort, 1996 | | | Euoniticellus cubiensis Castelnau, 1840 | Scarab beetle | | Euoniticellus fulvus Goeze, 1777 | | | Euoniticellus fumigatus Boucomont, 1923 | | | Euoniticellus inaequalis Reiche, 1850 | | | Euoniticellus intermedius Reiche, 1848 | Intermediate sandy dung beetle | | Euoniticellus kawanus Janssens, 1939 | | | Euoniticellus nasicornis Reiche, 1849 | | | Euoniticellus pallens Olivier, 1789 | | | Euoniticellus pallipes Fabricius, 1781 | Southern sandy dung beetle | | Euoniticellus parvus Kraatz, 1895 | | | Euoniticellus perniger Cambefort & ..., 1980 | | | Euoniticellus tai Cambefort, 1983 | | | Euoniticellus tibatensis Kolbe, 1904 | | | Euoniticellus triangulatus Harold, 1873 | | | Euoniticellus wittei Janssens, 1939 | | | Euoniticellus zumpti Janssens, 1953 | | | Euonthophagus aeneobrunneus Kabakov, 1977 | | | Euonthophagus amieti Cambefort, 1972 | | | Euonthophagus amyntas Olivier, 1789 | | | Euonthophagus amyntas alces Fabricius, 1792 | | | Euonthophagus amyntas amyntas Olivier, 1789 | | | Euonthophagus amyntas auchenia Redtenbacher, 1850 | | | Euonthophagus amyntas subviolaceus Ménétries, 1832 | | | Euonthophagus atramentarius Ménétriés, 1832 | | | Euonthophagus bedeli Reitter, 1891 | | | Euonthophagus carbonarius Klug, 1855 | | | Euonthophagus consanguineus Müller, 1944 | | | Euonthophagus consobrinus Péringuey, 1901 | | | Euonthophagus conterminus Petrovitz, 1971 | | | Euonthophagus crocatus Mulsant & Go..., 1870 | | | Euonthophagus crocatus clementi Baraud, 1960 | | | Euonthophagus crocatus crocatus | | | Euonthophagus dorbignyi Olsoufieff, 1900 | | | Euonthophagus dorbignyi neutto Olsoufieff, 1900 | | | Euonthophagus flavimargo Orbigny, 1902 | | | Euonthophagus gibbosus Scriba, 1790 | | | Euonthophagus hazariensis Kabakov, 1977 | | | Euonthophagus jeanneli diversiceps Frey, 1975 | | | Euonthophagus jeanneli jeanneli | | | Euonthophagus koshantschikoffi Reitter, 1891 | | | Euonthophagus laghmanicus Kabakov, 1977 | | | Euonthophagus loeffleri Petrovitz, 1965 | | | Euonthophagus maindroni d´Orbigny, 1897 | | | Euonthophagus mostafatsairii Palestrini, ..., 1979 | | | Euonthophagus pentaceros Olsoufieff, 1900 | | | Euonthophagus pertinax Balthasar, 1959 | | | Euonthophagus schnabeli Splichal, 1910 | | | Euonthophagus sulcicollis Reitter, 1892 | | | Euonthophagus tissoni Reitter, 1906 | | | Euonthophagus ustus d´Orbigny, 1905 | | | Euonthophagus verberatus Ziani, 2006 | | | Euonthophagus vicarius Péringuey, 1901 | | | Euonthophagus yama Ziani, 2006 | | | Euonthophagus yemenicus Balthasar, 1946 | | | Eurysternus aeneus Génier, 2009 | | | Eurysternus angustulus Harold, 1869 | | | Eurysternus arnaudi Génier, 2009 | | | Eurysternus atrosericus Génier, 2009 | | | Eurysternus balachowskyi Halffter & H..., 1976 | | | Eurysternus balthasari (Martínez, 1988) | ⇒ Eurysternus balachowskyi Halffter & H..., 1976 | | Eurysternus calligrammus Dalman, 1824 | | | Eurysternus cambeforti Génier, 2009 | | | Eurysternus caribaeus Herbst, 1789 | | | Eurysternus cavatus Génier, 2009 | | | Eurysternus cayennensis Castelnau, 1840 | | | Eurysternus claudicans (Kirsch, 1871) | ⇒ Eurysternus foedus Guérin-Ménev..., 1844 | | Eurysternus contractus Génier, 2009 | | | Eurysternus cyanescens Balthasar, 1939 | | | Eurysternus cyclops Génier, 2009 | | | Eurysternus deplanatus Germar, 1824 | | | Eurysternus fallaciosus Génier, 2009 | | | Eurysternus foedus Guérin-Ménev..., 1844 | | | Eurysternus francinae Génier, 2009 | | | Eurysternus gilli Génier, 2009 | | | Eurysternus gracilis Génier, 2009 | | | Eurysternus hamaticollis Balthasar, 1939 | | | Eurysternus harlequin Génier, 2009 | | | Eurysternus hirtellus Dalman, 1824 | | | Eurysternus howdeni Génier, 2009 | | | Eurysternus hypocrita Balthasar, 1939 | | | Eurysternus impressicollis Castelnau, 1840 | | | Eurysternus inca Génier, 2009 | | | Eurysternus inflexus Germar, 1824 | | | Eurysternus jessopi Martínez, 1988 | | | Eurysternus joffrei (Martínez, 1988) | ⇒ Eurysternus plebejus Harold, 1880 | | Eurysternus lanuginosus Génier, 2009 | | | Eurysternus magnus Castelnau, 1840 | | | Eurysternus marmoreus Castelnau, 1840 | | | Eurysternus martinsi Génier, 2009 | | | Eurysternus maya Génier, 2009 | | | Eurysternus mexicanus Harold, 1869 | | | Eurysternus nanus Génier, 2009 | | | Eurysternus navajasi Martínez, 1988 | | | Eurysternus nigrovirens Génier, 2009 | | | Eurysternus obliteratus Génier, 2009 | | | Eurysternus olivaceus Génier, 2009 | | | Eurysternus opatrinus (Perty, 1830) | ⇒ Eurysternus inflexus Germar, 1824 | | Eurysternus parallelus Castelnau, 1840 | | | Eurysternus plebejus Harold, 1880 | | | Eurysternus rivaldii Martínez, 1988 | | | Eurysternus sanbornei Gill, 1990 | | | Eurysternus squamosus Génier, 2009 | | | Eurysternus streblus Génier, 2009 | | | Eurysternus strigilatus Génier, 2009 | | | Eurysternus sulcifer Balthasar, 1939 | | | Eurysternus superbus Génier, 2009 | | | Eurysternus truncus Génier, 2009 | | | Eurysternus uniformis Génier, 2009 | | | Eurysternus vastiorum Martínez, 1988 | | | Eurysternus velutinus Bates, 1887 | | | Eurysternus ventricosus Gill, 1990 | | | Eurysternus wittmerorum Martínez, 1988 | | | Eusaproecius congolanus Balthasar, 1963 | | | Eusaproecius dudleyi Branco, 1992 | | | Eusaproecius laticeps d´Orbigny, 1908 | | | Eusaproecius magnigena d´Orbigny, 1913 | | | Eusaproecius optatus Péringuey, 1901 | | | Eusaproecius punctulatus Boucomont, 1928 | | | Eusaproecius tinantae Boucomont, 1923 | | | Eutrichillum arcus Solis & Kohl..., 2003 | | | Eutrichillum hirsutum Boucomont, 1928 | | | Eutrichillum hystrix Arrow, 1931 | | | Falsignambia grandis Paulian, 1987 | | | Feeridium woodruffi Vaz-de-Mello, 2008 | | | Frankenbergerius armatus Boheman, 1857 | | | Frankenbergerius armatus armatus | | | Frankenbergerius armatus tuberculatus Frolov & Sch..., 2005 | | | Frankenbergerius barratti Waterhouse, 1876 | | | Frankenbergerius forcipatus Harold, 1880 | | | Frankenbergerius gomesi Ferreira, 1954 | | | Frankenbergerius nanus Frolov & Sch..., 2005 | | | Frankenbergerius nitidus Frolov & Sch..., 2005 | | | Frankenbergerius opacus Frolov & Sch..., 2005 | | | Garreta annoyeri Moretto, 2021 | | | Garreta australugens Davis & Desc..., 2018 | | | Garreta azareus | | | Garreta azureus Fabricius, 1801 | | | Garreta basilewskyi Balthasar, 1961 | | | Garreta bechynei Pokorný & Zí..., 2018 | | | Garreta caffer Fahraeus, 1857 | | | Garreta crenulatus Kolbe, 1895 | | | Garreta dejeani Castelnau, 1840 | | | Garreta diffinis Waterhouse, 1890 | | | Garreta fastiditus Harold, 1867 | | | Garreta gilleti Garreta, 1914 | | | Garreta laetus Hope, 1842 | | | Garreta laetus laetus | | | Garreta laetus olivaceus Quedenfeldt, 1884 | | | Garreta lugens Fairmaire, 1891 | | | Garreta malleolus Kolbe, 1895 | | | Garreta matabelensis Janssens, 1938 | | | Garreta mombelgi Boucomont, 1929 | | | Garreta morosus Fairmaire, 1886 | | | Garreta mundus Wiedemann, 1819 | | | Garreta namalugens Davis & Desc..., 2018 | | | Garreta nitens Olivier, 1789 | | | Garreta nyassicus Kolbe, 1897 | | | Garreta opacus Redtenbacker, 1848 | | | Garreta ruficornis Motschulsky, 1854 | | | Garreta rutilans Castelnau, 1840 | | | Garreta smaragdifer Walker, 1858 | | | Garreta sumptuosus Castelnau, 1840 | | | Garreta sylvestris Mittal, 2011 | | | Garreta unicolor Fahraeus, 1857 | | | Garreta wahlbergi Fahraeus, 1857 | | | Garreta zumpti Frey, 1967 | | | Genieridium bidens Balthasar, 1938 | | | Genieridium bordoni Martínez, 1992 | | | Genieridium cryptops Arrow, 1913 | | | Genieridium margareteae Génier & Vaz..., 2002 | | | Genieridium medinae Gill & Vaz-d..., 2002 | | | Genieridium paranense Arrow, 1932 | | | Genieridium zanunciorum Vaz-de-Mello..., 1998 | | | Gilletellus collarti Janssens, 1937 | | | Gilletellus porculus Boheman, 1857 | | | Glauconia kefersteini | | | Glauconia undulata | | | Glyphoderus centralis Burmeister, 1873 | | | Glyphoderus monticola Burmeister, 1861 | | | Glyphoderus sterquilinus Westwood, 1837 | | | Gromphas aeruginosa Perty, 1830 | | | Gromphas amazonica Bates, 1870 | | | Gromphas dichroa Blanchard, 1846 | | | Gromphas inermis Harold, 1869 | | | Gromphas jardim Cupello & Va..., 2015 | | | Gromphas lacordairei (Brullé, 1837) | ⇒ Gromphas inermis Harold, 1869 | | Gromphas lemoinei (Waterhouse, 1891) | ⇒ Gromphas aeruginosa Perty, 1830 | | Gymnopleurus aciculatus Gebler, 1841 | | | Gymnopleurus aenescens Wiedemann, 1821 | | | Gymnopleurus aeruginosus Harold, 1867 | | | Gymnopleurus aethiops (Sharp, 1875) | ⇒ Paragymnopleurus melanarius Harold, 1867 | | Gymnopleurus andreaei Ferreira, 1954 | | | Gymnopleurus asperrimus Felsche, 1909 | | | Gymnopleurus assamensis (Waterhouse, 1890) | ⇒ Paragymnopleurus sinuatus assamensis Waterhouse, 1890 | | Gymnopleurus atratus Klug, 1845 | | | Gymnopleurus barovskyi (Kieseritsky, 1928) | ⇒ Gymnopleurus flagellatus Fabricius, 1787 | | Gymnopleurus bicallosus Felsche, 1909 | | | Gymnopleurus bicolor (Latreille, 1823) | ⇒ Gymnopleurus leei Fabricius, 1792 | | Gymnopleurus biharensis Arrow, 1931 | | | Gymnopleurus bombayensis Arrow, 1931 | | | Gymnopleurus coerulescens Olivier, 1789 | | | Gymnopleurus colmanti Janssens, 1938 | | | Gymnopleurus coriarius (Herbst, 1789) | ⇒ Gymnopleurus flagellatus Fabricius, 1787 | | Gymnopleurus cyaneus (Fabricius, 1798) | ⇒ Gymnopleurus sericeifrons Fairmaire, 1887 | | Gymnopleurus deperditus ♰ O. Heer, 1862 | | | Gymnopleurus elegans Klug, 1845 | | | Gymnopleurus eocaenicus ♰ F. Meunier, 1921 | | | Gymnopleurus flagellatus Fabricius, 1787 | | | Gymnopleurus flagellatus asperatus Motschulsky, 1849 | | | Gymnopleurus flagellatus calignosus Balthasar, 1934 | | | Gymnopleurus flagellatus flagellatus Fabricius, 1787 | | | Gymnopleurus flagellatus hornei Waterhouse, 1890 | | | Gymnopleurus flagellatus serratus Fischer von ..., 1821 | | | Gymnopleurus foricarius Garreta, 1914 | | | Gymnopleurus fulgidus Olivier, 1789 | | | Gymnopleurus gemmatus Harold, 1871 | | | Gymnopleurus geoffroyi Fuessly, 1775 | | | Gymnopleurus geoffroyi cyanescens Motschulsky, 1849 | | | Gymnopleurus geoffroyi geoffroyi | | | Gymnopleurus geoffroyi virescens Müller, 1902 | | | Gymnopleurus humanus MacLeay, 1821 | | | Gymnopleurus humeralis Klug, 1855 | | | Gymnopleurus hypocrita Balthasar, 1960 | | | Gymnopleurus ignitus Klug, 1855 | | | Gymnopleurus imitator Balthasar, 1963 | | | Gymnopleurus jacksoni Waterhouse, 1890 | | | Gymnopleurus koenigi Fabricius, 1775 | | | Gymnopleurus koenigii Fabricius, 1775 | | | Gymnopleurus lacunosus Klug, 1845 | | | Gymnopleurus laetus Hope, 1842 | | | Gymnopleurus laevicollis (Castelnau, 1840) | ⇒ Gymnopleurus purpureus Garreta, 1914 | | Gymnopleurus latreillei Castelnau, 1840 | | | Gymnopleurus leei Fabricius, 1792 | | | Gymnopleurus miliaris Fabricius, 1775 | | | Gymnopleurus mimus (Balthasar, 1934) | ⇒ Gymnopleurus geoffroyi Fuessly, 1775 | | Gymnopleurus moerens Kolbe, 1895 | | | Gymnopleurus mopsus Pallas, 1781 | | | Gymnopleurus mopsus mopsus | | | Gymnopleurus mopsus sinensis Balthasar, 1934 | | | Gymnopleurus naviauxi Montreuil, 2009 | | | Gymnopleurus nyankpalaensis Endrödi, 1976 | | | Gymnopleurus particolor Davis, 1986 | | | Gymnopleurus parvus MacLeay, 1821 | | | Gymnopleurus persianus (Reitter, 1909) | ⇒ Gymnopleurus mopsus Pallas, 1781 | | Gymnopleurus plicatulus Fairmaire, 1890 | | | Gymnopleurus profanus (Fabricius, 1792) | ⇒ Garreta nitens Olivier, 1789 | | Gymnopleurus pumilus (Reiche, 1847) | ⇒ Gymnopleurus virens Erichson, 1843 | | Gymnopleurus puncticollis (Gillet, 1909) | ⇒ Gymnopleurus virens Erichson, 1843 | | Gymnopleurus purpureus Garreta, 1914 | | | Gymnopleurus pustulatus Kolbe, 1895 | | | Gymnopleurus qurosh Montreuil, 2011 | | | Gymnopleurus reichei Waterhouse, 1890 | | | Gymnopleurus rhodesianus Balthasar, 1963 | | | Gymnopleurus rotundatus ♰ O. Heer, 1862 | | | Gymnopleurus ruandensis Janssens, 1938 | | | Gymnopleurus sericatus (Erichson, 1843) | ⇒ Gymnopleurus humanus MacLeay, 1821 | | Gymnopleurus sericeifrons Fairmaire, 1887 | | | Gymnopleurus serratus (Fischer von ..., 1821) | ⇒ Gymnopleurus flagellatus Fabricius, 1787 | | Gymnopleurus sindensis Arrow, 1931 | | | Gymnopleurus sinuatus (Geoffroy, 1785) | ⇒ Gymnopleurus geoffroyi Fuessly, 1775 | | Gymnopleurus sisyphus ♰ O. Heer, 1847 | | | Gymnopleurus smaragdinus (Fahraeus, 1857) | ⇒ Gymnopleurus ignitus Klug, 1855 | | Gymnopleurus somaliensis Lansberge, 1882 | | | Gymnopleurus sturmi (MacLeay, 1821) | ⇒ Gymnopleurus sturmii MacLeay, 1821 | | Gymnopleurus sturmii MacLeay, 1821 | | | Gymnopleurus thelwalli Waterhouse, 1890 | | | Gymnopleurus thoracicus Harold, 1868 | | | Gymnopleurus tristis Castelnau, 1840 | | | Gymnopleurus tuxeni Petrovitz, 1955 | | | Gymnopleurus virens Erichson, 1843 | | | Hammondantus psammophilus Cambefort, 1978 | | | Hansreia affinis Fabricius, 1801 | | | Hansreia coriacea Schmidt, 1922 | | | Hansreia grossii Valois, Vaz-..., 2015 | | | Hansreia krinskii Valois, Vaz-..., 2015 | | | Hansreia oxygona Perty, 1830 | | | Hansreia peugeoti Valois, Vaz-..., 2015 | | | Haroldius annandalai Silvestri, 1924 | | | Haroldius annandalei Silvestri, 1924 | | | Haroldius borneensis Paulian, 1993 | | | Haroldius brendelli Krikken & Hu..., 2009 | | | Haroldius cambeforti Krikken & Hu..., 2006 | | | Haroldius cardoni Boucomont, 1923 | | | Haroldius celebensis Krikken & Hu..., 2006 | | | Haroldius convexus Philips & Sc..., 2000 | | | Haroldius discoidalis Paulian, 1993 | | | Haroldius ennearthrus Janssens, 1949 | | | Haroldius fairmairei Boucomont, 1914 | | | Haroldius fleutiauxi Paulian, 1945 | | | Haroldius globosus Boucomont, 1925 | | | Haroldius heimi Wasmann, 1918 | | | Haroldius herrenorum Paulian, 1985 | | | Haroldius hwangi Masumoto, Le..., 2005 | | | Haroldius jechaer Král & Hájek, 2012 | | | Haroldius kawadai Masumoto, 1995 | | | Haroldius kolaka Krikken & Hu..., 2006 | | | Haroldius krali Utsunomiya &..., 2000 | | | Haroldius lassallei Cambefort, 1986 | | | Haroldius leleupi Janssens, 1953 | | | Haroldius loebli Paulian, 1987 | | | Haroldius lyleae Daniel, Strü..., 2021 | | | Haroldius maruyamai Utsunomiya &..., 2005 | | | Haroldius modestus Janssens, 1953 | | | Haroldius oharai Ochi, Kon & ..., 2009 | | | Haroldius pahangensis Kral, 2003 | | | Haroldius pauliani Scheuern, 1995 | | | Haroldius penelopae Krikken & Hu..., 2006 | | | Haroldius perroti Paulian, 1939 | | | Haroldius philippinensis Pereira, 1954 | | | Haroldius rugatulus Boucomont, 1914 | | | Haroldius stefanii Montreuil, 2010 | | | Haroldius stevensi Arrow, 1931 | | | Haroldius sumatranus Paulian & Sc..., 1994 | | | Haroldius tangkoko Krikken & Hu..., 2006 | | | Haroldius thailandensis Paulian & Sc..., 1994 | | | Haroldius turnai Kral, 2003 | | | Haroldius uenoi Masumoto & Yin, 1993 | | | Helictopleurus aeneoniger Lebis, 1960 | | | Helictopleurus ambiguus Paulian & Ca..., 1991 | | | Helictopleurus aurocupreus Fairmaire, 1901 | | | Helictopleurus basicornis Fairmaire, 1904 | | | Helictopleurus bivittatus d´Orbigny, 1915 | | | Helictopleurus cambeforti Montreuil, 2005 | | | Helictopleurus carbonarius Lebis, 1960 | | | Helictopleurus clouei Harold, 1869 | | | Helictopleurus coruscus d´Orbigny, 1915 | | | Helictopleurus cribricollis Lebis, 1960 | | | Helictopleurus dorbignyi Montreuil, 2005 | | | Helictopleurus fasciolatus Fairmaire, 1898 | | | Helictopleurus fasciolatus fasciolatus | | | Helictopleurus fasciolatus obscurus Lebis, 1960 | | | Helictopleurus fasciolatus pseudofasciolatus Montreuil, 2007 | | | Helictopleurus fasciolatus suturalis Montreuil, 2012 | | | Helictopleurus fissicollis Fairmaire, 1895 | | | Helictopleurus fissicollis australis Montreuil, 2007 | | | Helictopleurus fissicollis fissicollis | | | Helictopleurus fissicollis morettoi Montreuil, 2005 | | | Helictopleurus fulgens Lebis, 1960 | | | Helictopleurus fulgens fulgens | | | Helictopleurus fulgens purpuricollis Lebis, 1960 | | | Helictopleurus fungicola Fairmaire, 1899 | | | Helictopleurus fungicola peyrierasi Paulian & Ca..., 1991 | | | Helictopleurus furcicornis Lebis, 1960 | | | Helictopleurus gibbicollis Fairmaire, 1895 | | | Helictopleurus giganteus Harold, 1869 | | | Helictopleurus halffteri Balthasar, 1964 | | | Helictopleurus hanskii Montreuil, 2005 | | | Helictopleurus heidiae Montreuil, 2007 | | | Helictopleurus hypocrita Balthasar, 1941 | | | Helictopleurus infimus Fairmaire, 1901 | | | Helictopleurus littoralis Montreuil, 2005 | | | Helictopleurus marsyas Olivier, 1789 | | | Helictopleurus minutus Montreuil, 2007 | | | Helictopleurus neoamplicollis Krell, 2000 | | | Helictopleurus neuter Fairmaire, 1898 | | | Helictopleurus nicollei Lebis, 1960 | | | Helictopleurus niger d´Orbigny, 1915 | | | Helictopleurus nigricans d´Orbigny, 1915 | | | Helictopleurus nigritulus Lebis, 1960 | | | Helictopleurus nitidicollis Lebis, 1960 | | | Helictopleurus patricii Paulian, 1937 | | | Helictopleurus perpunctatus Balthasar, 1963 | | | Helictopleurus perrieri Fairmaire, 1898 | | | Helictopleurus peyrierasi (Paulian & Ca..., 1991) | ⇒ Helictopleurus fungicola peyrierasi Paulian & Ca..., 1991 | | Helictopleurus pluristriatus d´Orbigny, 1915 | | | Helictopleurus politicollis Fairmaire, 1902 | | | Helictopleurus praerugosus Fairmaire, 1898 | | | Helictopleurus pseudoniger Montreuil, 2005 | | | Helictopleurus quadripunctatus Olivier, 1789 | | | Helictopleurus rubricollis Lebis, 1960 | | | Helictopleurus rudicollis Fairmaire, 1898 | | | Helictopleurus semimetallicus Fairmaire, 1888 | | | Helictopleurus seminiger d´Orbigny, 1915 | | | Helictopleurus semivirens d´Orbigny, 1915 | | | Helictopleurus sericeus Lebis, 1960 | | | Helictopleurus sicardi Paulian, 1975 | | | Helictopleurus sinuaticornis Fairmaire, 1898 | | | Helictopleurus sinuatocornis Fairmaire, 1898 | | | Helictopleurus splendidicollis Fairmaire, 1893 | | | Helictopleurus splendidicollis similis Montreuil, 2007 | | | Helictopleurus splendidicollis splendidicollis | | | Helictopleurus splendidus d´Orbigny, 1915 | | | Helictopleurus steineri Paulian & Ca..., 1991 | | | Helictopleurus subretusus Fairmaire, 1895 | | | Helictopleurus sumptuosus Harold, 1873 | | | Helictopleurus tamatavensis Montreuil, 2005 | | | Helictopleurus tristis (d´Orbigny, 1915) | ⇒ Helictopleurus quadripunctatus Olivier, 1789 | | Helictopleurus undatus Olivier, 1789 | | | Helictopleurus unifasciatus Fairmaire, 1901 | | | Helictopleurus vadoni Lebis, 1960 | | | Helictopleurus viettei Paulian & Ca..., 1984 | | | Helictopleurus villiersi Paulian & Ca..., 1984 | | | Helictopleurus viridans Fairmaire, 1901 | | | Helictopleurus viridans pseudoviridans Montreuil, 2007 | | | Helictopleurus viridans viridans | | | Helictopleurus viridiflavus Fairmaire, 1898 | | | Heliocopris alatus Felsche, 1910 | | | Heliocopris anadematus Gillet, 1908 | | | Heliocopris andersoni Bates, 1868 | | | Heliocopris anguliceps Janssens, 1943 | | | Heliocopris antenor (Olivier, 1789) | ⇒ Heliocopris colossus Bates, 1868 | | Heliocopris antiquus ♰ I. Fujiyama, 1968 | | | Heliocopris ares Moretto & Mi..., 2022 | | | Heliocopris atropos Boheman, 1860 | | | Heliocopris beccarii Harold, 1871 | | | Heliocopris biimpressus Kolbe, 1893 | | | Heliocopris bucephalus Fabricius, 1775 | | | Heliocopris camerunus Pokorny & Zidek, 2009 | | | Heliocopris colossus Bates, 1868 | | | Heliocopris corniculatus Janssens, 1939 | | | Heliocopris coronatus Felsche, 1901 | | | Heliocopris cuneifer Lesne, 1906 | | | Heliocopris densissa Roth, 1851 | | | Heliocopris dianae (Hope, 1842) | ⇒ Heliocopris mutabilis Kolbe, 1893 | | Heliocopris dilloni Guérin-Ménev..., 1847 | | | Heliocopris dolosus Janssens, 1939 | | | Heliocopris dominus Bates, 1868 | | | Heliocopris erycoides Felsche, 1907 | | | Heliocopris eryx (Fabricius, 1801) | ⇒ Heliocopris neptunus Boheman, 1857 | | Heliocopris eryx eryx | | | Heliocopris eryx ikelenge Moretto, 2017 | | | Heliocopris eryx mutabilis (Kolbe, 1895) | ⇒ Heliocopris mutabilis Kolbe, 1893 | | Heliocopris eryx tridentatus Moretto, 2017 | | | Heliocopris faunus Boheman, 1857 | | | Heliocopris felschei Kolbe, 1904 | | | Heliocopris fischeri Schäfer, 2019 | | | Heliocopris fonsecai Ferreira, 1967 | | | Heliocopris furcithorax Müller, 1941 | | | Heliocopris gigas Linnaeus, 1758 | | | Heliocopris gigas erythraeus Müller, 1941 | | | Heliocopris gigas gigas | | | Heliocopris hamadryas Fabricius, 1775 | | | Heliocopris hamifer Harold, 1878 | | | Heliocopris haroldi Kolbe, 1893 | | | Heliocopris helleri Felsche, 1907 | | | Heliocopris hermes Gillet, 1911 | | | Heliocopris hunteri Waterhouse, 1891 | | | Heliocopris isidis (Boheman, 1857) | ⇒ Heliocopris andersoni Bates, 1868 | | Heliocopris japetus Klug, 1855 | | | Heliocopris kolbei Felsche, 1901 | | | Heliocopris letiranti Moretto, 2021 | | | Heliocopris maou Moretto, 2014 | | | Heliocopris marshalli Péringuey, 1901 | | | Heliocopris midas Fabricius, 1775 | | | Heliocopris mimus Janssens, 1939 | | | Heliocopris minos Gillet, 1907 | | | Heliocopris mutabilis Kolbe, 1893 | | | Heliocopris myrmidon Kolbe, 1893 | | | Heliocopris neptuniformis Felsche, 1907 | | | Heliocopris neptunoides Janssens, 1939 | | | Heliocopris neptunus Boheman, 1857 | | | Heliocopris pauliani Janssens, 1939 | | | Heliocopris pirmal Fabricius, 1798 | | | Heliocopris quinqueangulatus Janssens, 1939 | | | Heliocopris samson Harold, 1878 | | | Heliocopris sirius (Gillet, 1925) | ⇒ Heliocopris hamifer Harold, 1878 | | Heliocopris smithi Moretto, 2017 | | | Heliocopris solitarius Kolbe, 1893 | | | Heliocopris staudingeri Kolbe, 1893 | | | Heliocopris stroehlei Moretto, 2014 | | | Heliocopris sylvanus Gillet, 1925 | | | Heliocopris tyrannus Thomson, 1859 | | | Heliocopris werneri Moretto & Mi..., 2013 | | | Heteroclitopus annamariae Branco, 1991 | | | Heteroclitopus collarti Janssens, 1939 | | | Heteroclitopus ferreirae Branco, 1991 | | | Heteroclitopus foveatus Boucomont, 1923 | | | Heteroclitopus foveolatus Branco & Josso, 2007 | | | Heteroclitopus freyi Ferreira, 1969 | | | Heteroclitopus herteli Ferreira, 1969 | | | Heteroclitopus remipes Péringuey, 1901 | | | Heteroclitopus schulzei Branco, 1991 | | | Heteroclitopus simonettai Branco, 1991 | | | Heteronitis castelnaui Harold, 1862 | | | Heteronitis pauliani Janssens, 1942 | | | Heteronitis ragazzii Fairmaire, 1893 | | | Heteronitis tricornutus Fairmaire, 1891 | | | Heteronitis tridens Castelnau, 1840 | | | Heterosyphus sicardi Paulian, 1975 | | | Holocanthon fuscorubrus Blanchard, 1845 | | | Holocanthon giosilvai Sawaris, Cup..., 2019 | | | Holocanthon mateui Martinez & P..., 1956 | | | Holocephalus cristatus Gillet, 1907 | | | Holocephalus eridanus Olivier, 1789 | | | Holocephalus julieni Smith & Génier, 2001 | | | Holocephalus pokornyi Nunes & Vaz-..., 2016 | | | Holocephalus sculptus Gillet, 1907 | | | Holocephalus simoni Smith & Génier, 2001 | | | Homalotarsus impressus Janssens, 1932 | | | Homocopris achamas Harold, 1867 | | | Homocopris buckleyi Waterhouse, 1891 | | | Homocopris punctatissimus Curtis, 1844 | | | Homocopris torulosus Eschscholtz, 1822 | | | Homocopris torulosus minor Luederwaldt, 1925 | | | Homocopris torulosus valdivianus Philippi, 1859 | | | Hyalonthophagus alcedo Orbigny, 1913 | | | Hyalonthophagus alcyon Klug, 1855 | | | Hyalonthophagus alcyonides Orbigny, 1913 | | | Hyalonthophagus hyalcyon Palestrini &..., 1989 | | | Hyalonthophagus mixtifrons Orbigny, 1910 | | | Hyalonthophagus nigroviolaceus Orbigny, 1902 | | | Hyalonthophagus novalcedo Josso, 2021 | | | Hyalonthophagus pseudoalcyon Orbigny, 1913 | | | Hyalonthophagus pseudovirens Orbigny, 1913 | | | Hyalonthophagus pulcher Deschodt & D..., 2019 | | | Hyalonthophagus virens d´Orbigny, 1913 | | | Hyalonthophagus viridiceps Orbigny, 1910 | | | Hypocanthidium globulum Balthasar, 1938 | | | Isocopris foveolatus Luederwaldt, 1931 | | | Isocopris hypocrita Lucas, 1859 | | | Isocopris imitator Felsche, 1901 | | | Isocopris inhiatus Germar, 1824 | | | Isocopris nitidus Luederwaldt, 1922 | | | Isocopris tarsalis Luederwaldt, 1931 | | | Isocopris xacriaba Rossini & Va..., 2017 | | | Ixodina abyssinica Roth, 1851 | | | Ixodina abyssinica abyssinica | | | Ixodina abyssinica tangana Endrödi, 1971 | | | Ixodina abyssinica tanganus Endrodi, 1971 | | | Ixodina bos Endrödi, 1976 | | | Ixodina endroedyi Endrödi, 1976 | | | Ixodina freyi Janssens, 1953 | | | Ixodina kovacsi Endrödi, 1976 | | | Ixodina runica Arrow, 1909 | | | Ixodina saegeri Balthasar, 1963 | | | Ixodina szunyoghyi Endrödi, 1971 | | | Ixodina szunyoghyi occidentalis Endrödi, 1976 | | | Ixodina szunyoghyi szunyoghyi | | | Janssensantus alatus Josso, 2022 | | | Janssensantus inexpectatus Paulian, 1976 | | | Janssensantus pauliani Scholtz & Ho..., 1987 | | | Janssensellus bicaviventris Cambefort, 1975 | | | Janssensellus rasplusi Cambefort, 1995 | | | Kheper aegyptiorum Latreille, 1823 | | | Kheper aegyptiorum aegyptiorum | | | Kheper aegyptiorum purpurascens Gerstaecker, 1871 | | | Kheper aeratus Gerstaecker, 1871 | | | Kheper asceticus Gillet, 1907 | | | Kheper bonellii MacLeay, 1821 | | | Kheper clericus Boheman, 1857 | | | Kheper cupreus Castelnau, 1840 | | | Kheper cuvieri MacLeay, 1821 | | | Kheper devotus Redtenbacher, 1848 | | | Kheper erichsoni Harold, 1867 | | | Kheper festivus Harold, 1868 | | | Kheper festivus festivus | | | Kheper festivus nicolasi Moretto, 1998 | | | Kheper festivus simoni Moretto, 1998 | | | Kheper festivus wellsi Moretto, 1998 | | | Kheper intermedius Gillet, 1909 | | | Kheper kalaharicus Davis, Desch..., 2011 | | | Kheper laevistriatus Fairmaire, 1893 | | | Kheper lamarcki MacLeay, 1821 | | | Kheper nigroaeneus Boheman, 1857 | | | Kheper paganus Harold, 1878 | | | Kheper platynotus Bates, 1888 | | | Kheper porosus Bates, 1888 | | | Kheper prodigiosus Erichson, 1843 | | | Kheper pustulosus Gerstaecker, 1871 | | | Kheper rolciki Pokorný & Zí..., 2015 | | | Kheper sanctus Fabricius, 1798 | | | Kheper subaeneus Harold, 1869 | | | Kheper vanderkelleni Lansberge, 1886 | | | Kheper venerabilis Harold, 1871 | | | Kheper vethi Lansberge, 1886 | | | Kheper zurstrasseni Davis, 1986 | | | Kolbeellus ateuchoides Lansberge, 1875 | | | Krikkenius smeenki Krikken, 1982 | | | Labroma horrens Sharp, 1873 | | | Labroma tuberculata Waterhouse, 1874 | | | Labroma umbratilis Matthews, 1974 | | | Larhodius calcaratus Janssens, 1934 | | | Larhodius hashimi Masumoto & U..., 2003 | | | Larhodius maruyamai Masumoto & U..., 2003 | | | Latodrepanus caelatus Gerstaecker, 1871 | | | Latodrepanus laticollis Fahraeus, 1857 | | | Latodrepanus nicolasi Barbero, Pal..., 2009 | | | Latodrepanus pulvinarius Balthasar, 1963 | | | Latodrepanus schimperi Janssens, 1953 | | | Latodrepanus simonisi Barbero, Pal..., 2009 | | | Leotrichillum louzadaorum Vaz-de-Mello..., 1998 | | | Lepanus andersonorum Gunter & Weir, 2019 | | | Lepanus arator Matthews, 1974 | | | Lepanus australis Matthews, 1974 | | | Lepanus bidentatus Wilson, 1922 | | | Lepanus bournei Paulian, 1987 | | | Lepanus burnetti Gunter & Weir, 2019 | | | Lepanus cameroni Gunter & Weir, 2021 | | | Lepanus carbinensis Gunter & Weir, 2021 | | | Lepanus cardwellensis Gunter & Weir, 2021 | | | Lepanus crenidens Gunter & Weir, 2019 | | | Lepanus dichrous Gillet, 1925 | | | Lepanus dukungarri Gunter & Weir, 2019 | | | Lepanus eungella Gunter & Weir, 2019 | | | Lepanus feehani Gunter & Weir, 2019 | | | Lepanus fossulatus Paulian, 1985 | | | Lepanus furcifer Matthews, 1974 | | | Lepanus gelasinus Matthews, 1974 | | | Lepanus geoffi Gunter & Weir, 2019 | | | Lepanus glaber Matthews, 1974 | | | Lepanus globulus MacLeay, 1887 | | | Lepanus gressitti Balthasar, 1966 | | | Lepanus gubara Gunter & Weir, 2019 | | | Lepanus guthrieae Gunter & Weir, 2019 | | | Lepanus howdeni Paulian, 1985 | | | Lepanus illawarensis | | | Lepanus illawarrensis Matthews, 1974 | | | Lepanus kulki Gunter & Weir, 2021 | | | Lepanus latheticus Matthews, 1974 | | | Lepanus lemannae Gunter & Weir, 2021 | | | Lepanus lentil Gunter & Weir, 2019 | | | Lepanus lingziae Gunter & Weir, 2019 | | | Lepanus loftyensis Matthews & Weir, 2002 | | | Lepanus mckenziei Gunter & Weir, 2019 | | | Lepanus meierae Gunter & Weir, 2021 | | | Lepanus menendezae Gunter & Weir, 2019 | | | Lepanus mirabilis Paulian, 1985 | | | Lepanus monteithi Matthews, 1974 | | | Lepanus niger Lansberge, 1874 | | | Lepanus nitidus Matthews, 1974 | | | Lepanus occidentalis Matthews, 1974 | | | Lepanus ovatus Balthasar, 1966 | | | Lepanus oxleyi Gunter & Weir, 2019 | | | Lepanus palumensis Matthews, 1974 | | | Lepanus papuanus Paulian, 1985 | | | Lepanus parapisoniae Matthews, 1974 | | | Lepanus pecki Gunter & Weir, 2019 | | | Lepanus penelopae Matthews & Weir, 2002 | | | Lepanus pisoniae Lea, 1923 | | | Lepanus podocarp Gunter & Weir, 2019 | | | Lepanus politus Carter, 1936 | | | Lepanus pseudovillosus Gunter & Weir, 2019 | | | Lepanus pungalina Gunter & Weir, 2019 | | | Lepanus pygmaeus MacLeay, 1888 | | | Lepanus reidi Gunter & Weir, 2019 | | | Lepanus reticulatus Gunter & Weir, 2019 | | | Lepanus sauroni Gunter & Weir, 2019 | | | Lepanus storeyi Weir & Monteith, 2010 | | | Lepanus topend Gunter & Weir, 2019 | | | Lepanus tozerensis Gunter & Weir, 2019 | | | Lepanus ustulatus Lansberge, 1874 | | | Lepanus vangerweni Gunter & Weir, 2021 | | | Lepanus vestitus Matthews, 1974 | | | Lepanus villosus Matthews, 1974 | | | Lepanus williamsi Gunter & Weir, 2021 | | | Lepanus yorkensis Gunter & Weir, 2019 | | | Liatongus affinis Arrow, 1908 | | | Liatongus amitinus Boucomont, 1921 | | | Liatongus ancorifer Kral & Rejsek, 1999 | | | Liatongus appositicornis Kral & Rejsek, 1999 | | | Liatongus arrowi Boucomont, 1921 | | | Liatongus aterrimus Boucomont, 1921 | | | Liatongus bucerus Fairmaire, 1891 | | | Liatongus californicus Horn, 1882 | | | Liatongus capricornis Arrow, 1931 | | | Liatongus clypeocornis Scheuern, 1988 | | | Liatongus davidi Boucomont, 1919 | | | Liatongus denticornis (Fairmaire, 1887) | ⇒ Liatongus bucerus Fairmaire, 1891 | | Liatongus endroedii Balthasar, 1956 | | | Liatongus fairmairei Boucomont, 1921 | | | Liatongus femoratus Illiger, 1800 | | | Liatongus ferreirae Balthasar, 1964 | | | Liatongus fulvostriatus d´Orbigny, 1916 | | | Liatongus gagatinus Hope, 1831 | | | Liatongus hastatus Kral & Rejsek, 1999 | | | Liatongus imitator (Balthasar, 1938) | ⇒ Liatongus medius Fairmaire, 1889 | | Liatongus incurvicornis Fairmaire, 1887 | | | Liatongus indicus Arrow, 1908 | | | Liatongus interruptus (Quedenfeldt, 1884) | ⇒ Liatongus fulvostriatus d´Orbigny, 1916 | | Liatongus martialis Harold, 1879 | | | Liatongus medius Fairmaire, 1889 | | | Liatongus mergacerus Hope, 1831 | | | Liatongus militaris Castelnau, 1840 | Yellow Shouldered Dung Beetle | | Liatongus minutus Motschulsky, 1860 | | | Liatongus monstrosus (Bates, 1887) | ⇒ Attavicinus monstrosus Bates, 1887 | | Liatongus phanaeoides Westwood, 1839 | | | Liatongus phanaeoides fumiroi (Yohena, 1937) | ⇒ Liatongus minutus Motschulsky, 1860 | | Liatongus pugionatus Boheman, 1858 | | | Liatongus raffrayi Lansberge, 1886 | | | Liatongus rhadamistus Fabricius, 1775 | | | Liatongus rhinoceros Arrow, 1931 | | | Liatongus rhinocerulus Bates, 1889 | | | Liatongus schoutedeni Boucomont, 1920 | | | Liatongus sjostedti Felsche, 1904 | | | Liatongus spathulatus Roth, 1851 | | | Liatongus taurus Boucomont, 1920 | | | Liatongus testudo Lansberge, 1886 | | | Liatongus triacanthus Boucomont, 1920 | | | Liatongus tridentatus Boucomont, 1919 | | | Liatongus tuberculicollis Felsche, 1909 | | | Liatongus upembanus Janssens, 1953 | | | Liatongus urus Kolbe, 1914 | | | Liatongus venator Fabricius, 1801 | | | Liatongus vertagus Fabricius, 1798 | | | Liatongus vseteckai (Balthasar, 1933) | ⇒ Liatongus fairmairei Boucomont, 1921 | | Lithanomala crassa ♰ A. G. Ponoma..., 1990 | | | Lithanomala oblonga ♰ A. G. Ponoma..., 1990 | | | Lithanomala oblongus Ponomarenko, 1990 | | | Lithanomala sibirica ♰ A. G. Ponoma..., 1990 | | | Litocopris muticus Boheman, 1857 | | | Litocopris punctiventris Waterhouse, 1891 | | | Litocopris ruandanus Balthasar, 1961 | | | Litocopris simplex Harold, 1869 | | | Lobateuchus parisii ♰ O. Montreuil, 2010 | | | Lobidion punctatissimum Génier, 2010 | | | Lophodonitis carinatus Felsche, 1907 | | | Lophodonitis minettii Moretto, 2011 | | | Macroderes amplior Frolov & Sch..., 2004 | | | Macroderes arrowi Janssens, 1939 | | | Macroderes bias Olivier, 1789 | | | Macroderes cederbergensis Abdalla & De..., 2018 | | | Macroderes cornutus Frolov & Sch..., 2004 | | | Macroderes dubius Péringuey, 1901 | | | Macroderes endroedyi Frolov & Sch..., 2004 | | | Macroderes fornicatus Sharp, 1880 | | | Macroderes foveatus Frolov & Sch..., 2004 | | | Macroderes gifboomi Abdalla & De..., 2018 | | | Macroderes greeni Kirby, 1818 | | | Macroderes leipoldti Abdalla & De..., 2018 | | | Macroderes minutus Frolov & Sch..., 2004 | | | Macroderes mutilans Kolbe, 1908 | | | Macroderes mutilatus Kolbe, 1908 | | | Macroderes namakwanus Frolov & Sch..., 2004 | | | Macroderes nitidus Harold, 1877 | | | Macroderes oreatus Abdalla & De..., 2018 | | | Macroderes politulus (Preudhomme d..., 1880) | ⇒ Macroderes bias Olivier, 1789 | | Macroderes porselinus Abdalla & De..., 2018 | | | Macroderes pristinus Sharp, 1880 | | | Macroderes soleiana Abdalla & De..., 2018 | | | Macroderes spectabilis Péringuey, 1901 | | | Macroderes tortuosus Abdalla & De..., 2018 | | | Macroderes undulatus Preudhomme d..., 1880 | | | Macropanelus fujitai Ochi, Kon & ..., 2008 | | | Macropanelus karasuyamai Ochi, Kon & ..., 2008 | | | Macropanelus sumatrensis Ochi & Kon &..., 1998 | | | Macropocopris symbioticus (Arrow, 1920) | ⇒ Onthophagus symbioticus Arrow, 1920 | | Madateuchus viettei (Paulian, 1953) | ⇒ Scarabaeus viettei Paulian, 1953 | | Malagoniella argentina Gillet, 1911 | | | Malagoniella argentina argentina | | | Malagoniella argentina australis Halffter & M..., 1966 | | | Malagoniella argentina cuprea Gillet, 1911 | | | Malagoniella astyanax Olivier, 1789 | | | Malagoniella astyanax astyanax Olivier, 1789 | | | Malagoniella astyanax columbica Harold, 1867 | | | Malagoniella astyanax polita Halffter, Pe..., 1960 | | | Malagoniella astyanax punctatostriata Blanchard, 1846 | | | Malagoniella astyanax yucateca Harold, 1863 | | | Malagoniella bicolor Guérin-Ménev..., 1840 | | | Malagoniella chalybaea Blanchard, 1846 | | | Malagoniella coerulea Balthasar, 1939 | | | Malagoniella cupreicollis Waterhouse, 1890 | | | Malagoniella lanei Lange, 1945 | | | Malagoniella magnifica Balthasar, 1939 | | | Malagoniella puncticollis Blanchard, 1846 | | | Malagoniella puncticollis aeneicollis Waterhouse, 1890 | | | Malagoniella puncticollis puncticollis | | | Malagoniella puncticollis tubericeps Gillet, 1911 | | | Malagoniella virens Harold, 1869 | | | Martinezidium fulgens Arrow, 1913 | | | Martinezidium galileoae Génier & Vaz..., 2002 | | | Martinezidium martinsi Ferreira & G..., 1993 | | | Martinezidium maya Vaz-de-Mello..., 2004 | | | Megalonitis bohemani Lansberge, 1875 | | | Megalonitis bohemanni Lansberge, 1875 | | | Megalonitis gigas Bertoloni, 1855 | | | Megaponerophilus megaponerae Brauns, 1914 | | | Megatharsis buckleyi Waterhouse, 1891 | | | Megathopa argentina (Gillet, 1911) | ⇒ Malagoniella argentina Gillet, 1911 | | Megathopa magnifica (Balthasar, 1939) | ⇒ Malagoniella magnifica Balthasar, 1939 | | Megathopa puncticollis (Blanchard, 1845) | ⇒ Malagoniella puncticollis Blanchard, 1846 | | Megathopa villosa Eschscholtz, 1822 | | | Megathopa violacea Blanchard, 1846 | | | Megathopa yucateca (Harold, 1863) | ⇒ Malagoniella astyanax yucateca Harold, 1863 | | Megathoposoma candezei Harold, 1873 | | | Melanocanthon bispinatus Robinson, 1941 | | | Melanocanthon granulifer Schmidt, 1920 | Pearl blenny | | Melanocanthon nigricornis Say, 1823 | | | Melanocanthon punctaticollis Schaeffer, 1915 | | | Mentophilus hollandiae Boisduval, 1835 | | | Mentophilus subsulcatus Sharp, 1873 | | | Metacatharsius abortivus Fairmaire, 1891 | | | Metacatharsius anderseni Waterhouse, 1891 | | | Metacatharsius anomalus Felsche, 1910 | | | Metacatharsius auberti Fairmaire, 1891 | | | Metacatharsius basilewskyi Balthasar, 1961 | | | Metacatharsius bateke Walter, 1976 | | | Metacatharsius benadirensis Müller, 1947 | | | Metacatharsius bidentatus Ferreira, 1964 | | | Metacatharsius brevicosta Müller, 1947 | | | Metacatharsius clypeolatus Balthasar, 1940 | | | Metacatharsius convexiusculus Shipp, 1897 | | | Metacatharsius darfurensis Balthasar, 1965 | | | Metacatharsius dentinum Ferreira, 1964 | | | Metacatharsius duplicatus Müller, 1941 | | | Metacatharsius exiguiformis Ferreira, 1964 | | | Metacatharsius exiguus Boheman, 1860 | | | Metacatharsius ferreirae Balthasar, 1965 | | | Metacatharsius ferrugineus Olivier, 1789 | | | Metacatharsius freyi Ferreira, 1964 | | | Metacatharsius gardoensis Balthasar, 1941 | | | Metacatharsius holyi Balthasar, 1965 | | | Metacatharsius imitator Balthasar, 1965 | | | Metacatharsius inermis Castelnau, 1840 | | | Metacatharsius janssensi Frey, 1963 | | | Metacatharsius kindiai Paulian, 1939 | | | Metacatharsius latifrons Harold, 1868 | | | Metacatharsius lomii Müller, 1941 | | | Metacatharsius marani Balthasar, 1940 | | | Metacatharsius minutus Lansberge, 1882 | | | Metacatharsius modestus Balthasar, 1965 | | | Metacatharsius murati Paulian, 1939 | | | Metacatharsius nubiensis Harold, 1878 | | | Metacatharsius omoensis Müller, 1941 | | | Metacatharsius opacus Waterhouse, 1891 | | | Metacatharsius ovulum Balthasar, 1940 | | | Metacatharsius parcepunctatus Balthasar, 1965 | | | Metacatharsius patrizii Müller, 1947 | | | Metacatharsius peleus Olivier, 1789 | | | Metacatharsius pernitidus Müller, 1941 | | | Metacatharsius perpunctatus Balthasar, 1965 | | | Metacatharsius petrovitzi Balthasar, 1965 | | | Metacatharsius politus Paulian, 1934 | | | Metacatharsius pollicatus Harold, 1878 | | | Metacatharsius pseudoopacus Ferreira, 1965 | | | Metacatharsius pumilioniformis Ferreira, 1964 | | | Metacatharsius pumilionis Wallengren, 1881 | | | Metacatharsius pusio Kolbe, 1908 | | | Metacatharsius renaudi Frey, 1960 | | | Metacatharsius rochai Ferreira, 1964 | | | Metacatharsius rosinae Balthasar, 1970 | | | Metacatharsius rubidus Fairmaire, 1891 | | | Metacatharsius rugosipennis Frey, 1975 | | | Metacatharsius rusingae ♰ R. Paulian, 1976 | | | Metacatharsius seminulum Balthasar, 1940 | | | Metacatharsius simulator Balthasar, 1939 | | | Metacatharsius tchadensis Balthasar, 1961 | | | Metacatharsius tenellus Müller, 1947 | | | Metacatharsius togoensis Kolbe, 1893 | | | Metacatharsius transvaalensis Balthasar, 1968 | | | Metacatharsius trianguliceps Balthasar, 1965 | | | Metacatharsius tricarenatus Ferreira, 1964 | | | Metacatharsius troglodytes (Boheman, 1857) | ⇒ Metacatharsius anderseni Waterhouse, 1891 | | Metacatharsius tuberifrons Fairmaire, 1884 | | | Metacatharsius useramus Kolbe, 1893 | | | Metacatharsius wittei Ferreira, 1964 | | | Metacatharsius zuluanus Balthasar, 1965 | | | Milichus annae Nicolas & Ca..., 1994 | | | Milichus apicalis (Fahraeus, 1857) | ⇒ Milichus rhodesianus Péringuey, 1904 | | Milichus boucomonti Cambefort, 1996 | | | Milichus dudleyae Cambefort, 1996 | | | Milichus dudleyi Cambefort, 1996 | | | Milichus dudleyorum Cambefort, 1996 | | | Milichus durandi Cambefort, 1996 | | | Milichus inaequalis Boucomont, 1928 | | | Milichus inaequalis inaequalis | | | Milichus inaequalis lamottei Cambefort, 1996 | | | Milichus incisiceps Janssens, 1954 | | | Milichus jugatus d´Orbigny, 1907 | | | Milichus lecourti Nicolas & Ca..., 1994 | | | Milichus merzi Cambefort, 1983 | | | Milichus picticollis Gerstaecker, 1871 | | | Milichus rhodesianus Péringuey, 1904 | | | Milichus serratus d´Orbigny, 1907 | | | Mimonthophagus ambiguus Péringuey, 1901 | | | Mimonthophagus anomalus Klug, 1855 | | | Mimonthophagus apicehirtus d´Orbigny, 1915 | | | Mimonthophagus hinnulus Klug, 1832 | | | Mimonthophagus limbibasis d´Orbigny, 1905 | | | Mimonthophagus pictisternum d´Orbigny, 1902 | | | Mimonthophagus semicirculifer d´Orbigny, 1902 | | | Mimonthophagus semisetosus d´Orbigny, 1902 | | | Mnematidium multidentatum (Klug, 1845) | ⇒ Scarabaeus multidentatum Klug, 1845 | | Mnematidium multidentatus Klug, 1845 | | | Mnematium ritchiei (MacLeay, 1821) | ⇒ Scarabaeus ritchiei MacLeay, 1821 | | Mnematium silenus Gray, 1832 | | | Monoplistes curvipes Lea, 1923 | | | Monoplistes haroldi Lansberge, 1874 | | | Monoplistes leai Paulian, 1934 | | | Monoplistes occidentalis MacLeay, 1888 | | | Monoplistes phanophilus Lea, 1923 | | | Monoplistes tropicus Lea, 1923 | | | Namakwanus davisi Deschodt & S..., 2007 | | | Namakwanus endroedyi Deschodt, Da..., 2011 | | | Namakwanus irishi Scholtz & Ho..., 1987 | | | Namakwanus kamfferi Deschodt & D..., 2018 | | | Namakwanus minutus Deschodt & D..., 2018 | | | Namakwanus scholtzi Deschodt & D..., 2017 | | | Namakwanus streyi Frolov, 2005 | | | Nanos agaboides Boucomont, 1937 | | | Nanos andreiae Montreuil & ..., 2014 | | | Nanos ankaranae Paulian, 1976 | | | Nanos antsalovaensis Montreuil & ..., 2011 | | | Nanos antsihanakensis Lebis, 1953 | | | Nanos ater Castelnau, 1840 | | | Nanos bemarahaensis Montreuil & ..., 2014 | | | Nanos bicoloratus Lebis, 1953 | | | Nanos bimaculatus Kunckel, 1887 | | | Nanos binotatus Lebis, 1953 | | | Nanos clypeatus Castelnau, 1840 | | | Nanos constricticollis Montreuil & ..., 2014 | | | Nanos dubitatus Lebis, 1953 | | | Nanos fusconitens Fairmaire, 1899 | | | Nanos hanskii Montreuil & ..., 2007 | | | Nanos humbloti Lebis, 1953 | | | Nanos humeralis Paulian, 1975 | | | Nanos incertus Paulian, 1976 | | | Nanos magnus Montreuil & ..., 2014 | | | Nanos manomboensis Montreuil & ..., 2007 | | | Nanos manongorivoensis Montreuil & ..., 2014 | | | Nanos marojejyensis Montreuil & ..., 2014 | | | Nanos minutus Castelnau, 1840 | | | Nanos mirjae Montreuil & ..., 2014 | | | Nanos mixtus Montreuil & ..., 2014 | | | Nanos nitens Lebis, 1953 | | | Nanos nitens nitens | | | Nanos nitens pseudonitens Lebis, 1953 | | | Nanos occidentalis Paulian, 1976 | | | Nanos peyrierasi (Paulian, 1976) | ⇒ Apotolamprus peyrierasi Paulian, 1985 | | Nanos pseudofusconitens Montreuil & ..., 2014 | | | Nanos pseudominutus Montreuil & ..., 2014 | | | Nanos pseudorubromaculatus Montreuil & ..., 2014 | | | Nanos pseudoviettei Montreuil & ..., 2014 | | | Nanos punctatus Boucomont, 1937 | | | Nanos pygmaeus Castelnau, 1840 | | | Nanos ranomafanaensis Montreuil & ..., 2014 | | | Nanos rubromaculatus Kunckel d´He..., 1887 | | | Nanos rubrosignatus Lebis, 1953 | | | Nanos semicribrosus Fairmaire, 1898 | | | Nanos sinuatipes Boucomont, 1937 | | | Nanos vadoni Lebis, 1953 | | | Nanos viettei Paulian, 1976 | | | Nanos viridissimus Montreuil & ..., 2011 | | | Nebulasilvius insularis Deschodt & C..., 2008 | | | Nebulasilvius johani Deschodt & S..., 2008 | | | Neocanthidium bokermanni (Martinez, Ha..., 1964) | ⇒ Canthidium bokermanni Martinez, Ha..., 1964 | | Neocanthidium kelleri (Martinez, Ha..., 1964) | ⇒ Canthidium kelleri Martinez, Ha..., 1964 | | Neonitis nigritiae Gillet, 1918 | | | Neonitis rhodesiae Gillet, 1918 | | | Neosaproecius nyassalandicus Branco, 1990 | | | Neosaproecius trituberculatus Frey, 1958 | | | Neosisyphus angulicollis Felsche, 1909 | | | Neosisyphus armatus Gory, 1833 | | | Neosisyphus astriatus Schafer & Fi..., 2001 | | | Neosisyphus baoule Cambefort, 1984 | | | Neosisyphus barbarossa Wiedemann, 1823 | | | Neosisyphus basilewskyi Haaf, 1956 | | | Neosisyphus bowringi White, 1844 | | | Neosisyphus calcaratus Klug, 1855 | | | Neosisyphus confrater Kolbe, 1914 | | | Neosisyphus fortuitus Péringuey, 1901 | | | Neosisyphus gladiator Arrow, 1927 | | | Neosisyphus infuscatus Klug, 1855 | | | Neosisyphus kuehni Haaf, 1955 | | | Neosisyphus macrorubrus Paschalidis, 1974 | | | Neosisyphus mirabilis Arrow, 1927 | | | Neosisyphus paschalidisae Cambefort, 1984 | | | Neosisyphus penicillatus Harold, 1880 | | | Neosisyphus quadricollis Gory, 1833 | | | Neosisyphus rubripes Péringuey, 1901 | | | Neosisyphus rubrus Paschalidis, 1974 | | | Neosisyphus rugosus Gory, 1833 | | | Neosisyphus setiger Roth, 1851 | | | Neosisyphus spinipes Thunberg, 1818 | Grey dungball roller | | Neosisyphus tai Cambefort, 1984 | | | Neosisyphus tarantula Arrow, 1909 | | | Neosisyphus tibialis Raffray, 1877 | | | Neosisyphus trochantericus Fairmaire, 1891 | | | Nesosisyphus pygmaeus Vinson, 1946 | | | Nesosisyphus regnardi Alluaud, 1898 | | | Nesosisyphus rotundatus Vinson, 1946 | | | Nesosisyphus vicinus Vinson, 1939 | | | Nesovinsonia vinsoni Paulian, 1939 | | | Nitiocellus collarti Janssens, 1939 | | | Nitiocellus panthera Boucomont, 1921 | | | Nunoidium argentinum Arrow, 1913 | | | Ochicanthon besucheti Cuccodoro, 2011 | | | Ochicanthon cambeforti Ochi & Kon &..., 1997 | | | Ochicanthon ceylonicus Cuccodoro, 2011 | | | Ochicanthon cingalensis Arrow, 1931 | | | Ochicanthon crockermontis Krikken & Hu..., 2007 | | | Ochicanthon crypticus Krikken & Hu..., 2007 | | | Ochicanthon danum Krikken & Hu..., 2007 | | | Ochicanthon deplanatus Paulian, 1983 | | | Ochicanthon devagiriensis Sabu & Latha, 2011 | | | Ochicanthon dulitmontis Krikken & Hu..., 2007 | | | Ochicanthon dytiscoides Boucomont, 1914 | | | Ochicanthon edmondsi Krikken & Hu..., 2007 | | | Ochicanthon ernei Cuccodoro, 2011 | | | Ochicanthon fernandoi Sabu & Latha, 2011 | | | Ochicanthon gangkui Ochi & Kon &..., 1997 | | | Ochicanthon gauricola Cuccodoro, 2011 | | | Ochicanthon hanskii Krikken & Hu..., 2007 | | | Ochicanthon hikidai Ochi & Kon &..., 1997 | | | Ochicanthon javanus Krikken & Hu..., 2007 | | | Ochicanthon karasuyamai Ochi, Kon & ..., 2007 | | | Ochicanthon kikutai Ochi, Ueda &..., 2006 | | | Ochicanthon kimanis Krikken & Hu..., 2007 | | | Ochicanthon laetus Arrow, 1931 | | | Ochicanthon loebli Paulian, 1980 | | | Ochicanthon maryatiae Ochi, Ueda &..., 2006 | | | Ochicanthon masumotoi Ochi & Araya, 1996 | | | Ochicanthon mulu Krikken & Hu..., 2007 | | | Ochicanthon murthyi Vinod & Sabu, 2011 | | | Ochicanthon mussardi Cuccodoro, 2011 | | | Ochicanthon neglectus Krikken & Hu..., 2007 | | | Ochicanthon niinoii Ochi & Kon, 2014 | | | Ochicanthon niisatoi Ochi, Kon & ..., 2017 | | | Ochicanthon nitidus Paulian, 1980 | | | Ochicanthon obscurus Boucomont, 1920 | | | Ochicanthon ochii Hanboonsong ..., 2001 | | | Ochicanthon oharai Ochi, Kon, H..., 2008 | | | Ochicanthon okudai Ochi & Kon, 2014 | | | Ochicanthon parantisae Ochi & Kon &..., 1997 | | | Ochicanthon peninsularis Krikken & Hu..., 2007 | | | Ochicanthon philippinense Ochi, 1990 | | | Ochicanthon punctatus Boucomont, 1914 | | | Ochicanthon ranongensis Ochi, Kon & ..., 2020 | | | Ochicanthon rombauti Krikken & Hu..., 2007 | | | Ochicanthon takakui Ochi, Kon, H..., 2008 | | | Ochicanthon takuapaensis Ochi, Kon & ..., 2020 | | | Ochicanthon tambunan Krikken & Hu..., 2007 | | | Ochicanthon thai Paulian, 1987 | | | Ochicanthon thailandicum Masumoto, 1989 | | | Ochicanthon tristis Arrow, 1931 | | | Ochicanthon tristoides Paulian, 1983 | | | Ochicanthon uedai Ochi, Kon, H..., 2008 | | | Ochicanthon vazdemelloi (Latha & Sabu, 2011) | ⇒ Ochicanthon fernandoi Sabu & Latha, 2011 | | Ochicanthon woroae Ochi, Ueda &..., 2006 | | | Odontoloma apiculum Howden & Sch..., 1987 | | | Odontoloma cambeforti Howden & Sch..., 1987 | | | Odontoloma dentinum Harold, 1868 | | | Odontoloma disalatum Howden & Sch..., 1987 | | | Odontoloma doubei Howden & Sch..., 1987 | | | Odontoloma endroedyi Howden & Sch..., 1987 | | | Odontoloma louwi Howden & Sch..., 1987 | | | Odontoloma metasternale d´Orbigny, 1902 | | | Odontoloma multipunctatum Howden & Sch..., 1987 | | | Odontoloma obscurum Howden & Sch..., 1987 | | | Odontoloma pauliani Walter, 1976 | | | Odontoloma pauxillum Boheman, 1857 | | | Odontoloma peckorum Howden & Sch..., 1987 | | | Odontoloma planatum Howden & Sch..., 1987 | | | Odontoloma pusillum Howden & Sch..., 1987 | | | Odontoloma pygidiale Péringuey, 1901 | | | Odontoloma quadridens d´Orbigny, 1908 | | | Odontoloma relictum Cambefort, 1983 | | | Odontoloma sculpturatum Harold, 1868 | | | Odontoloma spinicaudum Howden & Sch..., 1987 | | | Oficanthon mirabilis Paulian, 1985 | | | Oniticellus (Paroniticellus) festivus Steven, 1809 | | | Oniticellus ambiguus (Lacordaire, 1856) | ⇒ Cyptochirus trogiformis Roth, 1851 | | Oniticellus amplicollis ♰ (O. Heer, 1862) | ⇒ Helictopleurus neoamplicollis Krell, 2000 | | Oniticellus caelatus (Gerstaecker, 1871) | ⇒ Latodrepanus caelatus Gerstaecker, 1871 | | Oniticellus californicus (Horn, 1882) | ⇒ Liatongus californicus Horn, 1882 | | Oniticellus cinctus Fabricius, 1775 | Bordered dung beetle | | Oniticellus cubiensis (Castelnau, 1840) | ⇒ Euoniticellus cubiensis Castelnau, 1840 | | Oniticellus egregius Klug, 1855 | | | Oniticellus falsus (Sharp, 1875) | ⇒ Sinodrepanus falsus Sharp, 1875 | | Oniticellus formosus (Chevrolat, 1830) | ⇒ Oniticellus pictus Hausmann, 1807 | | Oniticellus freyi Balthasar, 1941 | | | Oniticellus fulvus (Goeze, 1777) | ⇒ Euoniticellus fulvus Goeze, 1777 | | Oniticellus gayeni Biswas & Cha..., 1985 | | | Oniticellus inaequalis (Reiche, 1847) | ⇒ Euoniticellus inaequalis Reiche, 1850 | | Oniticellus intermedius (Reiche, 1848) | ⇒ Euoniticellus intermedius Reiche, 1848 | | Oniticellus militaris (Castelnau, 1840) | ⇒ Liatongus militaris Castelnau, 1840 | | Oniticellus namdaphaensis Biswas & Cha..., 1985 | | | Oniticellus nasicornis (Reiche, 1847) | ⇒ Euoniticellus nasicornis Reiche, 1849 | | Oniticellus pallens (Castelnau, 1840) | ⇒ Euoniticellus intermedius Reiche, 1848 | | Oniticellus pallipes (Fairmaire, 1887) | ⇒ Euoniticellus inaequalis Reiche, 1850 | | Oniticellus pictus Hausmann, 1807 | | | Oniticellus planatus Castelnau, 1840 | | | Oniticellus pseudoplanatus Balthasar, 1964 | | | Oniticellus puberulus Y.-W. Zhang, 1988 | | | Oniticellus rhadamistus Fabricius, 1775 | | | Oniticellus rhinocerulus (Bates, 1889) | ⇒ Liatongus rhinocerulus Bates, 1889 | | Oniticellus spinipes (Roth, 1851) | ⇒ Tiniocellus spinipes Roth, 1851 | | Oniticellus subhendui Biswas & Cha..., 1985 | | | Oniticellus tessellatus Harold, 1879 | | | Oniticellus yohenai (Matsumura, 1934) | ⇒ Liatongus phanaeoides Westwood, 1839 | | Onitis abyssinicus (Reiche, 1847) | ⇒ Onitis lycophron Klug, 1855 | | Onitis adelphus Kolbe, 1914 | | | Onitis adriani Ferreira, 1976 | | | Onitis aeneomicans Gillet, 1918 | | | Onitis aeneus Lansberge, 1875 | | | Onitis aerarius Harold, 1878 | | | Onitis aeruginosus Klug, 1855 | | | Onitis aethiops Lesne, 1906 | | | Onitis affinis Felsche, 1911 | | | Onitis albertcollarti Génier & Mor..., 2019 | | | Onitis alexis (Klug, 1835) | ⇒ Cheironitis ungaricus ungaricus Herbst, 1789 | | Onitis alexis alexis | | | Onitis alexis septentrionalis Balthasar, 1942 | | | Onitis androcles Janssens, 1937 | | | Onitis anthracinus Felsche, 1907 | | | Onitis archettii Janssens, 1943 | | | Onitis artuosus Gillet, 1909 | | | Onitis assamensis Biswas, 1980 | | | Onitis autumnalis Davis, 1986 | | | Onitis aygulus Fabricius, 1781 | | | Onitis bayanga Moretto, 2007 | | | Onitis belial Fabricius, 1798 | | | Onitis bilobatus Ferreira, 1976 | | | Onitis bispicticollis Krikken, 1977 | | | Onitis bocandei Gillet, 1911 | | | Onitis bordati Cambefort, 1988 | | | Onitis brahma Lansberge, 1875 | | | Onitis bredoi Janssens, 1949 | | | Onitis brevidens Bai, Yang & ..., 2006 | | | Onitis brucemillerorum Josso & Mont..., 2017 | | | Onitis caffer Boheman, 1857 | | | Onitis cambeforti Moretto, 2007 | | | Onitis camerunus (Gillet, 1933) | ⇒ Onitis cryptodus Gillet, 1918 | | Onitis camiadei Josso, 2012 | | | Onitis camiadeianus Josso & Mont..., 2017 | | | Onitis castaneus Redtenbacher, 1848 | | | Onitis cerrutii Müller, 1949 | | | Onitis chalceus Lansberge, 1875 | | | Onitis confusus Boheman, 1860 | | | Onitis consanguineus Felsche, 1911 | | | Onitis corydon Boisduval, 1835 | | | Onitis coxalis Gillet, 1918 | | | Onitis crassus Sharp, 1875 | | | Onitis crenatus Reiche, 1847 | | | Onitis cribratus Lansberge, 1875 | | | Onitis cryptodus Gillet, 1918 | | | Onitis cupreus Castelnau, 1840 | | | Onitis curvipes Lansberge, 1875 | | | Onitis damoetas Steven, 1806 | | | Onitis deceptor Péringuey, 1901 | | | Onitis denticoxa Müller, 1941 | | | Onitis dimidiatus Felsche, 1907 | | | Onitis dispar Péringuey, 1896 | | | Onitis endroedii Ferreira, 1976 | | | Onitis excavaticollis Balthasar, 1942 | | | Onitis excavatus Arrow, 1931 | | | Onitis ezechias Reiche, 1856 | | | Onitis fabricii Roth, 1851 | | | Onitis falcatus Wulfen, 1786 | | | Onitis feae Felsche, 1907 | | | Onitis femoralis Josso, 2012 | | | Onitis fractipes Boucomont, 1923 | | | Onitis fulgidus (Klug, 1855) | ⇒ Pleuronitis fulgidus Castelnau, 1840 | | Onitis fulmineus Janssens, 1943 | | | Onitis gazanus Arrow, 1909 | | | Onitis gilleti Boucomont, 1933 | | | Onitis granicollis Felsche, 1907 | | | Onitis guineensis Gillet, 1918 | | | Onitis hageni Lansberge, 1886 | | | Onitis humerosus Pallas, 1771 | | | Onitis inflaticollis Felsche, 1907 | | | Onitis insuetus (Kolbe, 1914) | ⇒ Onitis sulcipennis Felsche, 1907 | | Onitis intermedius Frivaldsky, 1892 | | | Onitis inversidens Lansberge, 1875 | | | Onitis ion Olivier, 1789 | | | Onitis janssenii Gomes Alves, 1954 | | | Onitis jeanneli Janssens, 1942 | | | Onitis jossoi Moretto, 2010 | | | Onitis kasanka Josso, 2012 | | | Onitis keniensis Gillet, 1933 | | | Onitis kingstoni Krikken, 1977 | | | Onitis lama Lansberge, 1875 | | | Onitis laminosus Ferreira, 1976 | | | Onitis lamnifer Gillet, 1909 | | | Onitis lansbergei Sharp, 1875 | | | Onitis laticollis Lansberge, 1875 | | | Onitis licitus Péringuey, 1901 | | | Onitis lobi Cambefort, 1984 | | | Onitis lobipes Felsche, 1907 | | | Onitis lognia Moretto, 2007 | | | Onitis longitibialis Ferreira, 1977 | | | Onitis ludekingi Lansberge, 1875 | | | Onitis lycophron Klug, 1855 | | | Onitis magus ♰ C. von Heyden, 1862 | | | Onitis malleatus Janssens, 1937 | | | Onitis marshalli Janssens, 1939 | | | Onitis mendax Gillet, 1918 | | | Onitis menieri Cambefort, 1981 | | | Onitis meruensis Ferreira, 1977 | | | Onitis meyeri Kolbe, 1891 | | | Onitis miesseni Gusakov & Kl..., 2009 | | | Onitis minutus Lansberge, 1875 | | | Onitis mniszechi Lansberge, 1875 | | | Onitis mniszechianus Janssens, 1951 | | | Onitis mnizechianus | | | Onitis monstrosus Felsche, 1910 | | | Onitis montreuili Josso, 2012 | | | Onitis mossambicensis Ferreira, 1976 | | | Onitis mraci Moretto, 2021 | | | Onitis multidentatus Gillet, 1918 | | | Onitis naviauxi Cambefort, 1988 | | | Onitis nemoralis Gillet, 1932 | | | Onitis niger (Lansberge, 1875) | ⇒ Liatongus femoratus Illiger, 1800 | | Onitis nigeriensis Ferreira, 1977 | | | Onitis nubiensis Lansberge, 1875 | | | Onitis numida Castelnau, 1840 | | | Onitis numida marginicollis Balthasar, 1928 | | | Onitis numida numida | | | Onitis obenbergeri Balthasar, 1942 | | | Onitis obscurus Lansberge, 1886 | | | Onitis occidentalis Gillet, 1933 | | | Onitis orthopus Lansberge, 1875 | | | Onitis overlaeti Janssens, 1938 | | | Onitis paraconfusus Ferreira, 1976 | | | Onitis paradoxus (Boheman, 1857) | ⇒ Tropidonitis paradoxus Boheman, 1857 | | Onitis parainflaticollis Ferreira, 1977 | | | Onitis paramniszechi Ferreira, 1976 | | | Onitis parvulus Balthasar, 1963 | | | Onitis pecuarius Lansberge, 1875 | | | Onitis perbrincki Ferreira, 1958 | | | Onitis perpunctatus Balthasar, 1963 | | | Onitis pers Gusakov & Kl..., 2009 | | | Onitis perturbator Péringuey, 1901 | | | Onitis phartopus Lansberge, 1875 | | | Onitis philemon (Fabricius, 1801) | ⇒ Onitis subopacus Arrow, 1931 | | Onitis picticollis Boheman, 1857 | | | Onitis pinheyi Ferreira, 1977 | | | Onitis podicinus Boucomont, 1932 | | | Onitis politus Lansberge, 1875 | | | Onitis popei Ferreira, 1977 | | | Onitis proximus Ferreira, 1977 | | | Onitis pseudojansenii Krikken, 1977 | | | Onitis pseudoorthopus Ferreira, 1976 | | | Onitis pseudosetosus Ferreira, 1976 | | | Onitis pumilio Balthasar, 1963 | | | Onitis punctatostriatus Janssens, 1937 | | | Onitis pyramus Gillet, 1933 | | | Onitis reichei Lansberge, 1875 | | | Onitis retrodentatus Gillet, 1932 | | | Onitis ringenbachi Keith, 2007 | | | Onitis robustus Boheman, 1857 | | | Onitis rothi Lansberge, 1875 | | | Onitis senegalensis Lansberge, 1875 | | | Onitis setosus Lansberge, 1875 | | | Onitis shoensis (Reiche, 1847) | ⇒ Onitis vicinus Lansberge, 1875 | | Onitis sibutensis Janssens, 1943 | | | Onitis similis Felsche, 1911 | | | Onitis singhalensis Lansberge, 1875 | | | Onitis siva Gillet, 1911 | | | Onitis sphinx Fabricius, 1775 | | | Onitis spinicrus Fairmaire, 1893 | | | Onitis spinipes Drury, 1770 | | | Onitis subcrenatus Kolbe, 1897 | | | Onitis subopacus Arrow, 1931 | | | Onitis sulcipennis Felsche, 1907 | | | Onitis sulcipennis naromorus Krikken, 1977 | | | Onitis sulcipennis sulcipennis | | | Onitis tanzaniensis Ferreira, 1977 | | | Onitis thalassinus Lansberge, 1875 | | | Onitis thoracicus Ferreira, 1976 | | | Onitis tingaudi Keith & Moretto, 2007 | | | Onitis tortuosus Houston, 1983 | | | Onitis trochantericus Ferreira, 1976 | | | Onitis tuberculifrons Josso, 2012 | | | Onitis tumidus Gillet, 1918 | | | Onitis uncinatoides Janssens, 1951 | | | Onitis uncinatus Klug, 1855 | | | Onitis unguiculatus Olivier, 1789 | | | Onitis upembanus Janssens, 1951 | | | Onitis vanderkelleni Lansberge, 1886 | Dung beetle | | Onitis vicinus Lansberge, 1875 | | | Onitis violaceus (Lansberge, 1875) | ⇒ Onitis humerosus Pallas, 1771 | | Onitis virens Lansberge, 1875 | | | Onitis viridulus (Boheman, 1857) | ⇒ Onitis pecuarius Lansberge, 1875 | | Onitis vrydaghi Janssens, 1937 | | | Onitis westermanni Lansberge, 1886 | | | Onitis wittei Janssens, 1938 | | | Onitis zambiensis Josso, 2012 | | | Onoreidium bottimeri Howden & Young, 1981 | | | Onoreidium cristatum Arrow, 1931 | | | Onoreidium howdeni Ferreira & G..., 1993 | | | Onoreidium ohausi Arrow, 1931 | | | Ontherus (Caelontherus) mexicanus Harold, 1868 | | | Ontherus aequatorius Bates, 1891 | | | Ontherus alexis Blanchard, 1846 | | | Ontherus amplector Génier, 1996 | | | Ontherus androgynus Génier, 1996 | | | Ontherus aphodioides Burmeister, 1874 | | | Ontherus appendiculatus Mannerheim, 1829 | | | Ontherus ashei Génier, 1996 | | | Ontherus atlantidis Génier, 1996 | | | Ontherus azteca Harold, 1869 | | | Ontherus brevicollis Kirsch, 1870 | | | Ontherus brevipennis Harold, 1867 | | | Ontherus bridgesi Waterhouse, 1891 | | | Ontherus cambeforti Génier, 1996 | | | Ontherus carinicollis Luederwaldt, 1930 | | | Ontherus carinifrons Luederwaldt, 1930 | | | Ontherus cephalotes Harold, 1869 | | | Ontherus compressicornis Luederwaldt, 1931 | | | Ontherus contractus (Burmeister, 1874) | ⇒ Ontherus appendiculatus Mannerheim, 1829 | | Ontherus dentatus Luederwaldt, 1930 | | | Ontherus diabolicus Génier, 1996 | | | Ontherus didymus (Erichson, 1847) | ⇒ Ontherus alexis Blanchard, 1846 | | Ontherus digitatus Harold, 1868 | | | Ontherus edentulus Génier, 1996 | | | Ontherus elegans Luederwaldt, 1930 | | | Ontherus erosioides Luederwaldt, 1930 | | | Ontherus erosus Harold, 1875 | | | Ontherus felicitae González & M..., 2015 | | | Ontherus gilli Génier, 1996 | | | Ontherus gladiator Génier, 1998 | | | Ontherus grandis Luederwaldt, 1931 | | | Ontherus hadros Génier, 1996 | | | Ontherus howdeni Génier, 1996 | | | Ontherus incisus Kirsch, 1870 | | | Ontherus insolitus Génier, 1996 | | | Ontherus irinus Balthasar, 1938 | | | Ontherus kirschii Harold, 1867 | | | Ontherus laminifer Balthasar, 1938 | | | Ontherus lichyi Martínez, 1947 | | | Ontherus lobifrons Génier, 1996 | | | Ontherus lunicollis Génier, 1996 | | | Ontherus magnus Génier, 1996 | | | Ontherus monilistriatus Génier, 1996 | | | Ontherus obliquus Génier, 1996 | | | Ontherus pilatus Génier, 1996 | | | Ontherus planus Génier, 1996 | | | Ontherus podiceps Harold, 1868 | | | Ontherus politus Génier, 1996 | | | Ontherus pseudodidymus Génier, 1996 | | | Ontherus pubens Génier, 1996 | | | Ontherus raptor Génier, 1996 | | | Ontherus rectangulidens Génier, 1996 | | | Ontherus rectus Génier, 1996 | | | Ontherus sanctaemartae Génier, 1996 | | | Ontherus sextuberculatus Génier, 1996 | | | Ontherus stridulator Génier, 1996 | | | Ontherus strius (Howden & Young, 1981) | ⇒ Ontherus azteca Harold, 1869 | | Ontherus sulcator Fabricius, 1775 | | | Ontherus tenuistriatus Génier, 1996 | | | Ontherus trituberculatus Balthasar, 1938 | | | Ontherus ulcopygus Génier, 1996 | | | Ontherus virescens Lucas, 1859 | | | Ontherus zikani Luederwaldt, 1930 | | | Onthobium asperatum Fauvel, 1903 | | | Onthobium boucheti Paulian, 1986 | | | Onthobium caledonicus Paulian, 1935 | | | Onthobium cookii Paulian, 1986 | | | Onthobium gutierrezi Paulian, 1984 | | | Onthobium howdeni Paulian, 1984 | | | Onthobium kuscheli Paulian, 1984 | | | Onthobium lerati Paulian, 1935 | | | Onthobium macleayi Montrouzier, 1860 | | | Onthobium montheiti Paulian, 1986 | | | Onthobium montrouzieri Paulian, 1984 | | | Onthobium paniei Paulian, 1986 | | | Onthobium tibialoides Paulian, 1986 | | | Onthobium tillieri Paulian, 1986 | | | Onthocharis ataenioides (Martínez, 1952) | ⇒ Anomiopus birai Canhedo, 2006 | | Onthocharis juanae (Martínez, 1952) | ⇒ Anomiopus juanae Martínez, 1952 | | Onthocharis melancholica (Martínez, 1955) | ⇒ Anomiopus virescens Westwood, 1842 | | Onthocharis nigrocoerulea (Martínez, 1955) | ⇒ Anomiopus nigrocoeruleus Martínez, 1955 | | Onthocharis panamensis (Paulian, 1939) | ⇒ Anomiopus panamensis Paulian, 1939 | | Onthocharis wittmeri (Martínez, 1952) | ⇒ Anomiopus germari Harold, 1867 | | Onthophagus (Afrostrandius) adspersus d´Orbigny, 1908 | | | Onthophagus (Afrostrandius) bimetallicus d´Orbigny, 1907 | | | Onthophagus (Afrostrandius) demeyeri Moretto, 2008 | | | Onthophagus (Afrostrandius) granulipennis (Lansberge, 1886) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus (Afrostrandius) loxodontaphilus Moretto, 2008 | | | Onthophagus (Afrostrandius) plebejus Klug, 1855 | | | Onthophagus (Afrostrandius) pseudoplebejus Moretto, 2008 | | | Onthophagus (Afrostrandius) tigrinus (d´Orbigny, 1908) | ⇒ Onthophagus pygmaeus Schaller, 1783 | | Onthophagus tigrinus kageraensis Moretto, 2009 | | | Onthophagus tigrinus tigrinus | | | Onthophagus (Altonthophagus) cervenkai Kabakov, 2008 | | | Onthophagus (Altonthophagus) inelegans Balthasar, 1935 | | | Onthophagus (Altonthophagus) kozlovi Kabakov, 1990 | | | Onthophagus (Altonthophagus) kukunorensis Kabakov, 1990 | | | Onthophagus (Altonthophagus) marmotae Kabakov, 1990 | | | Onthophagus (Altonthophagus) sibiricus Harold, 1877 | | | Onthophagus (Altonthophagus) turpidoides Kabakov, 2008 | | | Onthophagus (Amphionthophagus) falzonii Goidanich, 1926 | | | Onthophagus (Amphionthophagus) melitaeus Fabricius, 1798 | | | Onthophagus (Colobonthophagus) aenescens Wiedemann, 1823 | | | Onthophagus (Colobonthophagus) agnus Gillet, 1925 | | | Onthophagus (Colobonthophagus) arunensis Scheuern, 1995 | | | Onthophagus (Colobonthophagus) caprai Frey, 1956 | | | Onthophagus (Colobonthophagus) dama Fabricius, 1798 | | | Onthophagus (Colobonthophagus) metalliceps Arrow, 1931 | | | Onthophagus (Colobonthophagus) neocolobus Scheuern, 1996 | | | Onthophagus (Colobonthophagus) occipitalis Lansberge, 1885 | | | Onthophagus (Colobonthophagus) paliceps Arrow, 1931 | | | Onthophagus (Colobonthophagus) piceorufulus Kabakov, 1994 | | | Onthophagus (Colobonthophagus) poggii Scheuern, 1996 | | | Onthophagus (Colobonthophagus) quadridentatus (Fabricius, 1798) | ⇒ Onthophagus fissicornis Stevens, 1809 | | Onthophagus (Colobonthophagus) shillongensis Scheuern, 1995 | | | Onthophagus (Colobonthophagus) thai Kabakov, 1994 | | | Onthophagus (Colobonthophagus) tragus Fabricius, 1792 | | | Onthophagus (Colobonthophagus) transquadridentatus Scheuern, 1995 | | | Onthophagus (Digitonthophagus) bonasus Fabricius, 1775 | Dung beetle | | Onthophagus (Digitonthophagus) gazella (Fabricius, 1787) | ⇒ Digitonthophagus gazella Fabricius, 1787 | | Onthophagus (Euonthophagus) amyntas (Olivier, 1789) | ⇒ Euonthophagus amyntas Olivier, 1789 | | Onthophagus (Euonthophagus) atramentarius Ménétriés, 1832 | | | Onthophagus (Euonthophagus) carbonarius Klug, 1855 | | | Onthophagus (Euonthophagus) consobrinus Péringuey, 1901 | | | Onthophagus (Euonthophagus) crocatus Mulsant & Go..., 1870 | | | Onthophagus (Furconthophagus) amicus Gillet, 1925 | | | Onthophagus (Furconthophagus) boucomonti Paulian, 1931 | | | Onthophagus (Furconthophagus) furcatus (Fabricius, 1781) | ⇒ Onthophagus