Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles)
and region Botswana (Southern Africa)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 315
2340
4567 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 313
2338
4562 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 297
2276
4408 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 205
1928
3874 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 47
658
1476 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 43
537
1145 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 9
188
519 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 9
188
519 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 164
468 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 45
268 | Genera
Species |
Taxom
|
Genera for Botswana
45 Genera with 268 Species #ACDEGHKLMNOPST
| Allogymnopleurus Janssens, 1855 | | 2 | Species | Taxom | | Anachalcos Hope, 1857 | | 1 | Species | Taxom | | Caccobius Thomson, 1855 | | 5 | Species | Taxom | | Canthidium Erichson, 1867 | | 1 | Species | Taxom | | Catharsius Hope, 1857 | | 11 | Species | Taxom | | Cheironitis van Lansberge, 1776 | | 4 | Species | Taxom | | Cleptocaccobius Cambefort, 1857 | | 2 | Species | Taxom | | Copris Geoffroy, 1962 | Mexican dung beetle | 16 | Species | Taxom | | Coptorhina Hope, 1857 | | 2 | Species | Taxom | | Delopleurus Erichson, 1857 | | 2 | Species | Taxom | | Digitonthophagus Balthasar, 1787 | | 1 | Species | Taxom | | Epirinus Reiche, 1901 | | 1 | Species | Taxom | | Euoniticellus Janssens, 1873 | | 2 | Species | Taxom | | Euonthophagus Balthasar, 1902 | | 3 | Species | Taxom | | Garreta Janssens, 1857 | | 2 | Species | Taxom | | Gymnopleurus Illiger, 1843 | | 8 | Species | Taxom | | Heliocopris Hope, 1878 | | 8 | Species | Taxom | | Heteronitis Gillet, 1862 | | 1 | Species | Taxom | | Hyalonthophagus Palestrini &..., 1913 | | 3 | Species | Taxom | | Kheper Janssens, 1843 | | 5 | Species | Taxom | | Latodrepanus Krikken, 1857 | | 1 | Species | Taxom | | Megalonitis Janssens, 1875 | | 2 | Species | Taxom | | Metacatharsius Paulian, 1857 | | 14 | Species | Taxom | | Milichus Péringuey, 1904 | | 3 | Species | Taxom | | Mimonthophagus Balthasar, 1905 | | 2 | Species | Taxom | Neateuchus Gillet, 1911 | | 0 | Species | Taxom | | Neosisyphus Müller, 1818 | | 3 | Species | Taxom | | Odontoloma Boheman, 1868 | | 2 | Species | Taxom | | Oniticellus Serville, 1840 | | 4 | Species | Taxom | | Onitis Fabricius, 1886 | | 19 | Species | Taxom | | Onthophagus Latreille, 1913 | | 73 | Species | Taxom | | Pachylomera Griffith & P..., 1874 | | 2 | Species | Taxom | | Pedaria Laporte, 1857 | | 4 | Species | Taxom | | Phalops Erichson, 1867 | | 7 | Species | Taxom | | Pinacotarsus Harold, 1875 | | 1 | Species | Taxom | | Proagoderus Lansberge, 1901 | | 12 | Species | Taxom | | Pycnopanelus Arrow, 1978 | | 1 | Species | Taxom | | Sarophorus Erichson, 1857 | | 1 | Species | Taxom | | Scarabaeus Linnaeus, 1901 | | 27 | Species | Taxom | | Sceliages Westwood, 2002 | | 1 | Species | Taxom | Sebasteos Westwood, 1845 | | 0 | Species | Taxom | | Sisyphus Latreille, 1818 | | 7 | Species | Taxom | | Stiptopodius Harold, 2020 | | 1 | Species | Taxom | | Tiniocellus Péringuey, 1851 | | 2 | Species | Taxom | | Tragiscus Klug, 1855 | | 1 | Species | Taxom |