furcaticeps Masters, 1886 | | Onthophagus (Furconthophagus) karenensis Masumoto, Oc..., 2008 | | | Onthophagus (Furconthophagus) khonkaenus Masumoto, Oc..., 2008 | | | Onthophagus (Furconthophagus) lilliputanus Lansberge, 1883 | | | Onthophagus (Furconthophagus) monforti Josso & Prév..., 2006 | | | Onthophagus (Furconthophagus) papulatorius Kabakov, 2008 | | | Onthophagus (Furconthophagus) papulatus Boucomont, 1914 | | | Onthophagus (Furconthophagus) promontorii Boucomont, 1924 | | | Onthophagus (Furconthophagus) schoolmeestersi Ochi & Kon, 2007 | | | Onthophagus (Gibbonthophagus) apicetinctus (d´Orbigny, 1900) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus (Gibbonthophagus) atripennis Waterhouse, 1875 | | | Onthophagus (Gibbonthophagus) chineicus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) denticollis Lansberge, 1883 | | | Onthophagus (Gibbonthophagus) duporti Boucomont, 1914 | | | Onthophagus (Gibbonthophagus) euryceros Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) fujiii Ochi & Kon, 1995 | | | Onthophagus (Gibbonthophagus) gracilipes Boucomont, 1914 | | | Onthophagus (Gibbonthophagus) hiabunicus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) kentingensis Nomura, 1973 | | | Onthophagus (Gibbonthophagus) kimioi Ochi, Kon & ..., 2011 | | | Onthophagus (Gibbonthophagus) lenis Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) limbatus Herbst, 1789 | | | Onthophagus (Gibbonthophagus) luridipennis Boheman, 1858 | | | Onthophagus (Gibbonthophagus) nigriobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) palawanicus Ochi & Kon, 2004 | | | Onthophagus (Gibbonthophagus) parviobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) penmani Masumoto, Oc..., 2002 | | | Onthophagus (Gibbonthophagus) privus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) rufiobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) sasajii Ochi & Kon, 2001 | | | Onthophagus (Gibbonthophagus) semipersonatus Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) sunantaae Masumoto, 1989 | | | Onthophagus (Gibbonthophagus) taurinus White, 1844 | | | Onthophagus (Gibbonthophagus) viduus (Harold, 1875) | ⇒ Onthophagus orbus Boucomont, 1924 | | Onthophagus (Gibbonthophagus) viridicervicapra Ochi, Kon & ..., 2009 | | | Onthophagus (Hyalonthophagus) hyalcyon Palestrini &..., 1989 | | | Onthophagus (Indachorius) arai Masumoto, 1989 | | | Onthophagus (Indachorius) baykanus Kabakov, 1994 | | | Onthophagus (Indachorius) bonengus Kabakov, 1994 | | | Onthophagus (Indachorius) chinensis Balthasar, 1952 | | | Onthophagus (Indachorius) danumensis (Ochi, Kon & ..., 2009) | ⇒ Onthophagus danumicus Huijbregts &..., 2012 | | Onthophagus (Indachorius) doisuthepensis Masumoto, 1989 | | | Onthophagus (Indachorius) hsui Masumoto, Ch..., 2004 | | | Onthophagus (Indachorius) koshunensis Balthasar, 1941 | | | Onthophagus (Indachorius) lannamiibun Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) maesaensis Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) masaoi Ochi, 1992 | | | Onthophagus (Indachorius) mendeli Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) mongkhoni Masumoto, Ha..., 2002 | | | Onthophagus (Indachorius) nongkaiensis Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) octonaevus Kabakov, 1994 | | | Onthophagus (Indachorius) paramasaoi Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) phetchabunensis Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) phuquoci Paulian, 1945 | | | Onthophagus (Indachorius) platypus Zhang, 1997 | | | Onthophagus (Indachorius) pseudoworoae Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) scotti Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) semidanumensis Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) subsapaensis Kabakov, 1994 | | | Onthophagus (Indachorius) suginoi Ochi, 1984 | | | Onthophagus (Indachorius) tiamicus Kabakov, 1994 | | | Onthophagus (Indachorius) tongbantumi Masumoto, Oc..., 2002 | | | Onthophagus (Macronthophagus) cludtsi Ochi, 2003 | | | Onthophagus (Macronthophagus) curvicarinatus Boucomont, 1914 | | | Onthophagus (Macronthophagus) diabolicus Harold, 1877 | | | Onthophagus (Macronthophagus) manipurensis Arrow, 1907 | | | Onthophagus (Macronthophagus) menieri Ochi, 2003 | | | Onthophagus (Macronthophagus) nilgirensis Gillet, 1922 | | | Onthophagus (Macronthophagus) rubricollis Hope, 1831 | | | Onthophagus (Macronthophagus) uenoi Ochi, 1995 | | | Onthophagus uenoi borneomontanus Ochi, 2003 | | | Onthophagus uenoi uenoi Ochi, 1995 | | | Onthophagus (Matashia) anguliceps (Boucomont, 1914) | ⇒ Onthophagus avocetta Arrow, 1933 | | Onthophagus (Matashia) yubarinus (Matsumura, 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus (Micronthophagus) falsivigilans Masumoto, 1995 | | | Onthophagus (Micronthophagus) gigantivigilans Masumoto, Ha..., 2002 | | | Onthophagus (Micronthophagus) konoi Matsumura, 1938 | | | Onthophagus (Micronthophagus) quaestus Sharp, 1875 | | | Onthophagus (Micronthophagus) vigilans Boucomont, 1921 | | | Onthophagus (Micronthophagus) wangi Masumoto, Ch..., 2004 | | | Onthophagus (Onthophagiellus) kondaoensis Kabakov, 1994 | | | Onthophagus (Onthophagiellus) opacifalculatus Ochi, Kon & ..., 2009 | | | Onthophagus (Onthophagiellus) parafalculatus Ochi, Kon & ..., 2009 | | | Onthophagus (Onthophagiellus) popovi Kabakov, 1983 | | | Onthophagus (Onthophagiellus) taiyaruensis Masumoto, 1977 | | | Onthophagus (Onthophagiellus) thanwaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagiellus) unguiculatus Kabakov, 1994 | | | Onthophagus (Onthophagus) abacus Boucomont, 1921 | | | Onthophagus (Onthophagus) aciculatulus Blatchley, 1928 | | | Onthophagus (Onthophagus) acuticollis (Gillet, 1927) | ⇒ Onthophagus amphioxus Arrow, 1933 | | Onthophagus (Onthophagus) acuticornis Endrödi, 1973 | | | Onthophagus (Onthophagus) afghanus Petrovitz, 1961 | | | Onthophagus (Onthophagus) alluvius Howden and C..., 1963 | | | Onthophagus (Onthophagus) alternans (Raffray, 1877) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus (Onthophagus) amirus Kabakov, 1982 | | | Onthophagus (Onthophagus) anguicorius Boucomont, 1914 | | | Onthophagus (Onthophagus) angustatus Boucomont, 1914 | | | Onthophagus (Onthophagus) apilularius Masumoto, 1995 | | | Onthophagus (Onthophagus) arnetti Howden and C..., 1963 | | | Onthophagus (Onthophagus) arnoldii Kabakov, 1982 | | | Onthophagus (Onthophagus) barbieri Paulian, 1978 | | | Onthophagus (Onthophagus) batesi Howden and C..., 1963 | | | Onthophagus (Onthophagus) bokiaunus Masumoto, 1995 | | | Onthophagus (Onthophagus) brevifrons (Horn, 1881) | ⇒ Onthophagus abyssinicus Gillet, 1925 | | Onthophagus (Onthophagus) browni Howden and C..., 1963 | | | Onthophagus (Onthophagus) canelasensis Howden and G..., 2004 | | | Onthophagus (Onthophagus) cartwrighti Howden, 1973 | | | Onthophagus (Onthophagus) cavernicollis Howden and C..., 1963 | | | Onthophagus (Onthophagus) chaiyaphumensis Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) chevrolati Harold, 1869 | | | Onthophagus chevrolati chevrolati Harold, 1869 | | | Onthophagus chevrolati longecarinatus Zunino & Hal..., 1988 | | | Onthophagus chevrolati omiltemius Bates, 1887 | | | Onthophagus chevrolati retusus Harold, 1869 | | | Onthophagus (Onthophagus) coahuilae Zunino and H..., 1988 | | | Onthophagus (Onthophagus) cochisus Brown, 1927 | | | Onthophagus (Onthophagus) concinnus (Laporte, 1840) | ⇒ Onthophagus striatulus Palisot de B..., 1809 | | Onthophagus (Onthophagus) coproides Horn, 1881 | | | Onthophagus (Onthophagus) cribricollis (Horn, 1881) | ⇒ Onthophagus rugosicollis Gillet, 1925 | | Onthophagus (Onthophagus) cuevensis Howden, 1973 | | | Onthophagus (Onthophagus) cyanellus Bates, 1887 | | | Onthophagus (Onthophagus) cynomysi Brown, 1927 | | | Onthophagus (Onthophagus) dapcauensis Boucomont, 1921 | | | Onthophagus (Onthophagus) depressus Harold, 1871 | Flat African dung beetle | | Onthophagus (Onthophagus) doiinthanonensis Masumoto, 1989 | | | Onthophagus (Onthophagus) doipuiensis Masumoto, 1989 | | | Onthophagus (Onthophagus) doitungensis Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) dubernardi Boucomont, 1914 | | | Onthophagus (Onthophagus) embersoni Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) fissicornis Stevens, 1809 | | | Onthophagus (Onthophagus) formosanus Gillet, 1924 | | | Onthophagus (Onthophagus) fuscus Boucomont, 1932 | | | Onthophagus fuscus canescens Zunino & Hal..., 1988 | | | Onthophagus fuscus mycetorum Zunino & Hal..., 1988 | | | Onthophagus fuscus parafuscus Zunino & Hal..., 2005 | | | Onthophagus (Onthophagus) gibbulus Pallas, 1781 | | | Onthophagus (Onthophagus) gorochovi Kabakov, 1998 | | | Onthophagus (Onthophagus) gosoli Masumoto, 1989 | | | Onthophagus (Onthophagus) grandivigilans Masumoto, 1995 | | | Onthophagus (Onthophagus) hajimei Masumoto, 1984 | | | Onthophagus (Onthophagus) hastifer Lansberge, 1885 | | | Onthophagus (Onthophagus) hayashii Masumoto, 1991 | | | Onthophagus (Onthophagus) hecate Panzer, 1794 | Scooped Scarab | | Onthophagus hecate blatchleyi Brown, 1929 | | | Onthophagus hecate hecate Panzer, 1794 | | | Onthophagus (Onthophagus) hidalgus Zunino and H..., 1988 | | | Onthophagus (Onthophagus) hoepfneri Harold, 1869 | | | Onthophagus (Onthophagus) horii Ochi, Kon & ..., 2009 | | | Onthophagus (Onthophagus) illyricus Scopoli, 1763 | | | Onthophagus (Onthophagus) incensus Say, 1835 | Dung beetle | | Onthophagus (Onthophagus) ishiii Ochi & Kon, 1995 | | | Onthophagus (Onthophagus) javaecola Balthasar, 1959 | | | Onthophagus (Onthophagus) kanyaayonus Masumoto, 1992 | | | Onthophagus (Onthophagus) khonmiinitnoi Masumoto, 1990 | | | Onthophagus (Onthophagus) kinhthaicus Kabakov, 1998 | | | Onthophagus (Onthophagus) kiuchii Masumoto, 1995 | | | Onthophagus (Onthophagus) knausi Brown, 1927 | | | Onthophagus (Onthophagus) knulli Howden and C..., 1963 | | | Onthophagus (Onthophagus) koryoensis Kim, 1985 | | | Onthophagus (Onthophagus) krakadaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagus) kuatunensis Balthasar, 1942 | | | Onthophagus (Onthophagus) lakyim Masumoto, 1990 | | | Onthophagus (Onthophagus) landolti (Harold, 1880) | ⇒ Onthophagus schaefferi Howden and C..., 1963 | | Onthophagus landolti landolti Harold, 1880 | | | Onthophagus landolti texanus Schaeffer, 1914 | | | Onthophagus (Onthophagus) lecontei (Harold, 1871) | ⇒ Onthophagus subopacus Robinson, 1940 | | Onthophagus (Onthophagus) lindaae Masumoto, 1989 | | | Onthophagus (Onthophagus) lutosopictus Fairmaire, 1897 | | | Onthophagus (Onthophagus) maculatus Fabricius, 1801 | | | Onthophagus (Onthophagus) maleengnaafon Masumoto, 1990 | | | Onthophagus (Onthophagus) maleengnoi Masumoto, 1990 | | | Onthophagus (Onthophagus) maniti Masumoto, 1989 | | | Onthophagus (Onthophagus) margaritifer d´Orbigny, 1898 | | | Onthophagus (Onthophagus) masumotoi Ochi, 1985 | | | Onthophagus (Onthophagus) matsudai Ochi & Kon, 2004 | | | Onthophagus (Onthophagus) mcclevei Howden and G..., 2004 | | | Onthophagus (Onthophagus) medorensis Brown, 1929 | | | Onthophagus (Onthophagus) mendicus Gillet, 1924 | | | Onthophagus (Onthophagus) mextexus (Howden and C..., 1970) | ⇒ Onthophagus durangoensis Balthasar, 1939 | | Onthophagus (Onthophagus) mulleri van Lansberge, 1883 | | | Onthophagus (Onthophagus) naaroon Masumoto, 1990 | | | Onthophagus (Onthophagus) namnaoensis Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) nasonis Kabakov, 1998 | | | Onthophagus (Onthophagus) nuchicornis Linnaeus, 1758 | | | Onthophagus (Onthophagus) nurestanicus Kabakov, 1982 | | | Onthophagus (Onthophagus) oklahomensis Brown, 1927 | Oklahoma dung beetle | | Onthophagus (Onthophagus) orientalis (Harold, 1868) | ⇒ Onthophagus fissinasus Fairmaire, 1895 | | Onthophagus (Onthophagus) orpheus Panzer, 1794 | | | Onthophagus orpheus canadensis Fabricius, 1801 | | | Onthophagus orpheus orpheus Panzer, 1794 | | | Onthophagus orpheus pseudorpheus Howden & Car..., 1963 | | | Onthophagus (Onthophagus) pennsylvanicus Harold, 1871 | | | Onthophagus (Onthophagus) phraoensis Masumoto, 1990 | | | Onthophagus (Onthophagus) phrutsaphaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagus) polyphemi Hubbard, 1894 | Gopher Tortoise Onthophagus Beetle | | Onthophagus polyphemi polyphemi Hubbard, 1894 | | | Onthophagus polyphemi sparsisetosus Howden & Car..., 1963 | | | Onthophagus (Onthophagus) proletarius Harold, 1875 | | | Onthophagus (Onthophagus) pseudoarmatus Balthasar, 1944 | | | Onthophagus (Onthophagus) pseudofuscus Zunino and H..., 1988 | | | Onthophagus (Onthophagus) pseudohystrix Masumoto, 1995 | | | Onthophagus (Onthophagus) punctator Reitter, 1893 | | | Onthophagus (Onthophagus) punneeae Masumoto, 1989 | | | Onthophagus (Onthophagus) ratchasimaensis Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) roubali Balthasar, 1935 | | | Onthophagus (Onthophagus) rufimanus Kabakov, 1982 | | | Onthophagus (Onthophagus) rugulosus Harold, 1886 | | | Onthophagus (Onthophagus) sakaeratensis Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) sakainoi Masumoto, 1991 | | | Onthophagus (Onthophagus) salvazai Paulian, 1978 | | | Onthophagus (Onthophagus) sauteri Gillet, 1924 | | | Onthophagus (Onthophagus) schaefferi Howden and C..., 1963 | | | Onthophagus (Onthophagus) seabrai Gomes Alves, 1944 | | | Onthophagus (Onthophagus) setchan Masumoto, 1984 | | | Onthophagus (Onthophagus) singhaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagus) sobrius Balthasar, 1959 | | | Onthophagus (Onthophagus) striatulus Palisot de B..., 1809 | Surf blenny | | Onthophagus striatulus floridanus Blatchley, 1928 | | | Onthophagus striatulus striatulus Palisot de B..., 1809 | | | Onthophagus (Onthophagus) subaeneus (Palisot de B..., 1811) | ⇒ Onthophagus striatulus Palisot de B..., 1809 | | Onthophagus (Onthophagus) subalternans d´Orbigny, 1902 | | | Onthophagus (Onthophagus) subopacus Robinson, 1940 | | | Onthophagus (Onthophagus) subtropicus Howden and C..., 1963 | | | Onthophagus (Onthophagus) taayai Masumoto, 1995 | | | Onthophagus (Onthophagus) tatsienluensis Balthasar, 1942 | | | Onthophagus (Onthophagus) taurus Schreber, 1759 | Bull Headed Dung Beetle | | Onthophagus (Onthophagus) terminatus Eschscholtz, 1822 | | | Onthophagus (Onthophagus) tholaayi Masumoto, 1990 | | | Onthophagus (Onthophagus) tritinctus Boucomont, 1914 | | | Onthophagus (Onthophagus) trituber Wiedemann, 1823 | | | Onthophagus trituber jacobsoni Kabakov, 2006 | | | Onthophagus (Onthophagus) truncaticornis (Schaller, 1783) | ⇒ Onthophagus quadrinodosus Fahraeus, 1857 | | Onthophagus (Onthophagus) tuberculifrons Harold, 1871 | Dung beetle | | Onthophagus (Onthophagus) tumami Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) velutinus Horn, 1875 | | | Onthophagus (Onthophagus) vitulus Péringuey, 1901 | | | Onthophagus (Onthophagus) wangnamkhieoensis Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) yamaokai Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) yangi Masumoto, Ts..., 2006 | | | Onthophagus (Palaeonthophagus) afghanus Petrovitz, 1961 | | | Onthophagus (Palaeonthophagus) albarracinus Baraud, 1979 | | | Onthophagus (Palaeonthophagus) angorensis Petrovitz, 1963 | | | Onthophagus (Palaeonthophagus) baraudi Nicolas, 1964 | | | Onthophagus (Palaeonthophagus) bytinskii Balthasar, 1960 | | | Onthophagus (Palaeonthophagus) clitellifer Reitter, 1894 | | | Onthophagus (Palaeonthophagus) coenobita Herbst, 1783 | | | Onthophagus (Palaeonthophagus) conspersus Reitter, 1893 | | | Onthophagus (Palaeonthophagus) cruciatus (Ménétriés, 1832) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus (Palaeonthophagus) dacatrai Pittino, 2004 | | | Onthophagus (Palaeonthophagus) dellacasai Pittino & Ma..., 1981 | | | Onthophagus (Palaeonthophagus) eulaminicornis Pittino, 2006 | | | Onthophagus (Palaeonthophagus) excisus Reiche & Saulcy, 1856 | | | Onthophagus (Palaeonthophagus) excubitor Ziani & Gudenzi, 2006 | | | Onthophagus (Palaeonthophagus) finschi Harold, 1877 | | | Onthophagus (Palaeonthophagus) fissicornis Steven, 1809 | | | Onthophagus (Palaeonthophagus) fortigibber Reitter, 1909 | | | Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790) | ⇒ Onthophagus similis Scriba, 1790 | | Onthophagus (Palaeonthophagus) furciceps Marseul, 1869 | | | Onthophagus (Palaeonthophagus) gibbulus Pallas, 1781 | | | Onthophagus gibbulus gibbulus | | | Onthophagus gibbulus rostrifer Reitter, 1892 | | | Onthophagus (Palaeonthophagus) gorodinskii Kabakov, 2008 | | | Onthophagus (Palaeonthophagus) grossepunctatus Reitter, 1905 | | | Onthophagus (Palaeonthophagus) haroldi Ballion, 1870 | | | Onthophagus (Palaeonthophagus) hissariensis Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) isikdagensis Pittino, 2004 | | | Onthophagus (Palaeonthophagus) joannae Goljan, 1953 | | | Onthophagus (Palaeonthophagus) kindermanni Harold, 1877 | | | Onthophagus (Palaeonthophagus) kolenatii Reitter, 1893 | | | Onthophagus (Palaeonthophagus) laticornis Gebler, 1823 | | | Onthophagus (Palaeonthophagus) latigena d´Orbigny, 1897 | | | Onthophagus (Palaeonthophagus) lemur Fabricius, 1781 | | | Onthophagus (Palaeonthophagus) leucostigma Steven, 1811 | | | Onthophagus (Palaeonthophagus) lucidus Illiger, 1800 | | | Onthophagus (Palaeonthophagus) macedonicus Miksic, 1959 | | | Onthophagus (Palaeonthophagus) marginalis Gebler, 1817 | | | Onthophagus marginalis marginalis | | | Onthophagus marginalis przewalski Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) massai Baraud, 1975 | | | Onthophagus (Palaeonthophagus) medius (Kugelann, 1792) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus (Palaeonthophagus) medvedevi Kabakov, 1982 | | | Onthophagus (Palaeonthophagus) merdarius Chevrolat, 1865 | | | Onthophagus (Palaeonthophagus) muelleri (Lansberge, 1883) | ⇒ Onthophagus statzi Krell, 1990 | | Onthophagus (Palaeonthophagus) nikolajevi Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) novaki Boucomont, 1927 | | | Onthophagus (Palaeonthophagus) nuchicornis Linnaeus, 1758 | Mottled dung beetle | | Onthophagus (Palaeonthophagus) olsoufieffi Boucomont, 1924 | | | Onthophagus (Palaeonthophagus) opacicollis Reitter, 1893 | | | Onthophagus (Palaeonthophagus) ovatus Linnaeus, 1767 | | | Onthophagus (Palaeonthophagus) panici Petrovitz, 1964 | | | Onthophagus (Palaeonthophagus) parmatus Reitter, 1892 | | | Onthophagus (Palaeonthophagus) petrovitzi (Frey, 1975) | ⇒ Onthophagus petrovitzianus Pittino, 2006 | | Onthophagus (Palaeonthophagus) petrovitzianus Pittino, 2006 | | | Onthophagus (Palaeonthophagus) ponticus Harold, 1883 | | | Onthophagus (Palaeonthophagus) pseudocaccobius Reitter, 1889 | | | Onthophagus (Palaeonthophagus) psychopompus Ziani & Ghar..., 2010 | | | Onthophagus (Palaeonthophagus) pygargus Motschulsky, 1845 | | | Onthophagus (Palaeonthophagus) quadrinodus Reitter, 1896 | | | Onthophagus (Palaeonthophagus) rachelis Martín Piera, 1985 | | | Onthophagus (Palaeonthophagus) rakovici Pittino, 2004 | | | Onthophagus (Palaeonthophagus) rodentium Pittino, 2004 | | | Onthophagus (Palaeonthophagus) ruficapillus Brullé, 1832 | | | Onthophagus ruficapillus guilanensis (Pittino, 1982) | ⇒ Onthophagus ruficapillus Brullé, 1832 | | Onthophagus ruficapillus ruficapillus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) samai Ziani, 2011 | | | Onthophagus (Palaeonthophagus) scabriusculus Harold, 1873 | | | Onthophagus (Palaeonthophagus) semicornis Panzer, 1798 | | | Onthophagus (Palaeonthophagus) sericatus Reitter, 1893 | | | Onthophagus (Palaeonthophagus) shapovalovi Gusakov, 2012 | | | Onthophagus (Palaeonthophagus) similis (Scriba, 1790) | ⇒ Onthophagus fracticornis Preyssler, 1790 | | Onthophagus (Palaeonthophagus) speculifer Solsky, 1876 | | | Onthophagus (Palaeonthophagus) strabo Reitter, 1892 | | | Onthophagus (Palaeonthophagus) stylocerus Graells, 1851 | | | Onthophagus (Palaeonthophagus) suturellus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) tesquorum Semenov & Me..., 1927 | | | Onthophagus (Palaeonthophagus) trigibber Reitter, 1893 | | | Onthophagus (Palaeonthophagus) trispinus Reitter, 1892 | | | Onthophagus (Palaeonthophagus) truchmenus Kolenati, 1846 | | | Onthophagus truchmenus iranicus Kabakov, 2006 | | | Onthophagus truchmenus truchmenus | | | Onthophagus truchmenus turkmeniacus Kabakov, 2006 | | | Onthophagus truchmenus xerxes Petrovitz, 1965 | | | Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767 | | | Onthophagus (Palaeonthophagus) verticicornis (Laicharting, 1781) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus (Palaeonthophagus) viridis Ménétriés, 1832 | | | Onthophagus (Palaeonthophagus) vitulus (Fabricius, 1776) | ⇒ Onthophagus immundus Boheman, 1858 | | Onthophagus (Palaeonthophagus) vlasovi Medvedev, 1958 | | | Onthophagus (Palaeonthophagus) zuninoi Martín Piera, 1985 | | | Onthophagus (Paleonthophagus) andalusicus Waltl, 1835 | | | Onthophagus andalusicus andalusicus | | | Onthophagus andalusicus persianus Olsoufieff, 1900 | | | Onthophagus (Paleonthophagus) baraudi Nicolas, 1964 | | | Onthophagus (Paleonthophagus) coenobita Herbst, 1783 | | | Onthophagus (Paleonthophagus) fracticornis Preyssler, 1790 | | | Onthophagus (Paleonthophagus) gibbulus Pallas, 1781 | | | Onthophagus (Paleonthophagus) grossepunctatus Reitter, 1905 | | | Onthophagus (Paleonthophagus) joannae Goljan, 1953 | | | Onthophagus (Paleonthophagus) lemur Fabricius, 1781 | | | Onthophagus (Paleonthophagus) massai Baraud, 1975 | | | Onthophagus (Paleonthophagus) nuchicornis Linnaeus, 1758 | | | Onthophagus (Paleonthophagus) opacicollis Pierotti, 1959 | | | Onthophagus (Paleonthophagus) ovatus Linnaeus, 1767 | | | Onthophagus (Paleonthophagus) ruficapillus Brullé, 1832 | | | Onthophagus (Paleonthophagus) semicornis Panzer, 1798 | | | Onthophagus (Paleonthophagus) similis Scriba, 1790 | | | Onthophagus (Paleonthophagus) vacca Linnaeus, 1767 | | | Onthophagus (Paleonthophagus) verticicornis Laicharting, 1781 | | | Onthophagus (Paleonthophagus) vitulus Fabricius, 1776 | | | Onthophagus (Paraphanaeomorphus) argyropygus Gillet, 1927 | | | Onthophagus (Paraphanaeomorphus) carinensis Boucomont, 1914 | | | Onthophagus (Paraphanaeomorphus) comottoides Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) insignicollis Frey, 1954 | | | Onthophagus (Paraphanaeomorphus) janushevi Kabakov, 1983 | | | Onthophagus (Paraphanaeomorphus) napolovi Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) pseudojavanus Paulian, 1931 | | | Onthophagus (Paraphanaeomorphus) quadricolor Kabakov, 1983 | | | Onthophagus (Paraphanaeomorphus) semipiceus Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) strnadi Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) vaulogeri Boucomont, 1923 | | | Onthophagus (Parascatonomus) anceyi Boucomont, 1921 | | | Onthophagus (Parascatonomus) damaki Masumoto, Ha..., 2002 | | | Onthophagus (Parascatonomus) kiuchianus Masumoto, Ya..., 2004 | | | Onthophagus (Parascatonomus) koichii (Ochi & Araya, 1994) | ⇒ Caccobius jessoensis Harold, 1867 | | Onthophagus (Parascatonomus) leytensis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) miyakei Ochi & Araya, 1992 | | | Onthophagus (Parascatonomus) murasakianus Nomura, 1976 | | | Onthophagus (Parascatonomus) negrosensis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) politus Fabricius, 1798 | | | Onthophagus (Parascatonomus) pseudorudis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) quezonensis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) satoi Ochi, Shimad..., 2003 | | | Onthophagus (Parascatonomus) shizumui Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) takedai Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) takeshii Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) tricornis (Wiedemann, 1823) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus (Parascatonomus) yukae Masumoto, Oc..., 2002 | | | Onthophagus (Parentius) atricapillus d´Orbigny, 1908 | | | Onthophagus (Parentius) emarginatus Mulsant, 1842 | | | Onthophagus (Parentius) hispanicus Baraud, 1963 | | | Onthophagus (Parentius) kabakovi Martin-Piera, 1985 | | | Onthophagus (Parentius) punctatus Illiger, 1803 | | | Onthophagus (Paronthophagus) hidakai Ochi & Kon, 1995 | | | Onthophagus (Phalops) tricuspis (Semenov, 1900) | ⇒ Onthophagus fortigibber Reitter, 1909 | | Onthophagus (Phanaeomorphus) ater Waterhouse, 1875 | | | Onthophagus (Phanaeomorphus) cernyi (Balthasar, 1935) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus (Phanaeomorphus) johkii Ochi & Kon, 1994 | | | Onthophagus (Phanaeomorphus) lenzi Harold, 1875 | | | Onthophagus potanini hiurai Ochi, 1984 | | | Onthophagus (Phanaeomorphus) zetteli Masumoto, Oc..., 2002 | | | Onthophagus (Proagoderus) alcedo d´Orbigny, 1913 | | | Onthophagus (Proagoderus) alcyon (Klug, 1855) | ⇒ Onthophagus alcedo d´Orbigny, 1913 | | Onthophagus (Proagoderus) alcyonides d´Orbigny, 1913 | | | Onthophagus (Proagoderus) aureiceps d´Orbigny, 1902 | | | Onthophagus (Proagoderus) brucei Reiche, 1847 | | | Onthophagus brucei cyanopterus (d´Orbigny, 1905) | ⇒ Proagoderus cyanopterus d´Orbigny, 1905 | | Onthophagus (Proagoderus) dives Harold, 1877 | | | Onthophagus (Proagoderus) fossidorsis d´Orbigny, 1905 | | | Onthophagus (Proagoderus) mixtifrons d´Orbigny, 1910 | | | Onthophagus (Proagoderus) pegesimallus Marcus, 1917 | | | Onthophagus (Proagoderus) pseudoalcyon d´Orbigny, 1913 | | | Onthophagus (Proagoderus) pseudovirens d´Orbigny, 1913 | | | Onthophagus (Proagoderus) viridiceps d´Orbigny, 1910 | | | Onthophagus (Proagoderus) vividus Arrow, 1907 | | | Onthophagus (Pseudophanaeomorphus) parachandrai Ochi, Kon & ..., 2009 | | | Onthophagus (Relictonthophagus) emarginatus (Mulsant, 1842) | ⇒ Onthophagus rubrimaculatus MacLeay, 1864 | | Onthophagus (Relictonthophagus) nigellus (Illiger, 1803) | ⇒ Caccobius histeroides Ménétriés, 1832 | | Onthophagus (Relictonthophagus) punctatus (Illiger, 1803) | ⇒ Onthophagus emarginatus Mulsant, 1842 | | Onthophagus (Serrophorus) avocettoides Kabakov, 1983 | | | Onthophagus (Serrophorus) laevis (Harold, 1880) | ⇒ Onthophagus laevissimus d´Orbigny, 1905 | | Onthophagus laevis asiaticus Boucomont, 1919 | | | Onthophagus laevis laevis | | | Onthophagus laevis lampromelas Fairmaire, 1891 | | | Onthophagus laevis stevensi Arrow, 1931 | | | Onthophagus (Serrophorus) oblongus Zhang, 1997 | | | Onthophagus (Serrophorus) rectecornutus Lansberge, 1883 | | | Onthophagus (Serrophorus) sagittarius Fabricius, 1775 | Sagittarius dung beetle | | Onthophagus (Sinonthophagus) nampatensis Tarasov & Ka..., 2010 | | | Onthophagus (Strandius) changshouensis Zhang, 1997 | | | Onthophagus (Strandius) japonicus Harold, 1875 | | | Onthophagus (Strandius) kuraruanus (Matsumura, 1938) | ⇒ Onthophagus lenzi Harold, 1875 | | Onthophagus (Strandius) obliquus Olivier, 1789 | | | Onthophagus obliquus shimba (Cambefort, 1980) | ⇒ Onthophagus shimba Cambefort, 1980 | | Onthophagus (Strandius) potanini R. C. Brusca..., 1990 | | | Onthophagus potanini hiurai Ochi, 1984 | | | Onthophagus potanini potanini | | | Onthophagus (Strandius) subansiriensis Biswas, 1979 | | | Onthophagus (Strandius) taiwanus Nomura, 1973 | | | Onthophagus (Sunenaga) streltsovi Tarasov & Ka..., 2010 | | | Onthophagus (Trichonthophagus) dynastoides Arrow, 1931 | | | Onthophagus (Trichonthophagus) hirtus Illiger, 1803 | | | Onthophagus (Trichonthophagus) maki Illiger, 1803 | | | Onthophagus (Trichonthophagus) tarandus Fabricius, 1792 | | | Onthophagus abas Balthasar, 1946 | | | Onthophagus abei Ochi & Kon, 2015 | | | Onthophagus abeillei d´Orbigny, 1897 | | | Onthophagus abmisibilus Krikken & Hu..., 2012 | | | Onthophagus abreui Arrow, 1931 | | | Onthophagus abruptus d´Orbigny, 1913 | | | Onthophagus absyrtus Balthasar, 1946 | | | Onthophagus abyssinicus Gillet, 1925 | | | Onthophagus academus Balthasar, 1946 | | | Onthophagus acernorus Moctezuma, H..., 2021 | | | Onthophagus aceroides Krikken & Hu..., 2013 | | | Onthophagus acerus Gillet, 1930 | | | Onthophagus aciculatus (Fahraeus, 1857) | ⇒ Onthophagus aciculatulus Blatchley, 1928 | | Onthophagus acrisius Balthasar, 1946 | | | Onthophagus acuminatus (Harold, 1880) | ⇒ Onthophagus embrikianus Paulian, 1936 | | Onthophagus acutus d´Orbigny, 1908 | | | Onthophagus adelaidae Hope, 1847 | | | Onthophagus adelphus Gillet, 1930 | | | Onthophagus admetus Balthasar, 1946 | | | Onthophagus adornatus d´Orbigny, 1904 | | | Onthophagus aegrotus Balthasar, 1967 | | | Onthophagus aemulus Gillet, 1930 | | | Onthophagus aeneoniger d´Orbigny, 1913 | | | Onthophagus aeneopiceus d´Orbigny, 1902 | | | Onthophagus aequatus Péringuey, 1901 | | | Onthophagus aequepubens d´Orbigny, 1905 | | | Onthophagus aequiperus Matsumura & ..., 1937 | | | Onthophagus aerarius Reitter, 1892 | | | Onthophagus aereidorsis d´Orbigny, 1902 | | | Onthophagus aeremicans d´Orbigny, 1904 | | | Onthophagus aereomaculatus Boucomont, 1914 | | | Onthophagus aereopictus Boucomont, 1914 | | | Onthophagus aereopictus aereopictus | | | Onthophagus aereopictus rubropictus Boucomont, 1924 | | | Onthophagus aerestriatus d´Orbigny, 1913 | | | Onthophagus aeruginosus Roth, 1851 | | | Onthophagus aesopus Lansberge, 1882 | | | Onthophagus aethiopicus d´Orbigny, 1902 | | | Onthophagus affinis Gillet, 1930 | | | Onthophagus africanus Lansberge, 1886 | | | Onthophagus agaricophilus Arrow, 1931 | | | Onthophagus agilis (Matsumura, 1938) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus aguatus | | | Onthophagus ahenicollis d´Orbigny, 1902 | | | Onthophagus ahenomicans d´Orbigny, 1902 | | | Onthophagus akamineae Moretto & Gé..., 2020 | | | Onthophagus akhaus Masumoto, Oc..., 2007 | | | Onthophagus akinini Koenig, 1889 | | | Onthophagus akirai (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus alaindrumonti Krikken & Hu..., 2012 | | | Onthophagus alainmonforti Josso, 2022 | | | Onthophagus albicomus d´Orbigny, 1908 | | | Onthophagus albicornis Palisot de B..., 1805 | | | Onthophagus albipennis Péringuey, 1908 | | | Onthophagus albipodex d´Orbigny, 1902 | | | Onthophagus aleppensis Redtenbacher, 1843 | | | Onthophagus aleppensis aleppensis | | | Onthophagus aleppensis hyrcanicus Kabakov, 1982 | | | Onthophagus alfuricus Huijbregts &..., 2012 | | | Onthophagus aliceae Josso, 2019 | | | Onthophagus alienus Frey, 1958 | | | Onthophagus allojavanus Huijbregts &..., 2011 | | | Onthophagus alluaudi d´Orbigny, 1902 | | | Onthophagus aloysiellus Zunino, 1977 | | | Onthophagus alquirta Matthews, 1972 | | | Onthophagus altidorsis d´Orbigny, 1905 | | | Onthophagus altilamina d´Orbigny, 1905 | | | Onthophagus altivagans Howden & Gén..., 2004 | | | Onthophagus amamiensis Nomura, 1965 | | | Onthophagus amamiensis amamiensis | | | Onthophagus amamiensis matsumurai Ochi, Kawaha..., 2019 | | | Onthophagus ambang Krikken & Hu..., 2011 | | | Onthophagus ambang ambang | | | Onthophagus ambang morowali Krikken & Hu..., 2011 | | | Onthophagus ambiguus (Péringuey, 1901) | ⇒ Mimonthophagus ambiguus Péringuey, 1901 | | Onthophagus ambiquus | | | Onthophagus amenthus Matsumura, 1937 | | | Onthophagus amoenus d´Orbigny, 1908 | | | Onthophagus amphicoma Boucomont, 1914 | | | Onthophagus amphinasus Arrow, 1931 | | | Onthophagus amphioxus Arrow, 1933 | | | Onthophagus amplipennis d´Orbigny, 1905 | | | Onthophagus amycoides Kabakov, 2014 | | | Onthophagus anatolicus Petrovitz, 1962 | | | Onthophagus anchommatus Lea, 1923 | | | Onthophagus andersoni Howden & Gill, 1987 | | | Onthophagus andonarensis Boucomont, 1914 | | | Onthophagus andreji Prokofiev, 2014 | | | Onthophagus andrewesi Arrow, 1931 | | | Onthophagus andrewsmithi Krikken & Hu..., 2012 | | | Onthophagus androgynus d´Orbigny, 1905 | | | Onthophagus anewtoni Howden & Gén..., 2004 | | | Onthophagus angkhanensis Masumoto, Oc..., 2013 | | | Onthophagus angolensis Paulian, 1937 | | | Onthophagus angularis d´Orbigny, 1908 | | | Onthophagus angulicornis (Mulsant, 1842) | ⇒ Onthophagus semicornis Panzer, 1798 | | Onthophagus anisocerus Erichson, 1842 | | | Onthophagus annoyeri Moretto, 2010 | | | Onthophagus annulopunctatus Krikken & Hu..., 2017 | | | Onthophagus anogeissii Cambefort, 1984 | | | Onthophagus anomalicollis Frey, 1973 | | | Onthophagus anomalipes d´Orbigny, 1902 | | | Onthophagus antecornutus Josso, 2019 | | | Onthophagus antennalis Frey, 1961 | | | Onthophagus anthracinus (Harold, 1873) | ⇒ Caccobius histeroides Ménétriés, 1832 | | Onthophagus antillarum Arrow, 1903 | | | Onthophagus antilocapra Balthasar, 1937 | | | Onthophagus antoinei Walter, 1990 | | | Onthophagus antoineianus Josso, 2022 | | | Onthophagus aphodioides Lansberge, 1883 | | | Onthophagus apiciosus d´Orbigny, 1902 | | | Onthophagus apicosus | | | Onthophagus apomontanus Ochi, Kon & ..., 2023 | | | Onthophagus apterus Matthews, 1972 | Wingless dung beetle | | Onthophagus apunneea Masumoto, Oc..., 2007 | | | Onthophagus arayai Ochi & Kon, 2007 | | | Onthophagus arboreus Arrow, 1931 | | | Onthophagus arcifer d´Orbigny, 1904 | | | Onthophagus ardoini Frey, 1971 | | | Onthophagus areolatus d´Orbigny, 1905 | | | Onthophagus aries Kirby, 1825 | | | Onthophagus arizonensis (Schaeffer, 1909) | ⇒ Onthophagus hoepfneri Harold, 1869 | | Onthophagus arkoola Storey & Weir, 1990 | | | Onthophagus armatus Blanchard, 1853 | | | Onthophagus arrilla Matthews, 1972 | | | Onthophagus arunachalensis Biswas & Cha..., 1985 | | | Onthophagus arvernus ♰ L. Piton, 1939 | | | Onthophagus aschenborni Frey, 1975 | | | Onthophagus ashanticola Frey, 1973 | | | Onthophagus asiaticus Endrödi, 1973 | | | Onthophagus asimilis Péringuey, 1901 | | | Onthophagus asper MacLeay, 1864 | | | Onthophagus asperatus d´Orbigny, 1905 | | | Onthophagus aspericeps d´Orbigny, 1908 | | | Onthophagus aspericollis Lansberge, 1885 | | | Onthophagus asperipennis d´Orbigny, 1902 | | | Onthophagus aspernatus d´Orbigny, 1907 | | | Onthophagus asperodorsatus Howden & Gill, 1993 | | | Onthophagus asperrimus d´Orbigny, 1902 | | | Onthophagus asperulus d´Orbigny, 1905 | | | Onthophagus astigma d´Orbigny, 1904 | | | Onthophagus athiensis d´Orbigny, 1913 | | | Onthophagus atricolor d´Orbigny, 1907 | | | Onthophagus atridorsis d´Orbigny, 1902 | | | Onthophagus atriglabrus Howden & Gill, 1987 | | | Onthophagus atroaereus d´Orbigny, 1908 | | | Onthophagus atrofasciatus d´Orbigny, 1905 | | | Onthophagus atronitidus d´Orbigny, 1902 | | | Onthophagus atropolitus d´Orbigny, 1902 | | | Onthophagus atrosericeus Boucomont, 1932 | | | Onthophagus atrosplendens Frey, 1961 | | | Onthophagus atrostriatus d´Orbigny, 1913 | | | Onthophagus atrovirens d´Orbigny, 1905 | | | Onthophagus atrovittatus d´Orbigny, 1908 | | | Onthophagus atrox Harold, 1867 | | | Onthophagus aureofuscus Bates, 1887 | | | Onthophagus aureopilosus Boucomont, 1914 | | | Onthophagus auriculatus (Klug, 1855) | ⇒ Onthophagus latestriatus d´Orbigny, 1908 | | Onthophagus auritus Erichson, 1842 | | | Onthophagus australis Guérin-Ménev..., 1830 | Southern dung beetle | | Onthophagus avocetta Arrow, 1933 | | | Onthophagus axanicus Ochi, Kon & ..., 2021 | | | Onthophagus axillaris (Boheman, 1860) | ⇒ Onthophagus criniger d´Orbigny, 1904 | | Onthophagus aztecus Zunino & Hal..., 1988 | | | Onthophagus azusae Ochi & Kon, 2006 | | | Onthophagus babaulti d´Orbigny, 1915 | | | Onthophagus babirussa Eschscholtz, 1822 | | | Onthophagus baenzigeri Masumoto, Oc..., 2007 | | | Onthophagus baeri Boucomont, 1914 | | | Onthophagus baiyericus Krikken & Hu..., 2013 | | | Onthophagus bajacalifornianus Moctezuma & ..., 2019 | | | Onthophagus bakeri (Boucomont, 1919) | ⇒ Panelus bakeri Boucomont, 1921 | | Onthophagus bakweri Moretto, 2014 | | | Onthophagus balawaicus Scheuern, 1995 | | | Onthophagus baloghi Balthasar, 1967 | | | Onthophagus balthasari Vsetecka, 1939 | | | Onthophagus bambra Matthews, 1972 | | | Onthophagus bandamai Cambefort, 1984 | | | Onthophagus bangueyensis Boucomont, 1914 | | | Onthophagus baolocensis Ochi & Kon, 2015 | | | Onthophagus baoule Cambefort, 1984 | | | Onthophagus baramtagal Ochi, Kon & ..., 2016 | | | Onthophagus barretti Génier & How..., 1999 | | | Onthophagus barriorum Walter, 1990 | | | Onthophagus bartosi Balthasar, 1966 | | | Onthophagus basakata Walter & Cam..., 1977 | | | Onthophagus basicarinatus Rossini, Vaz..., 2018 | | | Onthophagus basicruentatus Goidanich, 1926 | | | Onthophagus basilewskyi Frey, 1961 | | | Onthophagus basipustulatus Heyden, 1879 | | | Onthophagus bassariscus Zunino & Hal..., 1988 | | | Onthophagus bateke Walter & Cam..., 1977 | | | Onthophagus batillifer Harold, 1875 | | | Onthophagus batui Huijbregts &..., 2009 | | | Onthophagus bayeri Balthasar, 1942 | | | Onthophagus bechynei Frey, 1953 | | | Onthophagus beelarong Storey & Weir, 1990 | | | Onthophagus beesoni Arrow, 1931 | | | Onthophagus begoniophilus Krikken & Hu..., 2017 | | | Onthophagus beiranus Péringuey, 1908 | | | Onthophagus belinga Walter, 1989 | | | Onthophagus bellus d´Orbigny, 1905 | | | Onthophagus belorhinus Bates, 1887 | | | Onthophagus benedictorum Walter & Cam..., 1977 | | | Onthophagus bengalensis Harold, 1886 | | | Onthophagus bengali Gordon & Opp..., 1977 | | | Onthophagus benguellianus Paulian, 1937 | | | Onthophagus bennigseni Paulian, 1937 | | | Onthophagus bequaerti d´Orbigny, 1915 | | | Onthophagus bergeri Frey, 1975 | | | Onthophagus bernaudi Cambefort & ..., 1991 | | | Onthophagus betschuanus Frey, 1975 | | | Onthophagus bicallifrons d´Orbigny, 1902 | | | Onthophagus bicallosus Klug, 1855 | | | Onthophagus bicarinaticeps Lea, 1923 | | | Onthophagus bicarinatus Gomes Alves, 1944 | | | Onthophagus bicavicollis Lea, 1923 | | | Onthophagus bicavifrons d´Orbigny, 1902 | | | Onthophagus bicolensis Ochi & Kon, 2006 | | | Onthophagus bicolensis asin Ochi & Kon, 2006 | | | Onthophagus bicolensis bicolensis | | | Onthophagus bicolor (Raffray, 1877) | ⇒ Onthophagus flavolimbatus Klug, 1855 | | Onthophagus biconifer d´Orbigny, 1905 | | | Onthophagus bicornis MacLeay, 1888 | | | Onthophagus bicristatus d´Orbigny, 1905 | | | Onthophagus bicristiger d´Orbigny, 1913 | | | Onthophagus bicuneus Kabakov, 2008 | | | Onthophagus bidens Olivier, 1789 | | | Onthophagus bidentatus (Drapiez, 1819) | ⇒ Caccobius schreberi Linnaeus, 1767 | | Onthophagus bidentifrons d´Orbigny, 1902 | | | Onthophagus biexcavatus (d´Orbigny, 1898) | ⇒ Onthophagus tarandus Fabricius, 1792 | | Onthophagus bifasciatus Fabricius, 1781 | | | Onthophagus bifidicornis d´Orbigny, 1902 | | | Onthophagus bifidus Reiche, 1849 | | | Onthophagus bifrons d´Orbigny, 1905 | | | Onthophagus bilingula Balthasar, 1965 | | | Onthophagus bimarginatus d´Orbigny, 1902 | | | Onthophagus bindaree Storey & Weir, 1990 | | | Onthophagus binodis Thunberg, 1818 | Humpbacked Dung Beetle | | Onthophagus binodosus d´Orbigny, 1908 | | | Onthophagus binodulus d´Orbigny, 1913 | | | Onthophagus binyana Storey & Weir, 1990 | | | Onthophagus biplagiatus Thomson, 1858 | | | Onthophagus birugatus d´Orbigny, 1902 | | | Onthophagus birugifer d´Orbigny, 1908 | | | Onthophagus bisbicornis d´Orbigny, 1909 | | | Onthophagus biscarinulatus Huijbregts &..., 2012 | | | Onthophagus bisectus Arrow, 1931 | | | Onthophagus bisignatus d´Orbigny, 1913 | | | Onthophagus bison Boucomont, 1919 | | | Onthophagus bisontinus ♰ Heer, 1862 | | | Onthophagus bisscrutator Krikken & Hu..., 2017 | | | Onthophagus bistiniocelloides Krikken, 1986 | | | Onthophagus bituber d´Orbigny, 1904 | | | Onthophagus bituberans d´Orbigny, 1905 | | | Onthophagus bituberculatus (Olivier, 1789) | ⇒ Onthophagus acutus d´Orbigny, 1908 | | Onthophagus bituberifrons Josso, 2018 | | | Onthophagus bituberoculus Krikken & Hu..., 2013 | | | Onthophagus bivertex Heyden, 1887 | | | Onthophagus blackburni Shipp, 1895 | | | Onthophagus blackwoodensis Blackburn, 1891 | | | Onthophagus blanchardi Harold, 1869 | | | Onthophagus blumei (Lansberge, 1883) | ⇒ Onthophagus avocetta Arrow, 1933 | | Onthophagus bocandei d´Orbigny, 1904 | | | Onthophagus bolivari Moctezuma, R..., 2016 | | | Onthophagus bomberaianus Balthasar, 1969 | | | Onthophagus bongkudai Krikken & Hu..., 2017 | | | Onthophagus bonorae Zunino, 1976 | | | Onthophagus bonsae Zunino, 1976 | | | Onthophagus boops d´Orbigny, 1905 | | | Onthophagus borassi Cambefort, 1984 | | | Onthophagus bordati Moretto, 2014 | | | Onthophagus borneensis Harold, 1877 | | | Onthophagus bornemisszai Matthews, 1972 | | | Onthophagus bornemisszanus Matthews, 1972 | | | Onthophagus borneotagal Ochi, Kon & ..., 2016 | | | Onthophagus boucomontianus Balthasar, 1935 | | | Onthophagus bourgognei Paulian, 1945 | | | Onthophagus bovimus | | | Onthophagus bovinus Péringuey, 1892 | | | Onthophagus brachypterus Zunino & Hal..., 1997 | | | Onthophagus brazzavillianus Balthasar, 1974 | | | Onthophagus breviceps d´Orbigny, 1902 | | | Onthophagus brevicollis Arrow, 1907 | | | Onthophagus breviconus Génier & How..., 1999 | | | Onthophagus brevigena d´Orbigny, 1902 | | | Onthophagus brevipennis d´Orbigny, 1902 | | | Onthophagus brevisetis d´Orbigny, 1915 | | | Onthophagus brittoni Paulian, 1948 | | | Onthophagus brivioi Frey, 1973 | | | Onthophagus bronzeus Arrow, 1907 | | | Onthophagus brooksi Matthews, 1972 | | | Onthophagus brunellii Müller, 1942 | | | Onthophagus brutus Arrow, 1931 | | | Onthophagus bubalus Harold, 1867 | | | Onthophagus buculus Mannerheim, 1829 | | | Onthophagus bufonidius Josso, 2019 | | | Onthophagus bufulus Arrow, 1941 | | | Onthophagus bulga Reid & Runag..., 2022 | | | Onthophagus bunamin Matthews, 1972 | | | Onthophagus bundara Storey & Weir, 1990 | | | Onthophagus burchelli d´Orbigny, 1908 | | | Onthophagus busiris Balthasar, 1969 | | | Onthophagus caelator Balthasar, 1967 | | | Onthophagus caesariatus Boucomont, 1921 | | | Onthophagus caffrarius d´Orbigny, 1902 | | | Onthophagus calamophilus Krikken, 1977 | | | Onthophagus calcaratus Boucomont, 1920 | | | Onthophagus calchas Balthasar, 1967 | | | Onthophagus calliger d´Orbigny, 1913 | | | Onthophagus callosipennis Boucomont, 1930 | | | Onthophagus cambrai Delgado & Curoe, 2014 | | | Onthophagus cameloides d´Orbigny, 1900 | | | Onthophagus camelus Fabricius, 1787 | | | Onthophagus camerunicus d´Orbigny, 1905 | | | Onthophagus camilleae Josso, 2022 | | | Onthophagus campestris Masumoto, Ki..., 2018 | | | Onthophagus cancer Lansberge, 1886 | | | Onthophagus canescens Zunino & Hal..., 1988 | | | Onthophagus capella (Kirby, 1818) | ⇒ Onthophagus ramosellus Bates, 1891 | | Onthophagus capelliformis Gillet, 1925 | | | Onthophagus capellinus Frey, 1963 | | | Onthophagus capitatus Castelnau, 1840 | | | Onthophagus capitosus Harold, 1867 | | | Onthophagus carayoni Cambefort, 1986 | | | Onthophagus carcharias Harold, 1875 | | | Onthophagus carinicollis Raffray, 1877 | | | Onthophagus carinidorsis d´Orbigny, 1908 | | | Onthophagus carinifer d´Orbigny, 1904 | | | Onthophagus carinulatus Harold, 1877 | | | Onthophagus carmodensis Blackburn, 1907 | | | Onthophagus carpanetoi Pittino, 1982 | | | Onthophagus carpophilus Pereira & Ha..., 1961 | | | Onthophagus castetsi Lansberge, 1887 | | | Onthophagus catenatus Lansberge, 1883 | | | Onthophagus catharinensis Paulian, 1936 | | | Onthophagus catherineae Josso, 2022 | | | Onthophagus catta Fabricius, 1787 | | | Onthophagus cavia Boucomont, 1914 | | | Onthophagus caviceps Frey, 1962 | | | Onthophagus cavifrons Harold, 1886 | | | Onthophagus cavivertex d´Orbigny, 1913 | | | Onthophagus centricornis Fabricius, 1798 | | | Onthophagus centurio Lansberge, 1885 | | | Onthophagus cephalophi Cambefort, 1984 | | | Onthophagus cervicapra Boucomont, 1914 | | | Onthophagus cervus (Fabricius, 1798) | ⇒ Onthophagus falsus Gillet, 1925 | | Onthophagus ceylonicus Harold, 1877 | | | Onthophagus chacoensis Rossini, Vaz..., 2018 | | | Onthophagus chalcostolus d´Orbigny, 1902 | | | Onthophagus championi Bates, 1887 | | | Onthophagus chandrai Ochi, 2007 | | | Onthophagus chebaicus Boucomont, 1919 | | | Onthophagus cheesmanae Arrow, 1941 | | | Onthophagus chepara Matthews, 1972 | | | Onthophagus chetroiensis Masumoto, Oc..., 2012 | | | Onthophagus cheyi Ochi & Kon, 2006 | | | Onthophagus chiangdaolekus Masumoto, Oc..., 2020 | | | Onthophagus chiapanecus Zunino & Hal..., 1988 | | | Onthophagus chibanus Matsumura & ..., 1937 | | | Onthophagus chilapensis Gasca-Ã�lvar..., 2018 | | | Onthophagus chimalapensis Delgado & Mo..., 2019 | | | Onthophagus chinantecus Moctezuma & ..., 2019 | | | Onthophagus chinbucephalus Ochi & Kon, 2018 | | | Onthophagus chinvigilans Ochi & Kon, 2018 | | | Onthophagus chirindaensis Josso, 2022 | | | Onthophagus chirindanus d´Orbigny, 1913 | | | Onthophagus chitipaensis Josso, 2019 | | | Onthophagus chloroderus d´Orbigny, 1905 | | | Onthophagus chlorophanus d´Orbigny, 1902 | | | Onthophagus choanicus d´Orbigny, 1904 | | | Onthophagus chokakurianus Matsumura, 1938 | | | Onthophagus chosensis Matsumura, 1937 | | | Onthophagus chremes Balthasar, 1969 | | | Onthophagus chryses Bates, 1887 | | | Onthophagus chrysoderus d´Orbigny, 1905 | | | Onthophagus chrysurus Arrow, 1931 | | | Onthophagus chumphonensis Masumoto, Oc..., 2008 | | | Onthophagus chuzenjianus (Matsumura & ..., 1937) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus cicer Balthasar, 1964 | | | Onthophagus cincticollis d´Orbigny, 1902 | | | Onthophagus cinctifrons d´Orbigny, 1902 | | | Onthophagus cinctipennis Quedenfeldt, 1884 | | | Onthophagus cineraceus d´Orbigny, 1902 | | | Onthophagus circulator Reitter, 1891 | | | Onthophagus circulifer Arrow, 1931 | | | Onthophagus circumdatus d´Orbigny, 1913 | | | Onthophagus citreum Boucomont, 1919 | | | Onthophagus civettae Cambefort, 1984 | | | Onthophagus civitasorum Moctezuma & ..., 2020 | | | Onthophagus clavijeroi Moctezuma, R..., 2016 | | | Onthophagus clavisetis d´Orbigny, 1905 | | | Onthophagus clermonti Paulian, 1931 | | | Onthophagus clitellarius d´Orbigny, 1908 | | | Onthophagus clivimerus Huijbregts &..., 2011 | | | Onthophagus clusifrons d´Orbigny, 1905 | | | Onthophagus clypealis Lea, 1923 | | | Onthophagus clypeatus Blanchard, 1846 | | | Onthophagus coeruleicollis Arrow, 1907 | | | Onthophagus cognatus Boucomont, 1921 | | | Onthophagus coiffaiti Frey, 1970 | | | Onthophagus colasi Paulian, 1937 | | | Onthophagus colffsi Lansberge, 1883 | | | Onthophagus collaris (Kabakov, 1983) | ⇒ Proagoderus loricatus Klug, 1855 | | Onthophagus collinsi Krikken & Hu..., 1987 | | | Onthophagus columbianus Boucomont, 1932 | | | Onthophagus comatulus d´Orbigny, 1905 | | | Onthophagus commottoides Kabakov, 1998 | | | Onthophagus comottoi Lansberge, 1885 | | | Onthophagus compactus Arrow, 1933 | | | Onthophagus comperei Blackburn, 1903 | Comperes dung beetle | | Onthophagus compositus Lea, 1923 | | | Onthophagus compressus Guérin-Ménev..., 1855 | | | Onthophagus concavifrons d´Orbigny, 1904 | | | Onthophagus concolor Sharp, 1878 | | | Onthophagus confertus Péringuey, 1908 | | | Onthophagus confluens (d´Orbigny, 1902) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus confusus Boucomont, 1932 | | | Onthophagus conradsi Balthasar, 1937 | | | Onthophagus consentaneus Harold, 1867 | Common northern dung beetle | | Onthophagus conspicuus MacLeay, 1864 | | | Onthophagus contiguicornis d´Orbigny, 1913 | | | Onthophagus contrapositus Génier, 2017 | | | Onthophagus convexicollis Boheman, 1858 | | | Onthophagus convexipennis Dierkens, 2022 | | | Onthophagus convexus d´Orbigny, 1908 | | | Onthophagus cooloola Storey & Weir, 1990 | | | Onthophagus coomani Paulian, 1931 | | | Onthophagus coorgensis Arrow, 1931 | | | Onthophagus coprimorphoides Huijbregts &..., 2012 | | | Onthophagus coprimorphus Gillet, 1930 | | | Onthophagus coptorhinodes Péringuey, 1901 | | | Onthophagus coptorhinoides Péringuey, 1901 | | | Onthophagus coracinoides Kabakov, 1983 | | | Onthophagus coracinus Boucomont, 1914 | | | Onthophagus coriaceoumbrosus Kohlmann & S..., 2001 | | | Onthophagus corniculatus (Reiche, 1847) | ⇒ Onthophagus senegalensis d´Orbigny, 1902 | | Onthophagus corniculiger d´Orbigny, 1913 | | | Onthophagus cornifrons Thomson, 1858 | | | Onthophagus cornutus Ferreira, 1971 | | | Onthophagus coronatus d´Orbigny, 1902 | | | Onthophagus corrosus Bates, 1887 | | | Onthophagus coscineus Bates, 1887 | | | Onthophagus costatus d´Orbigny, 1913 | | | Onthophagus costifer Gillet, 1930 | | | Onthophagus costiger d´Orbigny, 1915 | | | Onthophagus costilatus d´Orbigny, 1908 | | | Onthophagus costulicollis Frey, 1975 | | | Onthophagus crantor Balthasar, 1967 | | | Onthophagus crassicollis Boucomont, 1913 | | | Onthophagus crassus ♰ (Heer, 1862) | ⇒ Parascatonomus tricornis Wiedemann, 1823 | | Onthophagus cratippus Balthasar, 1969 | | | Onthophagus creber d´Orbigny, 1905 | | | Onthophagus cretus (Péringuey, 1901) | ⇒ Onthophagus setosus Fahraeus, 1857 | | Onthophagus cribellum d´Orbigny, 1902 | | | Onthophagus cribratus Lansberge, 1883 | | | Onthophagus cribripennis d´Orbigny, 1902 | | | Onthophagus criniger d´Orbigny, 1904 | | | Onthophagus crinitus (Harold, 1869) | ⇒ Onthophagus criniger d´Orbigny, 1904 | | Onthophagus crinitus crinitus | | | Onthophagus crinitus panamensis Bates, 1887 | | | Onthophagus cristatus d´Orbigny, 1905 | | | Onthophagus croesulus Bates, 1888 | | | Onthophagus crotchi Harold, 1871 | | | Onthophagus crucenotatus d´Orbigny, 1905 | | | Onthophagus cruciger MacLeay, 1888 | | | Onthophagus cruentatus Klug, 1855 | | | Onthophagus cryptodicranius Kohlmann & S..., 2001 | | | Onthophagus cryptogenus Boucomont, 1914 | | | Onthophagus cuboidalis (Bates, 1887) | ⇒ Onthophagus coproides Horn, 1881 | | Onthophagus cuneus Boucomont, 1924 | | | Onthophagus cuniculus MacLeay, 1864 | | | Onthophagus cupreiceps Arrow, 1907 | | | Onthophagus cupreopastillatus Ochi & Kon, 2006 | | | Onthophagus cupreopinguis Josso, 2022 | | | Onthophagus cupreotagal Ochi, Kon & ..., 2016 | | | Onthophagus cupreovirens d´Orbigny, 1904 | | | Onthophagus cupreus Harold, 1880 | | | Onthophagus cupricollis Péringuey, 1888 | | | Onthophagus curtipilis d´Orbigny, 1905 | | | Onthophagus curtulus Frey, 1961 | | | Onthophagus curvicornis Latreille, 1811 | | | Onthophagus curvifrons d´Orbigny, 1906 | | | Onthophagus curvilamina Müller, 1942 | | | Onthophagus curvispina Reitter, 1893 | | | Onthophagus cyaneiceps d´Orbigny, 1913 | | | Onthophagus cyaneoniger d´Orbigny, 1902 | | | Onthophagus cyaneus Kabakov, 2006 | | | Onthophagus cyanochlorus d´Orbigny, 1902 | | | Onthophagus cyclographus Bates, 1887 | | | Onthophagus cynomisi Brown, 1927 | | | Onthophagus dandalu Matthews, 1972 | | | Onthophagus danumicus Huijbregts &..., 2012 | | | Onthophagus dapitanensis Boucomont, 1919 | | | Onthophagus darlingtoni Matthews, 1972 | | | Onthophagus davisi Frey, 1975 | | | Onthophagus dayacus Boucomont, 1914 | | | Onthophagus daymanus Krikken & Hu..., 2013 | | | Onthophagus debilis d´Orbigny, 1905 | | | Onthophagus deccanensis Frey, 1962 | | | Onthophagus decedens Péringuey, 1904 | | | Onthophagus decedeus | | | Onthophagus decens Balthasar, 1964 | | | Onthophagus declivicollis d´Orbigny, 1902 | | | Onthophagus declivis Harold, 1869 | | | Onthophagus decolor d´Orbigny, 1902 | | | Onthophagus decoratus d´Orbigny, 1905 | | | Onthophagus decorsei d´Orbigny, 1908 | | | Onthophagus dedecor Wallengren, 1881 | | | Onthophagus deflexicollis Lansberge, 1883 | | | Onthophagus deflexus d´Orbigny, 1908 | | | Onthophagus delahayei Josso, 2011 | | | Onthophagus delicatulus Raffray, 1877 | | | Onthophagus delicatus d´Orbigny, 1915 | | | Onthophagus deliensis Lansberge, 1885 | | | Onthophagus deliensis bawangicus Ochi, Kon & ..., 2009 | | | Onthophagus deliensis deliensis | | | Onthophagus delphinensis d´Orbigny, 1913 | | | Onthophagus demarzi Frey, 1959 | | | Onthophagus densatus Frey, 1963 | | | Onthophagus densepunctatus Frey, 1963 | | | Onthophagus densipilis d´Orbigny, 1902 | | | Onthophagus denticornis Boucomont, 1914 | | | Onthophagus denticulatus d´Orbigny, 1902 | | | Onthophagus denudatus d´Orbigny, 1902 | | | Onthophagus denudatus denudatus | | | Onthophagus denudatus endroedii Frey, 1973 | | | Onthophagus depilatus d´Orbigny, 1913 | | | Onthophagus depilis d´Orbigny, 1902 | | | Onthophagus deplanatus Lansberge, 1883 | | | Onthophagus depressicollis Boucomont, 1914 | | | Onthophagus depressifrons d´Orbigny, 1915 | | | Onthophagus depressipennis Cambefort, 1975 | | | Onthophagus derasus d´Orbigny, 1902 | | | Onthophagus desaegeri Frey, 1961 | | | Onthophagus desectus MacLeay, 1871 | | | Onthophagus deterrens Péringuey, 1901 | | | Onthophagus devagiriensis Schoolmeeste..., 2006 | | | Onthophagus devexicornis Gillet, 1930 | | | Onthophagus devexus MacLeay, 1888 | | | Onthophagus dhanjuricus Frey, 1973 | | | Onthophagus diadematus d´Orbigny, 1902 | | | Onthophagus dicax Balthasar, 1966 | | | Onthophagus dicella Bates, 1888 | | | Onthophagus dicranius Bates, 1887 | | | Onthophagus dicranocerus Gillet, 1925 | | | Onthophagus dicranoides Balthasar, 1939 | | | Onthophagus difficilis (Le Guillou, 1844) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus digitatus Arrow, 1931 | | | Onthophagus digitifer (Boucomont, 1932) | ⇒ Onthophagus coscineus Bates, 1887 | | Onthophagus dignus Gillet, 1930 | | | Onthophagus dilutus d´Orbigny, 1905 | | | Onthophagus dinjeera Storey & Weir, 1990 | | | Onthophagus dinjerra Storey & Weir, 1990 | | | Onthophagus dinoderus d´Orbigny, 1913 | | | Onthophagus discolor (Hope, 1841) | ⇒ Onthophagus impurus Harold, 1868 | | Onthophagus discovirens d´Orbigny, 1902 | | | Onthophagus discretus Péringuey, 1901 | | | Onthophagus dispar Péringuey, 1901 | | | Onthophagus dissentaneus Balthasar, 1964 | | | Onthophagus dissidentatus Krikken & Hu..., 2013 | | | Onthophagus distichus Roth, 1851 | | | Onthophagus ditus (Péringuey, 1901) | ⇒ Onthophagus fungicola d´Orbigny, 1908 | | Onthophagus diversus Reiche, 1847 | | | Onthophagus dlabolai Balthasar, 1964 | | | Onthophagus dohertyi d´Orbigny, 1902 | | | Onthophagus doriae Harold, 1877 | | | Onthophagus dorsipilulus Howden & Gill, 1987 | | | Onthophagus dorsofasciatus Fairmaire, 1893 | | | Onthophagus dorsosignatus d´Orbigny, 1898 | | | Onthophagus dorsuosus d´Orbigny, 1902 | | | Onthophagus drescheri Paulian, 1939 | | | Onthophagus drumonti Moretto, 1999 | | | Onthophagus dryander Monteith & S..., 2013 | | | Onthophagus dubitabilis Howden & Gén..., 2004 | | | Onthophagus duboulayi Waterhouse, 1894 | | | Onthophagus ducorpsi d´Orbigny, 1913 | | | Onthophagus dudleyianus Josso, 2019 | | | Onthophagus dummal Matthews, 1972 | | | Onthophagus dunningi Harold, 1869 | | | Onthophagus durangoensis Balthasar, 1939 | | | Onthophagus duvivieri d´Orbigny, 1904 | | | Onthophagus ebenicolor d´Orbigny, 1902 | | | Onthophagus ebenus Péringuey, 1888 | | | Onthophagus eberus | | | Onthophagus eburneus d´Orbigny, 1913 | | | Onthophagus echinus Boucomont, 1914 | | | Onthophagus ecopas Josso & Prév..., 2006 | | | Onthophagus egenus (Harold, 1877) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus egurianus Matsumura, 1938 | | | Onthophagus elegans Klug, 1832 | | | Onthophagus eliptaminus Balthasar, 1969 | | | Onthophagus elongatus Frey, 1954 | | | Onthophagus embrikianus Paulian, 1936 | | | Onthophagus emeritus Péringuey, 1901 | | | Onthophagus endota Matthews, 1972 | | | Onthophagus endroedianus Walter & Cam..., 1977 | | | Onthophagus ensifer Boucomont, 1914 | | | Onthophagus enuguensis Frey, 1975 | | | Onthophagus ephippioderus Arrow, 1907 | | | Onthophagus epilamprus Bates, 1888 | | | Onthophagus erectinasus d´Orbigny, 1902 | | | Onthophagus erichsoni Hope, 1841 | | | Onthophagus escalerai d´Orbigny, 1902 | | | Onthophagus eschscholtzi Boucomont, 1924 | | | Onthophagus estelleae Josso, 2022 | | | Onthophagus etlaensis Kohlmann, Es..., 2019 | | | Onthophagus eulophus Bates, 1887 | | | Onthophagus euplynes Bates, 1888 | | | Onthophagus euzeti Walter & Cam..., 1977 | | | Onthophagus evae Balthasar, 1944 | | | Onthophagus evanidus Harold, 1869 | | | Onthophagus everestae ♰ Pierce, 1946 | | | Onthophagus exasperatus (Gerstaecker, 1871) | ⇒ Onthophagus latissimus d´Orbigny, 1902 | | Onthophagus excavatus (Redtenbacher, 1848) | ⇒ Liatongus phanaeoides Westwood, 1839 | | Onthophagus excisiceps d´Orbigny, 1902 | | | Onthophagus exiguus (Raffray, 1877) | ⇒ Onthophagus rubens d´Orbigny, 1902 | | Onthophagus exilis d´Orbigny, 1913 | | | Onthophagus expansicornis Bates, 1891 | | | Onthophagus exquisitus Arrow, 1931 | | | Onthophagus extensicollis d´Orbigny, 1907 | | | Onthophagus extensus Harold, 1878 | | | Onthophagus extraneus Josso, 2021 | | | Onthophagus fabricianus Goidanich, 1926 | | | Onthophagus fabricii Waterhouse, 1894 | | | Onthophagus falcarius Balthasar, 1966 | | | Onthophagus falcifer Harold, 1880 | | | Onthophagus falculatus Boucomont, 1914 | | | Onthophagus fallaciosus Raffray, 1877 | | | Onthophagus fallax d´Orbigny, 1913 | | | Onthophagus falsus Gillet, 1925 | | | Onthophagus fasciatus (Boucomont, 1914) | ⇒ Onthophagus fasciolatus Boucomont, 1924 | | Onthophagus fasciculiger d´Orbigny, 1902 | | | Onthophagus fasciolatus Boucomont, 1924 | | | Onthophagus favrei Boucomont, 1914 | | | Onthophagus feai d´Orbigny, 1905 | | | Onthophagus felix Arrow, 1931 | | | Onthophagus ferox Harold, 1867 | Western dung beetle | | Onthophagus ferrari Matthews, 1972 | | | Onthophagus filicornis Harold, 1873 | | | Onthophagus fimetarius Roth, 1851 | | | Onthophagus finchi | | | Onthophagus fissiceps MacLeay, 1888 | | | Onthophagus fissinasus Fairmaire, 1895 | | | Onthophagus fitiniensis Cambefort, 1984 | | | Onthophagus flagrans Reitter, 1893 | | | Onthophagus flavibasis d´Orbigny, 1904 | | | Onthophagus flaviclava d´Orbigny, 1902 | | | Onthophagus flavicornis Germar, 1824 | | | Onthophagus flavimargo d´Orbigny, 1902 | | | Onthophagus flavipennis d´Orbigny, 1905 | | | Onthophagus flavipleurus Dierkens, 2022 | | | Onthophagus flavoapicalis Lea, 1923 | | | Onthophagus flavocinctus Frey, 1975 | | | Onthophagus flavolimbatus Klug, 1855 | | | Onthophagus flavomaculatus Gillet, 1930 | | | Onthophagus flavorufus d´Orbigny, 1904 | | | Onthophagus fletcheri Blackburn, 1903 | | | Onthophagus flexicornis d´Orbigny, 1902 | | | Onthophagus flexifrons d´Orbigny, 1908 | | | Onthophagus fodiens Waterhouse, 1875 | | | Onthophagus foedus Boucomont, 1914 | | | Onthophagus foliaceus Lansberge, 1886 | Scarab beetle | | Onthophagus foliiceps Quedenfeldt, 1884 | | | Onthophagus foraminosus d´Orbigny, 1902 | | | Onthophagus formaneki Reitter, 1897 | | | Onthophagus fossibasis d´Orbigny, 1908 | | | Onthophagus fossifrons d´Orbigny, 1902 | | | Onthophagus fossor Arrow, 1931 | | | Onthophagus fossulatus d´Orbigny, 1908 | | | Onthophagus foulliouxi Cambefort, 1971 | | | Onthophagus foveatus Frey, 1957 | | | Onthophagus foveicollis Boucomont, 1930 | | | Onthophagus fradei Gomes Alves, 1956 | | | Onthophagus fragosus Génier & Ho..., 2014 | | | Onthophagus francoisgenieri Krikken & Hu..., 2012 | | | Onthophagus francoisi Josso, 2013 | | | Onthophagus frankenbergeri Balthasar, 1933 | | | Onthophagus frankrozendaali Huijbregts &..., 2012 | | | Onthophagus frenchi Blackburn, 1903 | | | Onthophagus fritschi d´Orbigny, 1902 | | | Onthophagus froggattellus Monteith & R..., 2017 | | | Onthophagus frontalis Raffray, 1877 | | | Onthophagus frugivorus Arrow, 1931 | | | Onthophagus fugitivus Péringuey, 1901 | | | Onthophagus fukuyamai Ochi, Kon & ..., 2018 | | | Onthophagus fuliginosus Erichson, 1842 | | | Onthophagus fulvocinctus d´Orbigny, 1902 | | | Onthophagus fulvus Sharp, 1875 | | | Onthophagus fumatus d´Orbigny, 1913 | | | Onthophagus funestus Moretto & Gé..., 2010 | | | Onthophagus fungicola d´Orbigny, 1908 | | | Onthophagus fungus Josso, 2022 | | | Onthophagus furcaticeps Masters, 1886 | | | Onthophagus furcatoides (Lansberge, 1886) | ⇒ Onthophagus neofurcatus Goidanich, 1926 | | Onthophagus furcicollis (Arrow, 1931) | ⇒ Onthophagus hecate Panzer, 1794 | | Onthophagus furcillifer Bates, 1891 | | | Onthophagus furculifer d´Orbigny, 1905 | | | Onthophagus furculus Fabricius, 1798 | | | Onthophagus fuscatus d´Orbigny, 1908 | | | Onthophagus fuscidorsis (d´Orbigny, 1902) | ⇒ Onthophagus fuscatus d´Orbigny, 1908 | | Onthophagus fuscivestis d´Orbigny, 1902 | | | Onthophagus fuscopunctatus (Fabricius, 1798) | ⇒ Onthophagus luridipennis Boheman, 1858 | | Onthophagus fuscostriatus Boucomont, 1914 | | | Onthophagus gabonensis Walter, 1989 | | | Onthophagus gaesatus Boucomont, 1925 | | | Onthophagus gagates Hope, 1831 | | | Onthophagus gagatinus (Gillet, 1930) | ⇒ Liatongus gagatinus Hope, 1831 | | Onthophagus gagatoides Kabakov, 2006 | | | Onthophagus gaillardi d´Orbigny, 1911 | | | Onthophagus gajo Krikken, 1977 | | | Onthophagus galeatus Boucomont, 1919 | | | Onthophagus ganalensis Gestro, 1895 | | | Onthophagus gandju Matthews, 1972 | | | Onthophagus gangeticus Gillet, 1925 | | | Onthophagus gangulu Matthews, 1972 | | | Onthophagus garambae Frey, 1961 | | | Onthophagus gazellinus Bates, 1887 | | | Onthophagus geelongensis Blackburn, 1891 | | | Onthophagus gemellatus Raffray, 1877 | | | Onthophagus geminatus d´Orbigny, 1907 | | | Onthophagus geminifrons d´Orbigny, 1905 | | | Onthophagus gemma Sharp, 1875 | | | Onthophagus genuinus Kohlmann & S..., 2001 | | | Onthophagus georgwerneri Josso, 2022 | | | Onthophagus germanus Gillet, 1927 | | | Onthophagus geryon Balthasar, 1969 | | | Onthophagus gestroi (Harold, 1877) | ⇒ Onthophagus bellus d´Orbigny, 1905 | | Onthophagus ghanensis Balthasar, 1966 | | | Onthophagus gibber d´Orbigny, 1905 | | | Onthophagus gibberosus d´Orbigny, 1905 | | | Onthophagus gibbicollis Lansberge, 1885 | | | Onthophagus gibbidorsis d´Orbigny, 1902 | | | Onthophagus gibbifrons d´Orbigny, 1902 | | | Onthophagus gibbus d´Orbigny, 1913 | | | Onthophagus gibsoni Howden & Gén..., 2004 | | | Onthophagus gidju Matthews, 1972 | | | Onthophagus gilli Delgado & Ho..., 2000 | | | Onthophagus ginyunensis Vsetecka, 1942 | | | Onthophagus giraffa Hausmann, 1807 | | | Onthophagus girardinae Walter, 1989 | | | Onthophagus giuseppecarpanetoi Tagliaferri ..., 2012 | | | Onthophagus glabratus Hope, 1842 | | | Onthophagus glasunovi Koshantschikov, 1894 | | | Onthophagus glasunowi D. Koshantsc..., 1894 | | | Onthophagus glaucinus Gillet, 1930 | | | Onthophagus gnu Frey, 1955 | | | Onthophagus godarra Storey & Weir, 1990 | | | Onthophagus gonipa Krikken & Hu..., 2011 | | | Onthophagus gonopygus d´Orbigny, 1902 | | | Onthophagus gonzaloi Moctezuma, H..., 2022 | | | Onthophagus gorokae Paulian, 1972 | | | Onthophagus gothicus Gillet, 1930 | | | Onthophagus gradivus Balthasar, 1966 | | | Onthophagus grandidorsis d´Orbigny, 1913 | | | Onthophagus grandifrons d´Orbigny, 1907 | | | Onthophagus graniceps d´Orbigny, 1908 | | | Onthophagus granosus d´Orbigny, 1913 | | | Onthophagus granulatus (Boheman, 1858) | ⇒ Onthophagus consentaneus Harold, 1867 | | Onthophagus granulifer (Harold, 1886) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus granulifrons Frey, 1961 | | | Onthophagus granulopygus Dierkens, 2022 | | | Onthophagus granulum d´Orbigny, 1904 | | | Onthophagus granum (Lansberge, 1885) | ⇒ Onthophagus granosus d´Orbigny, 1913 | | Onthophagus graphicus Wallengren, 1881 | | | Onthophagus grassei Walter, 1989 | | | Onthophagus grataehelenae Kohlmann & S..., 2001 | | | Onthophagus gratus Arrow, 1931 | | | Onthophagus gravearmatus Balthasar, 1944 | | | Onthophagus gravis Walker, 1858 | | | Onthophagus gravoti d´Orbigny, 1908 | | | Onthophagus griseoaeneus Lansberge, 1885 | | | Onthophagus griseosetosus Arrow, 1931 | | | Onthophagus guatemalensis (Bates, 1887) | ⇒ Onthophagus medorensis Brown, 1929 | | Onthophagus gulmarri Matthews, 1972 | | | Onthophagus gulo Arrow, 1931 | | | Onthophagus gurburra Storey & Weir, 1990 | | | Onthophagus guttatus (Boheman, 1860) | ⇒ Onthophagus quadrilunatus d´Orbigny, 1902 | | Onthophagus guttiger d´Orbigny, 1905 | | | Onthophagus haafi Frey, 1963 | | | Onthophagus haagi Harold, 1867 | | | Onthophagus haematopus Harold, 1875 | | | Onthophagus hageni Lansberge, 1883 | | | Onthophagus hagenmontis Krikken & Hu..., 2012 | | | Onthophagus halffteri Zunino, 1981 | | | Onthophagus hamaticeps Arrow, 1931 | | | Onthophagus hamaticornis Gillet, 1930 | | | Onthophagus hanboonsongae Masumoto, Oc..., 2021 | | | Onthophagus hartiniae Ochi & Kon, 2009 | | | Onthophagus helciatus Harold, 1871 | | | Onthophagus helmsi Blackburn, 1903 | | | Onthophagus hemichalceus d´Orbigny, 1915 | | | Onthophagus hemichlorus d´Orbigny, 1915 | | | Onthophagus hemipygus Frey, 1964 | | | Onthophagus henus | | | Onthophagus hericiniformis Moretto, 2010 | | | Onthophagus