Species list for Botswana
268 Species and 4 Subspecies | Allogymnopleurus splendidus Bertoloni, 1849 | | | Allogymnopleurus thalassinus Klug, 1855 | | | Anachalcos convexus Boheman, 1857 | | | Caccobius castaneus (Klug, 1855) | ⇒ Caccocnemus castaneus Klug, 1855 | | Caccobius cavatus d´Orbigny, 1908 | | | Caccobius ferrugineus Fahraeus, 1857 | | | Caccobius histerinus Fahraeus, 1857 | | | Caccobius nigritulus Klug, 1855 | | | Canthidium smaragdinum Harold, 1867 | | | Catharsius aegeus Génier, 2017 | | | Catharsius calaharicus Kolbe, 1893 | | | Catharsius disseptus (Péringuey, 1901) | ⇒ Catharsius calaharicus Kolbe, 1893 | | Catharsius heros Boheman, 1860 | | | Catharsius longiceps Gillet, 1907 | | | Catharsius marcellus Kolbe, 1893 | | | Catharsius melancholicus (Boheman, 1860) | ⇒ Catharsius longiceps Gillet, 1907 | | Catharsius pandion Harold, 1877 | | | Catharsius philus Kolbe, 1893 | | | Catharsius tricornutus DeGeer, 1778 | | | Catharsius ulysses Boheman, 1857 | | | Cheironitis hoplosternus Harold, 1868 | | | Cheironitis imitator Balthasar, 1963 | | | Cheironitis indicus (Lansberge, 1875) | ⇒ Cheironitis arrowi Janssens, 1937 | | Cheironitis scabrosus Fabricius, 1776 | | | Cleptocaccobius convexifrons Raffray, 1877 | | | Cleptocaccobius viridicollis Fahraeus, 1857 | | | Copris amyntor (Klug, 1855) | ⇒ Copris harrisi Waterhouse, 1891 | | Copris bootes Klug, 1855 | | | Copris cassius Péringuey, 1901 | | | Copris cornifrons Boheman, 1860 | | | Copris denticulatus Nguyen-phung, 1988 | | | Copris elphenor Klug, 1855 | | | Copris evanidus Klug, 1855 | | | Copris fallaciosus Gillet, 1907 | Kenyan dung beetle | | Copris fidius Olivier, 1789 | | | Copris harrisi Waterhouse, 1891 | | | Copris inhalatus perturbator Péringuey, 1901 | | | Copris mesacanthus Harold, 1878 | | | Copris misellus Péringuey, 1901 | | | Copris puncticollis Boheman, 1857 | | | Copris subsidens Péringuey, 1901 | | | Copris vilhenai Ferreira, 1962 | | | Coptorhina auspicata Péringuey, 1901 | | | Coptorhina nitidipennis Boheman, 1857 | | | Delopleurus darrenmanni Frolov, 2014 | | | Delopleurus pullus Boheman, 1857 | | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Epirinus gratus Péringuey, 1901 | | | Euoniticellus kawanus Janssens, 1939 | | | Euoniticellus triangulatus Harold, 1873 | | | Euonthophagus carbonarius Klug, 1855 | | | Euonthophagus consobrinus Péringuey, 1901 | | | Euonthophagus flavimargo Orbigny, 1902 | | | Garreta nitens Olivier, 1789 | | | Garreta wahlbergi Fahraeus, 1857 | | | Gymnopleurus aenescens Wiedemann, 1821 | | | Gymnopleurus asperrimus Felsche, 1909 | | | Gymnopleurus bicolor (Latreille, 1823) | ⇒ Gymnopleurus leei Fabricius, 1792 | | Gymnopleurus humanus MacLeay, 1821 | | | Gymnopleurus humeralis Klug, 1855 | | | Gymnopleurus ignitus Klug, 1855 | | | Gymnopleurus pumilus (Reiche, 1847) | ⇒ Gymnopleurus virens Erichson, 1843 | | Gymnopleurus virens Erichson, 1843 | | | Heliocopris andersoni Bates, 1868 | | | Heliocopris atropos Boheman, 1860 | | | Heliocopris faunus Boheman, 1857 | | | Heliocopris hamadryas Fabricius, 1775 | | | Heliocopris japetus Klug, 1855 | | | Heliocopris neptunus Boheman, 1857 | | | Heliocopris pirmal Fabricius, 1798 | | | Heliocopris samson Harold, 1878 | | | Heteronitis castelnaui Harold, 1862 | | | Hyalonthophagus alcedo Orbigny, 1913 | | | Hyalonthophagus alcyon Klug, 1855 | | | Hyalonthophagus alcyonides Orbigny, 1913 | | | Kheper cupreus Castelnau, 1840 | | | Kheper kalaharicus Davis, Desch..., 2011 | | | Kheper lamarcki MacLeay, 1821 | | | Kheper nigroaeneus Boheman, 1857 | | | Kheper prodigiosus Erichson, 1843 | | | Latodrepanus laticollis Fahraeus, 1857 | | | Megalonitis bohemani Lansberge, 1875 | | | Megalonitis bohemanni Lansberge, 1875 | | | Metacatharsius anderseni Waterhouse, 1891 | | | Metacatharsius dentinum Ferreira, 1964 | | | Metacatharsius exiguiformis Ferreira, 1964 | | | Metacatharsius exiguus Boheman, 1860 | | | Metacatharsius ferreirae Balthasar, 1965 | | | Metacatharsius freyi Ferreira, 1964 | | | Metacatharsius latifrons Harold, 1868 | | | Metacatharsius marani Balthasar, 1940 | | | Metacatharsius opacus Waterhouse, 1891 | | | Metacatharsius pumilioniformis Ferreira, 1964 | | | Metacatharsius pumilionis Wallengren, 1881 | | | Metacatharsius pusio Kolbe, 1908 | | | Metacatharsius rugosipennis Frey, 1975 | | | Metacatharsius troglodytes (Boheman, 1857) | ⇒ Metacatharsius anderseni Waterhouse, 1891 | | Milichus apicalis (Fahraeus, 1857) | ⇒ Milichus rhodesianus Péringuey, 1904 | | Milichus picticollis Gerstaecker, 1871 | | | Milichus rhodesianus Péringuey, 1904 | | | Mimonthophagus anomalus Klug, 1855 | | | Mimonthophagus limbibasis d´Orbigny, 1905 | | | Neosisyphus calcaratus Klug, 1855 | | | Neosisyphus rubrus Paschalidis, 1974 | | | Neosisyphus spinipes Thunberg, 1818 | Grey dungball roller | | Odontoloma louwi Howden & Sch..., 1987 | | | Odontoloma sculpturatum Harold, 1868 | | | Oniticellus egregius Klug, 1855 | | | Oniticellus formosus (Chevrolat, 1830) | ⇒ Oniticellus pictus Hausmann, 1807 | | Oniticellus pictus Hausmann, 1807 | | | Oniticellus planatus Castelnau, 1840 | | | Onitis adriani Ferreira, 1976 | | | Onitis aeruginosus Klug, 1855 | | | Onitis alexis (Klug, 1835) | ⇒ Cheironitis ungaricus ungaricus Herbst, 1789 | | Onitis alexis alexis | | | Onitis aygulus Fabricius, 1781 | | | Onitis cribratus Lansberge, 1875 | | | Onitis cupreus Castelnau, 1840 | | | Onitis curvipes Lansberge, 1875 | | | Onitis deceptor Péringuey, 1901 | | | Onitis fulgidus (Klug, 1855) | ⇒ Pleuronitis fulgidus Castelnau, 1840 | | Onitis inversidens Lansberge, 1875 | | | Onitis mendax Gillet, 1918 | | | Onitis obenbergeri Balthasar, 1942 | | | Onitis orthopus Lansberge, 1875 | | | Onitis picticollis Boheman, 1857 | | | Onitis robustus Boheman, 1857 | | | Onitis uncinatus Klug, 1855 | | | Onitis viridulus (Boheman, 1857) | ⇒ Onitis pecuarius Lansberge, 1875 | | Onitis westermanni Lansberge, 1886 | | | Onthophagus (Afrostrandius) plebejus Klug, 1855 | | | Onthophagus (Digitonthophagus) gazella (Fabricius, 1787) | ⇒ Digitonthophagus gazella Fabricius, 1787 | | Onthophagus (Onthophagus) taurus Schreber, 1759 | Bull Headed Dung Beetle | | Onthophagus (Onthophagus) trituber Wiedemann, 1823 | | | Onthophagus marginalis przewalski Kabakov, 2006 | | | Onthophagus (Proagoderus) alcedo d´Orbigny, 1913 | | | Onthophagus (Proagoderus) alcyon (Klug, 1855) | ⇒ Onthophagus alcedo d´Orbigny, 1913 | | Onthophagus (Proagoderus) alcyonides d´Orbigny, 1913 | | | Onthophagus (Proagoderus) aureiceps d´Orbigny, 1902 | | | Onthophagus (Proagoderus) pegesimallus Marcus, 1917 | | | Onthophagus acutus d´Orbigny, 