hericius d´Orbigny, 1902 | | | Onthophagus hermonensis Baraud, 1982 | | | Onthophagus herus Péringuey, 1901 | | | Onthophagus heteroclitus d´Orbigny, 1911 | | | Onthophagus heterorrhinus Lansberge, 1885 | | | Onthophagus heurni Gillet, 1930 | | | Onthophagus heydeni Harold, 1875 | | | Onthophagus heyrovskyi Vsetecka, 1943 | | | Onthophagus hielkemai Krikken & Hu..., 2011 | | | Onthophagus hikidai Ochi & Kon, 2006 | | | Onthophagus hikosanus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus hilaridis Cambefort, 1984 | | | Onthophagus hilaris d´Orbigny, 1905 | | | Onthophagus hildebrandti Harold, 1878 | | | Onthophagus hindu Arrow, 1931 | | | Onthophagus hingstoni Arrow, 1931 | | | Onthophagus hinnulus (Klug, 1832) | ⇒ Mimonthophagus hinnulus Klug, 1832 | | Onthophagus hippopotamus Harold, 1869 | | | Onthophagus hiranoyasutoshii Ochi & Kon, 2018 | | | Onthophagus hirculus (Mannerheim, 1829) | ⇒ Onthophagus mutatus Harold, 1859 | | Onthophagus hircus Billberg, 1815 | | | Onthophagus hiroyukii Ochi, 2007 | | | Onthophagus hirsutulus Lansberge, 1883 | | | Onthophagus hirsutus d´Orbigny, 1902 | | | Onthophagus hirtellus Frey, 1957 | | | Onthophagus hirticulus d´Orbigny, 1915 | | | Onthophagus hirtipodex d´Orbigny, 1904 | | | Onthophagus hirtuosus Gillet, 1930 | | | Onthophagus histeriformis Boucomont, 1914 | | | Onthophagus histrio d´Orbigny, 1902 | | | Onthophagus hoberlandti Balthasar, 1966 | | | Onthophagus hollowayi Krikken & Hu..., 2017 | | | Onthophagus holosericeus Harold, 1877 | | | Onthophagus holosericus Harold, 1877 | | | Onthophagus holzi Boucomont, 1914 | | | Onthophagus hoogstraali Saylor, 1943 | | | Onthophagus hopfneri | | | Onthophagus hoplocerus Lea, 1923 | | | Onthophagus hoplothorax Gillet, 1930 | | | Onthophagus horrens d´Orbigny, 1908 | | | Onthophagus horribilis Balthasar, 1969 | | | Onthophagus horridus d´Orbigny, 1908 | | | Onthophagus hosomai Ochi & Kon, 2014 | | | Onthophagus howdeni Zunino & Hal..., 1988 | | | Onthophagus howdenorum Zunino & Hal..., 1988 | | | Onthophagus hulstaerti Boucomont, 1932 | | | Onthophagus humboldti Kohlmann, So..., 2019 | | | Onthophagus humpatensis Lansberge, 1886 | | | Onthophagus hyaena Fabricius, 1801 | | | Onthophagus hystrix Boucomont, 1914 | | | Onthophagus ibex (Fabricius, 1792) | ⇒ Onthophagus tragus Fabricius, 1792 | | Onthophagus ibonus (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus ieti Cambefort, 1984 | | | Onthophagus ifugaoensis Ochi & Kon, 2006 | | | Onthophagus igneus Vigors, 1825 | | | Onthophagus igualensis Bates, 1887 | | | Onthophagus ikelengeensis Josso, 2018 | | | Onthophagus illotus Péringuey, 1901 | | | Onthophagus imbellis d´Orbigny, 1905 | | | Onthophagus imberbis d´Orbigny, 1902 | | | Onthophagus imbutus Sharp, 1875 | | | Onthophagus immundus Boheman, 1858 | | | Onthophagus imperator Castelnau, 1840 | | | Onthophagus importunus Péringuey, 1901 | | | Onthophagus impressicollis Boheman, 1860 | | | Onthophagus impunctatus d´Orbigny, 1904 | | | Onthophagus impurus Harold, 1868 | | | Onthophagus inaequalis d´Orbigny, 1902 | | | Onthophagus incantatus Balthasar, 1967 | | | Onthophagus incanus MacLeay, 1888 | | | Onthophagus incertus (d´Orbigny, 1897) | ⇒ Onthophagus asperrimus d´Orbigny, 1902 | | Onthophagus incisus Harold, 1877 | | | Onthophagus includens d´Orbigny, 1905 | | | Onthophagus inclusus d´Orbigny, 1905 | | | Onthophagus incollaris Kabakov, 2008 | | | Onthophagus incornutus MacLeay, 1871 | | | Onthophagus indigus Péringuey, 1901 | | | Onthophagus indosinicus Boucomont, 1921 | | | Onthophagus indutus d´Orbigny, 1902 | | | Onthophagus inecolorum Moctezuma & ..., 2020 | | | Onthophagus inediapterus Kohlmann & S..., 2001 | | | Onthophagus ineptus Harold, 1871 | | | Onthophagus inermiceps d´Orbigny, 1905 | | | Onthophagus inermicollis d´Orbigny, 1908 | | | Onthophagus inermivertex Boucomont, 1921 | | | Onthophagus infaustus Cambefort, 1984 | | | Onthophagus inflaticollis Bates, 1887 | | | Onthophagus inflatus Walter & Cam..., 1977 | | | Onthophagus infucatus Harold, 1877 | | | Onthophagus infuscatus Klug, 1845 | | | Onthophagus insignis (Péringuey, 1896) | ⇒ Proagoderus fossidorsis d´Orbigny, 1905 | | Onthophagus insularis Boheman, 1858 | | | Onthophagus insulindicus Boucomont, 1914 | | | Onthophagus insulsus Péringuey, 1901 | | | Onthophagus intermixtus d´Orbigny, 1902 | | | Onthophagus interruptus (Raffray, 1877) | ⇒ Onthophagus leanus Goidanich, 1926 | | Onthophagus interstitialis (Fahraeus, 1857) | ⇒ Euonthophagus carbonarius Klug, 1855 | | Onthophagus intonsus d´Orbigny, 1913 | | | Onthophagus intricatus Moretto, 2010 | | | Onthophagus investigator Lansberge, 1885 | | | Onthophagus investis d´Orbigny, 1904 | | | Onthophagus iodiellus Bates, 1887 | | | Onthophagus ioramaculatus Krikken & Hu..., 2012 | | | Onthophagus ioranus Krikken & Hu..., 2012 | | | Onthophagus irianus Balthasar, 1969 | | | Onthophagus iringaensis Dierkens, 2022 | | | Onthophagus iris Sharp, 1875 | | | Onthophagus isanus Masumoto, Oc..., 2008 | | | Onthophagus istmenus Moctezuma, S..., 2020 | | | Onthophagus itoi Nomura, 1976 | | | Onthophagus itoianus Masumoto, Oc..., 2022 | | | Onthophagus itonoborui Masumoto, Oc..., 2012 | | | Onthophagus iulicola Cambefort, 1984 | | | Onthophagus iumienus Masumoto, Oc..., 2007 | | | Onthophagus ixtepecorum Moctezuma & ..., 2020 | | | Onthophagus ixtlanensis Moctezuma & ..., 2020 | | | Onthophagus iyengari Arrow, 1931 | | | Onthophagus jacksoni d´Orbigny, 1913 | | | Onthophagus jacobeus Boucomont, 1924 | | | Onthophagus jalamari Matthews, 1972 | | | Onthophagus jalapensis Balthasar, 1939 | | | Onthophagus jaliscensis Zunino & Hal..., 1988 | | | Onthophagus jangga Matthews, 1972 | | | Onthophagus janssensi Frey, 1958 | | | Onthophagus janthinus (Harold, 1875) | ⇒ Onthophagus aeruginosus Roth, 1851 | | Onthophagus janus Panzer, 1794 | | | Onthophagus javanensis Balthasar, 1963 | | | Onthophagus javanoides Huijbregts &..., 2011 | | | Onthophagus jeanneli d´Orbigny, 1913 | | | Onthophagus jeanneli diversiceps Frey, 1975 | | | Onthophagus jeannelianus Paulian, 1945 | | | Onthophagus jeannelianus jeannelianus | | | Onthophagus jeannelianus shokhini Kabakov, 2014 | | | Onthophagus jedensis Matsumura, 1937 | | | Onthophagus jingping Masumoto, Oc..., 2007 | | | Onthophagus jiupengensis Masumoto, La..., 2017 | | | Onthophagus jocquei Josso, 2019 | | | Onthophagus joliveti Paulian, 1972 | | | Onthophagus jonathani Moretto & Gé..., 2020 | | | Onthophagus jubatus Harold, 1869 | | | Onthophagus jugicola d´Orbigny, 1902 | | | Onthophagus juheli Josso, 2022 | | | Onthophagus juncticornis d´Orbigny, 1908 | | | Onthophagus juniperensis Josso, 2019 | | | Onthophagus justei Walter, 1989 | | | Onthophagus juvencus Klug, 1835 | | | Onthophagus jwalae Karimbumkara..., 2016 | | | Onthophagus kachinicus Kabakov, 2006 | | | Onthophagus kaengkrachangus Masumoto, Oc..., 2008 | | | Onthophagus kakadu Storey & Weir, 1990 | | | Onthophagus kanarensis Arrow, 1931 | | | Onthophagus kandai (Matsumura, 1937) | ⇒ Onthophagus olsoufieffi Boucomont, 1924 | | Onthophagus kangeanus Paulian, 1936 | | | Onthophagus kaomaensis Josso, 2022 | | | Onthophagus kaosoidowensis Masumoto, Oc..., 2008 | | | Onthophagus kapangaensis Dierkens, 2022 | | | Onthophagus kapitensis Frey, 1971 | | | Onthophagus kapuri Endrödi, 1974 | | | Onthophagus karlwerneri Josso, 2022 | | | Onthophagus kashizakii Kon & Ochi, 2005 | | | Onthophagus kashmirensis Balthasar, 1966 | | | Onthophagus kassaicus d´Orbigny, 1908 | | | Onthophagus katangensis Frey, 1961 | | | Onthophagus katayamai Masumoto, Oc..., 2012 | | | Onthophagus kavirondi (d´Orbigny, 1913) | ⇒ Onthophagus kavirondus d´Orbigny, 1915 | | Onthophagus kavirondus d´Orbigny, 1915 | | | Onthophagus kawaharai Ochi & Kon, 2007 | | | Onthophagus kawarinus (Matsumura & ..., 1937) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus kchatriya Boucomont, 1914 | | | Onthophagus keikoae Ochi & Kon, 2014 | | | Onthophagus keiseri Frey, 1956 | | | Onthophagus keralensis Frey, 1975 | | | Onthophagus keralicus Biswas & Cha..., 1986 | | | Onthophagus kiambram Storey, 1977 | | | Onthophagus kindianus Frey, 1953 | | | Onthophagus kingstoni Cambefort, 1980 | | | Onthophagus kirki Frey, 1975 | | | Onthophagus kirokanus Frey, 1975 | | | Onthophagus kivuensis Walter, 2018 | | | Onthophagus kiyoshii Ochi & Kon, 2007 | | | Onthophagus klapperichi Balthasar, 1952 | | | Onthophagus kleinei Balthasar, 1935 | | | Onthophagus knapperti Krikken, 1981 | | | Onthophagus kochi Frey, 1957 | | | Onthophagus koebelei Blackburn, 1903 | | | Onthophagus koechlei Masumoto, Oc..., 2011 | | | Onthophagus kogatanus (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus kohlmanni Génier, 2017 | | | Onthophagus kokereka Matthews, 1972 | | | Onthophagus kokocellosus Krikken & Hu..., 2013 | | | Onthophagus kokodanus Krikken & Hu..., 2012 | | | Onthophagus kokodanus hagenaltus Krikken & Hu..., 2012 | | | Onthophagus kokodanus kokodanus | | | Onthophagus kokodentatus Krikken & Hu..., 2013 | | | Onthophagus kokoiorus Krikken & Hu..., 2012 | | | Onthophagus kokopygus Krikken & Hu..., 2013 | | | Onthophagus kokosquamatus Krikken & Hu..., 2012 | | | Onthophagus kolaka Huijbregts &..., 2009 | | | Onthophagus koma Matsumura, 1937 | | | Onthophagus komabellus Matsumura & ..., 1937 | | | Onthophagus komareki Balthasar, 1935 | | | Onthophagus kongkaewensis Masumoto, Oc..., 2012 | | | Onthophagus koni Ochi, 2007 | | | Onthophagus konsarnensis Masumoto, Oc..., 2008 | | | Onthophagus kontumicus Kabakov, 1983 | | | Onthophagus kora Storey, 1977 | | | Onthophagus kouassii Cambefort, 1984 | | | Onthophagus kozunonis (Matsumura, 1937) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus kraatzeanus Lansberge, 1883 | | | Onthophagus kudrnaianus Josso, 2022 | | | Onthophagus kukali Krikken & Hu..., 2013 | | | Onthophagus kulti Balthasar, 1952 | | | Onthophagus kuluensis Bates, 1891 | | | Onthophagus kulzeri Frey, 1958 | | | Onthophagus kumaonensis Arrow, 1931 | | | Onthophagus kumbaingeri Matthews, 1972 | | | Onthophagus kyleensis Frey, 1975 | | | Onthophagus kyokoae Masumoto & Ochi, 2015 | | | Onthophagus labdacus Balthasar, 1969 | | | Onthophagus laborans Arrow, 1931 | | | Onthophagus laceratus (Gerstaecker, 1871) | ⇒ Onthophagus depressus Harold, 1871 | | Onthophagus lacustris Harold, 1877 | | | Onthophagus laetus d´Orbigny, 1913 | | | Onthophagus laevatus d´Orbigny, 1902 | | | Onthophagus laeviceps d´Orbigny, 1902 | | | Onthophagus laevidorsis d´Orbigny, 1907 | | | Onthophagus laevigatus Fabricius, 1798 | | | Onthophagus laevissimus d´Orbigny, 1905 | | | Onthophagus lagnyi Paulian, 1937 | | | Onthophagus lahorensis Kabakov, 2008 | | | Onthophagus laichauensis Ochi, Kon & ..., 2021 | | | Onthophagus lamellicornis Frey, 1953 | | | Onthophagus lamelliger (Gerstaecker, 1871) | ⇒ Onthophagus ochropygus d´Orbigny, 1902 | | Onthophagus lamgalio Matthews, 1972 | | | Onthophagus laminatus (MacLeay, 1864) | ⇒ Onthophagus lunatus Harold, 1868 | | Onthophagus laminicornis (Lansberge, 1886) | ⇒ Onthophagus eulaminicornis Pittino, 2006 | | Onthophagus laminidorsis d´Orbigny, 1902 | | | Onthophagus laminosus d´Orbigny, 1905 | | | Onthophagus lamnifer d´Orbigny, 1902 | | | Onthophagus lamotellus Cambefort, 1980 | | | Onthophagus lamottei Frey, 1962 | | | Onthophagus lamtoensis Cambefort, 1984 | | | Onthophagus lamtoi Cambefort, 1980 | | | Onthophagus lamyi Paulian, 1945 | | | Onthophagus langkawiensis Ochi & Kon, 2015 | | | Onthophagus lanista (Castelnau, 1840) | ⇒ Proagoderus extensus Harold, 1878 | | Onthophagus lapillus Arrow, 1931 | | | Onthophagus laratinus Arrow, 1916 | | | Onthophagus lassulus Balthasar, 1964 | | | Onthophagus latenasutus Arrow, 1941 | | | Onthophagus latepunctatus d´Orbigny, 1913 | | | Onthophagus latestriatus d´Orbigny, 1908 | | | Onthophagus latevittatus d´Orbigny, 1905 | | | Onthophagus latigibber d´Orbigny, 1902 | | | Onthophagus latipennis d´Orbigny, 1897 | | | Onthophagus latissimus d´Orbigny, 1902 | | | Onthophagus latro Harold, 1877 | | | Onthophagus leai Blackburn, 1895 | | | Onthophagus leanus Goidanich, 1926 | | | Onthophagus lebasi Boucomont, 1932 | | | Onthophagus lefebvrei d´Orbigny, 1913 | | | Onthophagus lefiniensis Balthasar, 1967 | | | Onthophagus legendrei Walter & Cam..., 1977 | | | Onthophagus leichhardti Monteith & S..., 2013 | | | Onthophagus lekmaengtibanphrao Masumoto & Ochi, 2015 | | | Onthophagus leknamnaous Masumoto & Ochi, 2015 | | | Onthophagus lemagneni d´Orbigny, 1902 | | | Onthophagus lemekensis d´Orbigny, 1915 | | | Onthophagus lemniscatus Gillet, 1924 | | | Onthophagus lemuroides d´Orbigny, 1898 | | | Onthophagus lenzii Harold, 1874 | | | Onthophagus leomontanus Boucomont, 1914 | | | Onthophagus leroyi d´Orbigny, 1902 | | | Onthophagus leucopygus (Harold, 1867) | ⇒ Onthophagus albipodex d´Orbigny, 1902 | | Onthophagus leusermontis Huijbregts &..., 2011 | | | Onthophagus liberianus Lansberge, 1883 | | | Onthophagus lilongweensis Dierkens, 2022 | | | Onthophagus limonensis Kohlmann & S..., 2001 | | | Onthophagus lindu Huijbregts &..., 2009 | | | Onthophagus liodermus d´Orbigny, 1913 | | | Onthophagus lioides d´Orbigny, 1902 | | | Onthophagus liopterus Harold, 1880 | | | Onthophagus liothorax Koshantschikov, 1894 | | | Onthophagus lituratus Roth, 1851 | | | Onthophagus liui Masumoto, Oc..., 2014 | | | Onthophagus liwagensis Ochi & Kon, 2006 | | | Onthophagus liwondeensis Josso, 2019 | | | Onthophagus lobi Cambefort, 1984 | | | Onthophagus lojanus Balthasar, 1939 | | | Onthophagus lomii Müller, 1942 | | | Onthophagus longecarinatus Zunino & Hal..., 1988 | | | Onthophagus longegranus Frey, 1957 | | | Onthophagus longiceps d´Orbigny, 1904 | | | Onthophagus longimanus Bates, 1887 | | | Onthophagus longipes Paulian, 1937 | | | Onthophagus longipilis d´Orbigny, 1905 | | | Onthophagus lopeensis Josso, 2022 | | | Onthophagus lore Huijbregts &..., 2009 | | | Onthophagus lorianus Gillet, 1930 | | | Onthophagus loricatus Klug, 1855 | | | Onthophagus loroi Balthasar, 1941 | | | Onthophagus loudetiae Cambefort, 1984 | | | Onthophagus loveni Gillet, 1928 | | | Onthophagus loxodontae Cambefort, 1984 | | | Onthophagus luctuosus Boucomont, 1914 | | | Onthophagus ludicrus Balthasar, 1969 | | | Onthophagus ludio Boucomont, 1914 | | | Onthophagus lugens (Fahraeus, 1857) | ⇒ Onthophagus fimetarius Roth, 1851 | | Onthophagus lugubris (Fahraeus, 1857) | ⇒ Onthophagus lituratus Roth, 1851 | | Onthophagus luismargaritorum Delgado, 1995 | | | Onthophagus lumareti Walter & Cam..., 1977 | | | Onthophagus lunatus Harold, 1868 | | | Onthophagus lunulifer Boucomont, 1914 | | | Onthophagus lutaticollis d´Orbigny, 1907 | | | Onthophagus luteosignatus Lansberge, 1883 | | | Onthophagus luteus ♰ Oustalet, 1874 | | | Onthophagus luzonwallacei Ochi, Kawaha..., 2021 | | | Onthophagus mabuensis Daniel, Strü..., 2023 | | | Onthophagus macleayi Blackburn, 1903 | | | Onthophagus macrocephalus Kirby, 1818 | | | Onthophagus macroliberianus Moretto, 2010 | | | Onthophagus macrothorax d´Orbigny, 1902 | | | Onthophagus maculosipennis Gillet, 1930 | | | Onthophagus maculosus d´Orbigny, 1908 | | | Onthophagus madoqua Arrow, 1931 | | | Onthophagus maephaluangus Masumoto, Oc..., 2007 | | | Onthophagus maesalongensis Masumoto, Oc..., 2013 | | | Onthophagus magnini Paulian, 1933 | | | Onthophagus magnioculus Ochi & Kon, 2006 | | | Onthophagus magnipygus Boucomont, 1914 | | | Onthophagus makokou Walter, 1989 | | | Onthophagus malabarensis Boucomont, 1919 | | | Onthophagus malangensis Boucomont, 1914 | | | Onthophagus malasiacus Gillet, 1927 | | | Onthophagus malevolus Balthasar, 1964 | | | Onthophagus malthinus Gillet, 1930 | | | Onthophagus mamillatus Lea, 1923 | | | Onthophagus manantlanensis Sánchez-Huer..., 2021 | | | Onthophagus manguliensis Boucomont, 1914 | | | Onthophagus mankonoensis (Balthasar, 1966) | ⇒ Onthophagus mocquerysi d´Orbigny, 1902 | | Onthophagus manya Matthews, 1972 | | | Onthophagus marahouensis Cambefort, 1984 | | | Onthophagus margaretensis Blackburn, 1903 | | | Onthophagus marginatus (Castelnau, 1840) | ⇒ Onthophagus andalusicus Waltl, 1835 | | Onthophagus marginicollis Harold, 1880 | | | Onthophagus marginifer Frey, 1953 | | | Onthophagus marginipennis Josso, 2018 | | | Onthophagus mariozuninoi Delgado, Nav..., 1993 | | | Onthophagus marshalli d´Orbigny, 1908 | | | Onthophagus martaponsae | | | Onthophagus martelli Frey, 1972 | | | Onthophagus martellii Frey, 1972 | | | Onthophagus martialis Boucomont, 1914 | | | Onthophagus martinpierai Moctezuma, R..., 2016 | | | Onthophagus maruchanus Masumoto, Oc..., 2012 | | | Onthophagus maruyamai Ochi, Kon & ..., 2014 | | | Onthophagus maryatiae Ochi & Kon, 2005 | | | Onthophagus masaicus d´Orbigny, 1905 | | | Onthophagus matae Cambefort, 1984 | | | Onthophagus matanyo Huijbregts &..., 2009 | | | Onthophagus mateui Moctezuma & ..., 2020 | | | Onthophagus matsuii Ochi & Kon, 2006 | | | Onthophagus matsuokai (Matsumura & ..., 1937) | ⇒ Onthophagus sauteri Gillet, 1924 | | Onthophagus mauritii Boucomont, 1919 | | | Onthophagus maxwellianus Moretto, 2013 | | | Onthophagus maya Zunino, 1981 | | | Onthophagus mayeri Harold, 1876 | | | Onthophagus mediofuscatus d´Orbigny, 1908 | | | Onthophagus megapacificus Ochi & Kon, 2006 | | | Onthophagus megathorax Gillet, 1921 | | | Onthophagus mekara Huijbregts &..., 2009 | | | Onthophagus melanocephalus Klug, 1845 | | | Onthophagus menkaoensis Walter & Cam..., 1977 | | | Onthophagus mentaveiensis Boucomont, 1914 | | | Onthophagus merdrignaci Walter & Cam..., 1977 | | | Onthophagus meruanus d´Orbigny, 1908 | | | Onthophagus merus Péringuey, 1901 | | | Onthophagus metalliger d´Orbigny, 1913 | | | Onthophagus mexicanus Bates, 1887 | | | Onthophagus micellus Matsumura, 1937 | | | Onthophagus micropterus Zunino & Hal..., 1981 | | | Onthophagus micros d´Orbigny, 1902 | | | Onthophagus midorianus (Matsumura, 1938) | ⇒ Parascatonomus anguicorius Boucomont, 1914 | | Onthophagus miichunus Masumoto, Oc..., 2020 | | | Onthophagus mije Storey & Weir, 1990 | | | Onthophagus miles d´Orbigny, 1905 | | | Onthophagus militaris Boucomont, 1914 | | | Onthophagus millamilla Matthews, 1972 | | | Onthophagus millingeni d´Orbigny, 1900 | | | Onthophagus mimikanus Balthasar, 1969 | | | Onthophagus mimus d´Orbigny, 1913 | | | Onthophagus minans d´Orbigny, 1913 | | | Onthophagus minax d´Orbigny, 1904 | | | Onthophagus mindanaensis Boucomont, 1914 | | | Onthophagus mindanaoensis Frey, 1971 | | | Onthophagus minettianus Josso, 2019 | | | Onthophagus minoi Matsumura & ..., 1937 | | | Onthophagus minokuchianus (Matsumura, 1934) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus minotaurus Arrow, 1941 | | | Onthophagus minusculus (MacLeay, 1888) | ⇒ Onthophagus quadripustulatus Fabricius, 1775 | | Onthophagus minutissimus d´Orbigny, 1908 | | | Onthophagus minutulus Harold, 1875 | | | Onthophagus minutus (Hausmann, 1807) | ⇒ Onthophagus centricornis Fabricius, 1798 | | Onthophagus mirabilis Bates, 1887 | | | Onthophagus mirandus Arrow, 1931 | | | Onthophagus miricollis Frey, 1962 | | | Onthophagus miricornis d´Orbigny, 1902 | | | Onthophagus mirifrons d´Orbigny, 1905 | | | Onthophagus miscellaneus d´Orbigny, 1908 | | | Onthophagus miscellus d´Orbigny, 1905 | | | Onthophagus misellus d´Orbigny, 1907 | | | Onthophagus misujianus Matsumura & ..., 1937 | | | Onthophagus mixticeps d´Orbigny, 1905 | | | Onthophagus mixtidorsis d´Orbigny, 1905 | | | Onthophagus miyabei (Matsumura, 1937) | ⇒ Onthophagus punctator Reitter, 1893 | | Onthophagus miyazakianus Matsumura & ..., 1937 | | | Onthophagus mjobergi Gillet, 1925 | | | Onthophagus mniszechi Harold, 1869 | Kershaw´s burying beetle | | Onthophagus moajat Huijbregts &..., 2009 | | | Onthophagus mocquerysi d´Orbigny, 1902 | | | Onthophagus modestus Harold, 1862 | | | Onthophagus mogo Krikken & Hu..., 2008 | | | Onthophagus mokwamensis Krikken & Hu..., 2012 | | | Onthophagus monardi (Boucomont, 1936) | ⇒ Onthophagus monardiellus Frey, 1963 | | Onthophagus monardiellus Frey, 1963 | | | Onthophagus mongana Storey & Weir, 1990 | | | Onthophagus monodon Fahraeus, 1857 | | | Onthophagus monteithi Matthews, 1972 | | | Onthophagus monticolus (Howden & Car..., 1963) | ⇒ Onthophagus durangoensis Balthasar, 1939 | | Onthophagus montishannoniae Krikken & Hu..., 2008 | | | Onthophagus montivagus d´Orbigny, 1902 | | | Onthophagus montreuili Moretto & Gé..., 2010 | | | Onthophagus mopsus Fabricius, 1792 | | | Onthophagus mordindangus Masumoto, Oc..., 2008 | | | Onthophagus moreleti Baraud, 1980 | | | Onthophagus morenoi Moretto, 2013 | | | Onthophagus morettoi Josso, 2022 | | | Onthophagus morimotoi Masumoto, Oc..., 2021 | | | Onthophagus moroni Zunino & Hal..., 1988 | | | Onthophagus morosus Gerstaecker, 1871 | | | Onthophagus mouhoti Harold, 1875 | | | Onthophagus mpassa Walter, 1982 | | | Onthophagus mucronatus (Thomson, 1858) | ⇒ Pseudosaproecius validicornis Quedenfeldt, 1884 | | Onthophagus mucronifer d´Orbigny, 1905 | | | Onthophagus mugheseensis Josso, 2019 | | | Onthophagus mulgravei Paulian, 1937 | | | Onthophagus multipunctatus Boucomont, 1914 | | | Onthophagus mulutagal Ochi, Kon & ..., 2016 | | | Onthophagus mundill Matthews, 1972 | | | Onthophagus murchisoni Blackburn, 1892 | | | Onthophagus murgon Monteith & S..., 2013 | | | Onthophagus murphyi Josso, 2019 | | | Onthophagus musculus Frey, 1963 | | | Onthophagus musculus ganganensis Frey, 1963 | | | Onthophagus musculus musculus | | | Onthophagus mushanus Matsumura, 1938 | | | Onthophagus mushensis Matsumura, 1938 | | | Onthophagus mutatus Harold, 1859 | | | Onthophagus muticus MacLeay, 1864 | | | Onthophagus mwenseensis Dierkens, 2022 | | | Onthophagus mycetorum Zunino & Hal..., 1988 | | | Onthophagus nabeleki Balthasar, 1939 | | | Onthophagus naevius d´Orbigny, 1913 | | | Onthophagus naevuliger d´Orbigny, 1908 | | | Onthophagus nagasawai Matsumura, 1938 | | | Onthophagus nagpurensis Arrow, 1931 | | | Onthophagus nakatomii (Matsumura, 1937) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus nammuldi Matthews, 1972 | | | Onthophagus namnaolekus Masumoto, Oc..., 2020 | | | Onthophagus namnaonus Masumoto, Oc..., 2020 | | | Onthophagus namnaous Masumoto, Oc..., 2013 | | | Onthophagus nanus (Harold, 1878) | ⇒ Onthophagus waterhousei Gillet, 1927 | | Onthophagus nasalis Arrow, 1931 | | | Onthophagus nasicornis Harold, 1869 | | | Onthophagus nasicus Gillet, 1930 | | | Onthophagus naso Fahraeus, 1857 | | | Onthophagus nasutus Guérin-Ménev..., 1855 | | | Onthophagus natronis Cambefort, 1984 | | | Onthophagus navarretorum Delgado & Ca..., 1996 | | | Onthophagus nchilaensis Josso, 2022 | | | Onthophagus neavei d´Orbigny, 1914 | | | Onthophagus neboissi Frey, 1970 | | | Onthophagus nebulosus Reiche, 1864 | | | Onthophagus necessarius Reitter, 1893 | | | Onthophagus necrophagus Arrow, 1931 | | | Onthophagus neervoorti Boucomont, 1914 | | | Onthophagus nefarius Balthasar, 1941 | | | Onthophagus negligens Walker, 1858 | | | Onthophagus nemorivagus Kohlmann & S..., 2001 | | | Onthophagus neofurcatus Goidanich, 1926 | | | Onthophagus neofuscus Moctezuma & ..., 2020 | | | Onthophagus neomirabilis Howden, 1973 | | | Onthophagus neostenocerus Goidanich, 1926 | | | Onthophagus neyo Moretto, 2013 | | | Onthophagus nicobaricus Biswas, Chat..., 1999 | | | Onthophagus nigerianus d´Orbigny, 1913 | | | Onthophagus nigriceps Raffray, 1877 | | | Onthophagus nigricornis Fairmaire, 1887 | | | Onthophagus nigrinus Paulian, 1937 | | | Onthophagus nigripennis d´Orbigny, 1908 | | | Onthophagus nigriventris d´Orbigny, 1902 | Black-bottom dung beetle | | Onthophagus nigrivestis (d´Orbigny, 1902) | ⇒ Onthophagus laeviceps d´Orbigny, 1902 | | Onthophagus nigroflavus Josso, 2022 | | | Onthophagus nigroviolaceus d´Orbigny, 1902 | | | Onthophagus nilicola d´Orbigny, 1902 | | | Onthophagus niloticus Harold, 1879 | | | Onthophagus nimbatus d´Orbigny, 1905 | | | Onthophagus niokolokoba Moretto, 2007 | | | Onthophagus nitefactus Harold, 1877 | | | Onthophagus nitidifrons d´Orbigny, 1905 | | | Onthophagus nitidior (Bates, 1887) | ⇒ Onthophagus blackburni Shipp, 1895 | | Onthophagus nitidor (Blackburn, 1892) | ⇒ Onthophagus blackburni Shipp, 1895 | | Onthophagus nitidulus Klug, 1845 | | | Onthophagus nitidus Waterhouse, 1875 | | | Onthophagus nodieri d´Orbigny, 1908 | | | Onthophagus nodulifer Harold, 1867 | | | Onthophagus nonstriatus d´Orbigny, 1905 | | | Onthophagus notatus d´Orbigny, 1902 | | | Onthophagus notiodes Solis & Kohl..., 2003 | | | Onthophagus novaeirlandiae Balthasar, 1969 | | | Onthophagus nubilus Kohlmann & S..., 2001 | | | Onthophagus nudatus d´Orbigny, 1913 | | | Onthophagus nudifrons Balthasar, 1939 | | | Onthophagus nudus d´Orbigny, 1908 | | | Onthophagus numidicus d´Orbigny, 1908 | | | Onthophagus nummelini Moretto & Gé..., 2020 | | | Onthophagus nurubuan Matthews, 1972 | | | Onthophagus nutans (Roth, 1851) | ⇒ Onthophagus schaufussi Harold, 1867 | | Onthophagus nyctopus Bates, 1887 | | | Onthophagus nyikaensis Josso, 2018 | | | Onthophagus nymani Gillet, 1930 | | | Onthophagus oaxacanus Zunino & Hal..., 1988 | | | Onthophagus obenbergeri Balthasar, 1965 | | | Onthophagus oberthueri d´Orbigny, 1898 | | | Onthophagus oberthuri d´Orbigny, 1898 | | | Onthophagus obliguus | | | Onthophagus obliviosus Balthasar, 1935 | | | Onthophagus obscurior Boucomont, 1914 | | | Onthophagus obtusicornis (Fahraeus, 1857) | ⇒ Onthophagus marginalis przewalski Kabakov, 2006 | | Onthophagus obtusus Wiedemann, 1819 | | | Onthophagus obtutus Péringuey, 1901 | | | Onthophagus ocellatopunctatus Waterhouse, 1875 | | | Onthophagus ocellidorsis d´Orbigny, 1915 | | | Onthophagus ocellifer d´Orbigny, 1902 | | | Onthophagus ocelliger Harold, 1877 | | | Onthophagus ochii Masumoto, 1988 | | | Onthophagus ochreatus d´Orbigny, 1897 | | | Onthophagus ochromerus Harold, 1877 | | | Onthophagus ochropygus d´Orbigny, 1902 | Common snipe fly | | Onthophagus octogonus Frey, 1962 | | | Onthophagus oculatus Arrow, 1931 | | | Onthophagus ofianus Krikken & Hu..., 2013 | | | Onthophagus ohbayashii Nomura, 1939 | | | Onthophagus ohkuboi Ochi & Kon, 2006 | | | Onthophagus oishii Matsumura & ..., 1937 | | | Onthophagus okahandjanus Balthasar, 1974 | | | Onthophagus okushiranus Matsumura & ..., 1937 | | | Onthophagus okushirianus Matsumura, 1937 | | | Onthophagus olidus Balthasar, 1964 | | | Onthophagus omiltemius (Bates, 1887) | ⇒ Onthophagus chevrolati omiltemius Bates, 1887 | | Onthophagus omostigma d´Orbigny, 1902 | | | Onthophagus oniellus (Matsumura, 1937) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus onorei Zunino & Hal..., 1997 | | | Onthophagus onthochromus Arrow, 1913 | | | Onthophagus ontosatu Huijbregts &..., 2012 | | | Onthophagus opacihartiniae Ochi & Kon, 2015 | | | Onthophagus opacipennis (Lea, 1923) | ⇒ Onthophagus pugnax Harold, 1868 | | Onthophagus opacotaurus Krikken & Hu..., 2012 | | | Onthophagus ophion Erichson, 1847 | | | Onthophagus ophion confusus Boucomont, 1932 | | | Onthophagus ophtalmicus Frey, 1958 | | | Onthophagus orbicularis Lansberge, 1885 | | | Onthophagus orbus Boucomont, 1924 | | | Onthophagus orcas (Ménétriés, 1838) | ⇒ Onthophagus atramentarius Ménétriés, 1832 | | Onthophagus orissanus Arrow, 1931 | | | Onthophagus orizabensis Moctezuma, J..., 2019 | | | Onthophagus ornaticollis Gillet, 1930 | | | Onthophagus ornatulus d´Orbigny, 1908 | | | Onthophagus orphnoides (Bates, 1887) | ⇒ Onthophagus mirabilis Bates, 1887 | | Onthophagus orthocerus Thomson, 1858 | | | Onthophagus osculatii Guérin-Ménev..., 1855 | | | Onthophagus osellai Pittino, 1982 | | | Onthophagus oshimanus Nakane, 1960 | | | Onthophagus otai Ochi & Kon, 2006 | | | Onthophagus otjivarongus Balthasar, 1967 | | | Onthophagus ouratita Matthews, 1972 | | | Onthophagus ovatulus ♰ Heer, 1847 | | | Onthophagus ovigranosus Frey, 1971 | | | Onthophagus ovulum Gerstaecker, 1871 | | | Onthophagus pacificus Lansberge, 1885 | | | Onthophagus pactolus Fabricius, 1787 | | | Onthophagus padrinoi Delgado, 1999 | | | Onthophagus palamoui Biswas & Cha..., 1986 | | | Onthophagus palatus Boucomont, 1914 | | | Onthophagus palavanus (Huijbregts &..., 2011) | ⇒ Onthophagus javanensis Balthasar, 1963 | | Onthophagus palawantagal Ochi, Kon & ..., 2016 | | | Onthophagus pallens d´Orbigny, 1908 | | | Onthophagus pallidipennis (Fahraeus, 1857) | ⇒ Onthophagus rasipennis d´Orbigny, 1908 | | Onthophagus pallidus d´Orbigny, 1905 | | | Onthophagus paluma Matthews, 1972 | | | Onthophagus pangmaphanus Masumoto, Oc..., 2022 | | | Onthophagus papuater Krikken & Hu..., 2012 | | | Onthophagus papuensis Harold, 1877 | | | Onthophagus papuplicatus Krikken & Hu..., 2012 | | | Onthophagus papurugosus Krikken & Hu..., 2012 | | | Onthophagus paracenricornis Ochi & Kon, 2014 | | | Onthophagus paracentricornis Ochi & Kon, 2014 | | | Onthophagus parafasciatus Balthasar, 1974 | | | Onthophagus parafuscus Zunino & Hal..., 2005 | | | Onthophagus parallelicornis MacLeay, 1887 | | | Onthophagus parapalatus Krikken & Hu..., 1988 | | | Onthophagus parapedisequus Balthasar, 1969 | | | Onthophagus paravinctus Tagliaferri ..., 2012 | | | Onthophagus parceguttatus d´Orbigny, 1902 | | | Onthophagus parcenotatus d´Orbigny, 1913 | | | Onthophagus parcepictus d´Orbigny, 1908 | | | Onthophagus parcepilosus Balthasar, 1966 | | | Onthophagus pardalis Fabricius, 1798 | | | Onthophagus parenthesis Boucomont, 1913 | | | Onthophagus parisii (Balthasar, 1941) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus paroculus Krikken & Hu..., 1987 | | | Onthophagus parrumbal Matthews, 1972 | | | Onthophagus parryi Harold, 1869 | | | Onthophagus parumnotatus Fahraeus, 1857 | | | Onthophagus parvidens (Frey, 1971) | ⇒ Onthophagus clermonti Paulian, 1931 | | Onthophagus parvifrons (d´Orbigny, 1907) | ⇒ Onthophagus semiasper d´Orbigny, 1902 | | Onthophagus parvulus (Fabricius, 1798) | ⇒ Onthophagus frontalis Raffray, 1877 | | Onthophagus parvus Blanchard, 1853 | | | Onthophagus pasotagal Ochi, Kon & ..., 2016 | | | Onthophagus pastillatus Boucomont, 1919 | | | Onthophagus paucigranosus Lansberge, 1886 | | | Onthophagus paulianellus Frey, 1958 | | | Onthophagus pauliani (Balthasar, 1937) | ⇒ Onthophagus paulianellus Frey, 1958 | | Onthophagus pauper Boucomont, 1914 | | | Onthophagus pauxillus d´Orbigny, 1902 | | | Onthophagus pavidus Harold, 1877 | | | Onthophagus pedator Sharp, 1875 | | | Onthophagus pedester Howden & Gén..., 2004 | | | Onthophagus pedisequus Balthasar, 1969 | | | Onthophagus pendjarius Josso & Prév..., 2006 | | | Onthophagus penedwardsae Monteith & S..., 2013 | | | Onthophagus peninsularis Boucomont, 1914 | | | Onthophagus peninsulomerus Huijbregts &..., 2011 | | | Onthophagus pentacanthus Harold, 1867 | Five horned dung beetle | | Onthophagus peotoxus Krikken & Hu..., 2012 | | | Onthophagus perakensis Ochi, Kon & ..., 2014 | | | Onthophagus peramelinus Lea, 1923 | | | Onthophagus peringueyi Shipp, 1895 | | | Onthophagus perniger Boucomont, 1930 | | | Onthophagus perpilosus MacLeay, 1871 | | | Onthophagus personatus Boucomont, 1914 | | | Onthophagus petenensis Howden & Gill, 1993 | | | Onthophagus pexatus Harold, 1869 | | | Onthophagus peyrierasi Paulian, 1985 | | | Onthophagus phahompokus Masumoto & Ochi, 2015 | | | Onthophagus phakuaiensis Masumoto, Oc..., 2013 | | | Onthophagus phanaeicollis Lansberge, 1883 | | | Onthophagus phanaeides Frey, 1956 | | | Onthophagus phanaeiformis Boucomont, 1914 | | | Onthophagus phanaeoides (Westwood, 1839) | ⇒ Proagoderus lanista Castelnau, 1840 | | Onthophagus phanaeomorphus Janssens, 1954 | | | Onthophagus phatoensis Masumoto, Oc..., 2008 | | | Onthophagus philippemorettoi Josso, 2022 | | | Onthophagus philippinensis Boucomont, 1919 | | | Onthophagus phillippsorum Krikken & Hu..., 1987 | | | Onthophagus phoenicocerus Lea, 1923 | | | Onthophagus phrixus Balthasar, 1969 | | | Onthophagus phukhieoensis Masumoto, Oc..., 2007 | | | Onthophagus picatus d´Orbigny, 1902 | | | Onthophagus piceiceps d´Orbigny, 1904 | | | Onthophagus picipennis Hope, 1841 | | | Onthophagus pictipennis d´Orbigny, 1913 | | | Onthophagus pictipodex d´Orbigny, 1902 | | | Onthophagus picturatus d´Orbigny, 1908 | | | Onthophagus pictus (Reitter, 1893) | ⇒ Oniticellus tessellatus