1908 | | | Onthophagus aeruginosus Roth, 1851 | | | Onthophagus albipodex d´Orbigny, 1902 | | | Onthophagus apiciosus d´Orbigny, 1902 | | | Onthophagus asimilis Péringuey, 1901 | | | Onthophagus betschuanus Frey, 1975 | | | Onthophagus bicavifrons d´Orbigny, 1902 | | | Onthophagus bovinus Péringuey, 1892 | | | Onthophagus burchelli d´Orbigny, 1908 | | | Onthophagus chalcostolus d´Orbigny, 1902 | | | Onthophagus convexus d´Orbigny, 1908 | | | Onthophagus criniger d´Orbigny, 1904 | | | Onthophagus cyaneoniger d´Orbigny, 1902 | | | Onthophagus ebenicolor d´Orbigny, 1902 | | | Onthophagus ebenus Péringuey, 1888 | | | Onthophagus echinus Boucomont, 1914 | | | Onthophagus filicornis Harold, 1873 | | | Onthophagus fimetarius Roth, 1851 | | | Onthophagus flavolimbatus Klug, 1855 | | | Onthophagus fugitivus Péringuey, 1901 | | | Onthophagus granulifer (Harold, 1886) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus granulifrons Frey, 1961 | | | Onthophagus graphicus Wallengren, 1881 | | | Onthophagus guttatus (Boheman, 1860) | ⇒ Onthophagus quadrilunatus d´Orbigny, 1902 | | Onthophagus impressicollis Boheman, 1860 | | | Onthophagus juvencus Klug, 1835 | | | Onthophagus kochi Frey, 1957 | | | Onthophagus lamelliger (Gerstaecker, 1871) | ⇒ Onthophagus ochropygus d´Orbigny, 1902 | | Onthophagus leucopygus (Harold, 1867) | ⇒ Onthophagus albipodex d´Orbigny, 1902 | | Onthophagus macrothorax d´Orbigny, 1902 | | | Onthophagus merus Péringuey, 1901 | | | Onthophagus minutulus Harold, 1875 | | | Onthophagus minutus (Hausmann, 1807) | ⇒ Onthophagus centricornis Fabricius, 1798 | | Onthophagus obtusicornis (Fahraeus, 1857) | ⇒ Onthophagus marginalis przewalski Kabakov, 2006 | | Onthophagus pallidipennis (Fahraeus, 1857) | ⇒ Onthophagus rasipennis d´Orbigny, 1908 | | Onthophagus pauxillus d´Orbigny, 1902 | | | Onthophagus pilosus (Fahraeus, 1857) | ⇒ Onthophagus echinus Boucomont, 1914 | | Onthophagus praedatus Harold, 1862 | | | Onthophagus probus Péringuey, 1901 | | | Onthophagus pugionatus Fahraeus, 1857 | | | Onthophagus pullus (Roth, 1851) | ⇒ Onthophagus parenthesis Boucomont, 1913 | | Onthophagus quadraticeps Harold, 1867 | | | Onthophagus quadrilunatus d´Orbigny, 1902 | | | Onthophagus quadrimaculatus Raffray, 1877 | | | Onthophagus quadrinodosus Fahraeus, 1857 | | | Onthophagus rasipennis d´Orbigny, 1908 | | | Onthophagus rhyncophorus | | | Onthophagus rojkoffianus Josso, 2018 | | | Onthophagus rubens d´Orbigny, 1902 | | | Onthophagus semiflavus (Boheman, 1860) | ⇒ Onthophagus flavolimbatus Klug, 1855 | | Onthophagus signatus Fahraeus, 1857 | | | Onthophagus stellio Erichson, 1843 | | | Onthophagus suffusus Klug, 1855 | | | Onthophagus sugillatus Klug, 1855 | | | Onthophagus suturalis (Péringuey, 1888) | ⇒ Onthophagus trituber Wiedemann, 1823 | | Onthophagus tonsus d´Orbigny, 1902 | | | Onthophagus transcaspicus Koenig, 1888 | Gazella scarab | | Onthophagus tricorniger Boheman, 1860 | | | Onthophagus variegatus (Fabricius, 1798) | ⇒ Onthophagus filicornis Harold, 1873 | | Onthophagus verticalis (Fahraeus, 1857) | ⇒ Onthophagus praedatus Harold, 1862 | | Onthophagus vinctus Erichson, 1843 | | | Onthophagus virescens Harold, 1867 | | | Onthophagus vylderi d´Orbigny, 1913 | | | Pachylomera femoralis Kirby, 1828 | | | Pachylomera opaca Lansberge, 1874 | | | Pedaria aspera Péringuey, 1901 | | | Pedaria conformis Péringuey, 1901 | | | Pedaria cylindrica Fahraeus, 1857 | | | Pedaria picea Fahraeus, 1857 | | | Phalops ardea Klug, 1855 | | | Phalops boschas Klug, 1855 | | | Phalops dregei Harold, 1867 | | | Phalops flavocinctus Klug, 1855 | | | Phalops rufosignatus (Lansberge, 1885) | ⇒ Phalops dregei Harold, 1867 | | Phalops smaragdinus Harold, 1875 | | | Phalops wittei Harold, 1867 | | | Pinacotarsus dohrni Harold, 1875 | | | Proagoderus bicallosus Klug, 1855 | | | Proagoderus brucei (d´Orbigny, 1902) | ⇒ Onthophagus chalcostolus d´Orbigny, 1902 | | Proagoderus brucei brucei | | | Proagoderus chalcostolus d´Orbigny, 1902 | | | Proagoderus furcifer Boheman, 1860 | | | Proagoderus lanista Castelnau, 1840 | | | Proagoderus loricatus Klug, 1855 | | | Proagoderus marginidens d´Orbigny, 1904 | | | Proagoderus pyramidalis Klug, 1855 | | | Proagoderus rangifer Klug, 1855 | | | Proagoderus rectefurcatus Fairmaire, 1891 | | | Proagoderus sapphirinus (Péringuey, 1901) | ⇒ Onthophagus aureiceps d´Orbigny, 1902 | | Pycnopanelus krikkeni Cambefort, 1978 | | | Sarophorus costatus Fahraeus, 1857 | | | Scarabaeus ambiguus (Boheman, 1857) | ⇒ Gnorimus variabilis Linnaeus, 1758 | | Scarabaeus anderseni Waterhouse, 1890 | | | Scarabaeus bohemani Harold, 1868 | | | Scarabaeus carniphilus Davis & Desc..., 2015 | | | Scarabaeus costatus Wiedemann, 1823 | | | Scarabaeus damarensis Janssens, 1940 | | | Scarabaeus deludens zur Strassen, 1961 | | | Scarabaeus flavicornis Boheman, 1860 | | | Scarabaeus funebris Boheman, 1857 | | | Scarabaeus galenus Westwood, 1847 | | | Scarabaeus goryi Castelnau, 1840 | | | Scarabaeus indus (Linnaeus, 1758) | ⇒ Euphoria inda Linnaeus, 1758 | | Scarabaeus interstitialis Boheman, 1857 | | | Scarabaeus intricatus Fabricius, 1801 | | | Scarabaeus kochi Ferreira, 1952 | | | Scarabaeus lamarcki MacLeay, 1821 | | | Scarabaeus lucidulus Boheman, 1860 | | | Scarabaeus natalensis zur Strassen, 1962 | | | Scarabaeus palemo Olivier, 1789 | | | Scarabaeus proboscideus Guérin-Ménev..., 1844 | | | Scarabaeus profanus Fabricius, 1792 | | | Scarabaeus rusticus Boheman, 1857 | | | Scarabaeus satyrus (Boheman, 1860) | ⇒ Xyloryctes jamaicensis Drury, 1773 | | Scarabaeus sulcipennis Quedenfeldt, 1888 | | | Scarabaeus vicinus (Janssens, 1940) | ⇒ Lucanus cervus Linnaeus, 1758 | | Scarabaeus zambesianus Péringuey, 1901 | | | Scarabaeus zambezianus Péringuey, 1901 | | | Sceliages granulatus Forgie & Gre..., 2002 | | | Sisyphus calcaratus (Boucomont, 1937) | ⇒ Neosisyphus rugosus Gory, 1833 | | Sisyphus crispatus Gory, 1833 | | | Sisyphus goryi (Harold, 1859) | ⇒ Sisyphus caffer goryi Harold, 1859 | | Sisyphus impressipennis Lansberge, 1886 | | | Sisyphus nanniscus Péringuey, 1901 | | | Sisyphus rubrus (Paschalidis, 1974) | ⇒ Neosisyphus rubrus Paschalidis, 1974 | | Sisyphus spinipes (Thunberg, 1818) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Stiptopodius savuti Daniel & Des..., 2020 | | | Tiniocellus dolosus Branco, 2010 | | | Tiniocellus spinipes Roth, 1851 | | | Tragiscus dimidiatus Klug, 1855 | |
Subnodes of Scarabaeinae with occurrences in Botswana
Links, sources and further information Scarabaeinae at artfakta.se
Scarabaeinae at dyntaxa.se
Scarabaeinae at itis.gov
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