Harold, 1879 | | Onthophagus pierrei Josso, 2022 | | | Onthophagus piffli Petrovitz, 1961 | | | Onthophagus pilauco Tello, Verdú..., 2021 | | | Onthophagus pilicollis d´Orbigny, 1902 | | | Onthophagus pilipodex d´Orbigny, 1913 | | | Onthophagus pillara Matthews, 1972 | | | Onthophagus pilosus (Fahraeus, 1857) | ⇒ Onthophagus echinus Boucomont, 1914 | | Onthophagus pimpasaleei Masumoto, Oc..., 2008 | | | Onthophagus pinaroo Storey & Weir, 1990 | | | Onthophagus pinguis Gerstaecker, 1871 | | | Onthophagus pipitzi Ancey, 1883 | | | Onthophagus pisciphagus Walter & Cam..., 1977 | | | Onthophagus pithankithae Karimbumkara..., 2016 | | | Onthophagus placens Péringuey, 1904 | | | Onthophagus planaticeps d´Orbigny, 1913 | | | Onthophagus planiceps (MacLeay, 1886) | ⇒ Onthophagus pinguis Gerstaecker, 1871 | | Onthophagus planicollis Harold, 1880 | | | Onthophagus planidorsis Dierkens, 2022 | | | Onthophagus planifrons Frey, 1962 | | | Onthophagus platalea Arrow, 1941 | | | Onthophagus pleurogonus d´Orbigny, 1913 | | | Onthophagus plicatifrons d´Orbigny, 1908 | | | Onthophagus pljushtchi Ivanova, 2012 | | | Onthophagus poeophagus Kabakov, 1979 | | | Onthophagus politissimus d´Orbigny, 1908 | | | Onthophagus pollicatus Harold, 1879 | | | Onthophagus polyedrus d´Orbigny, 1905 | | | Onthophagus polyodon d´Orbigny, 1913 | | | Onthophagus polystigma d´Orbigny, 1902 | | | Onthophagus pooensis Cambefort & ..., 1991 | | | Onthophagus porcus Arrow, 1931 | | | Onthophagus poringensis Ochi & Kon, 2005 | | | Onthophagus possoi Walter, 1982 | | | Onthophagus posticicornis d´Orbigny, 1913 | | | Onthophagus posticus Erichson, 1842 | | | Onthophagus potosinus Moctezuma & ..., 2020 | | | Onthophagus praecellens Bates, 1887 | | | Onthophagus praedatus Harold, 1862 | | | Onthophagus praelaminatus Frey, 1957 | | | Onthophagus praestans Péringuey, 1901 | | | Onthophagus praetortus Péringuey, 1904 | | | Onthophagus prehensilis Arrow, 1920 | | | Onthophagus priscus | | | Onthophagus probus Péringuey, 1901 | | | Onthophagus procurvus Balthasar, 1935 | | | Onthophagus prodromus ♰ Heer, 1862 | | | Onthophagus producticollis d´Orbigny, 1908 | | | Onthophagus productus Arrow, 1907 | | | Onthophagus profanus Frey, 1958 | | | Onthophagus pronocarinatus Josso, 2019 | | | Onthophagus pronus Erichson, 1842 | | | Onthophagus propinquus (MacLeay, 1888) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus propraecellens Howden & Gill, 1987 | | | Onthophagus proteus d´Orbigny, 1902 | | | Onthophagus protuberans Frey, 1955 | | | Onthophagus proximus d´Orbigny, 1908 | | | Onthophagus pseudoaeneus d´Orbigny, 1908 | | | Onthophagus pseudoarai Ochi & Kon, 2018 | | | Onthophagus pseudobidens d´Orbigny, 1915 | | | Onthophagus pseudobrutus Kabakov, 1983 | | | Onthophagus pseudobufonidius Josso, 2019 | | | Onthophagus pseudoconvexicollis Bai & Yang, 2016 | | | Onthophagus pseudocoracinus Kabakov, 1983 | | | Onthophagus pseudocostifer Krikken & Hu..., 2012 | | | Onthophagus pseudocribripennis Josso, 2019 | | | Onthophagus pseudofimetarius Balthasar, 1946 | | | Onthophagus pseudoguatemalensis Moctezuma & ..., 2021 | | | Onthophagus pseudojaponicus Balthasar, 1941 | | | Onthophagus pseudojaponicus fukiensis Balthasar, 1941 | | | Onthophagus pseudojaponicus pseudojaponicus | | | Onthophagus pseudoliberianus Moretto, 2010 | | | Onthophagus pseudopilosus Frey, 1958 | | | Onthophagus pseudosanguineus Walter & Cam..., 1977 | | | Onthophagus pseudosellatus Balthasar, 1964 | | | Onthophagus pseudoundulans Zunino & Hal..., 1988 | | | Onthophagus pseudoundulans howdeni Zunino & Hal..., 1988 | | | Onthophagus pseudovinctus Tagliaferri ..., 2012 | | | Onthophagus ptox Erichson, 1842 | | | Onthophagus puberulus d´Orbigny, 1902 | | | Onthophagus pugionatus Fahraeus, 1857 | | | Onthophagus pugnacior Blackburn, 1903 | | | Onthophagus pugnax Harold, 1868 | | | Onthophagus pulchellus d´Orbigny, 1905 | | | Onthophagus pullatus d´Orbigny, 1905 | | | Onthophagus pullus (Roth, 1851) | ⇒ Onthophagus parenthesis Boucomont, 1913 | | Onthophagus pumilus Kabakov, 2014 | | | Onthophagus punthari Storey, 1977 | | | Onthophagus pupillatus Kolbe, 1886 | | | Onthophagus purifrons d´Orbigny, 1908 | | | Onthophagus purpurascens Boucomont, 1914 | | | Onthophagus purpureicollis MacLeay, 1864 | | | Onthophagus pusillus (Fabricius, 1798) | ⇒ Onthophagus modestus Harold, 1862 | | Onthophagus pusio (Fahraeus, 1857) | ⇒ Onthophagus confertus Péringuey, 1908 | | Onthophagus putealis (Matsumura, 1938) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus pygidialis Lansberge, 1883 | | | Onthophagus pygmaeus Schaller, 1783 | | | Onthophagus pyramidalis Klug, 1855 | | | Onthophagus quadraticeps Harold, 1867 | | | Onthophagus quadricallosus d´Orbigny, 1902 | | | Onthophagus quadricuspis d´Orbigny, 1908 | | | Onthophagus quadrilunatus d´Orbigny, 1902 | | | Onthophagus quadrimaculatus Raffray, 1877 | | | Onthophagus quadrinodosus Fahraeus, 1857 | | | Onthophagus quadrinotatus d´Orbigny, 1905 | | | Onthophagus quadripustulatus Fabricius, 1775 | | | Onthophagus quangnamensis Ochi, Kon & ..., 2021 | | | Onthophagus quasijohkii Ochi & Kon, 2005 | | | Onthophagus quasitagal Ochi & Kon, 2005 | | | Onthophagus queenslandicus Blackburn, 1903 | | | Onthophagus quetzalis Howden & Gill, 1993 | | | Onthophagus quinquetuberculatus MacLeay, 1871 | | | Onthophagus quiproquo Moretto & Gé..., 2010 | | | Onthophagus ragazzii d´Orbigny, 1904 | | | Onthophagus ramosellus Bates, 1891 | | | Onthophagus ramosus Wiedemann, 1823 | | | Onthophagus rana Arrow, 1931 | | | Onthophagus ranongensis Masumoto, Oc..., 2008 | | | Onthophagus ranongjohkii Ochi, Kon & ..., 2016 | | | Onthophagus ranunculus Arrow, 1913 | | | Onthophagus rasipennis d´Orbigny, 1908 | | | Onthophagus ratchataniensis Masumoto, Oc..., 2013 | | | Onthophagus rectestriatus d´Orbigny, 1915 | | | Onthophagus rectilamina d´Orbigny, 1902 | | | Onthophagus rectorispauliani Cambefort, 1980 | | | Onthophagus refulgens Arrow, 1931 | | | Onthophagus regalis Arrow, 1907 | | | Onthophagus reichardti (Medvedev, 1928) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus remotus Harold, 1879 | | | Onthophagus renejeanneli Dierkens, 2022 | | | Onthophagus reticollis MacLeay, 1886 | | | Onthophagus reticulatus d´Orbigny, 1902 | | | Onthophagus reticuliger Frey, 1960 | | | Onthophagus reticulundaticeps Josso, 2022 | | | Onthophagus retusus (Harold, 1869) | ⇒ Onthophagus chevrolati retusus Harold, 1869 | | Onthophagus reyesi Zunino & Hal..., 1988 | | | Onthophagus rhinocerus (Gomes Alves, 1944) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus rhinolophus Harold, 1869 | | | Onthophagus rhinophyllus Harold, 1868 | | | Onthophagus rhodesianus Frey, 1975 | | | Onthophagus rhynchophorus Péringuey, 1904 | | | Onthophagus rhyncophorus | | | Onthophagus ribbei Boucomont, 1914 | | | Onthophagus riparius Lansberge, 1885 | | | Onthophagus rizali Ochi, Kawaha..., 2021 | | | Onthophagus robertminettii Josso, 2022 | | | Onthophagus robertopoggii Ochi & Kon, 2006 | | | Onthophagus roessneri Ziani, 2016 | | | Onthophagus rohwedderi Moretto, 2017 | | | Onthophagus rojkoffianus Josso, 2018 | | | Onthophagus rorarius Harold, 1877 | | | Onthophagus rosemarieae Josso, 2022 | | | Onthophagus rosenbergi Krikken & Hu..., 2011 | | | Onthophagus rosenbergi divergens Krikken & Hu..., 2011 | | | Onthophagus rosenbergi rosenbergi | | | Onthophagus rosettae Frey, 1958 | | | Onthophagus rostratus (Harold, 1869) | ⇒ Onthophagus trinominatus Goidanich, 1926 | | Onthophagus rostrifer (Reitter, 1893) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus rotundatus d´Orbigny, 1905 | | | Onthophagus rotundibasis d´Orbigny, 1902 | | | Onthophagus rotundicollis Lansberge, 1883 | | | Onthophagus rougonorum Cambefort, 1984 | | | Onthophagus rouyeri Boucomont, 1914 | | | Onthophagus royi Biswas & Cha..., 1985 | | | Onthophagus ruandanus Frey, 1971 | | | Onthophagus rubefactus (d´Orbigny, 1913) | ⇒ Onthophagus tessulatus Harold, 1871 | | Onthophagus rubellus d´Orbigny, 1908 | | | Onthophagus rubens d´Orbigny, 1902 | | | Onthophagus rubenticollis d´Orbigny, 1902 | | | Onthophagus rubescens MacLeay, 1888 | | | Onthophagus rubicundulus MacLeay, 1871 | | | Onthophagus rubidus d´Orbigny, 1913 | | | Onthophagus rubrescens (Blanchard, 1843) | ⇒ Onthophagus stylocerus Graells, 1851 | | Onthophagus rubricatus d´Orbigny, 1902 | | | Onthophagus rubrimaculatus MacLeay, 1864 | | | Onthophagus rubripennis (Arrow, 1907) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus rufescens Bates, 1887 | | | Onthophagus ruficauda Arrow, 1931 | | | Onthophagus rufipodex d´Orbigny, 1913 | | | Onthophagus rufobasalis Fairmaire, 1887 | | | Onthophagus rufocastaneus Fairmaire, 1887 | | | Onthophagus rufolimbatus d´Orbigny, 1913 | | | Onthophagus rufonotatus d´Orbigny, 1902 | | | Onthophagus rufopygus Frey, 1957 | | | Onthophagus rufosignatus MacLeay, 1864 | | | Onthophagus rufostillans d´Orbigny, 1907 | | | Onthophagus rufovirens d´Orbigny, 1904 | | | Onthophagus rufulgens | | | Onthophagus rugicollis Harold, 1880 | | | Onthophagus rugidorsis d´Orbigny, 1913 | | | Onthophagus rugosicollis Gillet, 1925 | | | Onthophagus rugosipennis Frey, 1973 | | | Onthophagus rugosissimus d´Orbigny, 1913 | | | Onthophagus rugulipennis Fairmaire, 1887 | | | Onthophagus rupicapra Waterhouse, 1894 | | | Onthophagus rutilans (Sharp, 1875) | ⇒ Onthophagus borneensis Harold, 1877 | | Onthophagus rutilans aborneensis Ochi, Kon & ..., 2009 | | | Onthophagus rutilans rutilans | | | Onthophagus rutriceps Krikken & Hu..., 2012 | | | Onthophagus sabahensis Ochi & Kon, 2006 | | | Onthophagus sabai Masumoto, Oc..., 2012 | | | Onthophagus sacharovskii Olsoufieff, 1918 | | | Onthophagus saggitarius Fabricius, 1775 | | | Onthophagus sagittarus | | | Onthophagus sahai Biswas & Cha..., 1986 | | | Onthophagus saigonensis Boucomont, 1924 | | | Onthophagus saksphus Masumoto, Oc..., 2021 | | | Onthophagus salebrosus MacLeay, 1888 | | | Onthophagus salvadorensis Zunino & Hal..., 1988 | | | Onthophagus samarensis Ochi, Kon & ..., 2023 | | | Onthophagus samoengus Masumoto, Oc..., 2008 | | | Onthophagus sancristobalensis Moctezuma & ..., 2020 | | | Onthophagus sanggona Huijbregts &..., 2009 | | | Onthophagus sangirensis Boucomont, 1914 | | | Onthophagus sanguineus d´Orbigny, 1902 | | | Onthophagus sanguinolentus d´Orbigny, 1908 | | | Onthophagus sangwalus Masumoto, Oc..., 2008 | | | Onthophagus sanpabloetlorum Moctezuma & ..., 2020 | | | Onthophagus sansibaricus Harold, 1878 | | | Onthophagus santamariensis Moctezuma, S..., 2020 | | | Onthophagus sapphirinus (Fahraeus, 1857) | ⇒ Proagoderus aureiceps d´Orbigny, 1902 | | Onthophagus sarasinorum Krikken & Hu..., 2011 | | | Onthophagus savanicola Cambefort, 1984 | | | Onthophagus scaber Roth, 1851 | | | Onthophagus scaberrimus d´Orbigny, 1904 | | | Onthophagus scapularis d´Orbigny, 1902 | | | Onthophagus sceptrifer Boucomont, 1924 | | | Onthophagus schaefernai Balthasar, 1935 | | | Onthophagus schatzmayri (Pierotti, 1959) | ⇒ Onthophagus opacicollis Reitter, 1893 | | Onthophagus schaufussi Harold, 1867 | | | Onthophagus schawalleri Scheuern, 1996 | | | Onthophagus schillhammeri Kabakov, 2006 | | | Onthophagus schunckei Paulian, 1936 | | | Onthophagus sciron Balthasar, 1969 | | | Onthophagus scrutator Harold, 1877 | | | Onthophagus sculptilis Gerstaecker, 1871 | | | Onthophagus secundarius Roth, 1851 | | | Onthophagus sellatulus d´Orbigny, 1902 | | | Onthophagus sellatus Klug, 1845 | | | Onthophagus sembeli Krikken & Hu..., 2008 | | | Onthophagus semiaereopictus Ochi, Kawaha..., 2021 | | | Onthophagus semiaratus d´Orbigny, 1902 | | | Onthophagus semiasper d´Orbigny, 1902 | | | Onthophagus semichalcites d´Orbigny, 1902 | | | Onthophagus semicinctus d´Orbigny, 1897 | | | Onthophagus semicroceus d´Orbigny, 1915 | | | Onthophagus semiflavus (Boheman, 1860) | ⇒ Onthophagus flavolimbatus Klug, 1855 | | Onthophagus semigraniger d´Orbigny, 1905 | | | Onthophagus semigranosus Lansberge, 1883 | | | Onthophagus semilaevis d´Orbigny, 1913 | | | Onthophagus semimetallicus Lea, 1923 | | | Onthophagus seminitens d´Orbigny, 1902 | | | Onthophagus seminitidus d´Orbigny, 1908 | | | Onthophagus semiopacus (Harold, 1869) | ⇒ Onthophagus seminitens d´Orbigny, 1902 | | Onthophagus semipacificus Ochi & Kon, 2006 | | | Onthophagus semiperakensis Ochi, Kon & ..., 2014 | | | Onthophagus semivestitus Paulian, 1937 | | | Onthophagus semivirescens d´Orbigny, 1902 | | | Onthophagus semiviridis d´Orbigny, 1904 | | | Onthophagus semiworoae Ochi, Kon & ..., 2014 | | | Onthophagus senegalensis d´Orbigny, 1902 | | | Onthophagus senescens Péringuey, 1908 | | | Onthophagus senex Boucomont, 1914 | | | Onthophagus seniculus (Fabricius, 1781) | ⇒ Onthophagus senescens Péringuey, 1908 | | Onthophagus sepilokensis Ochi & Kon, 2006 | | | Onthophagus seramicus Huijbregts &..., 2012 | | | Onthophagus serdangensis Lansberge, 1886 | | | Onthophagus sericans Frey, 1975 | | | Onthophagus sericeicollis Boucomont, 1928 | | | Onthophagus serienotatus d´Orbigny, 1902 | | | Onthophagus seseba Krikken & Hu..., 2017 | | | Onthophagus setoculus Krikken & Hu..., 1987 | | | Onthophagus setosus Fahraeus, 1857 | | | Onthophagus sexdentatus Boucomont, 1919 | | | Onthophagus sexstriatus Montrouzier, 1855 | | | Onthophagus sharpi Harold, 1875 | | | Onthophagus shaykh Montanaro & ..., 2022 | | | Onthophagus shibatai Nakane, 1960 | | | Onthophagus shigeo (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus shigeoi Matsumura, 1937 | | | Onthophagus shimba Cambefort, 1980 | | | Onthophagus shinanensis (Matsumura, 1934) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus shinichii Ochi & Kon, 2018 | | | Onthophagus shirakii Nakane, 1960 | | | Onthophagus shoyozanus (Matsumura, 1937) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus shunichii Ochi, Kon & ..., 2021 | | | Onthophagus shurianus Matsumura & ..., 1937 | | | Onthophagus sibela Huijbregts &..., 2012 | | | Onthophagus sibuyanus Boucomont, 1924 | | | Onthophagus sidama Gestro, 1895 | | | Onthophagus sidangeaus Masumoto, Oc..., 2021 | | | Onthophagus sideki Krikken & Hu..., 1987 | | | Onthophagus signaticollis Frey, 1970 | | | Onthophagus signatus Fahraeus, 1857 | | | Onthophagus signifer Harold, 1877 | | | Onthophagus sihkahonoi Huijbregts &..., 2009 | | | Onthophagus sikkimensis Gillet, 1925 | | | Onthophagus silus Balthasar, 1960 | | | Onthophagus simboroni Ochi & Kon, 2006 | | | Onthophagus simillimus d´Orbigny, 1913 | | | Onthophagus simius Reitter, 1893 | | | Onthophagus simoni d´Orbigny, 1902 | | | Onthophagus simplex Raffray, 1877 | | | Onthophagus simpliciceps d´Orbigny, 1905 | | | Onthophagus simplicifrons Reitter, 1893 | | | Onthophagus simulator d´Orbigny, 1905 | | | Onthophagus sinagai Krikken & Hu..., 2008 | | | Onthophagus singulariformis Kohlmann & S..., 2001 | | | Onthophagus sinicus Zhang & Wang, 1997 | | | Onthophagus sinuosicollis d´Orbigny, 1905 | | | Onthophagus sinuosus d´Orbigny, 1913 | | | Onthophagus sipilouensis Cambefort, 1984 | | | Onthophagus sisyphoides Krikken, 1977 | | | Onthophagus sjoestethi d´Orbigny, 1904 | | | Onthophagus skelleyi Sánchez-Huer..., 2018 | | | Onthophagus sloanei Blackburn, 1903 | | | Onthophagus smeenki Cambefort, 1980 | | | Onthophagus smetanai Balthasar, 1952 | | | Onthophagus snoflaki Balthasar, 1944 | | | Onthophagus sobosanus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus socialis Arrow, 1931 | | | Onthophagus solidus (Gillet, 1927) | ⇒ Onthophagus compactus Arrow, 1933 | | Onthophagus solisi Howden & Gill, 1993 | | | Onthophagus solivagus Harold, 1886 | | | Onthophagus solmani Stebnicka, 1975 | | | Onthophagus solomonensis Krikken & Hu..., 2012 | | | Onthophagus somalicola Balthasar, 1941 | | | Onthophagus sonani Miwa, 1930 | | | Onthophagus songsokensis Biswas & Cha..., 1985 | | | Onthophagus sophiae Nicolas, 2006 | | | Onthophagus sopu Krikken & Hu..., 2017 | | | Onthophagus sparsepunctatus Frey, 1956 | | | Onthophagus sparsulus Reitter, 1893 | | | Onthophagus spathatus Boucomont, 1914 | | | Onthophagus spiculatus Boucomont, 1914 | | | Onthophagus spinicornis Gillet, 1930 | | | Onthophagus spinifex Fabricius, 1781 | | | Onthophagus spitsbergeniensis Krell, 2010 | | | Onthophagus spitsbergensis ♰ O. Heer, 1870 | | | Onthophagus splendidoides Josso, 2013 | | | Onthophagus splendidus Boucomont, 1932 | | | Onthophagus sprecherae Moretto, 2013 | | | Onthophagus spurcatus d´Orbigny, 1905 | | | Onthophagus squalidus Lea, 1923 | | | Onthophagus stanleyi Moretto & Gé..., 2010 | | | Onthophagus statzi ♰ Krell, 1990 | | | Onthophagus stehliki Balthasar, 1966 | | | Onthophagus steinheili Harold, 1880 | | | Onthophagus stellio Erichson, 1843 | | | Onthophagus stenocerus (Harold, 1867) | ⇒ Onthophagus neostenocerus Goidanich, 1926 | | Onthophagus sternalis Arrow, 1931 | | | Onthophagus sternax Balthasar, 1959 | | | Onthophagus sticticus Harold, 1867 | | | Onthophagus stictus d´Orbigny, 1913 | | | Onthophagus stigmosus d´Orbigny, 1902 | | | Onthophagus stillatus d´Orbigny, 1904 | | | Onthophagus stockwelli Howden & Young, 1956 | | | Onthophagus stomachosus Krell, 2009 | | | Onthophagus strandi Balthasar, 1935 | | | Onthophagus strictestriatus d´Orbigny, 1913 | | | Onthophagus stuhlmanni d´Orbigny, 1908 | | | Onthophagus subcancer Howden, 1973 | | | Onthophagus subcinctus d´Orbigny, 1913 | | | Onthophagus subcornutus Boucomont, 1914 | | | Onthophagus subdivisus d´Orbigny, 1908 | | | Onthophagus subhumeralis d´Orbigny, 1902 | | | Onthophagus subnudus d´Orbigny, 1902 | | | Onthophagus subocellatus d´Orbigny, 1905 | | | Onthophagus subocelliger (Blackburn, 1903) | ⇒ Onthophagus waterhousei Gillet, 1927 | | Onthophagus subplanus d´Orbigny, 1902 | | | Onthophagus subrugosus d´Orbigny, 1905 | | | Onthophagus subsulcatus d´Orbigny, 1908 | | | Onthophagus subulifer d´Orbigny, 1908 | | | Onthophagus suermelii Petrovitz, 1963 | | | Onthophagus suffusus Klug, 1855 | | | Onthophagus sugihartoi Ochi, 2007 | | | Onthophagus sugillatus Klug, 1855 | | | Onthophagus suginokoichii Ochi & Kon, 2008 | | | Onthophagus suillus Arrow, 1931 | | | Onthophagus sulawesiensis Krikken & Hu..., 2011 | | | Onthophagus sulawesijohkii Ochi, Kon & ..., 2016 | | | Onthophagus sulcatulus d´Orbigny, 1907 | | | Onthophagus sulci Balthasar, 1935 | | | Onthophagus sulcipennis d´Orbigny, 1902 | | | Onthophagus sumatramontanus Ochi & Kon, 2008 | | | Onthophagus sumatranus Lansberge, 1883 | | | Onthophagus sumptuosus Paulian, 1937 | | | Onthophagus sundanensis Lansberge, 1883 | | | Onthophagus surdus Boucomont, 1925 | | | Onthophagus susterai Balthasar, 1952 | | | Onthophagus sutiliceps d´Orbigny, 1902 | | | Onthophagus sutleinensis Splichal, 1910 | | | Onthophagus suturalis (Péringuey, 1888) | ⇒ Onthophagus trituber Wiedemann, 1823 | | Onthophagus sycophanta (Fairmaire, 1887) | ⇒ Euonthophagus gibbosus Scriba, 1790 | | Onthophagus sydneyensis Blackburn, 1903 | | | Onthophagus sylvestris Walter & Cam..., 1977 | | | Onthophagus sylvipapuanus Krikken & Hu..., 2012 | | | Onthophagus symbioticus Arrow, 1920 | | | Onthophagus synceri Cambefort, 1984 | | | Onthophagus tabellicornis MacLeay, 1864 | | | Onthophagus tabellifer Gillet, 1927 | | | Onthophagus tabidus Balthasar, 1935 | | | Onthophagus taboranus d´Orbigny, 1902 | | | Onthophagus tacanensis Chamé-Vázque..., 2022 | | | Onthophagus taeniatus Boucomont, 1914 | | | Onthophagus tagal Boucomont, 1924 | | | Onthophagus taiensis Cambefort, 1984 | | | Onthophagus takabayashii Matsumura, 1937 | | | Onthophagus takaoi Ochi & Kon, 2014 | | | Onthophagus talpa Fahraeus, 1857 | | | Onthophagus tambing Krikken & Hu..., 2011 | | | Onthophagus tamworthi Blackburn, 1903 | | | Onthophagus tanganus d´Orbigny, 1913 | | | Onthophagus taoi Ochi & Kon, 2006 | | | Onthophagus tapirus Sharp, 1877 | | | Onthophagus taprobanus Arrow, 1931 | | | Onthophagus tarascus Zunino & Hal..., 1988 | | | Onthophagus tarascus jaliscensis Zunino & Hal..., 1988 | | | Onthophagus tarasovi Krikken & Hu..., 2012 | | | Onthophagus tarsius Arrow, 1941 | | | Onthophagus tauroides Gillet, 1930 | | | Onthophagus taxillus Balthasar, 1969 | | | Onthophagus teitanicus d´Orbigny, 1902 | | | Onthophagus telegonus Balthasar, 1969 | | | Onthophagus telephus Balthasar, 1969 | | | Onthophagus temporalis d´Orbigny, 1902 | | | Onthophagus tenax Balthasar, 1964 | | | Onthophagus tenebrosus Harold, 1871 | | | Onthophagus tenuigraniger d´Orbigny, 1913 | | | Onthophagus tenuistriatus d´Orbigny, 1905 | | | Onthophagus terrara Storey, 1977 | | | Onthophagus tersicollis Müller, 1947 | | | Onthophagus tersipennis d´Orbigny, 1902 | | | Onthophagus tersus d´Orbigny, 1913 | | | Onthophagus tesseratus d´Orbigny, 1908 | | | Onthophagus tessulatus Harold, 1871 | | | Onthophagus testaceoviolaceus Paulian, 1937 | | | Onthophagus tetricus Harold, 1877 | | | Onthophagus thailaevis Masumoto, Oc..., 2008 | | | Onthophagus thainuaensis Masumoto, Oc..., 2008 | | | Onthophagus tharalithae Karimbumkara..., 2016 | | | Onthophagus thomsoni Bates, 1888 | | | Onthophagus thoreyi Harold, 1868 | | | Onthophagus thungphraensis Masumoto, Oc..., 2020 | | | Onthophagus tibetanus Arrow, 1907 | | | Onthophagus timorensis Boucomont, 1914 | | | Onthophagus tiniocelloides Boucomont, 1925 | | | Onthophagus tirapensis Biswas & Cha..., 1985 | | | Onthophagus tnai Nithya & Sabu, 2012 | | | Onthophagus togeman Matthews, 1972 | | | Onthophagus tonkineus Gillet, 1921 | | | Onthophagus tonsus d´Orbigny, 1902 | | | Onthophagus tonywhitteni Krikken & Hu..., 2011 | | | Onthophagus toopi Monteith & S..., 2013 | | | Onthophagus toraut Krikken & Hu..., 2011 | | | Onthophagus totonacus Moctezuma & ..., 2021 | | | Onthophagus totonicapamus Bates, 1887 | | | Onthophagus toxopeus Krikken & Hu..., 2012 | | | Onthophagus traginus Frey, 1972 | | | Onthophagus tragoides Boucomont, 1914 | | | Onthophagus transcaspicus Koenig, 1888 | Gazella scarab | | Onthophagus transisthmius Howden & Young, 1956 | | | Onthophagus transvestitus Huijbregts &..., 2009 | | | Onthophagus trapezicornis d´Orbigny, 1902 | | | Onthophagus traversii d´Orbigny, 1904 | | | Onthophagus trawalla Storey & Weir, 1990 | | | Onthophagus triacanthus Castelnau, 1840 | | | Onthophagus tricariniger d´Orbigny, 1902 | | | Onthophagus tricavicollis Lea, 1923 | | | Onthophagus triceratops Arrow, 1913 | | | Onthophagus trichopygus d´Orbigny, 1905 | | | Onthophagus tricolor Boucomont, 1914 | | | Onthophagus tricorniger Boheman, 1860 | | | Onthophagus tridenticeps d´Orbigny, 1902 | | | Onthophagus tridentitibialis Ochi & Kon, 2008 | | | Onthophagus trifidisetis d´Orbigny, 1905 | | | Onthophagus triimpressus d´Orbigny, 1905 | | | Onthophagus trinodosus Fahraeus, 1857 | | | Onthophagus trinominatus Goidanich, 1926 | | | Onthophagus tripartitus d´Orbigny, 1902 | | | Onthophagus tripolitanus Heyden, 1890 | | | Onthophagus triptolemus Balthasar, 1969 | | | Onthophagus tristis Harold, 1873 | | | Onthophagus triundulatus Balthasar, 1964 | | | Onthophagus trochilus Arrow, 1931 | | | Onthophagus troglodyta Wiedemann, 1823 | | | Onthophagus troniceki Balthasar, 1933 | | | Onthophagus tschadensis Balthasar, 1963 | | | Onthophagus tschoffeni d´Orbigny, 1904 | | | Onthophagus tshuapae Balthasar, 1964 | | | Onthophagus tsubakii Ochi & Kon, 2009 | | | Onthophagus tsutomui Ochi & Kon, 2016 | | | Onthophagus tubericollis (Raffray, 1877) | ⇒ Onthophagus bidens Olivier, 1789 | | Onthophagus tuckonie Matthews, 1972 | | | Onthophagus tumidulus Gerstaecker, 1871 | | | Onthophagus tungkamangensis Masumoto, Oc..., 2007 | | | Onthophagus tungkamungensis Masumoto, Oc..., 2008 | | | Onthophagus turbatus (Walker, 1858) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus turfanicus Balthasar, 1971 | | | Onthophagus turgidus Kohlmann & S..., 2012 | | | Onthophagus turneri Boucomont, 1936 | | | Onthophagus turpidus Reitter, 1887 | | | Onthophagus turrbal Matthews, 1972 | | | Onthophagus tuzetae Walter & Cam..., 1977 | | | Onthophagus tweedensis Blackburn, 1903 | | | Onthophagus ubaidillahi Krikken & Hu..., 2012 | | | Onthophagus uedai Ochi & Kon, 2006 | | | Onthophagus uedanus (Matsumura, 1937) | ⇒ Onthophagus olsoufieffi Boucomont, 1924 | | Onthophagus uelensis Frey, 1960 | | | Onthophagus ugandaensis Dierkens, 2022 | | | Onthophagus ugandicus Frey, 1975 | | | Onthophagus ugoi Moretto & Gé..., 2020 | | | Onthophagus ukerewensis Balthasar, 1937 | | | Onthophagus ulugombakensis Ochi, Kon & ..., 2014 | | | Onthophagus ulula Balthasar, 1966 | | | Onthophagus umbilicatus d´Orbigny, 1908 | | | Onthophagus umbilicopunctatus (Medvedev, 1959) | ⇒ Onthophagus trispinus Reitter, 1892 | | Onthophagus umbratus d´Orbigny, 1902 | | | Onthophagus undaticeps d´Orbigny, 1902 | | | Onthophagus undulans Bates, 1887 | | | Onthophagus undulans oaxacanus Zunino & Hal..., 1988 | | | Onthophagus unicarina d´Orbigny, 1904 | | | Onthophagus unidentatus Boucomont, 1927 | | | Onthophagus unifasciatus Schaller, 1783 | | | Onthophagus uniformis Heyden, 1886 | | | Onthophagus urellus Boucomont, 1919 | | | Onthophagus ursinus d´Orbigny, 1902 | | | Onthophagus ursus Boucomont, 1926 | | | Onthophagus urus (Harold, 1867) | ⇒ Onthophagus urusheeri Krell, 2000 | | Onthophagus urusheeri ♰ Krell, 2000 | | | Onthophagus usambaricus Paulian, 1937 | | | Onthophagus ushiodai (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus usurpator Balthasar, 1959 | | | Onthophagus valentineae Josso, 2019 | | | Onthophagus vanasseni Krikken & Hu..., 2008 | | | Onthophagus vanderblomi Krikken & Hu..., 2008 | | | Onthophagus vaneyeni Frey, 1960 | | | Onthophagus vanofwegeni Krikken & Hu..., 2008 | | | Onthophagus varianus Lea, 1923 | | | Onthophagus variatus d´Orbigny, 1902 | | | Onthophagus variegatus (Fabricius, 1798) | ⇒ Onthophagus filicornis Harold, 1873 | | Onthophagus variegranosus d´Orbigny, 1905 | | | Onthophagus variolaris Lansberge, 1883 | | | Onthophagus variolicollis Lea, 1923 | | | Onthophagus variolosus d´Orbigny, 1902 | | | Onthophagus varius d´Orbigny, 1913 | | | Onthophagus vassei d´Orbigny, 1908 | | | Onthophagus vatovai Müller, 1942 | | | Onthophagus velliger d´Orbigny, 1907 | | | Onthophagus ventralis Lansberge, 1883 | | | Onthophagus ventrosus d´Orbigny, 1905 | | | Onthophagus venustulus Erichson, 1843 | | | Onthophagus venzoi Balthasar, 1941 | | | Onthophagus veracruzensis Delgado & Pe..., 1998 | | | Onthophagus verae Josso, 2013 | | | Onthophagus vermiculatus Frey, 1970 | | | Onthophagus verrucosus d´Orbigny, 1902 | | | Onthophagus versutus Péringuey, 1901 | | | Onthophagus verticalis (Fahraeus, 1857) | ⇒ Onthophagus praedatus Harold, 1862 | | Onthophagus vesanus Balthasar, 1967 | | | Onthophagus vespertilio Howden, Cart..., 1956 | | | Onthophagus vespertilis | | | Onthophagus vestitus d´Orbigny, 1902 | | | Onthophagus vethi Krikken, 1977 | | | Onthophagus vicinus (Harold, 1886) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus victoriensis Blackburn, 1903 | | | Onthophagus vigens Péringuey, 1901 | | | Onthophagus vilis Harold, 1877 | | | Onthophagus villanuevai Delgado & De..., 1990 | | | Onthophagus villosus MacLeay, 1888 | | | Onthophagus vinctoides Frey, 1957 | | | Onthophagus vinctus Erichson, 1843 | | | Onthophagus violaceotinctus Gillet, 1925 | | | Onthophagus violaceoviridis Paulian, 1937 | | | Onthophagus violetae Zunino & Hal..., 1997 | | | Onthophagus virens (d´Orbigny, 1913) | ⇒ Onthophagus alcyonides d´Orbigny, 1913 | | Onthophagus virescens Harold, 1867 | | | Onthophagus viridiaereus d´Orbigny, 1908 | | | Onthophagus viridicatus d´Orbigny, 1902 | | | Onthophagus viridichevrolati Moctezuma & ..., 2020 | | | Onthophagus viridiperakensis Ochi, Kon & ..., 2014 | | | Onthophagus viriditinctus Reitter, 1893 | | | Onthophagus viridivinosus Kohlmann & S..., 2001 | | | Onthophagus viviensis d´Orbigny, 1905 | | | Onthophagus vladimiri Frey, 1957 | | | Onthophagus volsellatus (Boucomont, 1920) | ⇒ Onthophagus minax d´Orbigny, 1904 | | Onthophagus volucer Balthasar, 1959 | | | Onthophagus vuattouxi Cambefort, 1984 | | | Onthophagus vulpes Harold, 1877 | | | Onthophagus vulpinaris Schönfeldt, 1906 | | | Onthophagus vulpinus Arrow, 1931 | | | Onthophagus vultuosus d´Orbigny, 1902 | | | Onthophagus vultur Arrow, 1931 | | | Onthophagus vylderi d´Orbigny, 1913 | | | Onthophagus wagamen Matthews, 1972 | | | Onthophagus wakelbura Matthews, 1972 | | | Onthophagus wallacei Harold, 1871 | | | Onthophagus walteri Josso & Prév..., 2000 | | | Onthophagus waminda Matthews, 1972 | | | Onthophagus wanappe Storey, 1977 | | | Onthophagus wangnamkieous Masumoto, Oc..., 2008 | | | Onthophagus wangtaichieni Masumoto, Oc..., 2020 | | | Onthophagus waterhousei Gillet, 1927 | | | Onthophagus waterloti d´Orbigny, 1908 | | | Onthophagus waterstradti Boucomont, 1914 | | | Onthophagus watuwila Krikken & Hu..., 2008 | | | Onthophagus wayaua Huijbregts &..., 2012 | | | Onthophagus weisei (Reitter, 1891) | ⇒ Euonthophagus gibbosus Scriba, 1790 | | Onthophagus wensis Josso & Prév..., 2006 | | | Onthophagus weringerong Storey & Weir, 1990 | | | Onthophagus wiebesi Krikken, 1977 | | | Onthophagus wigmungan Matthews, 1972 | | | Onthophagus wilgi Matthews, 1972 | | | Onthophagus willameorum Walter & Cam..., 1977 | | | Onthophagus williamsi Storey & Weir, 1990 | | | Onthophagus wittei (Frey, 1958) | ⇒ Onthophagus witteianus Krell, 2009 | | Onthophagus witteianus Krell, 2009 | | | Onthophagus wombalano Matthews, 1972 | | | Onthophagus woroae Ochi & Kon, 2006 | | | Onthophagus woroae bruneiensis Ochi, Kon & ..., 2009 | | | Onthophagus woroae woroae | | | Onthophagus worooa Storey & Weir, 1990 | | | Onthophagus xanthochlorus Walter & Cam..., 1977 | | | Onthophagus xanthomerus Bates, 1887 | | | Onthophagus xanthopterus d´Orbigny, 1908 | | | Onthophagus xanthopygus d´Orbigny, 1908 | | | Onthophagus xiphias (Solis & Kohl..., 2003) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus yackatoon Storey & Weir, 1990 | | | Onthophagus yakuinsulanus Nakane, 1984 | | | Onthophagus yangmunensis Masumoto & Ochi, 2015 | | | Onthophagus yanoi Matsumura, 1938 | | | Onthophagus yaoi Masumoto, Oc..., 2014 | | | Onthophagus yaran Storey & Weir, 1990 | | | Onthophagus yarrumba Storey, 1977 | | | Onthophagus yasuhikoi Masumoto, Oc..., 2012 | | | Onthophagus yescaensis Moctezuma, H..., 2022 | | | Onthophagus yeyeko Matthews, 1972 | | | Onthophagus yifer Krikken & Hu..., 2012 | | | Onthophagus yiryoront Matthews, 1972 | | | Onthophagus yourula Storey & Weir, 1990 | | | Onthophagus yucatanus Delgado-Cast..., 2006 | | | Onthophagus yugianus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus yujii Masumoto, Oc..., 2012 | | | Onthophagus yumigatanus Matsumura, 1937 | | | Onthophagus yumotoi Ochi & Kon, 2006 | | | Onthophagus yungaburra Matthews, 1972 | | | Onthophagus yunkara Matthews, 1972 | | | Onthophagus zagrosicus Kabakov, 2006 | | | Onthophagus zairensis Walter & Cam..., 1977 | | | Onthophagus zambundaticeps Josso, 2022 | | | Onthophagus zapotecus Zunino & Hal..., 1988 | | | Onthophagus zavattarii Müller, 1947 | | | Onthophagus zavreli Balthasar, 1935 | | | Onthophagus zebra Arrow, 1931 | | | Onthophagus zebu Boucomont, 1921 | | | Onthophagus zicsii Balthasar, 1967 | | | Onthophagus zimmermanni Balthasar, 1959 | | | Onthophagus zinovskyi Qaryagdy, 1939 | | | Onthophagus zumpti Frey, 1954 | | | Onthophagus zuvandi Qaryagdy, 1939 | | | Onthophagus zymoticus Moxey, 1963 | | | Onychothecus ateuchoides Boucomont, 1912 | | | Onychothecus corniclypeus Zhang, 1980 | | | Onychothecus riekoae Ochi & Kon, 1998 | | | Onychothecus tridentigeris Zelenka, 1992 | | | Oruscatus davus Erichson, 1847 | | | Oruscatus opalescens Bates, 1870 | | | Outenikwanus tomentosus Scholtz & Ho..., 1987 | | | Oxysternon aeneum (Olsoufieff, 1924) | ⇒ Oxysternon silenus Castelnau, 1840 | | Oxysternon conspicillatum Weber, 1801 | | | Oxysternon durantoni Arnaud, 1984 | | | Oxysternon ebeninum Nevinson, 1890 | | | Oxysternon festivum Linnaeus, 1758 | | | Oxysternon festivum festivum | | | Oxysternon festivum nigerrimum Arnaud, 2002 | | | Oxysternon lautum MacLeay, 1819 | | | Oxysternon macleayi Nevison, 1892 | | | Oxysternon oberthuri (Olsoufieff, 1924) | ⇒ Oxysternon conspicillatum Weber, 1801 | | Oxysternon palaemon Castelnau, 1840 | | | Oxysternon pteroderum Nevinson, 1892 | | | Oxysternon sericeum (Olsoufieff, 1924) | ⇒ Oxysternon silenus Castelnau, 1840 | | Oxysternon silenum Castelnau, 1840 | | | Oxysternon silenus Castelnau, 1840 | | | Oxysternon spiniferum Castelnau, 1840 | | | Oxysternon striatopunctatum Olsoufieff, 1924 | | | Pachylomera femoralis Kirby, 1828 | | | Pachylomera opaca Lansberge, 1874 | | | Pachysoma aesculapius Olivier, 1789 | | | Pachysoma barbatus | | | Pachysoma bennigseni Felsche, 1907 | | | Pachysoma denticolle Péringuey, 1888 | | | Pachysoma denticollis Péringuey, 1888 | | | Pachysoma endroedyi Harrison, Sc..., 2003 | | | Pachysoma fitzsimonsi Ferreira, 1953 | | | Pachysoma gariepinum Ferreira, 1953 | | | Pachysoma gariepinus Ferreira, 1953 | | | Pachysoma glentoni Harrison, Sc..., 2003 | | | Pachysoma hippocrates MacLeay, 1821 | | | Pachysoma rodriguesi Ferreira, 1953 | | | Pachysoma rotundigena (Felsche, 1907) | ⇒ Scarabaeus rotundigenus Felsche, 1907 | | Pachysoma rotundigenus Felsche, 1907 | | | Pachysoma schinzi Fairmaire, 1888 | | | Pachysoma striata | | | Pachysoma striatum (Castelnau, 1840) | ⇒ Scarabaeus striatus Castelnau, 1840 | | Pachysoma striatus Castelnau, 1840 | | | Pachysoma valeflorae Ferreira, 1953 | | | Panelus assamensis Arrow, 1907 | | | Panelus bakeri Boucomont, 1921 | | | Panelus besucheti Paulian, 1980 | | | Panelus bidentata Wilson, 1922 | | | Panelus borneensis Paulian, 1992 | | | Panelus brendelli Ochi, Kon & ..., 2009 | | | Panelus ceylonicus Balthasar, 1972 | | | Panelus crenatus Nomura, 1973 | | | Panelus danumensis Ochi, Kon & ..., 2009 | | | Panelus deuvei Paulian, 1992 | | | Panelus fallax Balthasar, 1972 | | | Panelus fujitai Ochi, Kon & ..., 2008 | | | Panelus horni | | | Panelus imitator Balthasar, 1972 | | | Panelus jaccoudi Paulian, 1980 | | | Panelus kalimantanicus Ochi, Kon & ..., 2009 | | | Panelus karasuyamai Ochi, Kon & ..., 2008 | | | Panelus keralai Paulian, 1980 | | | Panelus kubotai Kawahara, In..., 2007 | | | Panelus maedai Nomura, 1973 | | | Panelus manasi Paulian, 1983 | | | Panelus manmiaoae Masumoto, Oc..., 2011 | | | Panelus mendeli Ochi, Kon & ..., 2009 | | | Panelus minutus Boucomont, 1914 | | | Panelus mussardi Paulian, 1980 | | | Panelus orousseti Paulian, 1981 | | | Panelus ovatus Nomura, 1973 | | | Panelus parvulus Waterhouse, 1874 | | | Panelus pernitidus Balthasar, 1972 | | | Panelus pisoniae (Lea, 1923) | ⇒ Lepanus pisoniae Lea, 1923 | | Panelus puncticollis Arrow, 1931 | | | Panelus quadridentatus Balthasar, 1952 | | | Panelus rhodesiensis Paulian, 1975 | | | Panelus rufocuprea | | | Panelus rufulus Nomura, 1973 | | | Panelus sabahi Paulian, 1992 | | | Panelus saigusai Ochi & Kon, 2016 | | | Panelus setosus Arrow, 1931 | | | Panelus sumatranus Lansberge, 1885 | | | Panelus sumatrensis Ochi, Kon & ..., 1998 | | | Panelus tonkinensis Paulian, 1939 | | | Panelus wangi Masumoto, Oc..., 2011 | | | Panelus watanabei Masumoto & Yin, 1993 | | | Panelus whitehousei | | | Panelus wiebesi | | | Paracanthon arnaudi Pacheco & Va..., 2020 | | | Paracanthon belloi Pacheco & Va..., 2020 | | | Paracanthon felipei Pacheco & Va..., 2020 | | | Paracanthon genieri Pacheco & Va..., 2020 | | | Paracanthon grossiorum Pacheco & Va..., 2020 | | | Paracanthon hirsutus Paulian, 1939 | | | Paracanthon laevinotus Pacheco & Va..., 2020 | | | Paracanthon lopesandradei Pacheco & Va..., 2020 | | | Paracanthon marinezae Pacheco & Va..., 2020 | | | Paracanthon millerorum Pacheco & Va..., 2020 | | | Paracanthon monteiroorum Pacheco & Va..., 2020 | | | Paracanthon muriloi Pacheco & Va..., 2020 | | | Paracanthon ocellatopunctatus Pacheco & Va..., 2020 | | | Paracanthon pereirai Andretta & M..., 1957 | | | Paracanthon rosinae Balthasar, 1942 | | | Paracanthon trichonotulus Balthasar, 1938 | | | Parachorius asymmetricus Tarasov, 2017 | | | Parachorius bolavensis Tarasov, 2017 | | | Parachorius fukiensis Balthasar, 1960 | | | Parachorius fungorum Kryzhanovsky..., 1966 | | | Parachorius globosus Arrow, 1931 | | | Parachorius gotoi Masumoto, 1986 | | | Parachorius hookeri Arrow, 1931 | | | Parachorius humeralis Arrow, 1907 | | | Parachorius javanus Boucomont, 1914 | | | Parachorius krali Utsunomiya &..., 2001 | | | Parachorius lannathai Hanboonsong ..., 2001 | | | Parachorius longipenis Tarasov, 2017 | | | Parachorius maruyamai Masumoto, Oc..., 2012 | | | Parachorius matsudai Ochi, Pham &..., 2016 | | | Parachorius newthayerae Tarasov, 2017 | | | Parachorius nudus Sharp, 1875 | | | Parachorius peninsularis Arrow, 1907 | | | Parachorius pseudojavanus Tarasov, 2017 | | | Parachorius schuelkei Tarasov, 2017 | | | Parachorius semsanganus Tarasov & Keith, 2011 | | | Parachorius solodovnikovi Tarasov, 2017 | | | Parachorius thomsoni Harold, 1873 | | | Paracryptocanthon borgmeieri Vulcano & Pe..., 1976 | | | Paracryptocanthon genieri Pacheco & Va..., 2017 | | | Paragymnopleurus ambiguus Janssens, 1945 | | | Paragymnopleurus brahminus Waterhouse, 1890 | | | Paragymnopleurus martinezi Balthasar, 1955 | | | Paragymnopleurus maurus Sharp, 1875 | | | Paragymnopleurus melanarius Harold, 1867 | | | Paragymnopleurus planus Sharp, 1875 | | | Paragymnopleurus rudis Sharp, 1875 | | | Paragymnopleurus sinuatus Olivier, 1789 | | | Paragymnopleurus sinuatus abax Sharp, 1875 | | | Paragymnopleurus sinuatus assamensis Waterhouse, 1890 | | | Paragymnopleurus sinuatus productus Sharp, 1875 | | | Paragymnopleurus sinuatus sinuatus | | | Paragymnopleurus sinuatus szechouanicus Balthasar, 1934 | | | Paragymnopleurus sparsus Sharp, 1875 | | | Paragymnopleurus sparsus arnoldi Krikken & Hu..., 1987 | | | Paragymnopleurus sparsus javanus Krikken & Hu..., 1987 | | | Paragymnopleurus sparsus palawanicus Ochi & Kon, 1997 | | | Paragymnopleurus sparsus sparsus | | | Paragymnopleurus spinotus Boucomont, 1914 | | | Paragymnopleurus stipes Sharp, 1875 | | | Paragymnopleurus striatus Sharp, 1875 | | | Paragymnopleurus sumatrensis Ochi & Kon, 1997 | | | Paraphacosomoides curvipes Balthasar, 1966 | | | Paraphytus africanus Boucomont, 1923 | | | Paraphytus andamanus Arrow, 1931 | | | Paraphytus andoi Ochi & Kon, 2015 | | | Paraphytus aphodioides Boucomont, 1928 | | | Paraphytus bechynei Balthasar, 1958 | | | Paraphytus dentifrons Lewis, 1895 | | | Paraphytus doriae Harold, 1877 | | | Paraphytus foveatus Boucomont, 1923 | | | Paraphytus harmandi Boucomont, 1923 | | | Paraphytus hindu Arrow, 1913 | | | Paraphytus ritsemae Harold, 1880 | | | Paraphytus sancyi Paulian, 1949 | | | Paraphytus sumatranus Paulian & Ca..., 1981 | | | Paraphytus tai Cambefort, 1984 | | | Paroniticellus festivus Steven, 1809 | | | Paronitis excavaticollis Balthasar, 1942 | | | Paronthobium adio Montreuil & ..., 2016 | | | Paronthobium caledonicum Paulian, 1935 | | | Paronthobium daghfousi Montreuil & ..., 2016 | | | Paronthobium dierkensi Montreuil & ..., 2016 | | | Paronthobium farino Montreuil & ..., 2016 | | | Paronthobium fauveli Paulian, 1984 | | | Paronthobium iac Montreuil & ..., 2016 | | | Paronthobium julieni Montreuil & ..., 2016 | | | Paronthobium memaoya Montreuil & ..., 2016 | | | Paronthobium micros Paulian, 1984 | | | Paronthobium minutum Montreuil & ..., 2016 | | | Paronthobium monteithi Paulian, 1987 | | | Paronthobium muticum Fauvel, 1903 | | | Paronthobium nasutum Montreuil & ..., 2016 | | | Paronthobium orientale Montreuil & ..., 2016 | | | Paronthobium peckorum Montreuil & ..., 2016 | | | Paronthobium petchecara Montreuil & ..., 2016 | | | Paronthobium poirieri Montreuil, 2021 | | | Paronthobium simplex Fauvel, 1903 | | | Paronthobium subdentatum Montreuil & ..., 2016 | | | Paronthobium taom Montreuil & ..., 2016 | | | Paronthobium tchingou Montreuil & ..., 2016 | | | Paronthobium tibiale Fauvel, 1903 | | | Parvuhowdenius harrisoni Deschodt & C..., 2008 | | | Peckolus alpinus Howden & Sch..., 1988 | | | Peckolus parvus Scholtz & Ho..., 1987 | | | Peckolus poenskopius Deschodt & C..., 2008 | | | Pedaria acuta Josso & Prév..., 2015 | | | Pedaria aethiopica Josso & Prév..., 2003 | | | Pedaria alternans Waterhouse, 1890 | | | Pedaria alternata (Lea, 1923) | ⇒ Demarziella alternata Lea, 1923 | | Pedaria angolana Boucomont, 1922 | | | Pedaria antoinei Josso & Prév..., 2009 | | | Pedaria armata Raffray, 1877 | | | Pedaria aspera Péringuey, 1901 | | | Pedaria barrei Josso & Prév..., 2006 | | | Pedaria biseria Josso & Prév..., 2003 | | | Pedaria brancoi Josso & Prév..., 2003 | | | Pedaria cambeforti Josso & Prév..., 2003 | | | Pedaria catherinae Josso & Prév..., 2009 | | | Pedaria cicatricosus (Péringuey, 1901) | ⇒ Sarophorus cicatricosus Péringuey, 1901 | | Pedaria conformis Péringuey, 1901 | | | Pedaria coprinarum Cambefort, 1982 | | | Pedaria criberrima Waterhouse, 1890 | | | Pedaria cylindrica Fahraeus, 1857 | | | Pedaria decorsei (Boucomont, 1922) | ⇒ Pedaria taylori Waterhouse, 1890 | | Pedaria dellacasai Josso & Prév..., 2009 | | | Pedaria demeyeri Josso & Prév..., 2003 | | | Pedaria dentata Raffray, 1877 | | | Pedaria dudleyi Josso & Prév..., 2003 | | | Pedaria durandi Paulian, Cam..., 1982 | | | Pedaria estellae Josso & Prév..., 2006 | | | Pedaria fernandezi Cambefort, 1982 | | | Pedaria genieri Josso & Prév..., 2003 | | | Pedaria genierorum Josso & Prév..., 2015 | | | Pedaria gigantea Josso & Prév..., 2003 | | | Pedaria granulosa Ferreira, 1962 | | | Pedaria hanae Balthasar, 1963 | | | Pedaria humana Cambefort, 1982 | | | Pedaria insularis Kolbe, 1914 | | | Pedaria intermedia Boucomont, 1922 | | | Pedaria jacksoni Waterhouse, 1890 | | | Pedaria jossoi Génier, 2017 | | | Pedaria juhellegrandi Josso & Prév..., 2003 | | | Pedaria manni Josso & Prév..., 2015 | | | Pedaria minetti Josso & Prév..., 2009 | | | Pedaria minima Josso & Prév..., 2003 | | | Pedaria montreuili Josso & Prév..., 2015 | | | Pedaria morettoi Josso & Prév..., 2003 | | | Pedaria murphyi Josso & Prév..., 2003 | | | Pedaria murphyi murphyi | | | Pedaria murphyi septentrionalis Josso & Prév..., 2009 | | | Pedaria nigra Laporte, 1832 | | | Pedaria oblonga Josso & Prév..., 2003 | | | Pedaria ouangoensis Cambefort, 1982 | | | Pedaria ovata Boucomont, 1922 | | | Pedaria picea Fahraeus, 1857 | | | Pedaria puncticollis Waterhouse, 1890 | | | Pedaria quirimbasensis Josso & Prév..., 2015 | | | Pedaria raffrayi Josso & Prév..., 2006 | | | Pedaria reichenbachi Josso & Prév..., 2006 | | | Pedaria renwarti Cambefort, 1982 | | | Pedaria reticulata Josso & Prév..., 2015 | | | Pedaria rohani Boucomont, 1922 | | | Pedaria rosemariae Josso & Prév..., 2009 | | | Pedaria segregis Péringuey, 1901 | | | Pedaria segrestis | | | Pedaria smrzi Josso & Prév..., 2003 | | | Pedaria sobrina (Péringuey, 1901) | ⇒ Pedaria picea Fahraeus, 1857 | | Pedaria spinipennis Josso & Prév..., 2003 | | | Pedaria spinithorax Paulian, Cam..., 1982 | | | Pedaria spinosa Boucomont, 1922 | | | Pedaria sudrei Josso & Prév..., 2003 | | | Pedaria tanganyika Josso & Prév..., 2003 | | | Pedaria taylori Waterhouse, 1890 | | | Pedaria tenebrosa Balthasar, 1937 | | | Pedaria tibialis Josso & Prév..., 2006 | | | Pedaria tuberculigera Waterhouse, 1890 | | | Pedaria walteri Josso & Prév..., 2003 | | | Pedaria werneri Josso & Prév..., 2003 | | | Pedaria wittei Janssens, 1939 | | | Pedaridium almeidai (Pereira, 1946) | ⇒ Pereiraidium almeidai Pereira, 1946 | | Pedaridium argentinum (Arrow, 1913) | ⇒ Nunoidium argentinum Arrow, 1913 | | Pedaridium bordoni (Martínez, 1992) | ⇒ Genieridium bordoni Martínez, 1992 | | Pedaridium brasiliensis (Ferreira & G..., 1993) | ⇒ Genieridium bidens Balthasar, 1938 | | Pedaridium caingua (Martínez, 1974) | ⇒ Trichillidium caingua Martínez, 1974 | | Pedaridium hirsutum Harold, 1859 | | | Pedaridium mansosotoi (Martínez, 1951) | ⇒ Genieridium cryptops Arrow, 1913 | | Pedaridium margaretae Génier, 2002 | | | Pedaridium maya (Vaz-de-Mello..., 2004) | ⇒ Martinezidium maya Vaz-de-Mello..., 2004 | | Pedaridium pilosum (Robinson, 1948) | ⇒ Trichillidium pilosum Robinson, 1948 | | Penalus waui Paulian, 1985 | | | Pereiraidium almeidai Pereira, 1946 | | | Phacosoma fallacilaetum (Masumoto, 1989) | ⇒ Ochicanthon thai Paulian, 1987 | | Phacosoma thailandicum (Masumoto, 1989) | ⇒ Ochicanthon thailandicum Masumoto, 1989 | | Phaedotrogus ceylonicus Balthasar, 1972 | | | Phalops ardea Klug, 1855 | | | Phalops aurifrons Fairmaire, 1884 | | | Phalops barbicornis Lansberge, 1883 | | | Phalops batesi Koenike, 1892 | | | Phalops batesi batesi | | | Phalops batesi pseudobatesi Moretto, 2007 | | | Phalops beccarii Harold, 1875 | | | Phalops boschas Klug, 1855 | | | Phalops bubalus Harold, 1867 | | | Phalops candezei Lansberge, 1883 | | | Phalops chalybeus Klug, 1845 | | | Phalops cyanescens d´Orbigny, 1897 | | | Phalops divisus Wiedemann, 1823 | | | Phalops djuricus Kuntzen, 1913 | | | Phalops dregei Harold, 1867 | | | Phalops faccai Müller, 1941 | | | Phalops fimbriatus Klug, 1835 | | | Phalops flavocinctus Klug, 1855 | | | Phalops gallanus d´Orbigny, 1904 | | | Phalops glauningi Kuntzen, 1913 | | | Phalops guttulatus Fairmaire, 1887 | | | Phalops inermis Lansberge, 1883 | | | Phalops iphis Olivier, 1789 | | | Phalops laminifrons Fairmaire, 1882 | | | Phalops lutatus d´Orbigny, 1902 | | | Phalops olivaceus Lansberge, 1883 | | | Phalops pauliani Barbero & Pa..., 2003 | | | Phalops plancus Erichson, 1843 | | | Phalops prasinus (Erichson, 1843) | ⇒ Phalops beccarii Harold, 1875 | | Phalops princeps d´Orbigny, 1908 | | | Phalops pyroides d´Orbigny, 1908 | | | Phalops rufosignatus (Lansberge, 1885) | ⇒ Phalops dregei Harold, 1867 | | Phalops sinuaticeps d´Orbigny, 1902 | | | Phalops smaragdinus Harold, 1875 | | | Phalops sulcatus Lansberge, 1883 | | | Phalops tricuspis d´Orbigny, 1908 | | | Phalops tuberosus d´Orbigny, 1905 | | | Phalops vanellus Lansberge, 1883 | | | Phalops wittei Harold, 1867 | | | Phalops zuninoi Barbero & Pa..., 2003 | | | Phanaeus (Notiophanaeus) achilles Boheman, 1858 | | | Phanaeus achilles achilles | | | Phanaeus achilles lydiae Arnaud, 2000 | | | Phanaeus (Notiophanaeus) bispinus Bates, 1868 | | | Phanaeus (Notiophanaeus) bordoni Arnaud, 1997 | | | Phanaeus (Notiophanaeus) cambeforti Arnaud, 1982 | | | Phanaeus (Notiophanaeus) chalcomelas Perty, 1830 | | | Phanaeus chalcomelas chalcomelas | | | Phanaeus chalcomelas grossii Arnaud, 2001 | | | Phanaeus (Notiophanaeus) dejeani Harold, 1868 | | | Phanaeus (Notiophanaeus) endymion Harold, 1863 | | | Phanaeus endymion porioni Arnaud, 2001 | | | Phanaeus (Notiophanaeus) halffterorum Edmonds, 1979 | | | Phanaeus (Notiophanaeus) haroldi Kirsch, 1871 | | | Phanaeus (Notiophanaeus) kirbyi Vigors, 1825 | | | Phanaeus kirbyi kirbyi | | | Phanaeus kirbyi ledezmai Arnaud, 2002 | | | Phanaeus (Notiophanaeus) lecourti Arnaud, 2000 | | | Phanaeus lecourti lecourti | | | Phanaeus lecourti peruanus Arnaud, 2000 | | | Phanaeus (Notiophanaeus) meleagris Blanchard, 1846 | | | Phanaeus meleagris minos Erichson, 1847 | | | Phanaeus (Notiophanaeus) melibaeus Blanchard, 1846 | | | Phanaeus (Notiophanaeus) pyrois Bates, 1887 | | | Phanaeus pyrois funereus Balthasar, 1939 | | | Phanaeus pyrois olsoufieffi Balthasar, 1939 | | | Phanaeus (Notiophanaeus) splendidulus Fabricius, 1781 | | | Phanaeus (Phanaeus) adonis Harold, 1863 | | | Phanaeus (Phanaeus) amethystinus Harold, 1863 | | | Phanaeus amethystinus amethystinus Harold, 1863 | | | Phanaeus amethystinus guatemalensis (Harold, 1871) | ⇒ Phanaeus guatemalensis Harold, 1871 | | Phanaeus amethystinus tepanensis Bates, 1889 | | | Phanaeus (Phanaeus) amithaon Harold, 1875 | | | Phanaeus (Phanaeus) difformis LeConte, 1847 | | | Phanaeus (Phanaeus) flohri Nevinson, 1892 | Collared blenny | | Phanaeus (Phanaeus) furiosus Bates, 1887 | | | Phanaeus furiosus pseudofurcosus Balthasar, 1939 | | | Phanaeus (Phanaeus) genieri Arnaud, 2001 | Scarface blenny | | Phanaeus (Phanaeus) igneus MacLeay, 1819 | Laguna Madre fiddler | | Phanaeus igneus floridanus Olsoufieff, 1924 | | | Phanaeus igneus igneus | | | Phanaeus (Phanaeus) palliatus Sturm, 1843 | | | Phanaeus (Phanaeus) quadridens Say, 1835 | | | Phanaeus quadridens borealis Olsoufieff, 1924 | | | Phanaeus (Phanaeus) triangularis Say, 1823 | | | Phanaeus triangularis niger Olsoufieff, 1924 | | | Phanaeus triangularis texensis (Edmonds, 1994) | ⇒ Phanaeus texensis Edmonds, 1994 | | Phanaeus triangularis triangularis Say, 1823 | | | Phanaeus (Phanaeus) vindex MacLeay, 1819 | Laguna Madre fiddler | | Phanaeus vindex cyanellus (Robinson, 1938) | ⇒ Phanaeus vindex MacLeay, 1819 | | Phanaeus (Phanaeus) yecoraensis Edmonds, 2004 | Fringed leopard blenny | | Phanaeus antiquus ♰ G. H. Horn, 1876 | | | Phanaeus arlettae Arnaud, 2018 | | | Phanaeus arnaudi Nogueira, Mo..., 2020 | | | Phanaeus auricollis (Harold, 1880) | ⇒ Sulcophanaeus auricollis Harold, 1880 | | Phanaeus balthasari Arnaud, 2001 | | | Phanaeus batesi (Harold, 1868) | ⇒ Sulcophanaeus batesi Harold, 1868 | | Phanaeus bellicosus Olivier, 1789 | | | Phanaeus beltianus Bates, 1887 | | | Phanaeus blackalleri Delgado-Cast..., 1991 | | | Phanaeus bogotensis (Kirsch, 1871) | ⇒ Phanaeus hermes Harold, 1868 | | Phanaeus borealis (Olsoufieff, 1924) | ⇒ Phanaeus quadridens borealis Olsoufieff, 1924 | | Phanaeus bravoensis Moctezuma, S..., 2017 | | | Phanaeus carnifex Linnaeus, 1758 | | | Phanaeus changdiazi Kohlmann & S..., 2001 | | | Phanaeus chiapanecus Moctezuma & ..., 2021 | | | Phanaeus chryseicollis (Harold, 1863) | ⇒ Sulcophanaeus chryseicollis Harold, 1863 | | Phanaeus coeruleus (Bates, 1887) | ⇒ Phanaeus daphnis coeruleus Bates, 1887 | | Phanaeus damocles Harold, 1863 | | | Phanaeus damon (Harold, 1863) | ⇒ Phanaeus demon Castelnau, 1840 | | Phanaeus daphnis Harold, 1863 | Beetle | | Phanaeus daphnis coeruleus Bates, 1887 | | | Phanaeus daphnis daphnis | | | Phanaeus daphnis herbeus Bates, 1887 | | | Phanaeus demon Castelnau, 1840 | | | Phanaeus demon excelsus Bates, 1887 | | | Phanaeus demon obliquans Bates, 1887 | | | Phanaeus dionysius Kohlmann, Ar..., 2018 | | | Phanaeus dzidoi Arnaud, 2000 | | | Phanaeus edmondsi Moctezuma, D..., 2019 | | | Phanaeus excelsus (Bates, 1887) | ⇒ Phanaeus demon excelsus Bates, 1887 | | Phanaeus eximius Bates, 1887 | | | Phanaeus funereus (Balthasar, 1939) | ⇒ Phanaeus pyrois funereus Balthasar, 1939 | | Phanaeus furcosus (Felsche, 1901) | ⇒ Phanaeus furiosus Bates, 1887 | | Phanaeus guatemalensis Harold, 1871 | | | Phanaeus herbeus (Bates, 1887) | ⇒ Phanaeus daphnis herbeus Bates, 1887 | | Phanaeus hermes Harold, 1868 | | | Phanaeus howdeni Arnaud, 1984 | | | Phanaeus huichol Moctezuma, S..., 2017 | | | Phanaeus imperator (Chevrolat, 1844) | ⇒ Sulcophanaeus imperator Chevrolat, 1844 | | Phanaeus jackenioi Moctezuma & ..., 2021 | | | Phanaeus jasion (Felsche, 1901) | ⇒ Coprophanaeus dardanus MacLeay, 1819 | | Phanaeus labreae ♰ W. D. Pierce, 1946 | | | Phanaeus lautus (MacLeay, 1819) | ⇒ Oxysternon lautum MacLeay, 1819 | | Phanaeus lunaris Taschenberg, 1870 | | | Phanaeus magnifescens | | | Phanaeus malyi Arnaud, 2002 | | | Phanaeus martinezorum Arnaud, 2000 | | | Phanaeus melampus Harold, 1863 | | | Phanaeus menelas (Castelnau, 1840) | ⇒ Sulcophanaeus menelas Castelnau, 1840 | | Phanaeus mexicanus Harold, 1863 | | | Phanaeus miles (Castelnau, 1840) | ⇒ Coprophanaeus lancifer Linnaeus, 1767 | | Phanaeus minutus (Motschulsky, 1860) | ⇒ Liatongus phanaeoides Westwood, 1839 | | Phanaeus moroni Arnaud, 2001 | | | Phanaeus nimrod Harold, 1863 | | | Phanaeus noctis (Bates, 1887) | ⇒ Sulcophanaeus noctis Bates, 1887 | | Phanaeus obliquans (Bates, 1887) | ⇒ Phanaeus demon obliquans Bates, 1887 | | Phanaeus olsoufieffi (Balthasar, 1939) | ⇒ Phanaeus pyrois olsoufieffi Balthasar, 1939 | | Phanaeus pacificus Moctezuma & ..., 2021 | | | Phanaeus palaeno Blanchard, 1846 | | | Phanaeus pilatei Harold, 1863 | | | Phanaeus pluto Harold, 1863 | | | Phanaeus porioni Arnaud, 2001 | | | Phanaeus prasinus Harold, 1868 | | | Phanaeus prasinus joliyi Arnaud, 2001 | | | Phanaeus prasinus jolyi Arnaud, 2001 | | | Phanaeus prasinus lugens Nevinson, 1889 | | | Phanaeus prasinus prasinus | | | Phanaeus prasinus trinidadens Arnaud, 2001 | | | Phanaeus prasinus trinidadensis Arnaud, 2001 | | | Phanaeus pseudofurcosus (Balthasar, 1939) | ⇒ Phanaeus furiosus pseudofurcosus Balthasar, 1939 | | Phanaeus rzedowskii Moctezuma & ..., 2021 | | | Phanaeus sallei Harold, 1863 | | | Phanaeus scutifer Bates, 1887 | | | Phanaeus sororibispinus Edmonds & Zidek, 2012 | | | Phanaeus substriolatus (Balthasar, 1939) | ⇒ Phanaeus daphnis Harold, 1863 | | Phanaeus texensis Edmonds, 1994 | | | Phanaeus torrens (LeConte, 1847) | ⇒ Phanaeus triangularis Say, 1823 | | Phanaeus tricornis (Olsoufieff, 1924) | ⇒ Phanaeus daphnis herbeus Bates, 1887 | | Phanaeus tridens Castelnau, 1840 | | | Phanaeus tridens balthasari Arnaud, 2001 | | | Phanaeus tridens moroni Arnaud, 2001 | | | Phanaeus tridens tridens | | | Phanaeus vicinus Martínez, 1944 | | | Phanaeus victoriae Moctezuma, 2021 | | | Phanaeus violae Halffter, Mo..., 2022 | | | Phanaeus violetae Zunino, 2013 | | | Phanaeus viridicollis Olsoufieff, 1924 | | | Phanaeus wagneri Harold, 1863 | | | Phanaeus wagneri pilatei Harold, 1863 | | | Phanaeus zapotecus Edmonds, 2006 | | | Phanaeus zoque Moctezuma & ..., 2017 | | | Pinacopodius meruensis Krikken, 1982 | | | Pinacopodius praestolatus Branco & Josso, 2007 | | | Pinacotarsus dohrni Harold, 1875 | | | Pinacotarsus proximus Ferreira, 1969 | | | Pinotus bicornis (Waterhouse, 1891) | ⇒ Dichotomius bicornis Waterhouse, 1891 | | Pinotus foveolatus (Luederwaldt, 1931) | ⇒ Isocopris foveolatus Luederwaldt, 1931 | | Pinotus gibbosus (Luederwaldt, 1928) | ⇒ Dichotomius triangulariceps Blanchard, 1846 | | Pinotus haroldi (Waterhouse, 1891) | ⇒ Dichotomius haroldi Waterhouse, 1891 | | Pinotus spadiceus (Luederwaldt, 1928) | ⇒ Dichotomius spadiceus Luederwaldt, 1928 | | Pinotus texanus (Felsche, 1901) | ⇒ Dichotomius texanus Felsche, 1901 | | Pinotus transiens (Luederwaldt, 1929) | ⇒ Dichotomius bicuspis Germar, 1824 | | Platyonitis bicuariensis Ferreira, 1976 | | | Platyonitis oberthueri Janssens, 1942 | | | Platyonitis smeenkorum Krikken, 1974 | | | Pleuronitis boucomonti Janssens, 1937 | | | Pleuronitis favareli Garreta, 1914 | | | Pleuronitis fulgidus Castelnau, 1840 | | | Pleuronitis minettii Josso, 2011 | | | Ponerotrogus annandalei (Silvestri, 1924) | ⇒ Haroldius annandalai Silvestri, 1924 | | Prionocephale deplanate ♰ Q. B. Lin, 1980 | | | Proagoderus aciculatus Fahraeus, 1857 | | | Proagoderus albicapillus d´Orbigny, 1902 | | | Proagoderus amblychromatus Marcus, 1917 | | | Proagoderus amplexus Sharp, 1875 | | | Proagoderus armicollis d´Orbigny, 1904 | | | Proagoderus atriclaviger d´Orbigny, 1905 | | | Proagoderus atrosetosus d´Orbigny, 1902 | | | Proagoderus auratus Fabricius, 1801 | | | Proagoderus aureiceps d´Orbigny, 1902 | | | Proagoderus badeni Harold, 1875 | | | Proagoderus biarmatus d´Orbigny, 1908 | | | Proagoderus bicallosus Klug, 1855 | | | Proagoderus boranus Gestro, 1895 | | | Proagoderus bottegi Gestro, 1895 | | | Proagoderus brucei (d´Orbigny, 1902) | ⇒ Onthophagus chalcostolus d´Orbigny, 1902 | | Proagoderus brucei brucei | | | Proagoderus brucei cyanostolus d´Orbigny, 1902 | | | Proagoderus brucei njombeensis Moretto, 2004 | | | Proagoderus cambeforti Palestrini, 1992 | | | Proagoderus camiadei Josso, 2014 | | | Proagoderus cavidorsis d´Orbigny, 1902 | | | Proagoderus ceratotherium Janssens, 1953 | | | Proagoderus chalcostolus d´Orbigny, 1902 | | | Proagoderus chrysopes Bates, 1888 | | | Proagoderus colmanti d´Orbigny, 1908 | | | Proagoderus cometes Bates, 1888 | | | Proagoderus confossus d´Orbigny, 1915 | | | Proagoderus cuspidatus d´Orbigny, 1913 | | | Proagoderus cyanopterus d´Orbigny, 1905 | | | Proagoderus dargei Moretto, 2013 | | | Proagoderus dives Harold, 1877 | | | Proagoderus dudleyi Cambefort, 1980 | | | Proagoderus edithae Josso, 2014 | | | Proagoderus elgoni d´Orbigny, 1915 | | | Proagoderus euchlorus d´Orbigny, 1902 | | | Proagoderus extensus Harold, 1878 | | | Proagoderus fastosus d´Orbigny, 1913 | | | Proagoderus flexicollis d´Orbigny, 1904 | | | Proagoderus fossidorsis d´Orbigny, 1905 | | | Proagoderus freyi Moretto, 2004 | | | Proagoderus furcifer Boheman, 1860 | | | Proagoderus furciramus d´Orbigny, 1902 | | | Proagoderus gangloffi Josso & Pré..., 2001 | | | Proagoderus gemmatus Péringuey, 1901 | | | Proagoderus gerstaeckeri Harold, 1867 | | | Proagoderus gibbiramus d´Orbigny, 1902 | | | Proagoderus gladiator Arrow, 1907 | | | Proagoderus harpax Fabricius, 1801 | | | Proagoderus hemicyaneus | | | Proagoderus hemicyanus d´Orbigny, 1902 | | | Proagoderus ignitus d´Orbigny, 1905 | | | Proagoderus imperator Castelnau, 1840 | | | Proagoderus incostatus d´Orbigny, 1902 | | | Proagoderus kachowskii Olsoufief, 1900 | | | Proagoderus kafueensis Josso, 2014 | | | Proagoderus katualensis Quedenfeldt, 1888 | | | Proagoderus kilimanus Kolbe, 1891 | | | Proagoderus kolbei Marcus, 1921 | | | Proagoderus kwaleensis Moretto & Ni..., 2002 | | | Proagoderus lallieri d´Orbigny, 1902 | | | Proagoderus lallierianus Janssens, 1953 | | | Proagoderus lanista Castelnau, 1840 | | | Proagoderus laticollis Klug, 1835 | | | Proagoderus longefossus d´Orbigny, 1905 | | | Proagoderus loricatus Klug, 1855 | | | Proagoderus louisi Josso, 2014 | | | Proagoderus lucasi d´Orbigny, 1902 | | | Proagoderus lujendae Bates, 1888 | | | Proagoderus marginidens d´Orbigny, 1904 | | | Proagoderus maringouensis Palestrini, 1992 | | | Proagoderus metallites (d´Orbigny, 1905) | ⇒ Proagoderus nigricornis Fairmaire, 1887 | | Proagoderus minettii Moretto & Ni..., 2002 | | | Proagoderus monochromus d´Orbigny, 1913 | | | Proagoderus moorei Moretto, 2013 | | | Proagoderus mouhoti Harold, 1875 | | | Proagoderus multicornis d´Orbigny, 1908 | | | Proagoderus nasidens d´Orbigny, 1905 | | | Proagoderus negus Raffray, 1882 | | | Proagoderus ngottoensis Moretto & Ni..., 2002 | | | Proagoderus nicolasi Moretto, 2004 | | | Proagoderus nigricornis Fairmaire, 1887 | | | Proagoderus nuba d´Orbigny, 1902 | | | Proagoderus opulentus d´Orbigny, 1902 | | | Proagoderus overlaeti Janssens, 1953 | | | Proagoderus pactolus Fabricius, 1787 | | | Proagoderus palestriniae Moretto & Ni..., 2004 | | | Proagoderus panoplus Bates, 1888 | | | Proagoderus paolae Palestrini, 1992 | | | Proagoderus patinatus Marcus, 1917 | | | Proagoderus pegesimallus Marcus, 1917 | | | Proagoderus plato Bates, 1888 | | | Proagoderus porrectus Reiche, 1847 | | | Proagoderus praecavatus d´Orbigny, 1913 | | | Proagoderus praedentatus d´Orbigny, 1904 | | | Proagoderus praefossus d´Orbigny, 1913 | | | Proagoderus prostans Reiche, 1847 | | | Proagoderus pseudoflexicollis Moretto, 2004 | | | Proagoderus pyramidalis Klug, 1855 | | | Proagoderus quadriarmatus Fairmaire, 1892 | | | Proagoderus quadricristatus d´Orbigny, 1908 | | | Proagoderus quadrispinosus d´Orbigny, 1908 | | | Proagoderus quadrituber d´Orbigny, 1908 | | | Proagoderus ramosicornis d´Orbigny, 1902 | | | Proagoderus rangifer Klug, 1855 | | | Proagoderus rarus Guérin-Mén..., 1829 | | | Proagoderus rasidorsis d´Orbigny, 1905 | | | Proagoderus rectefurcatus Fairmaire, 1891 | | | Proagoderus revoili Lansberge, 1882 | | | Proagoderus ritsemai Lansberge, 1883 | | | Proagoderus sapphirinus (Péringuey, 1901) | ⇒ Onthophagus aureiceps d´Orbigny, 1902 | | Proagoderus schwaneri Snellen van ..., 1864 | | | Proagoderus semiiris Thomson, 1858 | | | Proagoderus sexcornutus d´Orbigny, 1902 | | | Proagoderus somalicus d´Orbigny, 1902 | | | Proagoderus speculatus d´Orbigny, 1908 | | | Proagoderus speculicollis Quedenfeldt, 1884 | | | Proagoderus subextensus Kolbe, 1895 | | | Proagoderus superbus d´Orbigny, 1904 | | | Proagoderus taymansi d´Orbigny, 1908 | | | Proagoderus tersidorsis d´Orbigny, 1902 | | | Proagoderus trianguliceps d´Orbigny, 1905 | | | Proagoderus triarmatus d´Orbigny, 1904 | | | Proagoderus tricornifrons d´Orbigny, 1908 | | | Proagoderus uluguru Frey, 1975 | | | Proagoderus unidens Quedenfeldt, 1884 | | | Proagoderus versus d´Orbigny, 1913 | | | Proagoderus victor Sharp, 1875 | | | Proagoderus watanabei Ochi & Kon, 2002 | | | Proagoderus werneri Moretto & Ni..., 2002 | | | Proagoderus worsissa Roth, 1851 | | | Proagoderus yunnanus Boucomont, 1912 | | | Proagoderus yvescambeforti Moretto, 2004 | | | Proteroscarabaeus crassus Ponomarenko, 1990 | | | Proteroscarabaeus magnus ♰ G. V. Nikolajev, 1996 | | | Proteroscarabaeus nikritini ♰ G. V. Nikolajev, 1996 | | | Proteroscarabaeus robustus ♰ H. C. Zhang, 1997 | | | Proteroscarabaeus sibiricus Ponomarenko, 1990 | | | Proteroscarabaeus yeni ♰ Grabau, 1923 | | | Pseudignambia mimerops Matthews, 1974 | | | Pseudignambia squamata Matthews, 1974 | | | Pseudocanthon caeranus Matthews, 1966 | | | Pseudocanthon chaquensis Nazaré-Silva..., 2021 | | | Pseudocanthon chlorizans Bates, 1887 | | | Pseudocanthon iuanalaoi Matthews, 1966 | | | Pseudocanthon ivanalaoi | | | Pseudocanthon jamaicensis Matthews, 1966 | | | Pseudocanthon jamaicensis jamaicensis | | | Pseudocanthon jamaicensis viridescens Matthews, 1966 | | | Pseudocanthon pantanensis Nazaré-Silva..., 2021 | | | Pseudocanthon perplexus LeConte, 1847 | | | Pseudocanthon sylvaticus Matthews, 1966 | | | Pseudocanthon vazdemelloi Nazaré-Silva..., 2021 | | | Pseudocanthon vitraci Fleutiaux & ..., 1889 | | | Pseudocanthon xanthurus Blanchard, 1846 | | | Pseudochironitis bennigseni Felsche, 1907 | | | Pseudochironitis benningseni Felsche, 1907 | | | Pseudochironitis stuhlmanni Kolbe, 1897 | | | Pseudocopris lunaticeps Gillet, 1910 | | | Pseudocopris tubericollis Felsche, 1910 | | | Pseudonthobium doense Montreuil & ..., 2011 | | | Pseudonthobium foveolatum Montreuil & ..., 2011 | | | Pseudonthobium fracticolle Fauvel, 1903 | | | Pseudonthobium fracticolloides Paulian, 1984 | | | Pseudonthobium genieri Montreuil & ..., 2011 | | | Pseudonthobium humboldtense Montreuil & ..., 2011 | | | Pseudonthobium monteithianum Montreuil & ..., 2011 | | | Pseudonthobium paniense Montreuil & ..., 2011 | | | Pseudonthobium sinuatotibiale Montreuil & ..., 2011 | | | Pseudosaproecius baraudi Branco, 1995 | | | Pseudosaproecius bateke Cambefort & ..., 1977 | | | Pseudosaproecius binasus d´Orbigny, 1907 | | | Pseudosaproecius cambeforti Branco, 1995 | | | Pseudosaproecius comatus d´Orbigny, 1902 | | | Pseudosaproecius cylindroides d´Orbigny, 1908 | | | Pseudosaproecius dicerus d´Orbigny, 1915 | | | Pseudosaproecius falcatus Branco, 1995 | | | Pseudosaproecius mirepunctatus d´Orbigny, 1905 | | | Pseudosaproecius ocellatus Frey, 1958 | | | Pseudosaproecius portentosus Felsche, 1907 | | | Pseudosaproecius validicornis Quedenfeldt, 1884 | | | Pseuduroxys ohausi Balthasar, 1938 | | | Pteronyx dimidiatus Lansberge, 1874 | | | Pycnopanelus krikkeni Cambefort, 1978 | | | Pycnopanelus parvicollis d´Orbigny, 1913 | | | Pycnopanelus rotundus Arrow, 1931 | | | Saphobiamorpha maorianus Brookes, 1944 | | | Saphobius arrowi Paulian, 1935 | | | Saphobius brouni Paulian, 1935 | | | Saphobius curvipes Broun, 1893 | | | Saphobius edwardsi Sharp, 1873 | | | Saphobius fulvipes Broun, 1893 | | | Saphobius fuscus Broun, 1893 | | | Saphobius inflatipes Broun, 1893 | | | Saphobius laticollis Broun, 1914 | | | Saphobius lepidus Broun, 1912 | | | Saphobius lesnei Paulian, 1935 | | | Saphobius nitidulus (Broun, 1880) | ⇒ Saphobius edwardsi Sharp, 1873 | | Saphobius setosus Sharp, 1886 | | | Saphobius squamulosus Broun, 1886 | | | Saphobius tibialis Broun, 1895 | | | Saphobius wakefieldi Sharp, 1877 | | | Sarophorus angolensis Frolov, 2004 | | | Sarophorus bidentatus Frolov & Sch..., 2003 | | | Sarophorus carinatus Frolov & Sch..., 2003 | | | Sarophorus cicatricosus Péringuey, 1901 | | | Sarophorus costatus Fahraeus, 1857 | | | Sarophorus diabolus Roets, 2017 | | | Sarophorus frolovi Roets, 2017 | | | Sarophorus latus Frolov & Sch..., 2003 | | | Sarophorus nitidus Frolov & Sch..., 2003 | | | Sarophorus punctatus Frolov & Sch..., 2003 | | | Sarophorus striatus Frolov & Sch..., 2003 | | | Sarophorus tuberculatus Castelnau, 1840 | | | Sauvagesinella becki Paulian, 1934 | | | Sauvagesinella loftyensis Matthews & Weir, 2002 | | | Sauvagesinella monstrosa Paulian, 1934 | | | Sauvagesinella palustris Matthews, 1974 | | | Scaptocnemis segregis Péringuey, 1901 | | | Scarabaeus (Ateuchetus) armeniacus Ménétriés, 1832 | | | Scarabaeus (Ateuchetus) cicatricosus (Lucas, 1846) | ⇒ Scarabaeus palemo Olivier, 1789 | | Scarabaeus (Ateuchetus) laticollis Linnaeus, 1767 | | | Scarabaeus (Ateuchetus) puncticollis (Latreille, 1819) | ⇒ Geotrupes spiniger Marsham, 1802 | | Scarabaeus (Ateuchetus) semipunctatus Fabricius, 1792 | | | Scarabaeus (Ateuchetus) variolosus (Fabricius, 1787) | ⇒ Ateuchetus semipunctatus Fabricius, 1792 | | Scarabaeus (Scarabaeus) babori Balthasar, 1934 | | | Scarabaeus (Scarabaeus) bannuensis Janssens, 1940 | | | Scarabaeus (Scarabaeus) carinatus (Gebler, 1841) | ⇒ Calcodes carinatus Linnaeus, 1758 | | Scarabaeus (Scarabaeus) pius Illiger, 1803 | | | Scarabaeus (Scarabaeus) sacer (Linnaeus, 1758) | ⇒ Scarabaeus pius Illiger, 1803 | | Scarabaeus (Scarabaeus) typhon (Fischer von ..., 1823) | ⇒ Megasoma gyas Herbst, 1785 | | Scarabaeus (Scarabaeus) winkleri Stolfa, 1938 | | | Scarabaeus abbreviatus (Herbst, 1789) | ⇒ Catharsius molossus Linnaeus, 1758 | | Scarabaeus acuticollis Motschulsky, 1849 | | | Scarabaeus aegaeon Drury, 1773 | | | Scarabaeus aegyptiacus Stolfa, 1938 | | | Scarabaeus aegyptiorum Latreille, 1827 | | | Scarabaeus aegyptiorum purpurascens Gerstaecker, 1871 | | | Scarabaeus aesculapius Olivier, 1789 | | | Scarabaeus affinis (Sturm, 1800) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Scarabaeus alienus Péringuey, 1901 | | | Scarabaeus ambiguus (Boheman, 1857) | ⇒ Gnorimus variabilis Linnaeus, 1758 | | Scarabaeus anderseni Waterhouse, 1890 | | | Scarabaeus andreaei zur Strassen, 1963 | | | Scarabaeus andrewesi Felsche, 1907 | | | Scarabaeus auratus (Schrank, 1781) | ⇒ Gnorimus nobilis Linnaeus, 1758 | | Scarabaeus basuto zur Strassen, 1962 | | | Scarabaeus bellicosus (Olivier, 1789) | ⇒ Coprophanaeus bellicosus Olivier, 1789 | | Scarabaeus bennigseni Felsche, 1907 | | | Scarabaeus berbiceus (Herbst, 1789) | ⇒ Catharsius molossus Linnaeus, 1758 | | Scarabaeus bohemani Harold, 1868 | | | Scarabaeus bornemisszai zur Strassen, 1980 | | | Scarabaeus bornemizzai zur Strassen, 1980 | | | Scarabaeus brahminus Castelnau, 1840 | | | Scarabaeus caffer Boheman, 1857 | | | Scarabaeus canaliculatus Fairmaire, 1888 | | | Scarabaeus cancer Arrow, 1919 | | | Scarabaeus carniphilus Davis & Desc..., 2015 | | | Scarabaeus catenatus Gerstaecker, 1871 | | | Scarabaeus cineta Drury, 1782 | | | Scarabaeus clanceyi Ferreira, 1954 | | | Scarabaeus cognatus Péringuey, 1901 | | | Scarabaeus confusus Müller, 1942 | | | Scarabaeus convexus Hausmann, 1807 | | | Scarabaeus costatus Wiedemann, 1823 | | | Scarabaeus cristatus villiersi Baraud, 1973 | | | Scarabaeus damarensis Janssens, 1940 | | | Scarabaeus deludens zur Strassen, 1961 | | | Scarabaeus denticollis Péringuey, 1888 | | | Scarabaeus devotus Redtenbacher, 1848 | | | Scarabaeus dioscoridis Petrovitz, 1962 | | | Scarabaeus ebenus Klug, 1855 | | | Scarabaeus endroedyi Harrison & S..., 2003 | | | Scarabaeus erichsoni Harold, 1867 | | | Scarabaeus ermienae Deschodt & D..., 2015 | | | Scarabaeus festivus Harold, 1868 | | | Scarabaeus fitzsimonsi Ferreira, 1953 | | | Scarabaeus flavicornis Boheman, 1860 | | | Scarabaeus fraterculus Kolbe, 1895 | | | Scarabaeus fritschi Harold, 1868 | | | Scarabaeus funebris Boheman, 1857 | | | Scarabaeus galenus Westwood, 1847 | | | Scarabaeus gangeticus Castelnau, 1840 | | | Scarabaeus gariepinus Ferreira, 1953 | | | Scarabaeus geminogalenus Davis & Desc..., 2017 | | | Scarabaeus gilleti Janssens, 1940 | | | Scarabaeus glentoni Harrison & S..., 2003 | | | Scarabaeus goryi Castelnau, 1840 | | | Scarabaeus gracai Ferreira, 1952 | | | Scarabaeus heqvisti zur Strassen, 1962 | | | Scarabaeus hippocrates MacLeay, 1821 | | | Scarabaeus hottentorum Péringuey, 1901 | | | Scarabaeus indus (Linnaeus, 1758) | ⇒ Euphoria inda Linnaeus, 1758 | | Scarabaeus inguisitus | | | Scarabaeus inoportunus Ferreira, 1953 | | | Scarabaeus inquisitus Péringuey, 1908 | | | Scarabaeus interstitialis Boheman, 1857 | | | Scarabaeus intricatus Fabricius, 1801 | | | Scarabaeus irakensis Stolfa, 1938 | | | Scarabaeus isidis Castelnau, 1840 | | | Scarabaeus jalof Castelnau, 1840 | | | Scarabaeus janssensi Balthasar, 1940 | | | Scarabaeus janus (Olivier, 1789) | ⇒ Catharsius molossus Linnaeus, 1758 | | Scarabaeus karae Davis & Desc..., 2016 | | | Scarabaeus karlwerneri Nicolas & Mo..., 2002 | | | Scarabaeus karrooensis zur Strassen, 1961 | | | Scarabaeus knobeli Ferreira, 1958 | | | Scarabaeus kochi Ferreira, 1952 | | | Scarabaeus krutai Maly & Montr..., 2011 | | | Scarabaeus kwiluensis Janssens, 1940 | | | Scarabaeus laevifrons Fairmaire, 1884 | | | Scarabaeus laevigatus (Kolbe, 1887) | ⇒ Scarabaeus goryi Castelnau, 1840 | | Scarabaeus lamarcki MacLeay, 1821 | | | Scarabaeus leei (Donovan, 1798) | ⇒ Gymnopleurus leei Fabricius, 1792 | | Scarabaeus licitus Ferreira, 1954 | | | Scarabaeus lucidulus Boheman, 1860 | | | Scarabaeus megaparvulus Davis & Desc..., 2015 | | | Scarabaeus molossus (Linnaeus, 1758) | ⇒ Catharsius molossus Linnaeus, 1758 | | Scarabaeus montreuili meridionalis Moretto, 2023 | | | Scarabaeus montreuili montreuili | | | Scarabaeus multidentatum Klug, 1845 | | | Scarabaeus natalensis zur Strassen, 1962 | | | Scarabaeus nepos (Fairmaire, 1884) | ⇒ Scarabaeus goryi Castelnau, 1840 | | Scarabaeus niemandi Deschodt & D..., 2015 | | | Scarabaeus nigroaeneus Boheman, 1857 | | | Scarabaeus nitidicollis Lansberge, 1882 | | | Scarabaeus obsoletepunctatus Balthasar, 1940 | | | Scarabaeus opacipennis Fairmaire, 1884 | | | Scarabaeus osiridis Moretto, 2023 | | | Scarabaeus pabulator Péringuey, 1908 | | | Scarabaeus palemo Olivier, 1789 | | | Scarabaeus parvulus Boheman, 1860 | | | Scarabaeus piliventris zur Strassen, 1962 | | | Scarabaeus planifrons Fairmaire, 1884 | | | Scarabaeus planipennis Davis & Desc..., 2015 | | | Scarabaeus plansibilis | | | Scarabaeus platynotus Bates, 1888 | | | Scarabaeus plausibilis Péringuey, 1892 | | | Scarabaeus poggei Waterhouse, 1890 | | | Scarabaeus proboscideus Guérin-Ménev..., 1844 | | | Scarabaeus prodigiosus Erichson, 1843 | | | Scarabaeus profanus Fabricius, 1792 | | | Scarabaeus proserpinae O. Heer, 1862 | | | Scarabaeus proximus (Péringuey, 1901) | ⇒ Drepanopodus proximus Péringuey, 1901 | | Scarabaeus pseudogangeticus Moretto, 2023 | | | Scarabaeus pubiventris (Lansberge, 1874) | ⇒ Kheper prodigiosus Erichson, 1843 | | Scarabaeus radama Fairmaire, 1895 | | | Scarabaeus ritchiei MacLeay, 1821 | | | Scarabaeus rixosus Péringuey, 1901 | | | Scarabaeus rodriguesi Ferreira, 1953 | | | Scarabaeus rotundigenus Felsche, 1907 | | | Scarabaeus rubripennis Boheman, 1860 | | | Scarabaeus ruficornis | | | Scarabaeus rufus (DeGeer, 1778) | ⇒ Rhysothorax rufus Fabricius, 1792 | | Scarabaeus rugosus (Hausmann, 1807) | ⇒ Onthophagus taurus Schreber, 1759 | | Scarabaeus rusticus Boheman, 1857 | | | Scarabaeus sanctus Fabricius, 1798 | | | Scarabaeus satyrus (Boheman, 1860) | ⇒ Xyloryctes jamaicensis Drury, 1773 | | Scarabaeus savignyi MacLeay, 1821 | | | Scarabaeus schaefferi (Linnaeus, 1758) | ⇒ Sisyphus schaefferi schaefferi Linnaeus, 1758 | | Scarabaeus schinzi Fairmaire, 1888 | | | Scarabaeus scholtzi Mostert & Holm, 1982 | | | Scarabaeus schulzeae Ferreira, 1969 | | | Scarabaeus sennaariensis Castelnau, 1840 | | | Scarabaeus sevoistra Alluaud, 1902 | | | Scarabaeus sinuatus (Olivier, 1789) | ⇒ Paragymnopleurus sinuatus Olivier, 1789 | | Scarabaeus soutpansbergensis Deschodt & D..., 2015 | | | Scarabaeus spretus zur Strassen, 1962 | | | Scarabaeus striatus Castelnau, 1840 | | | Scarabaeus sulcipennis Quedenfeldt, 1888 | | | Scarabaeus suri Hausmann, 1807 | | | Scarabaeus thebanus (Herbst, 1779) | ⇒ Gametis versicolor Fabricius, 1775 | | Scarabaeus tonckeri Boucomont, 1923 | | | Scarabaeus transcaspicus Stolfa, 1938 | | | Scarabaeus vansoni Ferreira, 1958 | | | Scarabaeus venerabilis Harold, 1871 | | | Scarabaeus vethi Lansberge, 1886 | | | Scarabaeus viator Péringuey, 1901 | | | Scarabaeus vicinus (Janssens, 1940) | ⇒ Lucanus cervus Linnaeus, 1758 | | Scarabaeus viettei Paulian, 1953 | | | Scarabaeus westwoodi (Harold, 1869) | ⇒ Scarabaeus galenus Westwood, 1847 | | Scarabaeus wilsoni Waterhouse, 1890 | | | Scarabaeus xavieri Ferreira, 1968 | | | Scarabaeus zagrosensis Montreuil, 2006 | | | Scarabaeus zambesianus Péringuey, 1901 | | | Scarabaeus zambezianus Péringuey, 1901 | | | Scatimus cribrosus Génier & Koh..., 2003 | | | Scatimus cucullatus Erichson, 1847 | | | Scatimus erinnyos Kohlmann & S..., 1996 | | | Scatimus fernandezi Martínez, 1988 | | | Scatimus furcatus Balthasar, 1939 | | | Scatimus monstrosus Balthasar, 1939 | | | Scatimus onorei Génier & Koh..., 2003 | | | Scatimus ovatus Harold, 1862 | | | Scatimus pacificus Génier & Koh..., 2003 | | | Scatimus quadricuspis Génier & Koh..., 2003 | | | Scatimus quadridentatus (Balthasar, 1939) | ⇒ Scatimus ovatus Harold, 1862 | | Scatimus simulator Martínez, 1988 | | | Scatimus strandi Balthasar, 1939 | | | Scatimus strenua Martínez-Rev..., 2020 | | | Scatonomus barbatus (Erichson, 1835) | ⇒ Scatonomus fasciculatus Erichson, 1835 | | Scatonomus canhedoae Valois, Vaz-..., 2020 | | | Scatonomus chalybeus Erichson, 1835 | | | Scatonomus fasciculatus Erichson, 1835 | | | Scatonomus foresti Valois, Vaz-..., 2020 | | | Scatonomus insignis Harold, 1867 | | | Scatonomus janssensi Pereira, 1954 | | | Scatonomus lauropalui Vulcano & Pe..., 1973 | | | Scatonomus mitzae Valois, Vaz-..., 2020 | | | Scatonomus paulosawayai Vulcano & Pe..., 1973 | | | Scatonomus thalassinus Waterhouse, 1891 | | | Scatonomus viridis Erichson, 1835 | | | Scatonomus viridis viridis | | | Scatonomus viridis walzi Pereira & Ma..., 1959 | | | Scatonomus xanthopygus Pereira, 1954 | | | Scatrichus bicarinatus Harold, 1869 | | | Scatrichus goiasensis Génier & Koh..., 2003 | | | Scatrichus sulcifer Génier & Koh..., 2003 | | | Sceliages adamastor Lepeletier &..., 1828 | | | Sceliages augias Gillet, 1908 | | | Sceliages brittoni zur Strassen, 1965 | | | Sceliages curvipes (Gillet, 1911) | ⇒ Sceliages adamastor Lepeletier &..., 1828 | | Sceliages difficilis zur Strassen, 1965 | | | Sceliages gagates Shipp, 1895 | | | Sceliages granulatus Forgie & Gre..., 2002 | | | Sceliages hippias Westwood, 1844 | | | Scelocopris enertheus ♰ J. F. Zhang, 1989 | | | Scybalocanthon acrianus Silva & Valois, 2019 | | | Scybalocanthon adisi Silva & Valois, 2019 | | | Scybalocanthon aereus Schmidt, 1922 | | | Scybalocanthon arcabuquensis Molano & Medina, 2010 | | | Scybalocanthon arnaudi Silva & Valois, 2019 | | | Scybalocanthon ashei Silva & Génier, 2019 | | | Scybalocanthon balachowskyi Martinez & H..., 1972 | | | Scybalocanthon batesi Vaz-de-Mello..., 2017 | | | Scybalocanthon chamorroi Silva & Valois, 2019 | | | Scybalocanthon cyanocephalus Harold, 1868 | | | Scybalocanthon darlingtoni Paulian, 1939 | | | Scybalocanthon federicoescobari Silva & Valois, 2019 | | | Scybalocanthon haroldi Silva & Valois, 2019 | | | Scybalocanthon imitans Harold, 1868 | | | Scybalocanthon kaestneri Balthasar, 1939 | | | Scybalocanthon kelleri Pereira & Ma..., 1956 | | | Scybalocanthon korasakiae Silva, 2011 | | | Scybalocanthon luctuosus Harold, 1868 | | | Scybalocanthon maculatus Schmidt, 1920 | | | Scybalocanthon martinezi Silva & Valois, 2019 | | | Scybalocanthon moniliatus Bates, 1887 | | | Scybalocanthon nigellus Schmidt, 1922 | | | Scybalocanthon nigriceps Harold, 1868 | | | Scybalocanthon papaxibe Silva & Valois, 2019 | | | Scybalocanthon pinopterus Kirsch, 1873 | | | Scybalocanthon pygidialis Schmidt, 1922 | | | Scybalocanthon sexspilotus Guérin-Ménev..., 1855 | | | Scybalocanthon trimaculatus Schmidt, 1922 | | | Scybalocanthon uniplagiatus Schmidt, 1922 | | | Scybalocanthon zischkai Martínez, 1949 | | | Scybalophagus lacordairei Castelnau, 1840 | | | Scybalophagus patagonicus Martínez, 1953 | | | Scybalophagus pilosus Felsche, 1910 | | | Scybalophagus plicatipennis Blanchard, 1846 | | | Scybalophagus rugosus Blanchard, 1846 | | | Silvaphilus joselmae Daniel, Strü..., 2022 | | | Silvaphilus oubosiensis Roets & Ober..., 2010 | | | Silvinha unica Vaz-de-Mello, 2008 | | | Sinapisoma minuta Boucomont, 1928 | | | Sinapisoma minutum (Boucomont, 1928) | ⇒ Sinapisoma minuta Boucomont, 1928 | | Sinodrepanus besucheti Simonis, 1985 | | | Sinodrepanus exsul Sharp, 1875 | | | Sinodrepanus falsus Sharp, 1875 | | | Sinodrepanus rex Boucomont, 1912 | | | Sinodrepanus rosannae Simonis, 1985 | | | Sinodrepanus similis Simonis, 1985 | | | Sinodrepanus thailandicus Ochi, Kon & ..., 2004 | | | Sinodrepanus tsaii Masumoto, Ya..., 2004 | | | Sinodrepanus uenoi Ochi, Kon & ..., 2004 | | | Sisyphus (Sisyphus) struwei Schafer & Fi..., 1992 | | | Sisyphus struwei kivuensis Schafer & Fi..., 2001 | | | Sisyphus struwei struwei | | | Sisyphus aages Moxey, 1962 | | | Sisyphus africanus africanus | | | Sisyphus africanus occidentalis Montreuil, 2017 | | | Sisyphus africanus orientalis Montreuil, 2017 | | | Sisyphus alveatus Boucomont, 1935 | | | Sisyphus angulicollis (Felsche, 1909) | ⇒ Neosisyphus angulicollis Felsche, 1909 | | Sisyphus araneolus Arrow, 1927 | | | Sisyphus arboreus Walter, 1982 | | | Sisyphus armatus (Gory, 1833) | ⇒ Neosisyphus confrater Kolbe, 1914 | | Sisyphus astriatus Schafer & Fi..., 2001 | | | Sisyphus atratus (Klug, 1855) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Sisyphus auricomus Daniel & Davis, 2018 | | | Sisyphus australis Daniel & Davis, 2018 | | | Sisyphus baoule Cambefort, 1984 | | | Sisyphus barbarossa (Wiedemann, 1823) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Sisyphus basilewskyi (Haaf, 1956) | ⇒ Neosisyphus basilewskyi Haaf, 1956 | | Sisyphus bayanga Montreuil, 2016 | | | Sisyphus biarmatus Felsche, 1909 | | | Sisyphus bicuariensis Daniel & Davis, 2018 | | | Sisyphus bornemisszanus Endrödi, 1983 | | | Sisyphus bouyeri Montreuil, 2015 | | | Sisyphus bowringi White, 1844 | | | Sisyphus caffer Boheman, 1857 | | | Sisyphus caffer goryi Harold, 1859 | | | Sisyphus calcaratus (Boucomont, 1937) | ⇒ Neosisyphus rugosus Gory, 1833 | | Sisyphus cambeforti Montreuil, 2015 | | | Sisyphus costatus Thunberg, 1818 | | | Sisyphus crispatus Gory, 1833 | | | Sisyphus crispatus crispatus | | | Sisyphus crispatus hirtus Wiedemann, 1823 | | | Sisyphus crispatus mexicanus Harold, 1863 | | | Sisyphus denticrus (Fairmaire, 1886) | ⇒ Sisyphus neglectus Gory, 1833 | | Sisyphus desaegeri Haaf, 1959 | | | Sisyphus dierkensi Montreuil, 2015 | | | Sisyphus drumonti Montreuil, 2015 | | | Sisyphus eburneus Cambefort, 1984 | | | Sisyphus ethiopicus Montreuil, 2017 | | | Sisyphus fasciculatus Boheman, 1857 | | | Sisyphus felschei Montreuil, 2015 | | | Sisyphus fortuitus Péringuey, 1900 | | | Sisyphus gazanus Arrow, 1909 | | | Sisyphus genierorum Montreuil, 2015 | | | Sisyphus gladiator (Arrow, 1927) | ⇒ Neosisyphus gladiator Arrow, 1927 | | Sisyphus goryi (Harold, 1859) | ⇒ Sisyphus caffer goryi Harold, 1859 | | Sisyphus granulifer Montreuil, 2014 | | | Sisyphus hirtus (Gory, 1833) | ⇒ Sisyphus crispatus hirtus Wiedemann, 1823 | | Sisyphus ibaya Montreuil, 2017 | | | Sisyphus impressipennis Lansberge, 1886 | | | Sisyphus inconspicuus Daniel & Davis, 2018 | | | Sisyphus indicus Hope, 1831 | | | Sisyphus infuscatus (Klug, 1855) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Sisyphus janssensi Montreuil, 2015 | | | Sisyphus jossoi Montreuil, 2015 | | | Sisyphus juheli Montreuil, 2015 | | | Sisyphus kibale Montreuil, 2016 | | | Sisyphus kuehni Haaf, 1955 | | | Sisyphus laoticus Arrow, 1927 | | | Sisyphus latus (Boucomont, 1928) | ⇒ Sisyphus costatus Thunberg, 1818 | | Sisyphus lobi Cambefort, 1984 | | | Sisyphus longipes Olivier, 1789 | | | Sisyphus macrorubrus (Paschalidis, 1974) | ⇒ Neosisyphus macrorubrus Paschalidis, 1974 | | Sisyphus maniti Masumoto, 1988 | | | Sisyphus manni Montreuil, 2015 | | | Sisyphus mendicus Arrow, 1931 | | | Sisyphus mexicanus (Harold, 1863) | ⇒ Sisyphus crispatus mexicanus Harold, 1863 | | Sisyphus minettii Montreuil, 2014 | | | Sisyphus mirabilis Arrow, 1927 | | | Sisyphus mkomazi Montreuil, 2017 | | | Sisyphus moltaensis Vazirani, 1966 | | | Sisyphus moorei Montreuil, 2015 | | | Sisyphus morenoi Montreuil, 2015 | | | Sisyphus morettoi Montreuil, 2015 | | | Sisyphus muricatus Olivier, 1789 | | | Sisyphus nanniscus Péringuey, 1901 | | | Sisyphus natalensis Balthasar, 1968 | | | Sisyphus neglectus Gory, 1833 | | | Sisyphus negus Montreuil, 2015 | | | Sisyphus neobornemisszanus Daniel & Davis, 2016 | | | Sisyphus nodifer Gerstaecker, 1871 | | | Sisyphus ocellatus Reiche, 1847 | | | Sisyphus ocellatus nanniscus Péringuey, 1901 | | | Sisyphus ongo Montreuil, 2017 | | | Sisyphus oralensis Daniel & Davis, 2016 | | | Sisyphus paschalidisae Cambefort, 1984 | | | Sisyphus penicillatus Harold, 1880 | | | Sisyphus perissinottoi Montreuil, 2015 | | | Sisyphus popovi Kryzhanovski..., 1966 | | | Sisyphus quadricollis (Péringuey, 1901) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Sisyphus rubripes (Boheman, 1857) | ⇒ Neosisyphus confrater Kolbe, 1914 | | Sisyphus rubrus (Paschalidis, 1974) | ⇒ Neosisyphus rubrus Paschalidis, 1974 | | Sisyphus rugosus (Roth, 1851) | ⇒ Sisyphus costatus Thunberg, 1818 | | Sisyphus schaefferi Linnaeus, 1758 | | | Sisyphus schaefferi boschniaki Fischer von ..., 1823 | | | Sisyphus schaefferi morio Arrow, 1909 | | | Sisyphus schaefferi schaefferi Linnaeus, 1758 | | | Sisyphus seminulum (Gerstaecker, 1871) | ⇒ Sisyphus costatus Thunberg, 1818 | | Sisyphus setiger (Roth, 1851) | ⇒ Neosisyphus confrater Kolbe, 1914 | | Sisyphus sordidus (Boheman, 1857) | ⇒ Sisyphus caffer Boheman, 1857 | | Sisyphus spinipes (Thunberg, 1818) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Sisyphus splendidus Montreuil, 2015 | | | Sisyphus submonticolus Howden, 1965 | | | Sisyphus swazi Daniel & Davis, 2018 | | | Sisyphus tai Cambefort, 1984 | | | Sisyphus tarantula (Arrow, 1909) | ⇒ Neosisyphus tarantula Arrow, 1909 | | Sisyphus tembyi Daniel & Davis, 2018 | | | Sisyphus thoracicus Sharp, 1875 | | | Sisyphus thoracicus chaiyaphumensis Hanboonsong ..., 2000 | | | Sisyphus thoracicus thoracicus | | | Sisyphus tibialis Raffray, 1877 | | | Sisyphus transvaalensis (Péringuey, 1908) | ⇒ Sisyphus impressipennis Lansberge, 1886 | | Sisyphus trichodichromicus Montreuil, 2015 | | | Sisyphus trochantericus (Fairmaire, 1891) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Sisyphus umbraphilus Daniel & Davis, 2016 | | | Sisyphus usambaricus Montreuil, 2015 | | | Sisyphus walteri Montreuil, 2016 | | | Sisyphus werneri Montreuil, 2017 | | | Sisyphus youngai Endrödi, 1973 | | | Stiptocnemis gilleti Boucomont, 1923 | | | Stiptocnemis hiekei Branco, 1991 | | | Stiptopodius brancoi Strümpher & ..., 2021 | | | Stiptopodius doriae Harold, 1871 | | | Stiptopodius gaillardi Boucomont, 1923 | | | Stiptopodius glaber Branco, 1992 | | | Stiptopodius glabricollis Müller, 1942 | | | Stiptopodius granulosus Branco, 2009 | | | Stiptopodius krausei Branco, 1991 | | | Stiptopodius longipedis Branco, 1991 | | | Stiptopodius muellerae Daniel & Des..., 2020 | | | Stiptopodius nitidus Boucomont, 1923 | | | Stiptopodius nodieri Boucomont, 1923 | | | Stiptopodius patrizii Boucomont, 1923 | | | Stiptopodius peringueyi Daniel & Des..., 2020 | | | Stiptopodius savuti Daniel & Des..., 2020 | | | Stiptopodius singularis Péringuey, 1901 | | | Stiptotarsus latetarsatus Boucomont, 1923 | | | Stiptotarsus mocquerysi Boucomont, 1923 | | | Stiptotarsus tersus Balthasar, 1963 | | | Strandius obliquus (Cambefort, 1980) | ⇒ Onthophagus obliquus Olivier, 1789 | | Streblopus opatroides Lansberge, 1874 | | | Streblopus punctatus Balthasar, 1938 | | | Sukelus jessopi Branco, 1991 | | | Sulcodrepanus sulcicollis Castelnau, 1840 | | | Sulcophanaeus actaeon Erichson, 1847 | | | Sulcophanaeus auricollis Harold, 1880 | | | Sulcophanaeus auricollis auricollis | | | Sulcophanaeus auricollis joffrei Martínez, 1988 | | | Sulcophanaeus batesi Harold, 1868 | | | Sulcophanaeus carnifex Linnaeus, 1758 | | | Sulcophanaeus chryseicollis Harold, 1863 | | | Sulcophanaeus columbi MacLeay, 1819 | | | Sulcophanaeus faunus Fabricius, 1775 | | | Sulcophanaeus imperator Chevrolat, 1844 | | | Sulcophanaeus imperator alticollis Arnaud, 2002 | | | Sulcophanaeus imperator imperator | | | Sulcophanaeus imperator obscurus Arnaud, 2002 | | | Sulcophanaeus leander Waterhouse, 1891 | | | Sulcophanaeus menelas Castelnau, 1840 | | | Sulcophanaeus miyashitai Arnaud, 2002 | | | Sulcophanaeus miyashitai metallescens Arnaud, 2002 | | | Sulcophanaeus miyashitai miyashitai | | | Sulcophanaeus noctis Bates, 1887 | | | Sulcophanaeus noctis cupricollis Nevinson, 1891 | | | Sulcophanaeus noctis noctis | | | Sulcophanaeus rhadamanthus Harold, 1875 | | | Sulcophanaeus steinheili Harold, 1875 | | | Sulcophanaeus velutinus Murray, 1856 | | | Sylvicanthon aequinoctialis Harold, 1868 | | | Sylvicanthon attenboroughi Cupello & Va..., 2018 | | | Sylvicanthon bridarollii Martínez, 1949 | | | Sylvicanthon candezei Harold, 1869 | | | Sylvicanthon edmondsi Cupello & Va..., 2018 | | | Sylvicanthon enkerlini Cupello & Va..., 2018 | | | Sylvicanthon foveiventris Schmidt, 1920 | | | Sylvicanthon furvus Schmidt, 1920 | | | Sylvicanthon genieri Cupello & Va..., 2018 | | | Sylvicanthon machadoi Pereira & Ma..., 1967 | | | Sylvicanthon mayri Cupello & Va..., 2018 | | | Sylvicanthon monnei Cupello & Va..., 2018 | | | Sylvicanthon obscurus Schmidt, 1920 | | | Sylvicanthon proseni Martínez, 1949 | | | Sylvicanthon seag Cupello & Va..., 2018 | | | Sylvicanthon securus Schmidt, 1920 | | | Sylvicanthon xanthopus Blanchard, 1846 | | | Synapsis aquisgranensis Forster, 1869 | | | Synapsis birmanicus Gillet, 1907 | | | Synapsis boonlongi Hanboonsong ..., 1999 | | | Synapsis brahminus Hope, 1831 | | | Synapsis cambeforti (Ochi, Kon & ..., 2008) | ⇒ Synapsis ritsemae Lansberge, 1874 | | Synapsis davidi (Fairmaire, 1878) | ⇒ Synapsis davidis Fairmaire, 1878 | | Synapsis davidis Fairmaire, 1878 | | | Synapsis dickinsoni Hanboonsong ..., 1999 | | | Synapsis gilleti Arrow, 1931 | | | Synapsis horaki Zidek & Pokorny, 2010 | | | Synapsis kiuchii Hanboonsong ..., 1999 | | | Synapsis masumotoi Ochi, 1992 | | | Synapsis naxiorum Kral & Rejsek, 2000 | | | Synapsis ochii Masumoto, 1995 | | | Synapsis ovalis Boucomont, 1920 | | | Synapsis puluongensis Bui & Bonkowski, 2018 | | | Synapsis punctatus Ochi, Kon & ..., 2008 | | | Synapsis ritsemae Lansberge, 1874 | | | Synapsis roslihashimi Ochi, Kon & ..., 2008 | | | Synapsis satoi Ochi & Kon, 2007 | | | Synapsis simplex Sharp, 1875 | | | Synapsis strnadi Kral, 2002 | | | Synapsis thoas (Sharp, 1875) | ⇒ Synapsis ritsemae Lansberge, 1874 | | Synapsis tmolus Fischer von ..., 1821 | | | Synapsis tridens Sharp, 1881 | | | Synapsis vietnamicus Ochi, Kon & ..., 2020 | | | Synapsis yama Gillet, 1911 | | | Synapsis yunnanus Arrow, 1933 | | | Tanzanolus leleupi Balthasar, 1960 | | | Temnoplectron aeneolum Lansberge, 1885 | | | Temnoplectron aeneopiceum Matthews, 1974 | | | Temnoplectron arfakensis Ochi, Kon & ..., 2017 | | | Temnoplectron atropolitum Gillet, 1927 | | | Temnoplectron bornemisszai Matthews, 1974 | | | Temnoplectron boucomonti Paulian, 1934 | | | Temnoplectron cooki Reid & Storey, 2000 | | | Temnoplectron disruptum Matthews, 1974 | | | Temnoplectron diversicolle Blackburn, 1894 | | | Temnoplectron finnigani Reid & Storey, 2000 | | | Temnoplectron involucre Matthews, 1974 | | | Temnoplectron laeve Castelnau, 1840 | | | Temnoplectron lewisense Reid & Storey, 2000 | | | Temnoplectron major Paulian, 1985 | | | Temnoplectron monteithi Reid & Storey, 2000 | | | Temnoplectron politulum MacLeay, 1887 | | | Temnoplectron reyi Paulian, 1934 | | | Temnoplectron rotundum Westwood, 1841 | | | Temnoplectron subvolitans Matthews, 1974 | | | Temnoplectron waigeoensis Ochi, Kon & ..., 2017 | | | Temnoplectron wareo Reid & Storey, 2000 | | | Tesserodon angulatum Westwood, 1841 | | | Tesserodon elongatum (Lansberge, 1885) | ⇒ Tesserodon elongatus Lansberge, 1885 | | Tesserodon elongatus Lansberge, 1885 | | | Tesserodon erratum Storey, 1991 | | | Tesserodon feehani Storey, 1991 | | | Tesserodon gestroi Lansberge, 1885 | | | Tesserodon granulatum Matthews, 1974 | | | Tesserodon henryi Storey, 1991 | | | Tesserodon heurni Paulian, 1937 | | | Tesserodon hilleri Storey, 1991 | | | Tesserodon howdeni Paulian, 1985 | | | Tesserodon intricatum Lea, 1923 | | | Tesserodon novaehollandiae Fabricius, 1775 | | | Tesserodon papuanum Paulian, 1985 | | | Tesserodon pilicrepus Matthews, 1974 | | | Tesserodon seramicum Krikken & Hu..., 2009 | | | Tesserodon setulosum Balthasar, 1965 | | | Tesserodon simplicipunctatum Storey, 1991 | | | Tesserodon tenebroides Matthews, 1974 | | | Tesserodon variolosum MacLeay, 1888 | | | Tesserodon waigeoensis Ochi, Kon & ..., 2017 | | | Tesserodoniella elguetai Vaz-de-Mello..., 2006 | | | Tesserodoniella meridionalis Vaz-de-Mello..., 2006 | | | Tetraechma balteata Boheman, 1858 | | | Tetraechma chlorophana Mannerheim, 1828 | | | Tetraechma liturata Germar, 1813 | | | Tetraechma sanguineomaculata Blanchard, 1846 | | | Tetraechma tarsalis Balthasar, 1939 | | | Tetramereia convexa Harold, 1869 | | | Thyregis kershawi Blackburn, 1904 | | | Thyregis monteithi Matthews, 1976 | | | Thyregis relictus Matthews, 1976 | | | Thyregis tarsatus Paulian, 1933 | | | Tibiodrepanus hircus Wiedemann, 1823 | | | Tibiodrepanus kazirangensis Biswas, 1980 | | | Tibiodrepanus setosus Wiedemann, 1823 | | | Tibiodrepanus simplex Kabakov, 2006 | | | Tibiodrepanus sinicus Harold, 1868 | | | Tibiodrepanus sulcicollis Castelnau, 1840 | | | Tibiodrepanus tagliaferrii Barbero, Pal..., 2011 | | | Tiniocellus averyanovi Kabakov, 2014 | | | Tiniocellus dolosus Branco, 2010 | | | Tiniocellus eurypygus Branco, 2010 | | | Tiniocellus eurypygus eurypygus | | | Tiniocellus eurypygus transdrakensbergensis Branco, 2010 | | | Tiniocellus imbellis Bates, 1891 | | | Tiniocellus praetermissus Branco, 2010 | | | Tiniocellus setifer Kraatz, 1895 | | | Tiniocellus spinipes Roth, 1851 | | | Tomogonus cambeforti Nicolas, 2002 | | | Tomogonus chauvetorum Walter, 2012 | | | Tomogonus crassoides Boucomont, 1923 | | | Tomogonus crassus d´Orbigny, 1902 | | | Tomogonus implicatus Demange, 1978 | | | Tomogonus intortus Demange, 1971 | | | Tomogonus involutus Demange & Ma..., 1975 | | | Tomogonus lamottei Demange & Ma..., 1975 | | | Tomogonus samburu Walter, 2012 | | | Tomogonus schubarti Demange & Ma..., 1975 | | | Tomogonus subgrundus Demange, 1971 | | | Tragiscus dimidiatus Klug, 1855 | | | Trichillidium brevisetosum Howden & Young, 1981 | | | Trichillidium caingua Martínez, 1974 | | | Trichillidium pilosum Robinson, 1948 | | | Trichillidium quadridens Arrow, 1932 | | | Trichillum adjunctum Martínez, 1967 | | | Trichillum arrowi (Saylor, 1935) | ⇒ Trichillum pauliani Balthasar, 1939 | | Trichillum boucomonti (Saylor, 1935) | ⇒ Eutrichillum hirsutum Boucomont, 1928 | | Trichillum bradyporum (Boucomont, 1928) | ⇒ Bradypodidium bradyporum Boucomont, 1928 | | Trichillum cordobense Vaz-de-Mello..., 2005 | | | Trichillum depilatum Balthasar, 1942 | | | Trichillum externepunctatum Preudhomme d..., 1880 | | | Trichillum falsus | | | Trichillum halffteri Martínez, 1967 | | | Trichillum heydeni Harold, 1868 | | | Trichillum horacioi (Martínez, 1967) | ⇒ Besourenga horacioi Martínez, 1967 | | Trichillum minutum (Saylor, 1935) | ⇒ Besourenga minutum Saylor, 1935 | | Trichillum morellii Verdu & Galante, 1997 | | | Trichillum pauliani Balthasar, 1939 | | | Trichillum pereirai (Martínez, 1968) | ⇒ Trichillum heydeni Harold, 1868 | | Trichillum pseudoarrowi Vaz-de-Mello..., 2005 | | | Trichillum tishechkini Vaz-de-Mello..., 2005 | | | Trichillum woodruffi | | | Tropidonitis paradoxus Boheman, 1857 | | | Unidentis vwaza Josso & Prév..., 2008 | | | Uroxys angulatus Harold, 1868 | | | Uroxys angulicollis Boheman, 1858 | | | Uroxys aterrimus Harold, 1867 | | | Uroxys bahianus Boucomont, 1927 | | | Uroxys batesi Harold, 1868 | | | Uroxys besti Ratcliffe, 1980 | | | Uroxys bidentis (Howden & Young, 1981) | ⇒ Uroxys boneti Pereira & Ha..., 1961 | | Uroxys boneti Pereira & Ha..., 1961 | | | Uroxys bonplandi Kohlmann, So..., 2019 | | | Uroxys brachialis Arrow, 1933 | | | Uroxys brevis Waterhouse, 1891 | | | Uroxys catharinensis Balthasar, 1966 | | | Uroxys caucanus Arrow, 1933 | | | Uroxys chichanich Delgado & Ko..., 2007 | | | Uroxys coarctatus Harold, 1867 | | | Uroxys corniculatus Harold, 1880 | | | Uroxys corporaali Balthasar, 1940 | | | Uroxys cuprescens Westwood, 1842 | | | Uroxys deavilai Delgado & Ko..., 2007 | | | Uroxys depressifrons (Howden & Young, 1981) | ⇒ Uroxys pauliani Balthasar, 1940 | | Uroxys dilaticollis Blanchard, 1846 | | | Uroxys dureti (Martínez, 1947) | ⇒ Uroxys epipleurale Boucomont, 1928 | | Uroxys dybasi Howden & Young, 1981 | | | Uroxys elongatus Harold, 1868 | | | Uroxys epipleurale Boucomont, 1928 | | | Uroxys frankenbergeri Balthasar, 1940 | | | Uroxys gatunensis Howden & Young, 1981 | | | Uroxys gorgon Arrow, 1933 | | | Uroxys histeroides Arrow, 1933 | | | Uroxys inconspicuus Harold, 1868 | | | Uroxys kratochvili Balthasar, 1940 | | | Uroxys laevipennis Kirsch, 1870 | | | Uroxys latesulcatus Bates, 1891 | | | Uroxys latus Arrow, 1933 | | | Uroxys lobata | | | Uroxys lojanus Arrow, 1933 | | | Uroxys macrocularis (Howden & Young, 1981) | ⇒ Uroxys boneti Pereira & Ha..., 1961 | | Uroxys magnus Balthasar, 1940 | | | Uroxys metagorgon Howden & Young, 1981 | | | Uroxys metallescens Harold, 1868 | | | Uroxys microcularis Howden & Young, 1981 | | | Uroxys micros Bates, 1887 | | | Uroxys minutus Harold, 1868 | | | Uroxys monstrosus Balthasar, 1940 | | | Uroxys nebulinus Howden & Gill, 1987 | | | Uroxys ohausi Balthasar, 1938 | | | Uroxys pauliani Balthasar, 1940 | | | Uroxys peruanus Balthasar, 1940 | | | Uroxys platypyga Howden & Young, 1981 | | | Uroxys productus Arrow, 1933 | | | Uroxys pygmaeus Harold, 1883 | | | Uroxys rugatus Boucomont, 1928 | | | Uroxys simplex Waterhouse, 1891 | | | Uroxys spaethi Balthasar, 1940 | | | Uroxys striatus Harold, 1868 | | | Uroxys sulai Balthasar, 1940 | | | Uroxys sulcicollis Harold, 1880 | | | Uroxys tacanensis Delgado & Ko..., 2007 | | | Uroxys terminalis Waterhouse, 1891 | | | Uroxys thoracalis Balthasar, 1940 | | | Uroxys transversifrons Howden & Gill, 1987 | | | Uroxys trinitatis Arrow, 1933 | | | Uroxys tuberculatus Lansberge, 1874 | | | Uroxys variabilis Robinson, 1951 | | | Uroxys vincentiae Arrow, 1903 | | | Versicorpus daures Deschodt, 2019 | | | Versicorpus erongoense Deschodt, Da..., 2011 | | | Versicorpus erongoensis Deschodt, Da..., 2011 | | | Versicorpus streyi Frolov, 2005 | | | Vulcanocanthon seminulum Harold, 1867 | | | Walterantus rufoclypeatus Frey, 1960 | | | Xenocanthon sericans Schmidt, 1922 | | | Xenocanthon vianai (Martínez, 1952) | ⇒ Xenocanthon sericans Schmidt, 1922 | | Xinidium davisi Cambefort, 1985 | | | Xinidium dentilabris Harold, 1869 | | | Xinidium dewitzi Harold, 1878 | | | Xinidium howdeni Cambefort, 1993 | | | Yvescambefortius sarawacus Gillet, 1926 | | | Zonocopris gibbicollis Harold, 1868 | | | Zonocopris machadoi Vaz-de-Mello, 2007 | |
Subnodes of Scarabaeinae with occurrences in Continents | Tribe | Ateuchini
Montreuil, 1998
| 36
453 | Genera
Species |
Taxom
| | Tribe | Canthonini
Péringuey, 1900
| 81
1121 | Genera
Species |
Taxom
| | Tribe | Coprini
Kolbe, 1805
| 19
1188 | Genera
Species |
Taxom
| | Tribe | Deltochilini
Lacordaire, 1856
| 2
31 | Genera
Species |
Taxom
| | Tribe | Eucraniini
Brullé, 1834
| 4
19 | Genera
Species |
Taxom
| | Tribe | Gymnopleurini
Lacordaire, 1856
| 4
137 | Genera
Species |
Taxom
| | Tribe | Oniticellini
d´Orbigny, 1916
| 25
309 | Genera
Species |
Taxom
| | Tribe | Onitini
Hope, 1837
| 18
247 | Genera
Species |
Taxom
| | Tribe | Onthophagini
Lacordaire, 1856
| 35
3105 | Genera
Species |
Taxom
| | Tribe | Phanaeini
MacLeay, 1819
| 12
251 | Genera
Species |
Taxom
| | Tribe | Scarabaeini
Latreille, 1802
| 10
228 | Genera
Species |
Taxom
| | Tribe | Sisyphini
Mulsant, 1842
| 3
137 | Genera
Species |
Taxom
|
Links, sources and further information Scarabaeinae at artfakta.se
Scarabaeinae at dyntaxa.se
Scarabaeinae at itis.gov
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